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<document id="187F861437A186561443F3507229D1C4" ID-CLB-Dataset="4064" ID-DOI="10.11646/zootaxa.511.1.1" ID-GBIF-Dataset="d9f10a58-d4f7-4a74-900a-d5f2e041d181" ID-ISSN="1175­5334" ID-Zenodo-Dep="5030215" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624553966208" checkinUser="felipe" docAuthor="Holzenthal, Ralph W. &amp; Contents, Trond Andersen Table Of" docDate="2004" docId="D07687C5FF8AFFBA27320D235DAD7499" docLanguage="en" docName="zt00511.pdf" docOrigin="Zootaxa 511 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.511.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Triaenodes cuyotenango Holzenthal &amp; Contents 2004, new species" docType="treatment" docVersion="8" lastPageNumber="20" masterDocId="2C4FFFBDFF98FFA9263A0877597B7747" masterDocTitle="The caddisfly genus Triaenodes in the Neotropics (Trichoptera: Leptoceridae)" masterLastPageNumber="80" masterPageNumber="1" pageNumber="19" updateTime="1699020624569" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="7170260AD171DF36F977A9C5C6F3148D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D61E43EC3C0DB6E450578D958B2DC283">
<mods:title id="4554296C142C97EE82E985681065EB10">The caddisfly genus Triaenodes in the Neotropics (Trichoptera: Leptoceridae)</mods:title>
</mods:titleInfo>
<mods:name id="C7C1367180FE7E6BB680E3B343F4392F" type="personal">
<mods:role id="6400564A7C464BA0BB932AAD4C34D674">
<mods:roleTerm id="27F549BD3F28E74C5F8051EBD8D85AB9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ED89743251D61BA9DCF99E0F2C1654A4">Holzenthal, Ralph W.</mods:namePart>
<mods:affiliation id="7818A070A566096B52EC052C1FB78B6C">Department of Entomology, University of Minnesota, 1980 Folwell Ave., Room 219, St. Paul, Minnesota, 55108, U. S. A. (holze 001 @ umn. edu) Museum of Zoology, University of Bergen, Muséplass 3, N- 5007 Bergen, Norway (trond. andersen @ zmb. uib. no)</mods:affiliation>
</mods:name>
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<mods:role id="C16B7C61405B3166651C087294F78CBC">
<mods:roleTerm id="CDA4D6B352535AFD6447A2B79BF87619">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7F5CEA6AD06B79849D7EA383E815E516">Contents, Trond Andersen Table Of</mods:namePart>
</mods:name>
<mods:typeOfResource id="8559E7BA006506E704EFFA53C808D1C6">text</mods:typeOfResource>
<mods:relatedItem id="E7FC4260CB06CD7A2A6D2C1814CE1AB0" type="host">
<mods:titleInfo id="B86796D344111E1A3CB1673CD5DA18F9">
<mods:title id="87AECC27044391DF276FAA6655DE8A67">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7C7AC754B7E7AEF641669CEC5A7CCFE6">
<mods:date id="650C1AF23DD86876BAEDF2067E4BE392">2004</mods:date>
<mods:detail id="528BBC9F941ED93DE081722F61F3155E" type="pubDate">
<mods:number id="21FDEF9F985FCA305063104E864C91E9">2004-05-14</mods:number>
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<mods:detail id="D66B426D53E38C607556C28C26D0F313" type="volume">
<mods:number id="CEFBB33F23EC6FFAF3F730FF58C19E65">511</mods:number>
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<mods:detail id="B30ACDCB9080A9073B52AE8208D2815D" type="issue">
<mods:number id="6596DABC0D05C34B14965A612E3ED02A">1</mods:number>
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<mods:start id="1F23C4A422DDCF723172A1C202A7166D">1</mods:start>
<mods:end id="F06972383F8DDFCB81FC961F8660817B">80</mods:end>
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<mods:location id="1CA8E21BB3EE6CB43804B952F26C846B">
<mods:url id="AEE2199024F9C102696472B7FE073461">https://biotaxa.org/Zootaxa/article/view/zootaxa.511.1.1</mods:url>
</mods:location>
<mods:classification id="14ED3BEFFDE49E209778F86F8C4B6858">journal article</mods:classification>
<mods:identifier id="1B008173EC48A3E54783559341D63A22" type="CLB-Dataset">4064</mods:identifier>
<mods:identifier id="FDD3A4D0ADDB5B314E923F4564AB4657" type="DOI">10.11646/zootaxa.511.1.1</mods:identifier>
<mods:identifier id="9DEC4057E66DFEC4BA00BB834C99ADEF" type="GBIF-Dataset">d9f10a58-d4f7-4a74-900a-d5f2e041d181</mods:identifier>
<mods:identifier id="E5D229A92DF13E32790456E67791C076" type="ISSN">1175­5334</mods:identifier>
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<treatment id="D07687C5FF8AFFBA27320D235DAD7499" ID-DOI="http://doi.org/10.5281/zenodo.5560105" ID-GBIF-Taxon="188631171" ID-Zenodo-Dep="5560105" LSID="urn:lsid:plazi:treatment:D07687C5FF8AFFBA27320D235DAD7499" httpUri="http://treatment.plazi.org/id/D07687C5FF8AFFBA27320D235DAD7499" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<subSubSection id="10C56558FF8AFFBB27320D235BB47229" box="[264,719,1364,1390]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="586036D3FF8AFFBB27320D235BB47229" blockId="18.[264,719,1364,1425]" box="[264,719,1364,1390]" pageId="18" pageNumber="19">
<heading id="032881BFFF8AFFBB27320D235BB47229" bold="true" box="[264,719,1364,1390]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="6AABEAC1FF8AFFBB27320D235BB47229" bold="true" box="[264,719,1364,1390]" pageId="18" pageNumber="19">
<taxonomicName id="9FDF4D50FF8AFFBB27320D235B4B7229" authority="Holzenthal &amp; Contents, 2004" authorityName="Holzenthal &amp; Contents" authorityYear="2004" box="[264,560,1364,1390]" class="Insecta" family="Leptoceridae" genus="Triaenodes" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cuyotenango" status="sp. nov.">
<emphasis id="6AABEAC1FF8AFFBB27320D235B4B7229" bold="true" box="[264,560,1364,1390]" italics="true" pageId="18" pageNumber="19">Triaenodes cuyotenango</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="719857BAFF8AFFBB24050D235BB47229" box="[575,719,1364,1390]" pageId="18" pageNumber="19" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="10C56558FF8AFFBA27320D0058CC75A9" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
<paragraph id="586036D3FF8AFFBB27320D00582072D6" blockId="18.[264,719,1364,1425]" box="[264,347,1399,1425]" pageId="18" pageNumber="19">
<figureCitation id="C0E42A56FF8AFFBB27320D00582072D6" box="[264,347,1399,1425]" captionStart="FIGURE 10" captionStartId="51.[264,367,1444,1468]" captionTargetBox="[291,1259,318,1427]" captionTargetId="figure-38@51.[260,1327,305,1446]" captionTargetPageId="51" captionText="FIGURE 10. Triaenodes cuyotenango, new species. Male genitalia: A—lateral; B—dorsal; C— ventral; D—phallus, lateral, inset: apex of phalicata, ventral." figureDoi="http://doi.org/10.5281/zenodo.5030241" httpUri="https://zenodo.org/record/5030241/files/figure.png" pageId="18" pageNumber="19">Fig. 10</figureCitation>
</paragraph>
<paragraph id="586036D3FF8AFFBB27320DB05BA6711E" blockId="18.[264,1325,1479,1905]" pageId="18" pageNumber="19">
<taxonomicName id="9FDF4D50FF8AFFBB27320DB05B5372A7" authorityName="Holzenthal &amp; Contents" authorityYear="2004" box="[264,552,1479,1504]" class="Insecta" family="Leptoceridae" genus="Triaenodes" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cuyotenango">
<emphasis id="6AABEAC1FF8AFFBB27320DB05B5372A7" box="[264,552,1479,1504]" italics="true" pageId="18" pageNumber="19">Triaenodes cuyotenango</emphasis>
</taxonomicName>
and a number of other species have a digitate dorsomesal process of tergum X, with tergum X having a rounded or truncate apex (
<taxonomicName id="9FDF4D50FF8AFFBB25C90D985DF7714F" authorityName="Holzenthal &amp; Contents" authorityYear="2004" box="[1011,1164,1519,1545]" class="Insecta" family="Leptoceridae" genus="Triaenodes" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="6AABEAC1FF8AFFBB25C90D985D7A714F" box="[1011,1025,1519,1544]" italics="true" pageId="18" pageNumber="19">T</emphasis>
.
<emphasis id="6AABEAC1FF8AFFBB222A0D985DF7714F" box="[1040,1164,1519,1544]" italics="true" pageId="18" pageNumber="19">mexicanus</emphasis>
</taxonomicName>
,
<taxonomicName id="9FDF4D50FF8AFFBB22A00D985C58714F" authorityName="Flint &amp; Reyes." authorityYear="1991" box="[1178,1315,1519,1545]" class="Insecta" family="Leptoceridae" genus="Triaenodes" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="peruanus">
<emphasis id="6AABEAC1FF8AFFBB22A00D985DD3714F" box="[1178,1192,1519,1544]" italics="true" pageId="18" pageNumber="19">T</emphasis>
.
<emphasis id="6AABEAC1FF8AFFBB228C0D985C58714F" box="[1206,1315,1519,1544]" italics="true" pageId="18" pageNumber="19">peruanus</emphasis>
</taxonomicName>
,
<taxonomicName id="9FDF4D50FF8AFFBB27320E6058EF7177" authorityName="Holzenthal &amp; Contents" authorityYear="2004" box="[264,404,1559,1585]" class="Insecta" family="Leptoceridae" genus="Triaenodes" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="tuxtlensis">
<emphasis id="6AABEAC1FF8AFFBB27320E60586D7177" box="[264,278,1559,1584]" italics="true" pageId="18" pageNumber="19">T</emphasis>
.
<emphasis id="6AABEAC1FF8AFFBB271F0E6058EF7177" box="[293,404,1559,1584]" italics="true" pageId="18" pageNumber="19">tuxtlensis</emphasis>
</taxonomicName>
and variants of
<taxonomicName id="9FDF4D50FF8AFFBB24690E605BA17177" authorityName="Navas Dorsomesal" authorityYear="1924" box="[595,730,1559,1585]" class="Insecta" family="Leptoceridae" genus="Triaenodes" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis id="6AABEAC1FF8AFFBB24690E605B1A7177" box="[595,609,1559,1584]" italics="true" pageId="18" pageNumber="19">T</emphasis>
.
<emphasis id="6AABEAC1FF8AFFBB244A0E605BA17177" box="[624,730,1559,1584]" italics="true" pageId="18" pageNumber="19">delicatus</emphasis>
</taxonomicName>
). This new species can be separated from the others by its very long dorsomesal process.
</paragraph>
<paragraph id="586036D3FF8AFFBB27020E105AC171BE" blockId="18.[264,1325,1479,1905]" pageId="18" pageNumber="19">
Adult (male: n=2). Forewing length
<quantity id="9F279B36FF8AFFBB24D60E105A0471C6" box="[748,895,1639,1665]" metricMagnitude="-3" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.3" metricValueMin="5.6" pageId="18" pageNumber="19" unit="mm" value="5.949999999999999" valueMax="6.3" valueMin="5.6">5.6–6.3 mm</quantity>
, hind wing length
<quantity id="9F279B36FF8AFFBB225A0E105D8871C6" box="[1120,1267,1639,1665]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.2" metricValueMin="4.8" pageId="18" pageNumber="19" unit="mm" value="5.0" valueMax="5.2" valueMin="4.8">4.8–5.2 mm</quantity>
. Eye
<quantity id="9F279B36FF8AFFBB27320EF858C971EE" box="[264,434,1679,1705]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.3" metricValueMin="4.2" pageId="18" pageNumber="19" unit="mm" value="0.425" valueMax="0.43" valueMin="0.42">0.42–0.43 mm</quantity>
wide. Antennal scape
<quantity id="9F279B36FF8AFFBB24FB0EF85A1071EE" box="[705,875,1679,1705]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.699999999999999" metricValueMin="4.6000000000000005" pageId="18" pageNumber="19" unit="mm" value="0.46499999999999997" valueMax="0.47" valueMin="0.46">0.46–0.47 mm</quantity>
long. Maxillary palp segment lengths (in mm): 0.26–0.32, 0.35–0.39, 0.51–0.52, 0.27–0.29, 0.48–0.53. Forewing golden brown, with narrow line of cream colored hairs along anal margin.
</paragraph>
<paragraph id="586036D3FF8AFFBA27020F7058CC75A9" blockId="18.[264,1325,1479,1905]" lastBlockId="19.[264,1325,284,990]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">Male genitalia. Abdominal segment IX anterior margin slightly rounded, slightly produced anteroventrally; pleural region membranous, setose; tergum IX, in dorsal view, with posteromesal margin rounded, with pair of minute dorsomesal papillae; sternum IX in lateral view with posterior margin sinuate. Preanal appendage setose, about as long as wide, constricted basally, apex subacute. Dorsomesal process of tergum X digitate, originating basally, long, extending beyond apex of tergum X, with apical setae; tergum X, in dorsal view, with lateral margins subtriangular, apex slightly excavate medially; in lateral view, parallel sided, apex truncate, with lateral flange, apicolateral margins with microtrichia. Inferior appendage subquadrangular basally, setose, with long, tapering, apicoventral projection; recurved process gradually downturned, narrow, tapering throughout length, apex sharply attenuate; apicomesal lobe heavily setose, elongate, narrow, slightly broadened subbasally. Phallobase small, triangular, endothecal membranes prominent, highly convoluted, with pair of parameres, parameres subequal, long, gently curved, tapering; phalicata troughlike, straight, narrow throughout length, apex with shallow cleft, phallotremal sclerite not evident.</paragraph>
</subSubSection>
<subSubSection id="10C56558FF8BFFBA27020A8B5DAD7499" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="586036D3FF8BFFBA27020A8B5BAF7479" blockId="19.[264,1325,284,990]" pageId="19" pageNumber="20">
<materialsCitation id="E8B73C8EFF8BFFBA27020A8B5BAF7479" ID-GBIF-Occurrence="3383261314" collectingDate="1966-06-10" collectingDateMax="1966-06-20" collectingDateMin="1966-06-10" collectionCode="NMNH" collectorName="Flint &amp; Ortiz" country="Guatemala" location="Suchitepquez" municipality="Cuyotenango" pageId="19" pageNumber="20" specimenCode="UMSP000083650" specimenCount="1" typeStatus="holotype">
<emphasis id="6AABEAC1FF8BFFBA27020A8B5AFE7451" bold="true" box="[312,901,764,790]" pageId="19" pageNumber="20">
<typeStatus id="87648871FF8BFFBA27020A8B58D17451" box="[312,426,764,790]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="20C87643FF8BFFBA243B0A8B5BBE7451" box="[513,709,764,790]" name="Guatemala" pageId="19" pageNumber="20">GUATEMALA</collectingCountry>
:
<location id="5D006008FF8BFFBA24E90A8B5A047451" LSID="urn:lsid:plazi:treatment:D07687C5FF8AFFBA27320D235DAD7499:5D006008FF8BFFBA24E90A8B5A047451" box="[723,895,764,790]" country="Guatemala" municipality="Cuyotenango" name="Suchitepquez" pageId="19" pageNumber="20">Suchitepquez</location>
:
</emphasis>
<collectingMunicipality id="B804ACA9FF8BFFBA25B70A8B5D577451" box="[909,1068,764,790]" pageId="19" pageNumber="20">Cuyotenango</collectingMunicipality>
,
<date id="2C611013FF8BFFBA22000A8B5D9D7451" box="[1082,1254,764,790]" pageId="19" pageNumber="20" value="1966-06-10" valueMax="1966-06-20" valueMin="1966-06-10">
<collectingDate id="3C25E9FBFF8BFFBA22000A8B5D9D7451" box="[1082,1254,764,790]" pageId="19" pageNumber="20" value="1966-06-10" valueMax="1966-06-20" valueMin="1966-06-10">10–20.vi.1966</collectingDate>
</date>
,
<collectorName id="F52A5305FF8BFFBA22C90A8B5C517451" box="[1267,1322,764,790]" pageId="19" pageNumber="20">Flint</collectorName>
&amp;
<collectorName id="F52A5305FF8BFFBA271C0B5358197479" box="[294,354,804,830]" pageId="19" pageNumber="20">Ortiz</collectorName>
(
<specimenCode id="08799EA8FF8BFFBA27430B535B2B7479" box="[377,592,804,830]" collectionCode="UMSP" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="19" pageNumber="20" type="University or college">UMSP000083650</specimenCode>
) (
<collectionCode id="3ECEAE16FF8BFFBA245D0B535BBC7479" box="[615,711,804,830]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="20" type="Museum">NMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="586036D3FF8BFFBA27020B3B5B1C74C9" blockId="19.[264,1325,284,990]" pageId="19" pageNumber="20">
<materialsCitation id="E8B73C8EFF8BFFBA27020B3B5B1874C9" ID-GBIF-Occurrence="3383261344" collectingDate="1965-03-19" collectingDateMax="1965-03-20" collectingDateMin="1965-03-19" collectionCode="NMNH" collectorName="P J Spangler" country="Guatemala" location="Chimaltenango" pageId="19" pageNumber="20" specimenCount="1" specimenCount-male="1" stateProvince="Chimaltenango" typeStatus="paratype">
<emphasis id="6AABEAC1FF8BFFBA27020B3B5A1E7421" bold="true" box="[312,869,844,870]" pageId="19" pageNumber="20">
<typeStatus id="87648871FF8BFFBA27020B3B58D57421" box="[312,430,844,870]" pageId="19" pageNumber="20" type="paratype">Paratype</typeStatus>
:
<collectingCountry id="20C87643FF8BFFBA27F90B3B5BFC7421" box="[451,647,844,870]" name="Guatemala" pageId="19" pageNumber="20">GUATEMALA</collectingCountry>
:
<collectingRegion id="9A1BF831FF8BFFBA24A10B3B5A1B7421" box="[667,864,844,870]" country="Guatemala" name="Chimaltenango" pageId="19" pageNumber="20">Chimaltenango</collectingRegion>
:
</emphasis>
<location id="5D006008FF8BFFBA254E0B3B5D517421" LSID="urn:lsid:plazi:treatment:D07687C5FF8AFFBA27320D235DAD7499:5D006008FF8BFFBA254E0B3B5D517421" box="[884,1066,844,870]" country="Guatemala" name="Chimaltenango" pageId="19" pageNumber="20" stateProvince="Chimaltenango">Chimaltenango</location>
,
<date id="2C611013FF8BFFBA22040B3B5D977421" box="[1086,1260,844,870]" pageId="19" pageNumber="20" value="1965-03-19" valueMax="1965-03-20" valueMin="1965-03-19">
<collectingDate id="3C25E9FBFF8BFFBA22040B3B5D977421" box="[1086,1260,844,870]" pageId="19" pageNumber="20" value="1965-03-19" valueMax="1965-03-20" valueMin="1965-03-19">19–20.iii.1965</collectingDate>
</date>
,
<collectorName id="F52A5305FF8BFFBA22C50B3B580B74C9" pageId="19" pageNumber="20">P J Spangler</collectorName>
—
<specimenCount id="4ED9FD5AFF8BFFBA27A60B03589074C9" box="[412,491,884,910]" pageId="19" pageNumber="20" type="male">1 male</specimenCount>
(
<collectionCode id="3ECEAE16FF8BFFBA27C00B035B2174C9" box="[506,602,884,910]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="20" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="586036D3FF8BFFBA27020BEB5B2074F1" blockId="19.[264,1325,284,990]" box="[312,603,924,950]" pageId="19" pageNumber="20">
Distribution.
<collectingCountry id="20C87643FF8BFFBA27EE0BEB5B2C74F1" box="[468,599,924,950]" name="Guatemala" pageId="19" pageNumber="20">Guatemala</collectingCountry>
.
</paragraph>
<paragraph id="586036D3FF8BFFBA27020BB35DAD7499" blockId="19.[264,1325,284,990]" box="[312,1238,964,990]" pageId="19" pageNumber="20">
Etymology. Named for the
<typeStatus id="87648871FF8BFFBA24420BB35BD17499" box="[632,682,964,990]" pageId="19" pageNumber="20">type</typeStatus>
locality, the town of Cuyotenango,
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.
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