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<document id="7AD69654F1F52299EEC12432CDCDF3C6" ID-CLB-Dataset="5629" ID-DOI="10.11646/zootaxa.3788.1" ID-GBIF-Dataset="80d33caa-2379-4f5e-ba98-6d1cc29a7906" ID-ISSN="1175-5326" ID-Zenodo-Dep="4913761" ID-ZooBank="7484832A-025B-41FA-9696-DF7531C01AA4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623174507327" checkinUser="felipe" docAuthor="Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W. &amp; León-Cortés, Jorge L." docDate="2014" docId="C27E87FFFFAB613DFF14DD912730FA93" docLanguage="en" docName="zt03788p063.pdf" docOrigin="Zootaxa 3788 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tylodinus kuscheli Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés 2014, sp. nov." docType="treatment" docVersion="10" lastPageNumber="50" masterDocId="3E47FF87FF9B610CFF83DB32243FFFC6" masterDocTitle="A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico" masterLastPageNumber="63" masterPageNumber="1" pageNumber="49" updateTime="1699005316853" updateUser="ExternalLinkService">
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<mods:title id="6C595283568CE71F4845EF8D3EAD01AC">A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico</mods:title>
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<mods:namePart id="F85684EA2D836117439EDD74355D815B">Luna-Cozar, Jesús</mods:namePart>
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<mods:namePart id="705CC9C79B2D3E0CF18D3B9DB5F7F9EC">Anderson, Robert S.</mods:namePart>
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<mods:namePart id="A233C7A108209FFA165323AD271C355C">Jones, Robert W.</mods:namePart>
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<subSubSection id="02CD6562FFAB613CFF14DD91268FF97B" box="[151,688,1699,1725]" pageId="48" pageNumber="49" type="nomenclature">
<paragraph id="4A6836E9FFAB613CFF14DD91268FF97B" blockId="48.[151,688,1699,1758]" box="[151,688,1699,1725]" pageId="48" pageNumber="49">
<emphasis id="78A3EAFBFFAB613CFF14DD91268FF97B" bold="true" box="[151,688,1699,1725]" pageId="48" pageNumber="49">
29.
<taxonomicName id="8DD74D6AFFAB613CFF40DD912679F97B" authority="Luna-Cozar" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[195,582,1699,1725]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="kuscheli" status="sp. nov.">
<emphasis id="78A3EAFBFFAB613CFF40DD912599F97B" bold="true" box="[195,422,1699,1725]" italics="true" pageId="48" pageNumber="49">Tylodinus kuscheli</emphasis>
Luna-Cozar
</taxonomicName>
,
<taxonomicNameLabel id="63905780FFAB613CFDD0DD91268FF97B" box="[595,688,1699,1725]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
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<subSubSection id="02CD6562FFAB613CFF14DDF72585F918" box="[151,442,1733,1758]" pageId="48" pageNumber="49" type="description">
<paragraph id="4A6836E9FFAB613CFF14DDF72585F918" blockId="48.[151,688,1699,1758]" box="[151,442,1733,1758]" pageId="48" pageNumber="49">
(
<figureCitation id="D2EC2A6CFFAB613CFF1DDDF7257CF918" box="[158,323,1733,1758]" captionStart="FIGURES 7683" captionStartId="45.[151,264,1974,1996]" captionTargetBox="[205,1321,214,1948]" captionTargetId="figure-50@45.[196,1390,195,1949]" captionTargetPageId="45" captionText="FIGURES 7683. Dorsal and lateral habitus: 7677. Tylodinus rufus, 7879. Tylodinus dominicus, 8081. Tylodinus noctis, 8283. Tylodinus kuscheli." figureDoi="http://doi.org/10.5281/zenodo.4913785" httpUri="https://zenodo.org/record/4913785/files/figure.png" pageId="48" pageNumber="49">Figures 8283</figureCitation>
,
<figureCitation id="D2EC2A6CFFAB613CFECCDDF72544F918" box="[335,379,1733,1758]" captionStart="FIGURES 120129" captionStartId="57.[151,264,1917,1939]" captionTargetBox="[217,1371,209,1825]" captionTargetId="figure-67@57.[216,1371,195,1893]" captionTargetPageId="57" captionText="FIGURES 120129. Ventral and lateral habitus of median lobe: 120. Tylodinus dominicus, 121. Tylodinus noctis, 122. Tylodinus kuscheli, 123. Tylodinus pappi, 124. Tylodinus branstetteri, 125. Tylodinus immundus, 126. Tylodinus intzin, 127. Tylodinus gibbosus, 128. Tylodinus kissingeri, 129. Tylodinus pinguis." figureDoi="http://doi.org/10.5281/zenodo.4913797" httpUri="https://zenodo.org/record/4913797/files/figure.png" pageId="48" pageNumber="49">122</figureCitation>
,
<figureCitation id="D2EC2A6CFFAB613CFE04DDF7258DF918" box="[391,434,1733,1758]" captionStart="FIGURES 134135" captionStartId="60.[151,264,1876,1898]" captionTargetBox="[226,1363,215,1831]" captionTargetId="figure-62@60.[214,1372,195,1851]" captionTargetPageId="60" captionText="FIGURES 134135. Maps showing collecting sites. 134. Occurrences in Chiapas, México of: Tylodinus rufus, Tylodinus kuscheli, Tylodinus pappi, Tylodinus jonesi, Tylodinus coapillensis; 135. Occurrences in Chiapas, México of: Tylodinus spiniventris, Tylodinus lum, Tylodinus noctis, Tylodinus pinguis, Tylodinus rugosus." figureDoi="http://doi.org/10.5281/zenodo.4913803" httpUri="https://zenodo.org/record/4913803/files/figure.png" pageId="48" pageNumber="49">134</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="02CD6562FFAB613DFF14DC3F2796FCD3" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="diagnosis">
<paragraph id="4A6836E9FFAB613DFF14DC3F2796FCD3" blockId="48.[151,1437,1805,2010]" lastBlockId="49.[151,1436,151,1365]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<emphasis id="78A3EAFBFFAB613CFF14DC3F252EF8E0" bold="true" box="[151,273,1805,1830]" pageId="48" pageNumber="49">Diagnosis.</emphasis>
Length male,
<quantity id="8D2F9B0CFFAB613CFE35DC3F267EF8E3" box="[438,577,1805,1830]" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.2" metricValueMin="3.9" pageId="48" pageNumber="49" unit="mm" value="4.05" valueMax="4.2" valueMin="3.9">3.94.2 mm</quantity>
, female
<quantity id="8D2F9B0CFFAB613CFD21DC3F2712F8E3" box="[674,813,1805,1830]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.2" metricValueMin="4.1" pageId="48" pageNumber="49" unit="mm" value="4.15" valueMax="4.2" valueMin="4.1">4.14.2 mm</quantity>
. Width male,
<quantity id="8D2F9B0CFFAB613CFC4FDC3F2069F8E3" box="[972,1110,1805,1830]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="48" pageNumber="49" unit="mm" value="2.15" valueMax="2.2" valueMin="2.1">2.12.2 mm</quantity>
, female
<quantity id="8D2F9B0CFFAB613CFB34DC3F212DF8E3" box="[1207,1298,1805,1830]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="48" pageNumber="49" unit="mm" value="2.3">2.3 mm</quantity>
. Body 2.1x longer than wide, black. Head with punctures similar to punctures on rostrum, clothed with yellow scales; vertex with overlapping scales, uniformly distributed along vertex, similar in color and tonality to head scales; frons concave; rostrum moderately robust, moderately carinate, apical area puncticulate; eyes moderately convex. Pronotum in lateral view strongly convex, outline with anterior constriction, without posterior constriction (
<figureCitation id="D2EC2A6CFFAB613CFAD2DCAF248AF81C" captionStart="FIGURES 111. 15" captionStartId="8.[151,264,1743,1765]" captionTargetBox="[293,1295,205,1711]" captionTargetId="figure-208@8.[280,1300,195,1713]" captionTargetPageId="8" captionText="FIGURES 111. 15. Dorsal outline: 1. Generalized Tylodinus, indicating terminology used in this study. Key: letters indicate areas of pronotum. A, anterior area; B, median area; C, posterior area; a, anterior of lateral area; b, median section of lateral area; c, median section of dorsal area; d, anterior section of dorsal area; e, posterior section of lateral area; f, posterior section of dorsal area. 2. Tylodinus variabilis, note elytra basal margin straight. 3. Tylodinus ixchel, note elytra basal margin slightly sinuate. 4. Tylodinus immundus, note elytral basal margin sinuate. 5. Tylodinus elongatus, note elytra basal margin very sinuate. 611. Lateral outline: 6. Tylodinus complicatus, male, note ventrites 35 in same plane as ventrites 12. 7. Tylodinus variabilis, male, note ventrites 35 not in same plane as ventrites 12. 8. Tylodinus coapillensis, female, note ventrites 12 strongly convex, ventrites 35 not in same plane as ventrites 12. 9. Tylodinus zilchi (schematic). 10. Tylodinus sepulturaensis (schematic) 11. Tylodinus rugosus (schematic)." figureDoi="http://doi.org/10.5281/zenodo.4913763" httpUri="https://zenodo.org/record/4913763/files/figure.png" pageId="48" pageNumber="49">Figure 10</figureCitation>
,
<figureCitation id="D2EC2A6CFFAB613CFF41DCF324DEF81C" box="[194,225,1985,2010]" captionStart="FIGURES 7683" captionStartId="45.[151,264,1974,1996]" captionTargetBox="[205,1321,214,1948]" captionTargetId="figure-50@45.[196,1390,195,1949]" captionTargetPageId="45" captionText="FIGURES 7683. Dorsal and lateral habitus: 7677. Tylodinus rufus, 7879. Tylodinus dominicus, 8081. Tylodinus noctis, 8283. Tylodinus kuscheli." figureDoi="http://doi.org/10.5281/zenodo.4913785" httpUri="https://zenodo.org/record/4913785/files/figure.png" pageId="48" pageNumber="49">83</figureCitation>
); in dorsal view with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond middle; disc with longitudinal impression; surface with dense, deep punctures on flanks, becoming smaller, scattered towards middle, with yellow to light brown scales very dense on lateral area and flanks, anterior area forming a thick longitudinal fringe along middle and narrow strip that diverges from middle base to flanks, and with dark scales on middle and posterior areas of disc (
<figureCitation id="D2EC2A6CFFAA613DFCA1DA3627A9FEDB" box="[802,918,260,285]" captionStart="FIGURES 1219. 12" captionStartId="9.[151,264,1919,1941]" captionTargetBox="[219,1377,209,1898]" captionTargetId="figure-86@9.[219,1377,197,1898]" captionTargetPageId="9" captionText="FIGURES 1219. 12. Tylodinus pusillus head a) male, b) female. 1314. Dorsal habitus of pronotum showing the lateral side of pronotum 13. Tylodinus andersoni, 14. Tylodinus porvenirensis. 1517. Dorsal habitus: 15. Tylodinus rugosus, 16. Tylodinus exiguus, 17. Tylodinus jonesi. 18. Tylodinus coapillensis note tubercle on elytra declivity base, 19. Tylodinus elongatus showing metasternal tooth." figureDoi="http://doi.org/10.5281/zenodo.4913765" httpUri="https://zenodo.org/record/4913765/files/figure.png" pageId="49" pageNumber="50">Figure 14</figureCitation>
,
<figureCitation id="D2EC2A6CFFAA613DFC25DA3627F9FEDB" box="[934,966,260,285]" captionStart="FIGURES 7683" captionStartId="45.[151,264,1974,1996]" captionTargetBox="[205,1321,214,1948]" captionTargetId="figure-50@45.[196,1390,195,1949]" captionTargetPageId="45" captionText="FIGURES 7683. Dorsal and lateral habitus: 7677. Tylodinus rufus, 7879. Tylodinus dominicus, 8081. Tylodinus noctis, 8283. Tylodinus kuscheli." figureDoi="http://doi.org/10.5281/zenodo.4913785" httpUri="https://zenodo.org/record/4913785/files/figure.png" pageId="49" pageNumber="50">82</figureCitation>
); surface without granules. Elytra black, occasionally black with dark red color along intervals 12, basal margin slightly sinuate (
<figureCitation id="D2EC2A6CFFAA613DFB06DA1520D9FE86" box="[1157,1254,295,320]" captionStart="FIGURES 111. 15" captionStartId="8.[151,264,1743,1765]" captionTargetBox="[293,1295,205,1711]" captionTargetId="figure-208@8.[280,1300,195,1713]" captionTargetPageId="8" captionText="FIGURES 111. 15. Dorsal outline: 1. Generalized Tylodinus, indicating terminology used in this study. Key: letters indicate areas of pronotum. A, anterior area; B, median area; C, posterior area; a, anterior of lateral area; b, median section of lateral area; c, median section of dorsal area; d, anterior section of dorsal area; e, posterior section of lateral area; f, posterior section of dorsal area. 2. Tylodinus variabilis, note elytra basal margin straight. 3. Tylodinus ixchel, note elytra basal margin slightly sinuate. 4. Tylodinus immundus, note elytral basal margin sinuate. 5. Tylodinus elongatus, note elytra basal margin very sinuate. 611. Lateral outline: 6. Tylodinus complicatus, male, note ventrites 35 in same plane as ventrites 12. 7. Tylodinus variabilis, male, note ventrites 35 not in same plane as ventrites 12. 8. Tylodinus coapillensis, female, note ventrites 12 strongly convex, ventrites 35 not in same plane as ventrites 12. 9. Tylodinus zilchi (schematic). 10. Tylodinus sepulturaensis (schematic) 11. Tylodinus rugosus (schematic)." figureDoi="http://doi.org/10.5281/zenodo.4913763" httpUri="https://zenodo.org/record/4913763/files/figure.png" pageId="49" pageNumber="50">Figure 3</figureCitation>
), with tubercles as follows: I2 with tubercle slightly beyond declivity base; I3 with three tubercles, first subbasal, second on middle of elytra, similar to subbasal tubercle, third on middle of elytral declivity, similar to tubercle on I2; I4 with two tubercles, first very weakly developed between first two tubercles on I3, second on base of elytral declivity, longer than tubercle on I2; I5 with two tubercles at same position as first two tubercles on I3; I6 with two weak tubercles in same position as those on I4; I7 with two small tubercles in same position as tubercles on I5; elytral declivity with tubercle on I4 much longer than tubercle on I2. Metasternum glabrous, moderately concave, metasternal tooth absent. Male abdomen with ventrites 35 not in same plane as ventrites 12, ventrite 1 moderately concave, squamous, ventrite 2 as long as 34 combined, with a transverse line of scales along middle, ventrite 5 longer than ventrite 1, flat, with anterior impression. Femora armed, hind femur reaching elytral apex, premucro absent. Male genitalia (
<figureCitation id="D2EC2A6CFFAA613DFE8BD9BD25B7FD6E" box="[264,392,655,680]" captionStart="FIGURES 120129" captionStartId="57.[151,264,1917,1939]" captionTargetBox="[217,1371,209,1825]" captionTargetId="figure-67@57.[216,1371,195,1893]" captionTargetPageId="57" captionText="FIGURES 120129. Ventral and lateral habitus of median lobe: 120. Tylodinus dominicus, 121. Tylodinus noctis, 122. Tylodinus kuscheli, 123. Tylodinus pappi, 124. Tylodinus branstetteri, 125. Tylodinus immundus, 126. Tylodinus intzin, 127. Tylodinus gibbosus, 128. Tylodinus kissingeri, 129. Tylodinus pinguis." figureDoi="http://doi.org/10.5281/zenodo.4913797" httpUri="https://zenodo.org/record/4913797/files/figure.png" pageId="49" pageNumber="50">Figure 122</figureCitation>
) with median lobe in lateral view weakly curved, moderately stout; in ventral view longer than half the length of apodemes, swollen at 0.25 length from base of median lobe, convergent, apex acute; apical process in lateral view large (0.350.42 as long as median lobe), slightly curved; in ventral view not constricted beyond median orifice, lateral sides convergent. Female not dissected.
</paragraph>
</subSubSection>
<subSubSection id="02CD6562FFAA613DFF44D82D25FFFBF3" pageId="49" pageNumber="50" type="discussion">
<paragraph id="4A6836E9FFAA613DFF44D82D27B2FC46" blockId="49.[151,1436,151,1365]" pageId="49" pageNumber="50">
<emphasis id="78A3EAFBFFAA613DFF44D82D2570FCFE" bold="true" box="[199,335,799,824]" pageId="49" pageNumber="50">Comments.</emphasis>
This species is closely related to
<taxonomicName id="8DD74D6AFFAA613DFD5FD8132704FCF1" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[732,827,800,824]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pappi">
<emphasis id="78A3EAFBFFAA613DFD5FD8132704FCF1" box="[732,827,800,824]" italics="true" pageId="49" pageNumber="50">T. pappi</emphasis>
</taxonomicName>
and
<taxonomicName id="8DD74D6AFFAA613DFCF4D8132026FCFE" authorityName="Luna-Cozar &amp; Anderson &amp; Jones &amp; León-Cortés" authorityYear="2014" box="[887,1049,799,824]" class="Insecta" family="Curculionidae" genus="Tylodinus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="branstetteri">
<emphasis id="78A3EAFBFFAA613DFCF4D8132026FCFE" box="[887,1049,799,824]" italics="true" pageId="49" pageNumber="50">T. branstetteri</emphasis>
</taxonomicName>
based on the body being clothed mainly with yellow to fulvous scales, moderately dense, dark brown scales that are less sparse, and the median lobe in ventral view distinctly wider at the apical 1/3 (at median orifice).
</paragraph>
<paragraph id="4A6836E9FFAA613DFF44D8BE26F9FC63" blockId="49.[151,1436,151,1365]" box="[199,710,908,933]" pageId="49" pageNumber="50">
<emphasis id="78A3EAFBFFAA613DFF44D8BE25D8FC63" bold="true" box="[199,487,908,933]" pageId="49" pageNumber="50">Geographic distribution</emphasis>
.
<collectingCountry id="32C07679FFAA613DFE76D8BE2675FC63" box="[501,586,908,933]" name="Mexico" pageId="49" pageNumber="50">México</collectingCountry>
(
<collectingRegion id="8813F80BFFAA613DFDD9D8BE2685FC63" box="[602,698,908,933]" country="Mexico" name="Chiapas" pageId="49" pageNumber="50">Chiapas</collectingRegion>
).
</paragraph>
<paragraph id="4A6836E9FFAA613DFF44D89D2506FC2D" blockId="49.[151,1436,151,1365]" pageId="49" pageNumber="50">
<emphasis id="78A3EAFBFFAA613DFF44D89D25E2FC0E" bold="true" box="[199,477,943,968]" pageId="49" pageNumber="50">Habitat and elevation.</emphasis>
Specimens were collected from leaf litter in various wet forest
<typeStatus id="956C884BFFAA613DFB75D882210EFC0E" box="[1270,1329,944,968]" pageId="49" pageNumber="50">types</typeStatus>
between
<quantity id="8D2F9B0CFFAA613DFF14D8E6250AFC2D" box="[151,309,980,1005]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="49" pageNumber="50" unit="m" value="1450.0" valueMax="1600.0" valueMin="1300.0">13001600 m</quantity>
.
</paragraph>
<paragraph id="4A6836E9FFAA613DFF44D8C525FFFBF3" blockId="49.[151,1436,151,1365]" pageId="49" pageNumber="50">
<emphasis id="78A3EAFBFFAA613DFF44D8C5261CFBD6" bold="true" box="[199,547,1015,1040]" pageId="49" pageNumber="50">Derivation of specific name.</emphasis>
Patronym, named after Guillermo (Willy) Kuschel (
<collectingCountry id="32C07679FFAA613DFB13D8C5210DFBD6" box="[1168,1330,1015,1040]" name="New Zealand" pageId="49" pageNumber="50">New Zealand</collectingCountry>
), a wellknown worker on weevils.
</paragraph>
</subSubSection>
<subSubSection id="02CD6562FFAA613DFF44DF0D2730FA93" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="4A6836E9FFAA613DFF44DF0D2730FA93" blockId="49.[151,1436,151,1365]" pageId="49" pageNumber="50">
<emphasis id="78A3EAFBFFAA613DFF44DF0D2596FB9E" bold="true" box="[199,425,1087,1112]" pageId="49" pageNumber="50">Material examined</emphasis>
.
<materialsCitation id="FABF3CB4FFAA613DFE34DF0D2562FBBD" ID-GBIF-Occurrence="3319253360" collectionCode="CMNC, R" collectorName="Total" country="Mexico" location="Ejido Sierra Morena" municipality="Villa Corso" pageId="49" pageNumber="50" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Chiapas" typeStatus="holotype">
<collectorName id="E722533FFFAA613DFE34DF0D25CFFB9E" box="[439,496,1087,1112]" pageId="49" pageNumber="50">Total</collectorName>
<specimenCount id="5CD1FD60FFAA613DFE7BDF0D266EFB9E" box="[504,593,1087,1112]" pageId="49" pageNumber="50" type="male">4 males</specimenCount>
,
<specimenCount id="5CD1FD60FFAA613DFDDEDF0D26F1FB9E" box="[605,718,1087,1112]" pageId="49" pageNumber="50" type="female">2 females</specimenCount>
.
<typeStatus id="956C884BFFAA613DFD63DF0D2777FB9E" box="[736,840,1087,1112]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="2CC6AE2CFFAA613DFC16DF0D27D1FB9E" box="[917,1006,1087,1112]" pageId="49" pageNumber="50">CMNC</collectionCode>
):
<emphasis id="78A3EAFBFFAA613DFB82DF0D2067FB9E" bold="true" box="[1025,1112,1087,1112]" pageId="49" pageNumber="50">
<collectingCountry id="32C07679FFAA613DFB82DF0D2067FB9E" box="[1025,1112,1087,1112]" name="Mexico" pageId="49" pageNumber="50">México</collectingCountry>
</emphasis>
:
<collectingRegion id="8813F80BFFAA613DFBEBDF0D20FAFB9E" box="[1128,1221,1087,1112]" country="Mexico" name="Chiapas" pageId="49" pageNumber="50">Chiapas</collectingRegion>
,
<collectingMunicipality id="AA0CAC93FFAA613DFB53DF0D216CFB9E" box="[1232,1363,1087,1112]" pageId="49" pageNumber="50">Villa Corso</collectingMunicipality>
,
<location id="4F086032FFAA613DFADCDF0D2504FBBD" LSID="urn:lsid:plazi:treatment:C27E87FFFFAB613DFF14DD912730FA93:4F086032FFAA613DFADCDF0D2504FBBD" country="Mexico" municipality="Villa Corso" name="Ejido Sierra Morena" pageId="49" pageNumber="50" stateProvince="Chiapas">Ejido Sierra Morena</location>
,
<collectionCode id="2CC6AE2CFFAA613DFEC4DF562562FBBD" box="[327,349,1124,1147]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="49" pageNumber="50">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="FABF3CB4FFAA613DFEEBDF5624D8FB59" ID-GBIF-Occurrence="3319253623" collectingDate="2008-05-13" collectionCode="R" country="Mexico" elevation="1487" location="Biosfera La Sepultura" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Chiapas" typeStatus="holotype">
<location id="4F086032FFAA613DFEEBDF56265DFBBB" LSID="urn:lsid:plazi:treatment:C27E87FFFFAB613DFF14DD912730FA93:4F086032FFAA613DFEEBDF56265DFBBB" box="[360,610,1124,1149]" country="Mexico" latitude="16.162884" name="Biosfera La Sepultura" pageId="49" pageNumber="50" stateProvince="Chiapas">Biosfera La Sepultura</location>
, elev.
<quantity id="8D2F9B0CFFAA613DFD2ADF5626C0FBBD" box="[681,767,1124,1149]" metricMagnitude="3" metricUnit="m" metricValue="1.4869999999999999" pageId="49" pageNumber="50" unit="m" value="1487.0">
<elevation id="C1FAD1DAFFAA613DFD2ADF5626C0FBBD" box="[681,767,1124,1149]" metricMagnitude="3" metricUnit="m" metricValue="1.4869999999999999" pageId="49" pageNumber="50" unit="m" value="1487.0">1487 m</elevation>
</quantity>
(16°9.773N, 93°3536.211W),
<date id="3E691029FFAA613DFBEEDF5620E2FBBB" box="[1133,1245,1124,1149]" pageId="49" pageNumber="50" value="2008-05-13">
<collectingDate id="2E2DE9C1FFAA613DFBEEDF5620E2FBBB" box="[1133,1245,1124,1149]" pageId="49" pageNumber="50" value="2008-05-13">13.v.2008</collectingDate>
</date>
, oak forest litter, coll.
<collectionCode id="2CC6AE2CFFAA613DFF52DFBA24D8FB59" box="[209,231,1160,1183]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="49" pageNumber="50">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="FABF3CB4FFAA613DFF77DFB520B1FB59" ID-GBIF-Occurrence="3319253702" box="[244,1166,1159,1184]" collectionCode="R" collectorName="S. Anderson." country="Mexico" location="Ejido Sierra Morena" municipality="Villa Flores" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Chiapas" typeStatus="paratype">
<collectorName id="E722533FFFAA613DFF77DFB525B7FB66" box="[244,392,1159,1184]" pageId="49" pageNumber="50">S. Anderson.</collectorName>
<typeStatus id="956C884BFFAA613DFE19DFBA2634FB66" box="[410,523,1160,1184]" pageId="49" pageNumber="50" type="paratype">Paratypes</typeStatus>
:
<emphasis id="78A3EAFBFFAA613DFD94DFB52651FB66" bold="true" box="[535,622,1159,1184]" pageId="49" pageNumber="50">
<collectingCountry id="32C07679FFAA613DFD94DFB52651FB66" box="[535,622,1159,1184]" name="Mexico" pageId="49" pageNumber="50">México</collectingCountry>
</emphasis>
:
<collectingRegion id="8813F80BFFAA613DFDFCDFB526E3FB66" box="[639,732,1159,1184]" country="Mexico" name="Chiapas" pageId="49" pageNumber="50">Chiapas</collectingRegion>
,
<collectingMunicipality id="AA0CAC93FFAA613DFD69DFB5274DFB66" box="[746,882,1159,1184]" pageId="49" pageNumber="50">Villa Flores</collectingMunicipality>
,
<location id="4F086032FFAA613DFCFCDFB52054FB59" LSID="urn:lsid:plazi:treatment:C27E87FFFFAB613DFF14DD912730FA93:4F086032FFAA613DFCFCDFB52054FB59" box="[895,1131,1159,1184]" country="Mexico" municipality="Villa Flores" name="Ejido Sierra Morena" pageId="49" pageNumber="50" stateProvince="Chiapas">Ejido Sierra Morena</location>
,
<collectionCode id="2CC6AE2CFFAA613DFBFBDFBA20B1FB59" box="[1144,1166,1160,1183]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="49" pageNumber="50">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="FABF3CB4FFAA613DFB18DFB527AFFB05" ID-GBIF-Occurrence="3319253346" collectingDate="2003-07-19" collectingDateMax="2003-07-20" collectingDateMin="2003-07-19" collectionCode="R" country="Mexico" elevation="1400" location="Biosfera La Sepultura" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Chiapas" typeStatus="paratype">
<location id="4F086032FFAA613DFB18DFB521A7FB66" LSID="urn:lsid:plazi:treatment:C27E87FFFFAB613DFF14DD912730FA93:4F086032FFAA613DFB18DFB521A7FB66" box="[1179,1432,1159,1184]" country="Mexico" latitude="16.15289" longLatPrecision="1" longitude="-93.59031" name="Biosfera La Sepultura" pageId="49" pageNumber="50" stateProvince="Chiapas">Biosfera La Sepultura</location>
, elev.
<quantity id="8D2F9B0CFFAA613DFF50DF9E2516FB05" box="[211,297,1196,1221]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="49" pageNumber="50" unit="m" value="1400.0">
<elevation id="C1FAD1DAFFAA613DFF50DF9E2516FB05" box="[211,297,1196,1221]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="49" pageNumber="50" unit="m" value="1400.0">1400 m</elevation>
</quantity>
(16°9'10.4''N, 93°35'25.1''W),
<date id="3E691029FFAA613DFD0BDF9E270BFB03" box="[648,820,1196,1221]" pageId="49" pageNumber="50" value="2003-07-19" valueMax="2003-07-20" valueMin="2003-07-19">
<collectingDate id="2E2DE9C1FFAA613DFD0BDF9E270BFB03" box="[648,820,1196,1221]" pageId="49" pageNumber="50" value="2003-07-19" valueMax="2003-07-20" valueMin="2003-07-19">1920.vii.2003</collectingDate>
</date>
, coll.
<collectionCode id="2CC6AE2CFFAA613DFCF9DF9E27AFFB05" box="[890,912,1196,1219]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="49" pageNumber="50">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="FABF3CB4FFAA613DFC1FDF9E2689FB2E" ID-GBIF-Occurrence="3319253333" collectingDate="2008-05-12" collectingDateMax="2003-07-20" collectingDateMin="2003-07-19" collectionCode="ECOS, CMNC, LLAMA" collectorName="S. Anderson" country="Mexico" elevation="1600" location="Biosfera La Sepultura" pageId="49" pageNumber="50" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiapas" typeStatus="paratype">
<collectorName id="E722533FFFAA613DFC1FDF9E2014FB03" box="[924,1067,1196,1221]" pageId="49" pageNumber="50">S. Anderson</collectorName>
<specimenCount id="5CD1FD60FFAA613DFBCADF9E20AFFB03" box="[1097,1168,1196,1221]" pageId="49" pageNumber="50" type="male">1 male</specimenCount>
(
<collectionCode id="2CC6AE2CFFAA613DFB23DF9E20D2FB03" box="[1184,1261,1196,1221]" pageId="49" pageNumber="50">ECOS</collectionCode>
); same locality, elev.
<quantity id="8D2F9B0CFFAA613DFF59DFFD2509FB21" box="[218,310,1231,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="49" pageNumber="50" unit="m" value="1600.0">
<elevation id="C1FAD1DAFFAA613DFF59DFFD2509FB21" box="[218,310,1231,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="49" pageNumber="50" unit="m" value="1600.0">1600 m</elevation>
</quantity>
(16°9'10.6''N, 93°35'25.1''W)—
</materialsCitation>
<materialsCitation id="FABF3CB4FFAA613DFD35DFFD26EBFACB" ID-GBIF-Occurrence="3319253701" collectingDate="2008-05-12" collectingDateMax="2003-07-20" collectingDateMin="2003-07-19" collectionCode="ECOS, CMNC, LLAMA" collectorName="S. Anderson" country="Mexico" elevation="1600" location="Biosfera La Sepultura" pageId="49" pageNumber="50" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiapas" typeStatus="paratype">
<specimenCount id="5CD1FD60FFAA613DFD35DFFD2725FB2E" box="[694,794,1231,1256]" pageId="49" pageNumber="50" type="female">1 female</specimenCount>
(
<collectionCode id="2CC6AE2CFFAA613DFCB2DFFD27B5FB2E" box="[817,906,1231,1256]" pageId="49" pageNumber="50">CMNC</collectionCode>
); same locality, elev.
<quantity id="8D2F9B0CFFAA613DFB1BDFFD20CAFB21" box="[1176,1269,1231,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="49" pageNumber="50" unit="m" value="1360.0">
<elevation id="C1FAD1DAFFAA613DFB1BDFFD20CAFB21" box="[1176,1269,1231,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="49" pageNumber="50" unit="m" value="1360.0">1360 m</elevation>
</quantity>
(16.16028°N, 93.60507°N),
<date id="3E691029FFAA613DFEB5DFC62599FACB" box="[310,422,1268,1293]" pageId="49" pageNumber="50" value="2008-05-12">
<collectingDate id="2E2DE9C1FFAA613DFEB5DFC62599FACB" box="[310,422,1268,1293]" pageId="49" pageNumber="50" value="2008-05-12">12.v.2008</collectingDate>
</date>
,
<collectionCode id="2CC6AE2CFFAA613DFE31DFC62629FACB" box="[434,534,1268,1293]" pageId="49" pageNumber="50">LLAMA</collectionCode>
(Wa-A-01-1-40)
</materialsCitation>
<materialsCitation id="FABF3CB4FFAA613DFD77DFC62604FAF6" ID-GBIF-Occurrence="3319253331" collectingDate="2008-05-12" collectingDateMax="2003-07-20" collectingDateMin="2003-07-19" collectionCode="CMNC, LLAMA" collectorName="S. Anderson" country="Mexico" elevation="1600" location="Biosfera La Sepultura" pageId="49" pageNumber="50" specimenCount="1" specimenCount-female="1" stateProvince="Chiapas" typeStatus="paratype">
<specimenCount id="5CD1FD60FFAA613DFD77DFC62770FACB" box="[756,847,1268,1293]" pageId="49" pageNumber="50" type="female">1 female</specimenCount>
(
<collectionCode id="2CC6AE2CFFAA613DFCDDDFC62788FACB" box="[862,951,1268,1293]" pageId="49" pageNumber="50">CMNC</collectionCode>
); same locality (16.15969°N, 93.60516°N),
<date id="3E691029FFAA613DFF14DE252538FAF6" box="[151,263,1303,1328]" pageId="49" pageNumber="50" value="2008-05-12">
<collectingDate id="2E2DE9C1FFAA613DFF14DE252538FAF6" box="[151,263,1303,1328]" pageId="49" pageNumber="50" value="2008-05-12">12.v.2008</collectingDate>
</date>
,
<collectionCode id="2CC6AE2CFFAA613DFE95DE252545FAF6" box="[278,378,1303,1328]" pageId="49" pageNumber="50">LLAMA</collectionCode>
(Wa-A-01-1-23)
</materialsCitation>
<materialsCitation id="FABF3CB4FFAA613DFDD8DE252609FA93" ID-GBIF-Occurrence="3319253800" collectingDate="2008-05-12" collectingDateMax="2003-07-20" collectingDateMin="2003-07-19" collectionCode="ECOS, LLAMA" collectorName="S. Anderson" country="Mexico" elevation="1310" location="Biosfera La Sepultura" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Chiapas" typeStatus="paratype">
<specimenCount id="5CD1FD60FFAA613DFDD8DE25269BFAF6" box="[603,676,1303,1328]" pageId="49" pageNumber="50" type="male">1 male</specimenCount>
(
<collectionCode id="2CC6AE2CFFAA613DFD34DE25273BFAF6" box="[695,772,1303,1328]" pageId="49" pageNumber="50">ECOS</collectionCode>
); same locality, elev.
<quantity id="8D2F9B0CFFAA613DFC7CDE252068FAE9" box="[1023,1111,1303,1328]" metricMagnitude="3" metricUnit="m" metricValue="1.31" pageId="49" pageNumber="50" unit="m" value="1310.0">
<elevation id="C1FAD1DAFFAA613DFC7CDE252068FAE9" box="[1023,1111,1303,1328]" metricMagnitude="3" metricUnit="m" metricValue="1.31" pageId="49" pageNumber="50" unit="m" value="1310.0">1310 m</elevation>
</quantity>
(16.15404°N, 93.60080°N),
<date id="3E691029FFAA613DFF14DE0E2538FA93" box="[151,263,1340,1365]" pageId="49" pageNumber="50" value="2008-05-12">
<collectingDate id="2E2DE9C1FFAA613DFF14DE0E2538FA93" box="[151,263,1340,1365]" pageId="49" pageNumber="50" value="2008-05-12">12.v.2008</collectingDate>
</date>
,
<collectionCode id="2CC6AE2CFFAA613DFE90DE0E2548FA93" box="[275,375,1340,1365]" pageId="49" pageNumber="50">LLAMA</collectionCode>
(Wa-A-01-2-17)
</materialsCitation>
<materialsCitation id="FABF3CB4FFAA613DFDD6DE0E2734FA93" ID-GBIF-Occurrence="3319253575" box="[597,779,1340,1365]" collectionCode="CMNC" country="Mexico" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Chiapas" typeStatus="paratype">
<specimenCount id="5CD1FD60FFAA613DFDD6DE0E26A4FA93" box="[597,667,1340,1365]" pageId="49" pageNumber="50" type="male">1 male</specimenCount>
(
<collectionCode id="2CC6AE2CFFAA613DFD28DE0E273BFA93" box="[683,772,1340,1365]" pageId="49" pageNumber="50">CMNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>