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<document ID-DOI="10.35929/RSZ.0049" ID-GBIF-Dataset="172fa5c5-c0c4-4bd7-b710-d608237b8458" ID-ISSN="0035-418" ID-Zenodo-Dep="5639938" checkinTime="1635892563566" checkinUser="felipe" docAuthor="Schuchert, Peter &amp; Collins, Richard" docDate="2021" docId="D0118A7C5B1E0006FF38F9B5FAD87F00" docLanguage="en" docName="RevSuisseZool.128.2.237-356.pdf.imf" docOrigin="Revue suisse de Zoologie 128 (2)" docStyle="DocumentStyle:EA5207814960044D272DF5BE47DFF256.2:RevSuisseZool.2020-.journal_article.1cover" docStyleId="EA5207814960044D272DF5BE47DFF256" docStyleName="RevSuisseZool.2020-.journal_article.1cover" docStyleVersion="2" docTitle="Larsonia pterophylla" docType="treatment" docVersion="6" lastPageNumber="254" masterDocId="2C28F2045B0C0014FFCFFF96FFDB7820" masterDocTitle="Hydromedusae observed during night dives in the Gulf Stream" masterLastPageNumber="356" masterPageNumber="237" pageNumber="254" updateTime="1683579073239" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>Hydromedusae observed during night dives in the Gulf Stream</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schuchert, Peter</mods:namePart>
<mods:affiliation>Muséum dhistoire naturelle, C. P. 6434, CH- 1211 Genève 6, Switzerland</mods:affiliation>
<mods:nameIdentifier type="email">peter.schuchert@ville-ge.ch</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Collins, Richard</mods:namePart>
<mods:affiliation>880 NE 33 rd Street, Boca Raton, Florida, USA</mods:affiliation>
<mods:nameIdentifier type="email">rc6684@icloud.com</mods:nameIdentifier>
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<mods:title>Revue suisse de Zoologie</mods:title>
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<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-10-21</mods:number>
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<mods:detail type="volume">
<mods:number>128</mods:number>
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<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:identifier type="ISSN">0035-418</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5639986" ID-GBIF-Taxon="190510857" ID-Zenodo-Dep="5639986" LSID="urn:lsid:plazi:treatment:D0118A7C5B1E0006FF38F9B5FAD87F00" httpUri="http://treatment.plazi.org/id/D0118A7C5B1E0006FF38F9B5FAD87F00" lastPageNumber="254" pageId="18" pageNumber="254">
<subSubSection box="[247,660,1571,1595]" pageId="18" pageNumber="254" type="nomenclature">
<paragraph blockId="18.[247,660,1571,1627]" box="[247,660,1571,1595]" pageId="18" pageNumber="254">
<heading bold="true" box="[247,660,1571,1595]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="254" reason="2">
<taxonomicName ID-CoL="3SBNX" ID-ENA="498529" authority="(Haeckel, 1879)" baseAuthorityName="Haeckel" baseAuthorityYear="1879" box="[247,660,1571,1595]" class="Hydrozoa" family="Pandeidae" genus="Larsonia" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="pterophylla">
<emphasis bold="true" box="[247,660,1571,1595]" pageId="18" pageNumber="254">
<emphasis bold="true" box="[247,477,1571,1595]" italics="true" pageId="18" pageNumber="254">Larsonia pterophylla</emphasis>
(
<bibRefCitation author="Haeckel" box="[491,653,1571,1595]" firstAuthor="Haeckel" pageId="18" pageNumber="254" pagination="1 - 360" refId="ref68751" refString="Haeckel E. 1879. Das System der Medusen. Erster Teil einer Monographie der Medusen. Denkschriften der Medicinisch- Naturwissenschaftlichen Gesellschaft zu Jena 1: I-XX, 1 - 360, 20 pls." type="book chapter" year="1879">Haeckel, 1879</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[392,515,1603,1627]" pageId="18" pageNumber="254" type="description">
<paragraph blockId="18.[247,660,1571,1627]" box="[392,515,1603,1627]" pageId="18" pageNumber="254">
<figureCitation box="[392,466,1603,1627]" captionStart="Fig" captionStartId="19.[151,186,1535,1557]" captionTargetBox="[151,1436,257,1482]" captionTargetId="figure-160@19.[151,1436,257,1482]" captionTargetPageId="19" captionText="Fig. 10. Larsonia pterophylla. (A) BFLA4453, lateral view of animal, size 27 mm. (B) BFLA4453, close up of manubrium in lateral view. (C) Animal photographed 08-AUG-20181 part of bell margin in aboral view showing rudimentary tentacle bulbs (warts)." figureDoi="http://doi.org/10.5281/zenodo.5639964" httpUri="https://zenodo.org/record/5639964/files/figure.png" pageId="18" pageNumber="254">Fig. 10</figureCitation>
A-D
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="254" type="reference_group">
<paragraph blockId="18.[151,757,1649,2077]" pageId="18" pageNumber="254">
<treatmentCitationGroup pageId="18" pageNumber="254">
<treatmentCitation author="Haeckel E." box="[151,752,1649,1671]" page="52" pageId="18" pageNumber="254" year="1879">
<taxonomicName ID-CoL="6ZWVR" authority="Haeckel, 1879: 52" authorityName="Haeckel" authorityPageNumber="52" authorityYear="1879" box="[151,752,1649,1671]" class="Hydrozoa" family="Pandeidae" genus="Stomotoca" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="pterophylla" subGenus="Stomotocanna">
<emphasis box="[151,544,1649,1670]" italics="true" pageId="18" pageNumber="254">Stomotoca (Stomotocanna) pterophylla</emphasis>
<bibRefCitation author="Haeckel" box="[558,752,1649,1671]" firstAuthor="Haeckel" page="52" pageId="18" pageNumber="254" pagination="1 - 360" refId="ref68751" refString="Haeckel E. 1879. Das System der Medusen. Erster Teil einer Monographie der Medusen. Denkschriften der Medicinisch- Naturwissenschaftlichen Gesellschaft zu Jena 1: I-XX, 1 - 360, 20 pls." type="book chapter" year="1879">Haeckel, 1879: 52</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 4 fig. 10.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="18.[151,757,1649,2077]" pageId="18" pageNumber="254">
<treatmentCitationGroup pageId="18" pageNumber="254">
<taxonomicName ID-CoL="6ZWVR" authorityName="Haeckel" authorityYear="1879" box="[151,367,1707,1728]" class="Hydrozoa" family="Pandeidae" genus="Stomotoca" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="pterophylla">
<emphasis box="[151,367,1707,1728]" italics="true" pageId="18" pageNumber="254">Stomotoca pterophylla</emphasis>
</taxonomicName>
.
<treatmentCitation author="Mayer A. G." box="[389,556,1707,1728]" page="113" pageId="18" pageNumber="254" year="1910">
<bibRefCitation author="Mayer" box="[389,556,1707,1728]" firstAuthor="Mayer" page="113" pageId="18" pageNumber="254" pagination="1 - 76" refId="ref71665" refString="Mayer A. G. 1910. Medusae of the world. Hydromedusae, Vols. I &amp; II. Scyphomedusae, Vol III. Carnegie Institution, Washington, pp. 735, plates 1 - 76." type="journal article" year="1910">Mayer, 1910: 113</bibRefCitation>
</treatmentCitation>
, pl. 29 figs 3-5, pl. 30 fig. 7.
<treatmentCitation author="Bigelow H. B." box="[311,505,1736,1758]" page="372" pageId="18" pageNumber="254" year="1918">
<bibRefCitation author="Bigelow" box="[311,505,1736,1758]" firstAuthor="Bigelow" page="372" pageId="18" pageNumber="254" pagination="363 - 442" refId="ref65028" refString="Bigelow H. B. 1918. Some Medusae and Siphonophora from the western Atlantic. Bulletin of the Museum of comparative Zoology of Harvard College 62: 363 - 442, pls 1 - 8." type="journal article" year="1918">Bigelow, 1918: 372</bibRefCitation>
</treatmentCitation>
, synonymy, discussion of species.
<treatmentCitation author="Uchida T." box="[330,513,1765,1787]" page="284" pageId="18" pageNumber="254" year="1940">
<bibRefCitation author="Uchida" box="[330,513,1765,1787]" firstAuthor="Uchida" page="284" pageId="18" pageNumber="254" pagination="277 - 297" refId="ref75662" refString="Uchida T. 1940. The Fauna of Akkesi Bay. XI. Medusae. Journal of the Faculty of Science, Hokkaido Imperial University Series VI. Zoology 7 (3): 277 - 297." type="journal article" year="1940">Uchida, 1940: 284</bibRefCitation>
</treatmentCitation>
, fig. 3.
<treatmentCitation author="Kramp P. L." page="119" pageId="18" pageNumber="254" year="1959">
<bibRefCitation author="Kramp" firstAuthor="Kramp" page="119" pageId="18" pageNumber="254" pagination="1 - 283" refId="ref69812" refString="Kramp P. L. 1959 a. The Hydromedusae of the Atlantic Ocean and adjacent waters. Dana Report 46: 1 - 283." type="book chapter" year="1959" yearSuffix="a">Kramp, 1959a: 119</bibRefCitation>
</treatmentCitation>
, fig. 115.
<treatmentCitation author="Kramp P. L." box="[379,557,1794,1816]" page="115" pageId="18" pageNumber="254" year="1961">
<bibRefCitation author="Kramp" box="[379,557,1794,1816]" firstAuthor="Kramp" page="115" pageId="18" pageNumber="254" pagination="1 - 469" refId="ref69868" refString="Kramp P. L. 1961. Synopsis of the medusae of the world. Journal of the Marine Biological Association of the U. K. 40: 1 - 469." type="journal article" year="1961">Kramp, 1961: 115</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation author="Kramp P. L." box="[585,752,1794,1816]" page="44" pageId="18" pageNumber="254" year="1968">
<bibRefCitation author="Kramp" box="[585,752,1794,1816]" firstAuthor="Kramp" page="44" pageId="18" pageNumber="254" pagination="1 - 200" refId="ref69950" refString="Kramp P. L. 1968. The hydromedusae of the Pacific and Indian oceans. Sections II and III. Dana Report 72: 1 - 200." type="book chapter" year="1968">Kramp, 1968: 44</bibRefCitation>
</treatmentCitation>
, fig. 113.
<treatmentCitation author="Larson R. J." box="[342,528,1823,1844]" page="433" pageId="18" pageNumber="254" year="1982">
<bibRefCitation author="Larson" box="[342,528,1823,1844]" firstAuthor="Larson" page="433" pageId="18" pageNumber="254" pagination="433 - 439" refId="ref70347" refString="Larson R. J. 1982. Life history of the hydromedusa Stomotoca pterophylla Haeckel and its ichtyoparasitic hydroid. Smithsonian contributions to the marine sciences 12: 433 - 439." type="journal article" year="1982">Larson, 1982: 433</bibRefCitation>
</treatmentCitation>
, fig. 183.
<treatmentCitation author="Wedler E. &amp; Larson R." page="97" pageId="18" pageNumber="254" year="1986">
<bibRefCitation author="Wedler &amp; Larson" firstAuthor="Wedler" page="97" pageId="18" pageNumber="254" pagination="69 - 101" refId="ref76329" refString="Wedler E., Larson R. 1986. Athecate hydroids from Puerto Rico and the Virgin Islands. Studies on Neotropical Fauna and Environment 21 (1 - 2): 69 - 101." type="journal article" year="1986">Wedler &amp; Larson, 1986: 97</bibRefCitation>
</treatmentCitation>
, fig. 11a-b, hydroid, young medusa.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="18.[151,757,1649,2077]" pageId="18" pageNumber="254">
<treatmentCitationGroup pageId="18" pageNumber="254">
<treatmentCitation author="Maas O." box="[151,479,1881,1902]" page="11" pageId="18" pageNumber="254" year="1897">
<taxonomicName ID-CoL="6ZX7N" authority="Maas, 1897: 11" authorityName="Maas" authorityPageNumber="11" authorityYear="1897" box="[151,479,1881,1902]" class="Hydrozoa" family="Pandeidae" genus="Stomotoca" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="divisa">
<emphasis box="[151,317,1881,1902]" italics="true" pageId="18" pageNumber="254">Stomotoca divisa</emphasis>
<bibRefCitation author="Maas" box="[325,479,1881,1902]" firstAuthor="Maas" page="11" pageId="18" pageNumber="254" pagination="1 - 92" refId="ref70983" refString="Maas O. 1897. Die Medusen. 21 st Reports on an exploration off the West Coast of Mexico, Central and South America and of the Galapagos Islands, in charge of A. Agassiz by the U. S. Fish Commission Steamer &quot; Albatross &quot; during 1891. Memoirs of the Museum of Comparative Zoology at Harvard College 23 (1): 1 - 92, pls 1 - 15." type="journal article" year="1897">Maas, 1897: 11</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 1 figs 1-9.
<treatmentCitation author="Vanhoffen E." page="14" pageId="18" pageNumber="254" year="1913">
<bibRefCitation author="Vanhoffen" firstAuthor="Vanhoffen" page="14" pageId="18" pageNumber="254" pagination="1 - 34" refId="ref75972" refString="Vanhoffen E. 1913 b. Die Craspedoten Medusen der &quot; Vettor Pisani &quot;. Zoologica, Stuttart 26 (67): 1 - 34." type="journal article" year="1913" yearSuffix="b">Vanhöffen, 1913b: 14</bibRefCitation>
</treatmentCitation>
, synonym.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="18.[151,757,1649,2077]" pageId="18" pageNumber="254">
<treatmentCitationGroup pageId="18" pageNumber="254">
<emphasis box="[151,402,1939,1961]" italics="true" pageId="18" pageNumber="254">
<taxonomicName ID-CoL="3SBNX" baseAuthorityName="Haeckel" baseAuthorityYear="1879" box="[151,369,1939,1961]" class="Hydrozoa" family="Pandeidae" genus="Larsonia" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="pterophylla">Larsonia pterophylla</taxonomicName>
.
</emphasis>
<treatmentCitation author="Boero F. &amp; Bouillon J. &amp; Gravili C." box="[421,694,1939,1961]" page="198" pageId="18" pageNumber="254" year="1991">
<bibRefCitation author="Boero" box="[421,694,1939,1961]" etAl="et al." firstAuthor="Boero" page="198" pageId="18" pageNumber="254" pagination="189 - 199" refId="ref65317" refString="Boero F., Bouillon J., Gravili C. 1991. The life cycle of Hydrichthys mirus (Cnidaria: Hydrozoa: Anthomedusae: Pandeidae). Zoological Journal of the Linnean Society 101 (2): 189 - 199." type="journal article" year="1991">
Boero
<emphasis box="[498,552,1939,1961]" italics="true" pageId="18" pageNumber="254">et al</emphasis>
., 1991: 198
</bibRefCitation>
</treatmentCitation>
, new combination.
<bibRefCitation author="Woodstock" box="[376,602,1968,1990]" etAl="et al." firstAuthor="Woodstock" pageId="18" pageNumber="254" pagination="10" refId="ref76493" refString="Woodstock M. S., Golightly C., Fenolio D., Moore J. A. 2019. Larsonia pterophylla (Cnidaria, Pandeidae) Parasitic on Two Leptocephali: Paraconger sp. (Congridae) and Callechelyini gen. sp. (Ophichthidae) in the Gulf of Mexico. Gulf and Caribbean Research 3 (1): SC 7 - SC 10. DOI: 10.18785 / gcr. 3001.05" type="journal article" year="2019">
Woodstock
<emphasis box="[490,543,1968,1990]" italics="true" pageId="18" pageNumber="254">et al.,</emphasis>
2019
</bibRefCitation>
: fig. 1, hydroid.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="18.[151,757,1649,2077]" pageId="18" pageNumber="254">
<treatmentCitationGroup pageId="18" pageNumber="254">
<taxonomicName ID-CoL="6ZX7G" authorityName="L. Agassiz" authorityYear="1862" box="[151,302,1997,2019]" class="Hydrozoa" family="Pandeidae" genus="Stomotoca" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="atra">
<emphasis box="[151,302,1997,2019]" italics="true" pageId="18" pageNumber="254">Stomotoca atra</emphasis>
</taxonomicName>
.
<treatmentCitation author="Vanhoffen E." box="[340,557,1997,2019]" page="14" pageId="18" pageNumber="254" year="1913">
<bibRefCitation author="Vanhoffen" box="[340,557,1997,2019]" firstAuthor="Vanhoffen" page="14" pageId="18" pageNumber="254" pagination="1 - 34" refId="ref75972" refString="Vanhoffen E. 1913 b. Die Craspedoten Medusen der &quot; Vettor Pisani &quot;. Zoologica, Stuttart 26 (67): 1 - 34." type="journal article" year="1913" yearSuffix="b">Vanhöffen, 1913b: 14</bibRefCitation>
</treatmentCitation>
, pl. 2 figs 12-16.
<treatmentCitation author="Stampar S. N. &amp; Kodja G." box="[227,505,2026,2048]" page="55" pageId="18" pageNumber="254" year="2007">
<bibRefCitation author="Stampar &amp; Kodja" box="[227,505,2026,2048]" firstAuthor="Stampar" page="55" pageId="18" pageNumber="254" pagination="55 - 57" refId="ref75086" refString="Stampar S. N., Kodja G. 2007. Cnidaria, Hydrozoa, Anthoathecata, Pandeidae, Stomotoca atra: distribution extension. Check List 3 (1): 55 - 57." type="journal article" year="2007">Stampar &amp; Kodja, 2007: 55</bibRefCitation>
</treatmentCitation>
, figs 2-3. [not
<taxonomicName authority="L. Agassiz, 1862" authorityName="L. Agassiz" authorityYear="1862" class="Hydrozoa" family="Pandeidae" genus="Stomotoca" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="atra">
<emphasis italics="true" pageId="18" pageNumber="254">Stomotoca atra</emphasis>
L.
<bibRefCitation author="Agassiz A." box="[298,436,2054,2077]" pageId="18" pageNumber="254" refId="ref64002" refString="Agassiz A. 1862. In: Agassiz L. 1862." title="Agassiz L" type="book chapter" year="1862">Agassiz, 1862</bibRefCitation>
</taxonomicName>
]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="254" type="materials_examined">
<paragraph blockId="18.[831,1437,257,409]" pageId="18" pageNumber="254">
<emphasis bold="true" box="[831,1086,257,281]" pageId="18" pageNumber="254">Examined material:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3410574477" collectingDate="2020-06-08" collectionCode="BFLA, FU" pageId="18" pageNumber="254" specimenCode="BFLA4453, FU-014064" specimenCount="1">
<specimenCode box="[1114,1242,257,281]" collectionCode="BFLA" pageId="18" pageNumber="254">BFLA4453</specimenCode>
; one formalin fixed specimen deposited under catalogue number
<specimenCode box="[831,960,321,345]" pageId="18" pageNumber="254">FU-014064</specimenCode>
; collected
<date box="[1091,1244,321,345]" pageId="18" pageNumber="254" value="2020-06-08">
<collectingDate box="[1091,1244,321,345]" pageId="18" pageNumber="254" value="2020-06-08">08-JUN-2020</collectingDate>
</date>
; height
<quantity box="[1345,1432,321,345]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="18" pageNumber="254" unit="mm" value="27.0">27 mm</quantity>
.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3410574446" collectingDate="2018-08-08" pageId="18" pageNumber="254" specimenCount="1">
<date box="[868,1030,353,377]" pageId="18" pageNumber="254" value="2018-08-08">
<collectingDate box="[868,1030,353,377]" pageId="18" pageNumber="254" value="2018-08-08">08-AUG-2018</collectingDate>
</date>
,
<specimenCount box="[1057,1197,353,377]" pageId="18" pageNumber="254" type="generic">1 specimen</specimenCount>
photographed; not collected...
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="254" type="discussion">
<paragraph blockId="18.[831,1437,432,902]" pageId="18" pageNumber="254">
<emphasis bold="true" box="[831,994,432,456]" pageId="18" pageNumber="254">Observations:</emphasis>
Medusa
<quantity box="[1112,1200,432,456]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="18" pageNumber="254" unit="mm" value="27.0">27 mm</quantity>
, with low umbrella and big, conical, pointed apical process; subumbrella shallow; with broad gastric peduncle reaching to velum level; 4-5 ribbon-like radial canals; two opposite tentacles, long, whitish, with swollen base clasping bell margin, no ocelli; about 80-100 atentaculate bulbs along bell margin, small, wart-like, all about the same size, without ocelli (
<figureCitation box="[1083,1192,655,679]" captionStart="Fig" captionStartId="19.[151,186,1535,1557]" captionTargetBox="[151,1436,257,1482]" captionTargetId="figure-160@19.[151,1436,257,1482]" captionTargetPageId="19" captionText="Fig. 10. Larsonia pterophylla. (A) BFLA4453, lateral view of animal, size 27 mm. (B) BFLA4453, close up of manubrium in lateral view. (C) Animal photographed 08-AUG-20181 part of bell margin in aboral view showing rudimentary tentacle bulbs (warts)." figureDoi="http://doi.org/10.5281/zenodo.5639964" httpUri="https://zenodo.org/record/5639964/files/figure.png" pageId="18" pageNumber="254">Fig. 10C</figureCitation>
); manubrium barrelshaped, with mouth drawn out into four long perradial lips, margin crenulated; gonads large, covering stomach in eight adradial series of about 10 branched folds directed towards interradial, pairs of adradial fold series connected perradially by a thick, vertical fold; adradial folds branching with up to four ends (
<figureCitation box="[1237,1334,846,870]" captionStart="Fig" captionStartId="19.[151,186,1535,1557]" captionTargetBox="[151,1436,257,1482]" captionTargetId="figure-160@19.[151,1436,257,1482]" captionTargetPageId="19" captionText="Fig. 10. Larsonia pterophylla. (A) BFLA4453, lateral view of animal, size 27 mm. (B) BFLA4453, close up of manubrium in lateral view. (C) Animal photographed 08-AUG-20181 part of bell margin in aboral view showing rudimentary tentacle bulbs (warts)." figureDoi="http://doi.org/10.5281/zenodo.5639964" httpUri="https://zenodo.org/record/5639964/files/figure.png" pageId="18" pageNumber="254">Fig. 10B</figureCitation>
). Gonads and manubrium colour golden-brown or brown.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="254" type="distribution">
<paragraph blockId="18.[831,1437,924,1108]" pageId="18" pageNumber="254">
<emphasis bold="true" box="[831,981,924,948]" pageId="18" pageNumber="254">Distribution:</emphasis>
American coasts from Gulf of Maine to
<collectingCountry box="[831,899,957,981]" name="Brazil" pageId="18" pageNumber="254">Brazil</collectingCountry>
, West Africa, Pacific coasts of
<collectingCountry box="[1254,1339,957,980]" name="Panama" pageId="18" pageNumber="254">Panama</collectingCountry>
to
<collectingCountry box="[1381,1433,957,980]" name="Peru" pageId="18" pageNumber="254">Peru</collectingCountry>
,
<collectingCountry box="[831,895,989,1012]" name="Japan" pageId="18" pageNumber="254">Japan</collectingCountry>
,
<collectingCountry box="[918,1151,988,1012]" name="Papua New Guinea" pageId="18" pageNumber="254">Papua New Guinea</collectingCountry>
, Indian Ocean (
<bibRefCitation author="Kramp" firstAuthor="Kramp" pageId="18" pageNumber="254" pagination="1 - 200" refId="ref69950" refString="Kramp P. L. 1968. The hydromedusae of the Pacific and Indian oceans. Sections II and III. Dana Report 72: 1 - 200." type="book chapter" year="1968">Kramp, 1968</bibRefCitation>
;
<bibRefCitation author="Bouillon" box="[900,1064,1020,1044]" firstAuthor="Bouillon" pageId="18" pageNumber="254" pagination="307 - 344" refId="ref65707" refString="Bouillon J. 1980. Hydromeduses de la Mer de Bismarck. (Papouasie Nouvelle-Guinee). Partie 3: Anthomedusae - Filifera (Hydrozoa - Cnidaria). Cahiers de Biologie Marine 21 (3): 307 - 344." type="journal article" year="1980">Bouillon, 1980</bibRefCitation>
;
<bibRefCitation author="Navas-Pereira &amp; Vannucci" box="[1076,1431,1020,1044]" firstAuthor="Navas-Pereira" pageId="18" pageNumber="254" pagination="25 - 60" refId="ref72799" refString="Navas-Pereira D., Vannucci M. 1991. The Hydromedusae and water masses of the Indian Ocean. Boletim do Instituto Oceanografico 39 (1): 25 - 60." type="journal article" year="1991">Navas-Pereira &amp; Vannucci, 1991</bibRefCitation>
;
<bibRefCitation author="Stampar &amp; Kodja" box="[831,1119,1052,1076]" firstAuthor="Stampar" pageId="18" pageNumber="254" pagination="55 - 57" refId="ref75086" refString="Stampar S. N., Kodja G. 2007. Cnidaria, Hydrozoa, Anthoathecata, Pandeidae, Stomotoca atra: distribution extension. Check List 3 (1): 55 - 57." type="journal article" year="2007">Stampar &amp; Kodja, 2007</bibRefCitation>
, as
<taxonomicName authorityName="L. Agassiz" authorityYear="1862" box="[1177,1352,1053,1076]" class="Hydrozoa" family="Pandeidae" genus="Stomotoca" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="atra">
<emphasis box="[1177,1352,1053,1076]" italics="true" pageId="18" pageNumber="254">Stomotoca atra</emphasis>
</taxonomicName>
).
<typeStatus box="[1383,1436,1053,1077]" pageId="18" pageNumber="254">Type</typeStatus>
Locality: Caribbean Sea,
<geoCoordinate box="[1103,1193,1084,1108]" degrees="20.60" direction="north" orientation="latitude" pageId="18" pageNumber="254" precision="555" value="20.6">20.60°N</geoCoordinate>
<geoCoordinate box="[1199,1296,1084,1108]" degrees="79.00" direction="west" orientation="longitude" pageId="18" pageNumber="254" precision="555" value="-79.0">79.00°W</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="254" type="discussion">
<paragraph blockId="18.[831,1437,1131,1824]" pageId="18" pageNumber="254">
<emphasis bold="true" box="[831,945,1131,1155]" pageId="18" pageNumber="254">Remarks:</emphasis>
The observed specimens agreed well with existing descriptions of
<taxonomicName baseAuthorityName="Haeckel" baseAuthorityYear="1879" box="[1111,1345,1163,1187]" class="Hydrozoa" family="Pandeidae" genus="Larsonia" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="pterophylla">
<emphasis box="[1111,1345,1163,1187]" italics="true" pageId="18" pageNumber="254">Larsonia pterophylla</emphasis>
</taxonomicName>
, except that one had five radial canals instead of four. This surplus radial canal was interpreted as a developmental aberration or variation without taxonomic significance. Also other tetraradial hydromedusa species occasionally show three or five radial canals (see
<taxonomicName box="[1220,1350,1322,1346]" class="Hydrozoa" family="Campanulariidae" genus="Clytia" kingdom="Animalia" order="Leptothecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="undetermined">
<emphasis box="[1220,1285,1322,1346]" italics="true" pageId="18" pageNumber="254">Clytia</emphasis>
spec.
</taxonomicName>
).
</paragraph>
<paragraph blockId="18.[831,1437,1131,1824]" pageId="18" pageNumber="254">
<taxonomicName authority="L. Agassiz, 1862" authorityName="L. Agassiz" authorityYear="1862" box="[831,1219,1354,1378]" class="Hydrozoa" family="Pandeidae" genus="Stomotoca" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="atra">
<emphasis box="[831,1004,1354,1378]" italics="true" pageId="18" pageNumber="254">Stomotoca atra</emphasis>
L.
<bibRefCitation author="Agassiz A." box="[1059,1219,1354,1378]" pageId="18" pageNumber="254" refId="ref64002" refString="Agassiz A. 1862. In: Agassiz L. 1862." title="Agassiz L" type="book chapter" year="1862">Agassiz, 1862</bibRefCitation>
</taxonomicName>
is a very similar species and
<bibRefCitation author="Vanhoffen" box="[989,1205,1386,1410]" firstAuthor="Vanhoffen" pageId="18" pageNumber="254" pagination="1 - 34" refId="ref75972" refString="Vanhoffen E. 1913 b. Die Craspedoten Medusen der &quot; Vettor Pisani &quot;. Zoologica, Stuttart 26 (67): 1 - 34." type="journal article" year="1913" yearSuffix="b">Vanhöffen (1913b)</bibRefCitation>
thought that they could be conspecific. However,
<taxonomicName authorityName="L. Agassiz" authorityYear="1862" box="[1185,1258,1418,1442]" class="Hydrozoa" family="Pandeidae" genus="Stomotoca" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="atra">
<emphasis box="[1185,1258,1418,1442]" italics="true" pageId="18" pageNumber="254">S. atra</emphasis>
</taxonomicName>
is distinct from
<taxonomicName baseAuthorityName="Haeckel" baseAuthorityYear="1879" box="[831,984,1450,1474]" class="Hydrozoa" family="Pandeidae" genus="Larsonia" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="pterophylla">
<emphasis box="[831,984,1450,1474]" italics="true" pageId="18" pageNumber="254">L. pterophylla</emphasis>
</taxonomicName>
, notably they have different polyp stages (comp.
<bibRefCitation author="Larson" box="[914,1055,1482,1506]" firstAuthor="Larson" pageId="18" pageNumber="254" pagination="433 - 439" refId="ref70347" refString="Larson R. J. 1982. Life history of the hydromedusa Stomotoca pterophylla Haeckel and its ichtyoparasitic hydroid. Smithsonian contributions to the marine sciences 12: 433 - 439." type="journal article" year="1982">Larson, 1982</bibRefCitation>
and
<bibRefCitation author="Boero &amp; Bouillon" box="[1110,1376,1482,1506]" firstAuthor="Boero" pageId="18" pageNumber="254" pagination="1 - 7" refId="ref65280" refString="Boero F., Bouillon J. 1989. The life cycles of Octotiara russelli and Stomotoca atra (Cnidaria, Anthomedusae, Pandeidae). Zoologica Scripta 18 (1): 1 - 7." type="journal article" year="1989">Boero &amp; Bouillon, 1989</bibRefCitation>
). The medusae are distinguishable: in
<taxonomicName baseAuthorityName="Haeckel" baseAuthorityYear="1879" box="[1173,1323,1513,1537]" class="Hydrozoa" family="Pandeidae" genus="Larsonia" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="pterophylla">
<emphasis box="[1173,1323,1513,1537]" italics="true" pageId="18" pageNumber="254">L. pterophylla</emphasis>
</taxonomicName>
the mouth margin is crenulated and perradially drawn out into long lips, in
<taxonomicName authorityName="L. Agassiz" authorityYear="1862" box="[914,987,1578,1601]" class="Hydrozoa" family="Pandeidae" genus="Stomotoca" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="atra">
<emphasis box="[914,987,1578,1601]" italics="true" pageId="18" pageNumber="254">S. atra</emphasis>
</taxonomicName>
it is not crenulated and the perradial lips are short. Additionally, the transverse gonad folds are branched several times in
<taxonomicName baseAuthorityName="Haeckel" baseAuthorityYear="1879" box="[1116,1267,1641,1665]" class="Hydrozoa" family="Pandeidae" genus="Larsonia" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="pterophylla">
<emphasis box="[1116,1267,1641,1665]" italics="true" pageId="18" pageNumber="254">L. pterophylla</emphasis>
</taxonomicName>
, while they are mostly unbranched in
<taxonomicName authorityName="L. Agassiz" authorityYear="1862" box="[1074,1147,1673,1696]" class="Hydrozoa" family="Pandeidae" genus="Stomotoca" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="atra">
<emphasis box="[1074,1147,1673,1696]" italics="true" pageId="18" pageNumber="254">S. atra</emphasis>
</taxonomicName>
(comp.
<bibRefCitation author="Schuchert" box="[1239,1429,1673,1697]" firstAuthor="Schuchert" pageId="18" pageNumber="254" refId="ref74677" refString="Schuchert P. 2017 b. Specimen photos of DNA sample MHNG Hydrozoa DNA 884. Zenodo. DOI: 10.5281 / zenodo. 998731" type="book" year="2017" yearSuffix="b">Schuchert, 2017b</bibRefCitation>
) or in loops in very mature animals (comp.
<bibRefCitation author="Bigelow" box="[1275,1431,1705,1729]" firstAuthor="Bigelow" pageId="18" pageNumber="254" pagination="363 - 442" refId="ref65028" refString="Bigelow H. B. 1918. Some Medusae and Siphonophora from the western Atlantic. Bulletin of the Museum of comparative Zoology of Harvard College 62: 363 - 442, pls 1 - 8." type="journal article" year="1918">Bigelow, 1918</bibRefCitation>
;
<bibRefCitation author="Arai &amp; Brinckmann-Voss" box="[831,1174,1736,1761]" firstAuthor="Arai" pageId="18" pageNumber="254" pagination="1 - 192" refId="ref64599" refString="Arai M. N., Brinckmann-Voss A. 1980. Hydromedusae of British Columbia and Puget Sound. Canadian Bulletin of Fisheries and Aquatic Sciences 204: 1 - 192." type="journal article" year="1980">Arai &amp; Brinckmann-Voss, 1980</bibRefCitation>
). The Brazilian medusae depicted in
<bibRefCitation author="Stampar &amp; Kodja" box="[964,1242,1768,1792]" firstAuthor="Stampar" pageId="18" pageNumber="254" pagination="55 - 57" refId="ref75086" refString="Stampar S. N., Kodja G. 2007. Cnidaria, Hydrozoa, Anthoathecata, Pandeidae, Stomotoca atra: distribution extension. Check List 3 (1): 55 - 57." type="journal article" year="2007">Stampar &amp; Kodja (2007)</bibRefCitation>
and identified as
<taxonomicName authorityName="L. Agassiz" authorityYear="1862" box="[831,902,1801,1824]" class="Hydrozoa" family="Pandeidae" genus="Stomotoca" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="atra">
<emphasis box="[831,902,1801,1824]" italics="true" pageId="18" pageNumber="254">S. atra</emphasis>
</taxonomicName>
must thus belong to
<taxonomicName baseAuthorityName="Haeckel" baseAuthorityYear="1879" box="[1126,1276,1800,1824]" class="Hydrozoa" family="Pandeidae" genus="Larsonia" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="254" phylum="Cnidaria" rank="species" species="pterophylla">
<emphasis box="[1126,1276,1800,1824]" italics="true" pageId="18" pageNumber="254">L. pterophylla</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>