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<document id="B2277CE2A91EBAFC59CA856EDAD21AD5" ID-CLB-Dataset="3142" ID-DOI="10.11646/zootaxa.5072.5.2" ID-GBIF-Dataset="cdde4106-d3d5-4294-9f6c-1e82afd8d580" ID-ISSN="1175-5326" ID-Zenodo-Dep="5748997" ID-ZooBank="B1D30BB4-8CAB-419E-98DE-02D97726137C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638425477373" checkinUser="plazi" docAuthor="Hutchinson, Paul M. &amp; Allsopp, Peter G." docDate="2021" docId="79268507FFCAFFA2CBE5D4CCFA15F99A" docLanguage="en" docName="zootaxa.5072.5.2.pdf" docOrigin="Zootaxa 5072 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enracius Hutchinson &amp; Allsopp 2021" docType="key" docVersion="7" lastPageNumber="446" masterDocId="851FFD7FFFCDFFA5CB72D024FF89FFA0" masterDocTitle="Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina)" masterLastPageNumber="462" masterPageNumber="439" pageNumber="446" updateTime="1699471722501" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="6B172959BEE6B59550D0AF5FA7B8AB5F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6482DAFE691D49A977F6CBF9B81F1D41">
<mods:title id="6E5605ED24853CE1C88D8182046626B4">Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina)</mods:title>
</mods:titleInfo>
<mods:name id="D19FDEC6FC9FB1C69D7FB070EA91AB31" type="personal">
<mods:role id="F5BCAAAA850748786503E3B186EEA50B">
<mods:roleTerm id="03F6796517B6BABE4876DAD62681542A">Author</mods:roleTerm>
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<mods:namePart id="9B7FC0B0CEEF36D8BA72F8C1A51F8201">Hutchinson, Paul M.</mods:namePart>
<mods:affiliation id="261A9087F9EFFFBEFD8A766D156AE1F5">Department of Primary Industries and Regional Development (Quarantine WA), Locked Bag 69, Welshpool DC, Western Australia 6986, Australia.</mods:affiliation>
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</mods:role>
<mods:namePart id="DE7AEC586C18E25B758750E4A5932E06">Allsopp, Peter G.</mods:namePart>
<mods:affiliation id="5223DAA5A96E6AFD0C31BE8EF97A74F1">Queensland Museum, PO Box 3300, South Brisbane, Queensland 4101, Australia.</mods:affiliation>
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<mods:typeOfResource id="0170E97DBB156C94926D70F6E08611B1">text</mods:typeOfResource>
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<mods:title id="A9663C856F3F398F9BD1FDF5A493260C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2CEF719A1741352755C206EFE1ED0632">
<mods:date id="66DE6D62D25D4F27D214C7B18130BE1C">2021</mods:date>
<mods:detail id="8E68E7CAD66B8B94CD6528A26B332992" type="pubDate">
<mods:number id="62134553BEA627A59846CE6A6D7CB2F2">2021-12-02</mods:number>
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<mods:number id="2C8405C730C958EADF9D89043B1534A2">5072</mods:number>
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<mods:number id="CF9C75DEDFFABD7F66C86C71BF432170">5</mods:number>
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<mods:classification id="F54C54BB2A07098E677EAA00E7AEB014">journal article</mods:classification>
<mods:identifier id="03310BAD70C9389C41C6911DA1251D65" type="CLB-Dataset">3142</mods:identifier>
<mods:identifier id="9CC3318AB5B29D4048C5CD191E362AD1" type="DOI">10.11646/zootaxa.5072.5.2</mods:identifier>
<mods:identifier id="2457D7992C0DC769ABB5E576A9831CF0" type="GBIF-Dataset">cdde4106-d3d5-4294-9f6c-1e82afd8d580</mods:identifier>
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<treatment id="79268507FFCAFFA2CBE5D4CCFA15F99A" ID-DOI="http://doi.org/10.5281/zenodo.5750350" ID-GBIF-Taxon="190874935" ID-Zenodo-Dep="5750350" LSID="urn:lsid:plazi:treatment:79268507FFCAFFA2CBE5D4CCFA15F99A" httpUri="http://treatment.plazi.org/id/79268507FFCAFFA2CBE5D4CCFA15F99A" lastPageNumber="446" pageId="7" pageNumber="446">
<subSubSection id="B995679AFFCAFFA2CBE5D4CCFE54FAA2" box="[151,477,1256,1282]" pageId="7" pageNumber="446" type="nomenclature">
<paragraph id="F1303411FFCAFFA2CBE5D4CCFE54FAA2" blockId="7.[151,477,1256,1282]" box="[151,477,1256,1282]" pageId="7" pageNumber="446">
<heading id="AA78837DFFCAFFA2CBE5D4CCFE54FAA2" bold="true" box="[151,477,1256,1282]" fontSize="11" level="1" pageId="7" pageNumber="446" reason="1">
<emphasis id="C3FBE803FFCAFFA2CBE5D4CCFE54FAA2" bold="true" box="[151,477,1256,1282]" pageId="7" pageNumber="446">
Key to species of
<taxonomicName id="368F4F92FFCAFFA2CA1CD4CCFE54FAA2" authorityName="Hutchinson &amp; Allsopp" authorityYear="2021" box="[366,477,1256,1282]" class="Insecta" family="Dynastidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="446" phylum="Arthropoda" rank="genus">
<emphasis id="C3FBE803FFCAFFA2CA1CD4CCFE54FAA2" bold="true" box="[366,477,1256,1282]" italics="true" pageId="7" pageNumber="446">Enracius</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B995679AFFCAFFA2CBE5D514FD3FFAEA" box="[151,694,1328,1355]" pageId="7" pageNumber="446" type="multiple">
<paragraph id="F1303411FFCAFFA2CBE5D514FD3FFAEA" blockId="7.[151,694,1328,1355]" box="[151,694,1328,1355]" pageId="7" pageNumber="446">
Males of
<taxonomicName id="368F4F92FFCAFFA2CA73D515FE0FFAEA" authority="Hutchinson &amp; Allsopp, 2021" authorityName="Hutchinson &amp; Allsopp" authorityYear="2021" box="[257,390,1328,1354]" class="Insecta" family="Dynastidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="446" phylum="Arthropoda" rank="species" species="richardsi" status="sp. nov.">
<emphasis id="C3FBE803FFCAFFA2CA73D515FE0FFAEA" box="[257,390,1328,1354]" italics="true" pageId="7" pageNumber="446">E. richardsi</emphasis>
</taxonomicName>
<emphasis id="C3FBE803FFCAFFA2CAFFD514FD9FFAEA" bold="true" box="[397,534,1328,1354]" pageId="7" pageNumber="446">
<taxonomicNameLabel id="D8C85578FFCAFFA2CAFFD514FD9FFAEA" box="[397,534,1328,1354]" pageId="7" pageNumber="446" rank="species">new species</taxonomicNameLabel>
</emphasis>
are unknown.
</paragraph>
</subSubSection>
<subSubSection id="B995679AFFCAFFA2CBE5D551FA15F99A" pageId="7" pageNumber="446" type="key">
<keyStep id="4A7B2CB4FFCAFFA2CBE5D551FA15F99A" pageId="7" pageNumber="446">
<key id="851E6320FFCAFFA2CBE5D551FA15F99A" pageId="7" pageNumber="446">
<paragraph id="F1303411FFCAFFA2CBE5D551FA14FA43" blockId="7.[151,1437,1397,1595]" pageId="7" pageNumber="446">
<materialsCitation id="41E73E4CFFCAFFA2CBE5D551FA14FA43" ID-GBIF-Occurrence="3413046305" location="Mount Ragged" municipality="Fitzgerald River" pageId="7" pageNumber="446" specimenCount="1" stateProvince="Western Australia">
<keyLead id="4A7E9724FFCAFFA2CBE5D551FA14FA43" pageId="7" pageNumber="446">
1 Clypeus glabrous (
<figureCitation id="69B42894FFCAFFA2CAEBD551FE5CFA2B" box="[409,469,1397,1419]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[173,1415,219,1920]" captionTargetId="figure-26@6.[173,1415,181,1920]" captionTargetPageId="6" captionText="FIGURES 8–13. Enracius richardsi Hutchinson &amp; Allsopp new species, holotype female. 8, Dorsal view; 9, ventral view; 10, lateral view; 11, head dorsal view; 12, head ventral view; 13, labels and left antennomeres 2–10." figureDoi="http://doi.org/10.5281/zenodo.5749007" httpUri="https://zenodo.org/record/5749007/files/figure.png" pageId="7" pageNumber="446">Fig. 8</figureCitation>
); clypeofrontal ridge arcuate (
<figureCitation id="69B42894FFCAFFA2C98DD551FCCFFA2B" box="[767,838,1397,1419]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[173,1415,219,1920]" captionTargetId="figure-26@6.[173,1415,181,1920]" captionTargetPageId="6" captionText="FIGURES 8–13. Enracius richardsi Hutchinson &amp; Allsopp new species, holotype female. 8, Dorsal view; 9, ventral view; 10, lateral view; 11, head dorsal view; 12, head ventral view; 13, labels and left antennomeres 2–10." figureDoi="http://doi.org/10.5281/zenodo.5749007" httpUri="https://zenodo.org/record/5749007/files/figure.png" pageId="7" pageNumber="446">Fig. 11</figureCitation>
); ocular canthus with lateral margin continuous with clypeus (
<figureCitation id="69B42894FFCAFFA2CB9BD5B6FEB9FA08" box="[233,304,1426,1448]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[173,1415,219,1920]" captionTargetId="figure-26@6.[173,1415,181,1920]" captionTargetPageId="6" captionText="FIGURES 8–13. Enracius richardsi Hutchinson &amp; Allsopp new species, holotype female. 8, Dorsal view; 9, ventral view; 10, lateral view; 11, head dorsal view; 12, head ventral view; 13, labels and left antennomeres 2–10." figureDoi="http://doi.org/10.5281/zenodo.5749007" httpUri="https://zenodo.org/record/5749007/files/figure.png" pageId="7" pageNumber="446">Fig. 11</figureCitation>
); elytral striae deeply impressed (
<figureCitation id="69B42894FFCAFFA2C908D5B6FD3CFA08" box="[634,693,1426,1448]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[173,1415,219,1920]" captionTargetId="figure-26@6.[173,1415,181,1920]" captionTargetPageId="6" captionText="FIGURES 8–13. Enracius richardsi Hutchinson &amp; Allsopp new species, holotype female. 8, Dorsal view; 9, ventral view; 10, lateral view; 11, head dorsal view; 12, head ventral view; 13, labels and left antennomeres 2–10." figureDoi="http://doi.org/10.5281/zenodo.5749007" httpUri="https://zenodo.org/record/5749007/files/figure.png" pageId="7" pageNumber="446">Fig. 8</figureCitation>
); apical ridge of pygidium glabrous; body
<quantity id="367799F4FFCAFFA2CF2BD5B6FB48FA08" box="[1113,1217,1426,1448]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="7" pageNumber="446" unit="mm" value="20.0" valueMax="21.0" valueMin="19.0">19–21 mm</quantity>
long; central southern coastal
<collectingRegion id="334BFAF3FFCAFFA2CA58D594FE5CFA66" box="[298,469,1456,1478]" country="Australia" name="Western Australia" pageId="7" pageNumber="446">Western Australia</collectingRegion>
from
<collectingMunicipality id="1154AE6BFFCAFFA2C97DD594FD25FA66" box="[527,684,1456,1478]" pageId="7" pageNumber="446">Fitzgerald River</collectingMunicipality>
to
<location id="F45062CAFFCAFFA2C9B9D594FCD3FA66" LSID="urn:lsid:plazi:treatment:79268507FFCAFFA2CBE5D4CCFA15F99A:F45062CAFFCAFFA2C9B9D594FCD3FA66" box="[715,858,1456,1478]" municipality="Fitzgerald River" name="Mount Ragged" pageId="7" pageNumber="446" stateProvince="Western Australia">Mount Ragged</location>
(
<figureCitation id="69B42894FFCAFFA2C815D594FC24FA66" box="[871,941,1456,1478]" captionStart="FIGURE 14" captionStartId="7.[151,250,1114,1138]" captionTargetBox="[151,1435,309,1090]" captionTargetId="figure-82@7.[151,1435,309,1090]" captionTargetPageId="7" captionText="FIGURE 14. Known distribution of Enracius longipes Dechambre, 1999 and E. richardsi Hutchinson &amp; Allsopp new species in Western Australia." figureDoi="http://doi.org/10.5281/zenodo.5749009" httpUri="https://zenodo.org/record/5749009/files/figure.png" pageId="7" pageNumber="446">Fig. 14</figureCitation>
)...............................................................................................
<taxonomicName id="368F4F92FFCAFFA2C8FAD5EAFA95FA43" authority="Hutchinson &amp; Allsopp" authorityName="Hutchinson &amp; Allsopp" authorityYear="2021" box="[904,1308,1485,1507]" class="Insecta" family="Dynastidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="446" phylum="Arthropoda" rank="species" species="richardsi" status="sp. nov.">
<emphasis id="C3FBE803FFCAFFA2C8FAD5EAFBB5FA42" box="[904,1084,1485,1507]" italics="true" pageId="7" pageNumber="446">Enracius richardsi</emphasis>
Hutchinson &amp; Allsopp
</taxonomicName>
,
<emphasis id="C3FBE803FFCAFFA2CE54D5E9FA14FA43" bold="true" box="[1318,1437,1485,1507]" pageId="7" pageNumber="446">
<taxonomicNameLabel id="D8C85578FFCAFFA2CE54D5E9FA14FA43" box="[1318,1437,1485,1507]" pageId="7" pageNumber="446" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</materialsCitation>
</paragraph>
<paragraph id="F1303411FFCAFFA2CBE5D5CFFA15F99A" blockId="7.[151,1437,1397,1595]" pageId="7" pageNumber="446">
<keyLead id="4A7E9724FFCAFFA2CBE5D5CFFA15F99A" pageId="7" pageNumber="446">
- Clypeus setose (
<figureCitation id="69B42894FFCAFFA2CA0ED5CEFE37FA5F" box="[380,446,1514,1536]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1304,1328]" captionTargetBox="[196,1432,179,1277]" captionTargetId="figure-19@3.[156,1432,179,1277]" captionTargetPageId="3" captionText="FIGURES 1–4. Enracius longipes Dechambre, 1999, paratype male. 1, Dorsal view; 2, pygidium; 3, phallobase and parameres, lateral view; 4, labels (photograph by Fabien Dupuis)." figureDoi="http://doi.org/10.5281/zenodo.5749001" httpUri="https://zenodo.org/record/5749001/files/figure.png" pageId="7" pageNumber="446">Figs. 1</figureCitation>
,
<figureCitation id="69B42894FFCAFFA2CABBD5CEFE5EF9A0" box="[457,471,1514,1536]" captionStart="FIGURES 5–7" captionStartId="4.[151,264,2000,2024]" captionTargetBox="[174,1423,460,1977]" captionTargetId="figure-154@4.[162,1426,453,1977]" captionTargetPageId="4" captionText="FIGURES 5–7. Enracius longipes Dechambre, 1999, female. 5, Dorsal view; 6, ventral view; 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5749003" httpUri="https://zenodo.org/record/5749003/files/figure.png" pageId="7" pageNumber="446">5</figureCitation>
); clypeofrontal ridge linear (
<figureCitation id="69B42894FFCAFFA2C994D5CEFCA1FA5F" box="[742,808,1514,1536]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1304,1328]" captionTargetBox="[196,1432,179,1277]" captionTargetId="figure-19@3.[156,1432,179,1277]" captionTargetPageId="3" captionText="FIGURES 1–4. Enracius longipes Dechambre, 1999, paratype male. 1, Dorsal view; 2, pygidium; 3, phallobase and parameres, lateral view; 4, labels (photograph by Fabien Dupuis)." figureDoi="http://doi.org/10.5281/zenodo.5749001" httpUri="https://zenodo.org/record/5749001/files/figure.png" pageId="7" pageNumber="446">Figs. 1</figureCitation>
,
<figureCitation id="69B42894FFCAFFA2C841D5CEFCC8F9A0" box="[819,833,1514,1536]" captionStart="FIGURES 5–7" captionStartId="4.[151,264,2000,2024]" captionTargetBox="[174,1423,460,1977]" captionTargetId="figure-154@4.[162,1426,453,1977]" captionTargetPageId="4" captionText="FIGURES 5–7. Enracius longipes Dechambre, 1999, female. 5, Dorsal view; 6, ventral view; 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5749003" httpUri="https://zenodo.org/record/5749003/files/figure.png" pageId="7" pageNumber="446">5</figureCitation>
); ocular canthus with lateral margin angular to clypeus (
<figureCitation id="69B42894FFCAFFA2CE27D5CEFA1FFA5F" box="[1365,1430,1514,1536]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1304,1328]" captionTargetBox="[196,1432,179,1277]" captionTargetId="figure-19@3.[156,1432,179,1277]" captionTargetPageId="3" captionText="FIGURES 1–4. Enracius longipes Dechambre, 1999, paratype male. 1, Dorsal view; 2, pygidium; 3, phallobase and parameres, lateral view; 4, labels (photograph by Fabien Dupuis)." figureDoi="http://doi.org/10.5281/zenodo.5749001" httpUri="https://zenodo.org/record/5749001/files/figure.png" pageId="7" pageNumber="446">Figs. 1</figureCitation>
,
<figureCitation id="69B42894FFCAFFA2CB90D62CFF79F9BE" box="[226,240,1544,1566]" captionStart="FIGURES 5–7" captionStartId="4.[151,264,2000,2024]" captionTargetBox="[174,1423,460,1977]" captionTargetId="figure-154@4.[162,1426,453,1977]" captionTargetPageId="4" captionText="FIGURES 5–7. Enracius longipes Dechambre, 1999, female. 5, Dorsal view; 6, ventral view; 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5749003" httpUri="https://zenodo.org/record/5749003/files/figure.png" pageId="7" pageNumber="446">5</figureCitation>
); elytral striae finely impressed (
<figureCitation id="69B42894FFCAFFA2C959D62CFDE4F9BD" box="[555,621,1544,1566]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1304,1328]" captionTargetBox="[196,1432,179,1277]" captionTargetId="figure-19@3.[156,1432,179,1277]" captionTargetPageId="3" captionText="FIGURES 1–4. Enracius longipes Dechambre, 1999, paratype male. 1, Dorsal view; 2, pygidium; 3, phallobase and parameres, lateral view; 4, labels (photograph by Fabien Dupuis)." figureDoi="http://doi.org/10.5281/zenodo.5749001" httpUri="https://zenodo.org/record/5749001/files/figure.png" pageId="7" pageNumber="446">Figs. 1</figureCitation>
,
<figureCitation id="69B42894FFCAFFA2C90BD62CFD0EF9BE" box="[633,647,1544,1566]" captionStart="FIGURES 5–7" captionStartId="4.[151,264,2000,2024]" captionTargetBox="[174,1423,460,1977]" captionTargetId="figure-154@4.[162,1426,453,1977]" captionTargetPageId="4" captionText="FIGURES 5–7. Enracius longipes Dechambre, 1999, female. 5, Dorsal view; 6, ventral view; 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5749003" httpUri="https://zenodo.org/record/5749003/files/figure.png" pageId="7" pageNumber="446">5</figureCitation>
); apical ridge of pygidium setose; body
<quantity id="367799F4FFCAFFA2CF7BD62CFB1AF9BD" box="[1033,1171,1544,1566]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.75" metricValueMin="1.35" pageId="7" pageNumber="446" unit="mm" value="15.5" valueMax="17.5" valueMin="13.5">13.5–17.5 mm</quantity>
long; southeastern
<collectingRegion id="334BFAF3FFCAFFA2CE3CD62CFEB3F99B" country="Australia" name="Western Australia" pageId="7" pageNumber="446">Western Australia</collectingRegion>
from Caiguna to the border with
<collectingRegion id="334BFAF3FFCAFFA2C90ED601FC99F99B" box="[636,784,1573,1595]" country="Australia" name="South Australia" pageId="7" pageNumber="446">South Australia</collectingRegion>
(
<figureCitation id="69B42894FFCAFFA2C86CD601FCEAF99B" box="[798,867,1573,1595]" captionStart="FIGURE 14" captionStartId="7.[151,250,1114,1138]" captionTargetBox="[151,1435,309,1090]" captionTargetId="figure-82@7.[151,1435,309,1090]" captionTargetPageId="7" captionText="FIGURE 14. Known distribution of Enracius longipes Dechambre, 1999 and E. richardsi Hutchinson &amp; Allsopp new species in Western Australia." figureDoi="http://doi.org/10.5281/zenodo.5749009" httpUri="https://zenodo.org/record/5749009/files/figure.png" pageId="7" pageNumber="446">Fig. 14</figureCitation>
)................
<taxonomicName id="368F4F92FFCAFFA2CF4CD602FA15F99A" authority="Dechambre, 1999" authorityName="Dechambre" authorityYear="1999" box="[1086,1436,1573,1595]" class="Insecta" family="Dynastidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="446" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="C3FBE803FFCAFFA2CF4CD602FB62F99A" box="[1086,1259,1573,1595]" italics="true" pageId="7" pageNumber="446">Enracius longipes</emphasis>
<bibRefCitation id="951E49E0FFCAFFA2CF80D601FA15F99A" author="Dechambre, R. - P." box="[1266,1436,1573,1595]" pageId="7" pageNumber="446" pagination="37 - 38" refId="ref10492" refString="Dechambre, R. - P. (1999) Enracius longipes n. gen., n. sp., nouveau Dynastidae d'Australie (Col. Scarabaeoidea Dynastidae). Bulletin de la Societe Entomologique de France, 104, 37 - 38. https: // doi. org / 10.3406 / bsef. 1999.16541" type="journal article" year="1999">Dechambre, 1999</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</key>
</keyStep>
</subSubSection>
</treatment>
</document>
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