205 lines
20 KiB
XML
205 lines
20 KiB
XML
<document ID-DOI="10.11646/zootaxa.5254.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="7727424" ID-ZooBank="5D189B4F-87E1-4F6D-9C8E-9E3F1663512C" approvalRequired="2" approvalRequired_for_textStreams="2" checkinTime="1678695648311" checkinUser="plazi" docAuthor="Bian, Dongju & Zhang, Yi" docDate="2023" docId="E1624D40FFE3FFE56CEDFDD2FEA2B7BC" docLanguage="en" docName="zootaxa.5254.2.6.pdf" docOrigin="Zootaxa 5254 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5254.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Grouvellinus spnaericus Bian & Zhang 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="258" masterDocId="1D5B3538FFE2FFE46C7AFFE9FFF1B04B" masterDocTitle="Descriptions of eight new species of Grouvellinus from China (Coleoptera, Elmidae)" masterLastPageNumber="277" masterPageNumber="257" pageNumber="258" updateTime="1678696102211" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Descriptions of eight new species of Grouvellinus from China (Coleoptera, Elmidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Bian, Dongju</mods:namePart>
|
||
<mods:affiliation>Key Laboratory of Forest Ecology and Management, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang, 110016, China University of Chinese Academy of Sciences, Beijing 100049, China.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Zhang, Yi</mods:namePart>
|
||
<mods:affiliation>Key Laboratory of Forest Ecology and Management, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang, 110016, China University of Chinese Academy of Sciences, Beijing 100049, China.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2023</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2023-03-10</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>5254</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>257</mods:start>
|
||
<mods:end>277</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.11646/zootaxa.5254.2.6</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.11646/zootaxa.5254.2.6</mods:identifier>
|
||
<mods:identifier type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">7727424</mods:identifier>
|
||
<mods:identifier type="ZooBank">5D189B4F-87E1-4F6D-9C8E-9E3F1663512C</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7727432" ID-Zenodo-Dep="7727432" LSID="urn:lsid:plazi:treatment:E1624D40FFE3FFE56CEDFDD2FEA2B7BC" httpUri="http://treatment.plazi.org/id/E1624D40FFE3FFE56CEDFDD2FEA2B7BC" lastPageNumber="258" pageId="1" pageNumber="258">
|
||
<subSubSection box="[151,548,571,598]" pageId="1" pageNumber="258" type="nomenclature">
|
||
<paragraph blockId="1.[151,548,571,634]" box="[151,548,571,598]" pageId="1" pageNumber="258">
|
||
<heading bold="true" box="[151,548,571,598]" fontSize="11" level="1" pageId="1" pageNumber="258" reason="1">
|
||
<emphasis bold="true" box="[151,548,571,598]" pageId="1" pageNumber="258">
|
||
<taxonomicName authority="Bian & Zhang, 2023" authorityName="Bian & Zhang" authorityYear="2023" box="[151,447,571,598]" class="Insecta" family="Elmidae" genus="Grouvellinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="spnaericus" status="sp. nov.">
|
||
<emphasis bold="true" box="[151,447,571,598]" italics="true" pageId="1" pageNumber="258">Grouvellinus spnaericus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[454,548,572,598]" pageId="1" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[151,472,608,634]" pageId="1" pageNumber="258" type="description">
|
||
<paragraph blockId="1.[151,548,571,634]" box="[151,472,608,634]" pageId="1" pageNumber="258">
|
||
<heading bold="true" box="[151,472,608,634]" fontSize="11" level="1" pageId="1" pageNumber="258" reason="1">
|
||
<emphasis bold="true" box="[151,472,608,634]" pageId="1" pageNumber="258">
|
||
(
|
||
<figureCitation box="[159,285,608,634]" captionStart="FIGURE 1" captionStartId="2.[151,250,1655,1679]" captionTargetBox="[170,1422,197,1612]" captionTargetId="figure-31@2.[151,1436,179,1629]" captionTargetPageId="2" captionText="FIGURE 1. Habitus A–B) Grouvellinus spnaericus sp. nov. C–D) G. elongatus sp. nov. A, C) dorsal view B, D) ventral view." figureDoi="http://doi.org/10.5281/zenodo.7727426" httpUri="https://zenodo.org/record/7727426/files/figure.png" pageId="1" pageNumber="258">Figs 1A–B</figureCitation>
|
||
,
|
||
<figureCitation box="[298,368,608,634]" captionStart="FIGURE 5" captionStartId="12.[151,250,979,1003]" captionTargetBox="[265,1323,236,954]" captionTargetId="figure-52@12.[265,1323,236,955]" captionTargetPageId="12" captionText="FIGURE 5. Grouvellinus spnaericus sp. nov. A) anterior part of body, dorsal view; B) same, ventral view; C) posterior part of body, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7727440" httpUri="https://zenodo.org/record/7727440/files/figure.png" pageId="1" pageNumber="258">5A–C</figureCitation>
|
||
,
|
||
<figureCitation box="[381,465,608,634]" captionStart="FIGURE 13" captionStartId="16.[151,250,1577,1601]" captionTargetBox="[320,1267,181,1553]" captionTargetId="figure-31@16.[320,1267,181,1553]" captionTargetPageId="16" captionText="FIGURE 13 Aedeagus A–C) Grouvellinus spnaericus sp. nov. D–F) G. elongatus sp. nov. A, D) ventral view B, E) dorsal view C, F) lateral view." figureDoi="http://doi.org/10.5281/zenodo.7727466" httpUri="https://zenodo.org/record/7727466/files/figure.png" pageId="1" pageNumber="258">13A–C</figureCitation>
|
||
)
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="1" pageNumber="258" type="materials_examined">
|
||
<paragraph blockId="1.[151,1437,680,2039]" pageId="1" pageNumber="258">
|
||
<emphasis bold="true" box="[151,452,680,706]" pageId="1" pageNumber="258">Type materiral examined</emphasis>
|
||
:
|
||
<materialsCitation collectingDate="2019-06-21" collectionCode="IAECAS" country="China" county="Yingjiang County" elevation="925" latitude="24.641666" location="Yingjiang County" longLatPrecision="92" longitude="97.586334" pageId="1" pageNumber="258" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
|
||
<typeStatus box="[466,567,680,706]" pageId="1" pageNumber="258">holotype</typeStatus>
|
||
, male (
|
||
<collectionCode box="[647,755,680,706]" pageId="1" pageNumber="258">IAECAS</collectionCode>
|
||
): “
|
||
<collectingCountry box="[790,885,680,706]" name="China" pageId="1" pageNumber="258">CHINA</collectingCountry>
|
||
:
|
||
<collectingRegion box="[894,980,681,707]" country="China" name="Yunnan" pageId="1" pageNumber="258">Yunnan</collectingRegion>
|
||
/
|
||
<collectingCounty box="[1000,1199,680,706]" pageId="1" pageNumber="258">Yingjiang County</collectingCounty>
|
||
/ Tongbiguan Vill.ļ
|
||
<geoCoordinate box="[151,286,716,743]" degrees="97" direction="east" minutes="35.18" orientation="longitude" pageId="1" pageNumber="258" precision="9" value="97.586334">97°35.18’ E</geoCoordinate>
|
||
<geoCoordinate box="[294,417,716,743]" degrees="24" direction="north" minutes="38.5" orientation="latitude" pageId="1" pageNumber="258" precision="92" value="24.641666">24°38.5’ N</geoCoordinate>
|
||
/
|
||
<quantity box="[441,515,716,743]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="1" pageNumber="258" unit="m" value="925.0">
|
||
<elevation box="[441,515,716,743]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="1" pageNumber="258" unit="m" value="925.0">925 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date box="[528,655,716,742]" pageId="1" pageNumber="258" value="2019-06-21">
|
||
<collectingDate box="[528,655,716,742]" pageId="1" pageNumber="258" value="2019-06-21">21.VI.2019</collectingDate>
|
||
</date>
|
||
/ Leg. Peng
|
||
<emphasis box="[802,859,716,742]" italics="true" pageId="1" pageNumber="258">et al.</emphasis>
|
||
(2)”
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation collectingDate="2019-06-21" collectionCode="IAECAS" country="China" county="Yingjiang County" elevation="925" latitude="24.641666" location="Yingjiang County" longLatPrecision="92" longitude="97.586334" pageId="1" pageNumber="258" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
|
||
<typeStatus box="[927,1039,717,743]" pageId="1" pageNumber="258">Paratypes</typeStatus>
|
||
,
|
||
<specimenCount box="[1051,1141,716,742]" pageId="1" pageNumber="258" type="male">3 males</specimenCount>
|
||
,
|
||
<specimenCount box="[1153,1251,716,742]" pageId="1" pageNumber="258" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode box="[1267,1374,716,742]" pageId="1" pageNumber="258">IAECAS</collectionCode>
|
||
), the same data as holotype
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="1" pageNumber="258" type="diagnosis">
|
||
<paragraph blockId="1.[151,1437,680,2039]" pageId="1" pageNumber="258">
|
||
<emphasis bold="true" box="[199,322,788,814]" pageId="1" pageNumber="258">Diagnosis.</emphasis>
|
||
Body broadly obovate, dark brown to black; disc of pronotum without median carina or sulcus; sublateral carinae present in basal 0.4; pronotum hemispherically vaulted; elytra short, sub-cordiform. This species can be easily distinguished from other known species in the genus
|
||
<taxonomicName authorityName="Champion" authorityYear="1923" box="[915,1060,860,886]" class="Insecta" family="Elmidae" genus="Grouvellinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[915,1060,860,886]" italics="true" pageId="1" pageNumber="258">Grouvellinus</emphasis>
|
||
</taxonomicName>
|
||
by its pronotum hemispherically vaulted; elytra short, sub-cordiform, and also by its penis broad in basal 0.8, and distal 0.2 suddenly narrowed, apex narrowly rounded.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="1" pageNumber="258" type="description">
|
||
<paragraph blockId="1.[151,1437,680,2039]" pageId="1" pageNumber="258">
|
||
<emphasis bold="true" box="[199,337,968,994]" pageId="1" pageNumber="258">Description</emphasis>
|
||
. Habitus see
|
||
<figureCitation box="[492,618,968,994]" captionStart="FIGURE 1" captionStartId="2.[151,250,1655,1679]" captionTargetBox="[170,1422,197,1612]" captionTargetId="figure-31@2.[151,1436,179,1629]" captionTargetPageId="2" captionText="FIGURE 1. Habitus A–B) Grouvellinus spnaericus sp. nov. C–D) G. elongatus sp. nov. A, C) dorsal view B, D) ventral view." figureDoi="http://doi.org/10.5281/zenodo.7727426" httpUri="https://zenodo.org/record/7727426/files/figure.png" pageId="1" pageNumber="258">Figs 1A–B</figureCitation>
|
||
. Body broadly obovate. BL
|
||
<quantity box="[939,1028,968,995]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="258" unit="mm" value="2.5">2.5 mm</quantity>
|
||
, BW
|
||
<quantity box="[1093,1183,968,995]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="258" unit="mm" value="1.4">1.4 mm</quantity>
|
||
. Dark brown to black, legs dark brown except tarsi light brown.
|
||
</paragraph>
|
||
<paragraph blockId="1.[151,1437,680,2039]" pageId="1" pageNumber="258">Head. Labrum black, but yellowish brown at the anterior margin, with dense long yellowish setae anteriorlaterally; surface smooth and shining, fine setiferous punctures sparsely distributed. Clypeus and frons black, smooth and shining, medium-sized setiferous punctures densely distributed.</paragraph>
|
||
<paragraph blockId="1.[151,1437,680,2039]" pageId="1" pageNumber="258">
|
||
Pronotum (
|
||
<figureCitation box="[329,420,1148,1175]" captionStart="FIGURE 5" captionStartId="12.[151,250,979,1003]" captionTargetBox="[265,1323,236,954]" captionTargetId="figure-52@12.[265,1323,236,955]" captionTargetPageId="12" captionText="FIGURE 5. Grouvellinus spnaericus sp. nov. A) anterior part of body, dorsal view; B) same, ventral view; C) posterior part of body, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7727440" httpUri="https://zenodo.org/record/7727440/files/figure.png" pageId="1" pageNumber="258">Fig. 5A</figureCitation>
|
||
) hemispherically vaulted, PL
|
||
<quantity box="[773,865,1148,1175]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="258" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, PW
|
||
<quantity box="[933,1025,1148,1175]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="258" unit="mm" value="1.1">1.1 mm</quantity>
|
||
, broadest at basal 0.3, lateral sides slightly attenuated posteriorly and distinctly attenuated anteriorly. Lateral margin denticulated in anterior half. Anterior angles sharp, slightly produced; posterior angles nearly right-angled. Surface coarse, densely punctate and pubescent, setae yellow and short. Sublateral carina present in basal 0.4, median carina absent.
|
||
</paragraph>
|
||
<paragraph blockId="1.[151,1437,680,2039]" pageId="1" pageNumber="258">
|
||
EL
|
||
<quantity box="[242,333,1292,1319]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="258" unit="mm" value="1.7">1.7 mm</quantity>
|
||
, EW
|
||
<quantity box="[399,490,1292,1319]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="258" unit="mm" value="1.4">1.4 mm</quantity>
|
||
, subparallel in anterior half, then distinctly attenuated posteriorly. Strial punctures small and deep, separated by 0.5 to 1 time of their diameters in basal half, then well separated in distal half, separated by 2–3 times of their diameters. Intervals coarse and convex, sparsely punctate and pubescent. Intervals 3, 5, 7, 8 carinate. Carinae on intervals 3 present from base to basal half. Carinae on intervals 5, 7, 8 extend from base to subapical part.
|
||
</paragraph>
|
||
<paragraph blockId="1.[151,1437,680,2039]" pageId="1" pageNumber="258">
|
||
Prosternum densely pubescent, prosternal process (
|
||
<figureCitation box="[771,857,1472,1499]" captionStart="FIGURE 5" captionStartId="12.[151,250,979,1003]" captionTargetBox="[265,1323,236,954]" captionTargetId="figure-52@12.[265,1323,236,955]" captionTargetPageId="12" captionText="FIGURE 5. Grouvellinus spnaericus sp. nov. A) anterior part of body, dorsal view; B) same, ventral view; C) posterior part of body, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7727440" httpUri="https://zenodo.org/record/7727440/files/figure.png" pageId="1" pageNumber="258">Fig. 5B</figureCitation>
|
||
) slightly narrowed in basal 0.6, subparallel in distal 0.4, apex broadly rounded. Surface smooth and shining, sparsely punctate and pubescent. Disc slightly elevated, lateral margin distinctly rimmed. Metaventrite (
|
||
<figureCitation box="[677,761,1544,1571]" captionStart="FIGURE 5" captionStartId="12.[151,250,979,1003]" captionTargetBox="[265,1323,236,954]" captionTargetId="figure-52@12.[265,1323,236,955]" captionTargetPageId="12" captionText="FIGURE 5. Grouvellinus spnaericus sp. nov. A) anterior part of body, dorsal view; B) same, ventral view; C) posterior part of body, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7727440" httpUri="https://zenodo.org/record/7727440/files/figure.png" pageId="1" pageNumber="258">Fig. 5C</figureCitation>
|
||
) broadly impressed, disc coarse with large punctures densely impressed, an oblique row of large punctures behind of metacoxa and a groove in front of metacoxa. Lateral area densely pubescent with small punctures sparsely distributed.
|
||
</paragraph>
|
||
<paragraph blockId="1.[151,1437,680,2039]" pageId="1" pageNumber="258">
|
||
Ventrites I–II with disc wrinkled and densely punctate. Ventrites III–IV and anterior third of ventrite V with disc smooth and shining, sparsely punctate, only with a few pubescence. A pair of ad-median carinae present on ventrite I. Ventrite V densely granulate and pubescent in distal 2/3; lateral area with dense pubescence (
|
||
<figureCitation box="[1215,1304,1724,1751]" captionStart="FIGURE 5" captionStartId="12.[151,250,979,1003]" captionTargetBox="[265,1323,236,954]" captionTargetId="figure-52@12.[265,1323,236,955]" captionTargetPageId="12" captionText="FIGURE 5. Grouvellinus spnaericus sp. nov. A) anterior part of body, dorsal view; B) same, ventral view; C) posterior part of body, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7727440" httpUri="https://zenodo.org/record/7727440/files/figure.png" pageId="1" pageNumber="258">Fig. 5C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="1.[151,1437,680,2039]" pageId="1" pageNumber="258">
|
||
Aedeagus (
|
||
<figureCitation box="[324,467,1760,1786]" captionStart="FIGURE 13" captionStartId="16.[151,250,1577,1601]" captionTargetBox="[320,1267,181,1553]" captionTargetId="figure-31@16.[320,1267,181,1553]" captionTargetPageId="16" captionText="FIGURE 13 Aedeagus A–C) Grouvellinus spnaericus sp. nov. D–F) G. elongatus sp. nov. A, D) ventral view B, E) dorsal view C, F) lateral view." figureDoi="http://doi.org/10.5281/zenodo.7727466" httpUri="https://zenodo.org/record/7727466/files/figure.png" pageId="1" pageNumber="258">Figs 13A–C</figureCitation>
|
||
). 1.0 mm long. Penis is about 2.7 times as long as phallobase. Penis broad, about three times as wide as the paramere, slightly broad from base to basal 0.8, then distinctly sharpened, apex narrowly rounded. Apophyses developed, reaching the subapical part of the phallobase. Parameres (in dorsal view) slim, about 1/3 as wide as penis, apices of the parameres narrowly rounded, not overreaching the penis.
|
||
</paragraph>
|
||
<paragraph blockId="1.[151,1437,680,2039]" box="[199,1214,1904,1931]" pageId="1" pageNumber="258">
|
||
Males: BL
|
||
<quantity box="[324,475,1904,1931]" metricMagnitude="-3" metricUnit="m" metricValue="2.4749999999999996" metricValueMax="2.55" metricValueMin="2.4" pageId="1" pageNumber="258" unit="mm" value="2.4749999999999996" valueMax="2.55" valueMin="2.4">2.4–2.55 mm</quantity>
|
||
, BW
|
||
<quantity box="[538,673,1904,1931]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="1" pageNumber="258" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4–1.5 mm</quantity>
|
||
(n=4); females: BL
|
||
<quantity box="[901,989,1904,1931]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="258" unit="mm" value="2.5">2.5 mm</quantity>
|
||
, BW
|
||
<quantity box="[1052,1138,1904,1931]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="258" unit="mm" value="1.4">1.4 mm</quantity>
|
||
(n=1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,533,1940,1967]" pageId="1" pageNumber="258" type="distribution">
|
||
<paragraph blockId="1.[151,1437,680,2039]" box="[199,533,1940,1967]" pageId="1" pageNumber="258">
|
||
<emphasis bold="true" box="[199,353,1940,1966]" pageId="1" pageNumber="258">Distribution.</emphasis>
|
||
<collectingCountry box="[359,430,1940,1966]" name="China" pageId="1" pageNumber="258">China</collectingCountry>
|
||
:
|
||
<collectingRegion box="[440,529,1941,1967]" country="China" name="Yunnan" pageId="1" pageNumber="258">Yunnan</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="1" pageNumber="258" type="etymology">
|
||
<paragraph blockId="1.[151,1437,680,2039]" pageId="1" pageNumber="258">
|
||
<emphasis bold="true" box="[199,334,1976,2002]" pageId="1" pageNumber="258">Etymology.</emphasis>
|
||
The epithet is derived from the Latin adjective “spnaericus” = “globose”, and refers to the shape of the pronotum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |