673 lines
54 KiB
XML
673 lines
54 KiB
XML
<document ID-DOI="10.24349/z72f-jdlc" ID-ISSN="2107-7207" ID-Zenodo-Dep="7159892" approvalRequired="14" approvalRequired_for_textStreams="14" checkinTime="1665194714052" checkinUser="felipe" docAuthor="Ermilov, Sergey G. & Martens, Jochen" docDate="2021" docId="7242476B884AF048FE2A7549F47E93F9" docLanguage="en" docName="Acarologia.61.4.995-1014.pdf.pdf" docOrigin="Acarologia 61 (4)" docSource="https://www1.montpellier.inrae.fr/CBGP/acarologia/article.php?id=4480" docStyle="DocumentStyle:0387F17C4AF9F8D6336E46CC5E5664D0.4:Acarologia.2018-.journal_article" docStyleId="0387F17C4AF9F8D6336E46CC5E5664D0" docStyleName="Acarologia.2018-.journal_article" docStyleVersion="4" docTitle="Leobodes becki Ermilov & Martens 2021, n. sp." docType="treatment" docVersion="2" lastPageNumber="1004" masterDocId="8E7B3F138849F042FFC17710F17B911B" masterDocTitle="Contribution to the knowledge of the oribatid mite genus Leobodes (Acari, Oribatida, Nippobodidae)" masterLastPageNumber="1014" masterPageNumber="995" pageNumber="997" updateTime="1665195551819" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Contribution to the knowledge of the oribatid mite genus Leobodes (Acari, Oribatida, Nippobodidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ermilov, Sergey G.</mods:namePart>
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<mods:affiliation>Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Martens, Jochen</mods:namePart>
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<mods:affiliation>Johannes Gutenberg-Universität, Institut für Organismische und Molekulare Evolutionsbiologie (iomE), Mainz, Germany ; Senckenberg Research Institute, Arachnology, Frankfurt am Main, Germany. & Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Acarologia</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-12-10</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>61</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>995</mods:start>
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<mods:end>1014</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>https://www1.montpellier.inrae.fr/CBGP/acarologia/article.php?id=4480</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.24349/z72f-jdlc</mods:identifier>
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<mods:identifier type="ISSN">2107-7207</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7159892</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:7242476B884AF048FE2A7549F47E93F9" httpUri="http://treatment.plazi.org/id/7242476B884AF048FE2A7549F47E93F9" lastPageId="10" lastPageNumber="1004" pageId="3" pageNumber="997">
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<subSubSection pageId="3" pageNumber="997" type="nomenclature">
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||
<paragraph blockId="3.[491,765,601,627]" box="[491,765,601,627]" pageId="3" pageNumber="997">
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||
<heading bold="true" box="[491,765,601,627]" fontSize="11" level="2" pageId="3" pageNumber="997" reason="3">
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<emphasis bold="true" box="[491,765,601,627]" pageId="3" pageNumber="997">
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<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[491,690,601,627]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="997" phylum="Arthropoda" rank="species" species="becki" status="n. sp.">
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<emphasis bold="true" box="[491,690,601,627]" italics="true" pageId="3" pageNumber="997">Leobodes becki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[698,765,601,627]" pageId="3" pageNumber="997" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="3.[491,1544,657,2045]" box="[491,1000,657,676]" pageId="3" pageNumber="997">
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Zoobank:
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<uuid box="[580,1000,657,676]" pageId="3" pageNumber="997">60B6A947-D356-424F-8C47-F7F7A0A8D718</uuid>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[531,1073,688,712]" pageId="3" pageNumber="997" type="description">
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||
<paragraph blockId="3.[491,1544,657,2045]" box="[531,1073,688,712]" pageId="3" pageNumber="997">
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(
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<figureCitation box="[539,640,688,712]" captionStart="Figure 1" captionStartId="4.[119,186,1962,1983]" captionTargetBox="[103,1484,241,1880]" captionTargetId="figure-2@4.[103,1494,221,1908]" captionTargetPageId="4" captionText="Figure 1 Leobodes becki n. sp., adult: a – dorsal view (legs omitted); b – ventral view (gnathosoma and legs omitted); c – lateral view (gnathosoma and legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7159894" httpUri="https://zenodo.org/record/7159894/files/figure.png" pageId="3" pageNumber="997">Figures 1</figureCitation>
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a-c, 2a, 2c, 2d, 3a-d, 4a-d, 5a-d, 6a, 6b)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="997" type="diagnosis">
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<paragraph blockId="3.[491,1544,657,2045]" pageId="3" pageNumber="997">
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<emphasis bold="true" box="[531,640,722,746]" pageId="3" pageNumber="997">Diagnosis</emphasis>
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— Adult. Body size: 531–630 × 332–431. Body surface with dense granulate microsculpturing; foveolate ornamentation absent. Prodorsal processes connected medially at a relatively large distance; their anterior parts forming an oval, transversely oriented aperture located anterior to connecting region. Rostral, lamellar and interlamellar setae slightly thickened;
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<emphasis box="[610,631,859,882]" italics="true" pageId="3" pageNumber="997">in</emphasis>
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shortest, directed posterolaterad. Bothridial seta with distinctly lanceolate head having short attenuate tip. Anteromedian part of notogaster depressed. Notogastral tubercle absent. Distance between medial notogastral condyles comparatively long. Ten pairs of notogastral setae present: seta
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<emphasis box="[823,835,962,985]" italics="true" pageId="3" pageNumber="997">c</emphasis>
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long, with slightly phylliform basal part and bent posteriad attenuate mediodistal part; other setae short, thickened. Subcapitular seta
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<emphasis box="[1287,1300,995,1019]" italics="true" pageId="3" pageNumber="997">h</emphasis>
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setiform. Seta
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<emphasis box="[1468,1481,995,1019]" italics="true" pageId="3" pageNumber="997">l′</emphasis>
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</paragraph>
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<paragraph blockId="3.[491,1544,657,2045]" pageId="3" pageNumber="997">
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on trochanter III of medium length, setiform;
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<emphasis box="[979,997,1030,1054]" italics="true" pageId="3" pageNumber="997">v′</emphasis>
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on leg trochanters III, IV comparatively short, setiform.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="999" pageId="3" pageNumber="997" type="description">
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<paragraph blockId="3.[491,1544,657,2045]" box="[531,1489,1098,1122]" pageId="3" pageNumber="997">
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<emphasis bold="true" box="[531,758,1098,1122]" pageId="3" pageNumber="997">Description of adult</emphasis>
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— Measurements – Body length: 630 (
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<typeStatus box="[1195,1295,1098,1122]" pageId="3" pageNumber="997">holotype</typeStatus>
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: female), 531–630
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</paragraph>
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<paragraph blockId="3.[491,1544,657,2045]" box="[491,1500,1132,1157]" pageId="3" pageNumber="997">
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(
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<specimenCount box="[499,636,1132,1157]" pageId="3" pageNumber="997" type="generic" typeStatus="paratypes">16 paratypes</specimenCount>
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:
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<specimenCount box="[647,745,1132,1156]" pageId="3" pageNumber="997" type="male">six males</specimenCount>
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and
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<specimenCount box="[802,922,1132,1156]" pageId="3" pageNumber="997" type="female">10 females</specimenCount>
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); body width: 415 (
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<typeStatus box="[1144,1244,1132,1156]" pageId="3" pageNumber="997">holotype</typeStatus>
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), 332–431 (
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<specimenCount box="[1351,1489,1132,1156]" pageId="3" pageNumber="997" type="generic" typeStatus="paratypes">16 paratypes</specimenCount>
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).
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</paragraph>
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<paragraph blockId="3.[491,1544,657,2045]" pageId="3" pageNumber="997">Integument – Body color brown. Body surface with dense granulate microsculpturing. Prodorsal processes close to prodorsal aperture sparsely tuberculate (diameter of tubercle up to 10). Region between bothridium and acetabula I–III densely tuberculate (diameter of tubercle up to 4). Region between bothridium and pedotecta I with some longitudinal folds.</paragraph>
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<paragraph blockId="3.[491,1544,657,2045]" pageId="3" pageNumber="997">Prodorsum – Rostrum rounded. Lateral carinae fused medially before rostral margin. Lamella slightly developed. Prolamella present, but inconspicuous. Tutorium and prodorsal hollow of prodorsum well developed. Prodorsal processes connected medially at a relatively large distance. Anterior parts of prodorsal processes forming oval, transversely oriented prodorsal aperture located anteriorly to connecting region. Rostral (86–90), lamellar (90–94)</paragraph>
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<paragraph blockId="3.[491,1544,657,2045]" box="[491,1468,1474,1498]" pageId="3" pageNumber="997">
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and interlamellar (57–61) setae slightly thickened, barbed;
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<emphasis box="[1139,1162,1475,1498]" italics="true" pageId="3" pageNumber="997">ro</emphasis>
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with slightly attenuate tip;
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</paragraph>
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<paragraph blockId="3.[491,1544,657,2045]" pageId="3" pageNumber="997">
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<emphasis box="[491,510,1508,1532]" italics="true" pageId="3" pageNumber="997">le</emphasis>
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inserted on distinct tubercle;
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<emphasis box="[836,857,1509,1532]" italics="true" pageId="3" pageNumber="997">in</emphasis>
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directed posterolaterad (in dorsal view). Distance
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<emphasis box="[1413,1466,1508,1532]" italics="true" pageId="3" pageNumber="997">in–le</emphasis>
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distinctly longer than
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<emphasis box="[730,785,1543,1567]" italics="true" pageId="3" pageNumber="997">le–ro</emphasis>
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. Bothridial seta (98–106) with long stalk and short, distinctly lanceolate head having short attenuate tip, roughened.
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</paragraph>
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<paragraph blockId="3.[491,1544,657,2045]" pageId="3" pageNumber="997">
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Notogaster – Anteromedian part of notogaster depressed, fused with prodorsal hollow. Notogastral tubercle absent. Distance between medial notogastral condyles comparatively long (about as long as length between setae
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<emphasis box="[915,982,1679,1703]" italics="true" pageId="3" pageNumber="997">lm–lm</emphasis>
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). Ten pairs of notogastral setae present: seta
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</paragraph>
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<paragraph blockId="3.[491,1544,657,2045]" pageId="3" pageNumber="997">
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<emphasis box="[491,503,1715,1738]" italics="true" pageId="3" pageNumber="997">c</emphasis>
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(123–131) with slightly phylliform basal part and bent posteriad attenuate mediodistal part, barbed; other setae (32–41) thickened, barbed. Opisthonotal gland opening and all lyrifissures distinct.
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</paragraph>
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<paragraph blockId="3.[491,1544,657,2045]" pageId="3" pageNumber="997">
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Gnathosoma – Subcapitulum size: 135–139 × 114–123. Subcapitular setae (
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<emphasis box="[1350,1363,1817,1840]" italics="true" pageId="3" pageNumber="997">a</emphasis>
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: 12–14;
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<emphasis box="[1463,1482,1817,1840]" italics="true" pageId="3" pageNumber="997">m</emphasis>
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: 45–49;
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<emphasis box="[571,584,1850,1874]" italics="true" pageId="3" pageNumber="997">h</emphasis>
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: 36–41) with attenuate tip, slightly barbed;
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<emphasis box="[1050,1069,1851,1874]" italics="true" pageId="3" pageNumber="997">a</emphasis>
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thinnest. Palp length: 77–82. Postpalpal seta (8) spiniform, roughened. Chelicera length: 155–164. Cheliceral setae
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<emphasis box="[1311,1349,1885,1909]" italics="true" pageId="3" pageNumber="997">cha</emphasis>
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(: 49–53;
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<emphasis box="[1448,1486,1885,1909]" italics="true" pageId="3" pageNumber="997">chb</emphasis>
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: 20–24) setiform, barbed;
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<emphasis box="[762,807,1919,1943]" italics="true" pageId="3" pageNumber="997">cha</emphasis>
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thicker than
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<emphasis box="[939,977,1919,1943]" italics="true" pageId="3" pageNumber="997">chb</emphasis>
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.
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</paragraph>
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<paragraph blockId="3.[491,1544,657,2045]" box="[531,1470,1953,1977]" pageId="3" pageNumber="997">
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Epimeral and lateral podosomal regions – Epimeral setae
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<emphasis box="[1161,1173,1953,1977]" italics="true" pageId="3" pageNumber="997">1</emphasis>
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(
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<emphasis box="[1173,1188,1953,1977]" italics="true" pageId="3" pageNumber="997">b</emphasis>
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,
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<emphasis box="[1202,1227,1953,1977]" italics="true" pageId="3" pageNumber="997">3c</emphasis>
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: 32–41; others: 28–36)
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</paragraph>
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<paragraph blockId="3.[491,1544,657,2045]" pageId="3" pageNumber="997">thin, with attenuate tip, slightly barbed. Ventrolateral tubercle on epimere II distinct. Discidium slightly developed.</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7159894" ID-Zenodo-Dep="7159894" httpUri="https://zenodo.org/record/7159894/files/figure.png" pageId="4" pageNumber="998" startId="4.[119,186,1962,1983]" targetBox="[103,1484,241,1880]" targetPageId="4">
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<paragraph blockId="4.[118,1525,1962,2006]" pageId="4" pageNumber="998">
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<emphasis bold="true" box="[119,207,1962,1983]" pageId="4" pageNumber="998">Figure 1</emphasis>
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<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[224,374,1962,1984]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="998" phylum="Arthropoda" rank="species" species="becki" status="n. sp.">
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<emphasis box="[224,374,1962,1984]" italics="true" pageId="4" pageNumber="998">Leobodes becki</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[382,439,1962,1983]" pageId="4" pageNumber="998">
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<taxonomicNameLabel box="[382,439,1962,1983]" pageId="4" pageNumber="998" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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, adult: a – dorsal view (legs omitted); b – ventral view (gnathosoma and legs omitted); c – lateral view (gnathosoma and legs omitted). Scale bar 100 μm.
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</paragraph>
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</caption>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7159896" ID-Zenodo-Dep="7159896" httpUri="https://zenodo.org/record/7159896/files/figure.png" pageId="5" pageNumber="999" startId="5.[119,186,1069,1090]" targetBox="[136,1477,251,998]" targetPageId="5">
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<paragraph blockId="5.[119,1488,1068,1135]" pageId="5" pageNumber="999">
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<emphasis bold="true" box="[119,205,1069,1090]" pageId="5" pageNumber="999">Figure 2</emphasis>
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Gnathosoma of
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<taxonomicName box="[372,512,1068,1090]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="999" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis box="[372,464,1068,1090]" italics="true" pageId="5" pageNumber="999">Leobodes</emphasis>
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spp.
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</taxonomicName>
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, dissected adults: a – subcapitulum of
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<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[886,962,1068,1090]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="999" phylum="Arthropoda" rank="species" species="becki" status="n. sp.">
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<emphasis box="[886,962,1068,1090]" italics="true" pageId="5" pageNumber="999">L. becki</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[969,1024,1069,1090]" pageId="5" pageNumber="999">
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<taxonomicNameLabel box="[969,1024,1069,1090]" pageId="5" pageNumber="999" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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, ventral view; b – subcapitulum of
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<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[1368,1486,1068,1090]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="999" phylum="Arthropoda" rank="species" species="schalleri" status="n. sp.">
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<emphasis box="[1368,1486,1068,1090]" italics="true" pageId="5" pageNumber="999">L. schalleri</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[119,173,1091,1112]" pageId="5" pageNumber="999">
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<taxonomicNameLabel box="[119,173,1091,1112]" pageId="5" pageNumber="999" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
|
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, ventral view; c – palp of
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<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[423,498,1091,1113]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="999" phylum="Arthropoda" rank="species" species="becki" status="n. sp.">
|
||
<emphasis box="[423,498,1091,1113]" italics="true" pageId="5" pageNumber="999">L. becki</emphasis>
|
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</taxonomicName>
|
||
<emphasis bold="true" box="[505,559,1091,1112]" pageId="5" pageNumber="999">
|
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<taxonomicNameLabel box="[505,559,1091,1112]" pageId="5" pageNumber="999" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
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, left, paraxial view; d – chelicera of
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<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[910,985,1091,1113]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="999" phylum="Arthropoda" rank="species" species="becki" status="n. sp.">
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<emphasis box="[910,985,1091,1113]" italics="true" pageId="5" pageNumber="999">L. becki</emphasis>
|
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</taxonomicName>
|
||
<emphasis bold="true" box="[991,1046,1091,1112]" pageId="5" pageNumber="999">
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||
<taxonomicNameLabel box="[991,1046,1091,1112]" pageId="5" pageNumber="999" rank="species">n. sp.</taxonomicNameLabel>
|
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</emphasis>
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, right, antiaxial view. Scale bars 20 μm (a, b,
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</paragraph>
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</caption>
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<paragraph blockId="5.[119,1488,1068,1135]" box="[119,256,1113,1135]" pageId="5" pageNumber="999">d), 10 μm (c).</paragraph>
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<paragraph blockId="5.[491,1551,1253,2037]" pageId="5" pageNumber="999">Anogenital region – Genital seta (24–28) thin, with attenuate tip, slightly barbed. Aggenital, anal and adanal setae (28–36) thickened, barbed. Adanal lyrifissure distinct.</paragraph>
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<paragraph blockId="5.[491,1551,1253,2037]" pageId="5" pageNumber="999">Legs – Claw of all legs strong, inconspicuously roughened on dorsal side. Dorsoparaxial porose area on femora I–IV distinct; porose area on trochanters III, IV not observed. Formulas of leg setation and solenidia: I (1–4–2–4–16) [1–2–2], II (1–4–2–2–15) [1–1–2], III (2–3–0–</paragraph>
|
||
<paragraph blockId="5.[491,1551,1253,2037]" pageId="5" pageNumber="999">
|
||
2–15) [1–1–0], IV (1–2–1–2–12) [0–1–0]; homology of setae and solenidia indicated in
|
||
<tableCitation captionStart="Table 1" captionStartId="10.[119,175,248,269]" captionText="Table 1 Leg setation and solenidia of adultLeobodes becki n. sp., L. schallerin. sp. andL. schawallerin. sp." pageId="5" pageNumber="999">Table 1</tableCitation>
|
||
. Famulus on tarsus I short, erect, slightly swollen distally, inserted posterior to solenidion
|
||
<subScript attach="right" box="[1484,1493,1473,1490]" fontSize="7" pageId="5" pageNumber="999">1</subScript>
|
||
ω. Seta
|
||
<emphasis box="[545,555,1496,1519]" italics="true" pageId="5" pageNumber="999">s</emphasis>
|
||
on tarsus I eupathidial, located before setae
|
||
<emphasis box="[1033,1061,1495,1519]" italics="true" pageId="5" pageNumber="999">a′</emphasis>
|
||
and
|
||
<emphasis box="[1106,1131,1495,1519]" italics="true" pageId="5" pageNumber="999">a″</emphasis>
|
||
. Setae
|
||
<emphasis box="[1213,1232,1495,1519]" italics="true" pageId="5" pageNumber="999">u′</emphasis>
|
||
,
|
||
<emphasis box="[1246,1270,1495,1519]" italics="true" pageId="5" pageNumber="999">u″</emphasis>
|
||
on tarsi I and
|
||
<emphasis box="[1427,1451,1495,1519]" italics="true" pageId="5" pageNumber="999">u″</emphasis>
|
||
on tarsus II spiniform;
|
||
<emphasis box="[702,721,1530,1554]" italics="true" pageId="5" pageNumber="999">u′</emphasis>
|
||
on tarsus II and
|
||
<emphasis box="[901,920,1530,1554]" italics="true" pageId="5" pageNumber="999">u′</emphasis>
|
||
,
|
||
<emphasis box="[934,958,1530,1554]" italics="true" pageId="5" pageNumber="999">u″</emphasis>
|
||
on tarsi III, IV spiniform. Seta
|
||
<emphasis box="[1299,1312,1530,1554]" italics="true" pageId="5" pageNumber="999">l′</emphasis>
|
||
on trochanter III of medium length, setiform;
|
||
<emphasis box="[796,814,1564,1588]" italics="true" pageId="5" pageNumber="999">v′</emphasis>
|
||
on trochanters III, IV comparatively short, setiform.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="999" type="materials_examined">
|
||
<paragraph blockId="5.[491,1551,1253,2037]" pageId="5" pageNumber="999">
|
||
<emphasis bold="true" box="[531,746,1599,1623]" pageId="5" pageNumber="999">Material examined</emphasis>
|
||
—
|
||
<collectingCountry box="[786,852,1599,1623]" name="Nepal" pageId="5" pageNumber="999">Nepal</collectingCountry>
|
||
, Sankhua Sabha District, Arun Valley, Chichila,
|
||
<geoCoordinate box="[1388,1487,1599,1623]" degrees="27" direction="north" minutes="27,02" orientation="latitude" pageId="5" pageNumber="999" precision="9" value="27.450333">27°27,02′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[491,594,1633,1657]" degrees="87" direction="east" minutes="13,13" orientation="longitude" pageId="5" pageNumber="999" precision="9" value="87.218834">87°13,13′E</geoCoordinate>
|
||
,
|
||
<quantity box="[605,742,1633,1657]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="5" pageNumber="999" unit="m" value="1950.0" valueMax="2000.0" valueMin="1900.0">1900–2000 m</quantity>
|
||
a.s.l., leaf litter and upper layer of soil under bushes in degraded
|
||
<taxonomicName box="[491,580,1668,1691]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="999" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[491,580,1668,1691]" italics="true" pageId="5" pageNumber="999">Quercus</emphasis>
|
||
</taxonomicName>
|
||
forest near village (sample No. 414),
|
||
<date box="[993,1130,1668,1692]" pageId="5" pageNumber="999" value="1988-06-18" valueMax="1988-06-20" valueMin="1988-06-18">
|
||
18–20.
|
||
<collectionCode box="[1054,1080,1668,1692]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="999">VI</collectionCode>
|
||
.1988
|
||
</date>
|
||
(collected by J. Martens and W. Schawaller).
|
||
</paragraph>
|
||
<paragraph blockId="5.[491,1551,1253,2037]" pageId="5" pageNumber="999">
|
||
<emphasis bold="true" box="[531,711,1737,1761]" pageId="5" pageNumber="999">Type deposition</emphasis>
|
||
— The
|
||
<typeStatus box="[807,901,1737,1761]" pageId="5" pageNumber="999">holotype</typeStatus>
|
||
and
|
||
<specimenCount box="[961,1094,1738,1762]" pageId="5" pageNumber="999" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
||
are deposited in the collection of the
|
||
<collectionCode box="[491,571,1771,1795]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="5" pageNumber="999">SMNH</collectionCode>
|
||
;
|
||
<specimenCount box="[582,712,1771,1796]" pageId="5" pageNumber="999" type="generic" typeStatus="paratypes">15 paratypes</specimenCount>
|
||
are deposited in the collection of the
|
||
<collectionCode box="[1136,1230,1771,1795]" pageId="5" pageNumber="999">TSUMZ</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="999" type="etymology">
|
||
<paragraph blockId="5.[491,1551,1253,2037]" pageId="5" pageNumber="999">
|
||
<emphasis bold="true" box="[531,654,1806,1830]" pageId="5" pageNumber="999">Etymology</emphasis>
|
||
— The specific name is dedicated to Prof. Dr. Ludwig Beck (State Museum of Natural History, Karlsruhe,
|
||
<collectingCountry box="[786,885,1840,1864]" name="Germany" pageId="5" pageNumber="999">Germany</collectingCountry>
|
||
), the well-known acarologist.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="10" lastPageNumber="1004" pageId="5" pageNumber="999" type="discussion">
|
||
<paragraph blockId="5.[491,1551,1253,2037]" pageId="5" pageNumber="999">
|
||
<emphasis bold="true" box="[531,634,1875,1899]" pageId="5" pageNumber="999">Remarks</emphasis>
|
||
— In main morphological traits (prodorsal processes have prodorsal aperture; notogastral seta
|
||
<emphasis box="[664,676,1910,1933]" italics="true" pageId="5" pageNumber="999">c</emphasis>
|
||
setiform, with mediodistal part bent and directed posteriad, other notogastral setae thickened; distance between medial prodorsal condyles equal
|
||
<emphasis box="[1241,1268,1944,1968]" italics="true" pageId="5" pageNumber="999">lm</emphasis>
|
||
to–
|
||
<emphasis box="[1281,1308,1944,1968]" italics="true" pageId="5" pageNumber="999">lm</emphasis>
|
||
)
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[1323,1488,1944,1968]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="999" phylum="Arthropoda" rank="species" species="becki" status="n. sp.">
|
||
<emphasis box="[1323,1488,1944,1968]" italics="true" pageId="5" pageNumber="999">Leobodes becki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[491,551,1979,2003]" pageId="5" pageNumber="999">
|
||
<taxonomicNameLabel box="[491,551,1979,2003]" pageId="5" pageNumber="999" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is morphologically most similar to
|
||
<taxonomicName authority="Chen and Wang, 2007" authorityName="Chen and Wang" authorityYear="2007" box="[932,1435,1979,2003]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="999" phylum="Arthropoda" rank="species" species="praeconcavus">
|
||
<emphasis box="[932,1194,1979,2003]" italics="true" pageId="5" pageNumber="999">Leobodes praeconcavus</emphasis>
|
||
<bibRefCitation author="Chen J. & Wang H. F." box="[1194,1435,1979,2003]" pageId="5" pageNumber="999" pagination="45 - 63" refId="ref7356" refString="Chen J., Wang H. F. 2007. Taxonomic study on the family Nippobodidae (Acari: Oribatida) from China. Zootaxa, 1464: 45 - 63. https: // doi. org / 10.11646 / zootaxa. 1464.1.2" type="journal article" year="2007">Chen and Wang, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Aoki, 2000" authorityName="Aoki" authorityYear="2000" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="999" phylum="Arthropoda" rank="species" species="yinae">
|
||
<emphasis italics="true" pageId="5" pageNumber="999">L. yinae</emphasis>
|
||
<bibRefCitation author="Aoki J." box="[556,674,2013,2037]" pageId="5" pageNumber="999" pagination="1 - 6" refId="ref7284" refString="Aoki J. 2000. Two new species of the family Nippobodidae (Acari: Oribatida) from Yunnan, southwest China, with a key to the species. In: Aoki J., Yin W., Imadate G. (Eds). Taxonomical Studies on the Soil Fauna of Yunnan Province in Southwest China. Tokyo: Tokai University Press. pp. 1 - 6." type="book chapter" year="2000">Aoki, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
. However, the new species differs from both by the presence of interlamellar
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7159900" ID-Zenodo-Dep="7159900" httpUri="https://zenodo.org/record/7159900/files/figure.png" pageId="6" pageNumber="1000" startId="6.[119,186,1985,2006]" targetBox="[220,1366,247,1866]" targetPageId="6">
|
||
<paragraph blockId="6.[118,1486,1985,2075]" pageId="6" pageNumber="1000">
|
||
<emphasis bold="true" box="[119,205,1985,2006]" pageId="6" pageNumber="1000">Figure 3</emphasis>
|
||
Legs of
|
||
<taxonomicName box="[297,437,1985,2007]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="1000" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis box="[297,389,1985,2007]" italics="true" pageId="6" pageNumber="1000">Leobodes</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
, dissected adults: a – leg I of
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[725,801,1985,2007]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="1000" phylum="Arthropoda" rank="species" species="becki" status="n. sp.">
|
||
<emphasis box="[725,801,1985,2007]" italics="true" pageId="6" pageNumber="1000">L. becki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[807,862,1985,2006]" pageId="6" pageNumber="1000">
|
||
<taxonomicNameLabel box="[807,862,1985,2006]" pageId="6" pageNumber="1000" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, right, antiaxial view; b – leg II of
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[1201,1277,1985,2007]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="1000" phylum="Arthropoda" rank="species" species="becki" status="n. sp.">
|
||
<emphasis box="[1201,1277,1985,2007]" italics="true" pageId="6" pageNumber="1000">L. becki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1284,1338,1985,2006]" pageId="6" pageNumber="1000">
|
||
<taxonomicNameLabel box="[1284,1338,1985,2006]" pageId="6" pageNumber="1000" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, left, antiaxial view; c – leg III of
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[307,383,2007,2029]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="1000" phylum="Arthropoda" rank="species" species="becki" status="n. sp.">
|
||
<emphasis box="[307,383,2007,2029]" italics="true" pageId="6" pageNumber="1000">L. becki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[390,445,2008,2029]" pageId="6" pageNumber="1000">
|
||
<taxonomicNameLabel box="[390,445,2008,2029]" pageId="6" pageNumber="1000" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, right, paraxial view; d – leg IV of
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[791,867,2007,2029]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="1000" phylum="Arthropoda" rank="species" species="becki" status="n. sp.">
|
||
<emphasis box="[791,867,2007,2029]" italics="true" pageId="6" pageNumber="1000">L. becki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[874,929,2008,2029]" pageId="6" pageNumber="1000">
|
||
<taxonomicNameLabel box="[874,929,2008,2029]" pageId="6" pageNumber="1000" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, left, antiaxial view; e – leg trochanter III of
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[1368,1486,2007,2029]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="1000" phylum="Arthropoda" rank="species" species="schalleri" status="n. sp.">
|
||
<emphasis box="[1368,1486,2007,2029]" italics="true" pageId="6" pageNumber="1000">L. schalleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[119,174,2030,2051]" pageId="6" pageNumber="1000">
|
||
<taxonomicNameLabel box="[119,174,2030,2051]" pageId="6" pageNumber="1000" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, left, antiaxial view; f – leg trochanter IV of
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[615,732,2030,2052]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="1000" phylum="Arthropoda" rank="species" species="schalleri" status="n. sp.">
|
||
<emphasis box="[615,732,2030,2052]" italics="true" pageId="6" pageNumber="1000">L. schalleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[731,787,2030,2051]" pageId="6" pageNumber="1000">
|
||
<taxonomicNameLabel box="[731,787,2030,2051]" pageId="6" pageNumber="1000" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, left, antiaxial view; g – leg trochanter III of
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[1230,1375,2030,2052]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="1000" phylum="Arthropoda" rank="species" species="schawalleri" status="n. sp.">
|
||
<emphasis box="[1230,1375,2030,2052]" italics="true" pageId="6" pageNumber="1000">L. schawalleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1374,1430,2030,2051]" pageId="6" pageNumber="1000">
|
||
<taxonomicNameLabel box="[1374,1430,2030,2051]" pageId="6" pageNumber="1000" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, left, antiaxial view; h – leg trochanter IV of
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[497,641,2053,2075]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="1000" phylum="Arthropoda" rank="species" species="schawalleri" status="n. sp.">
|
||
<emphasis box="[497,641,2053,2075]" italics="true" pageId="6" pageNumber="1000">L. schawalleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[640,695,2053,2074]" pageId="6" pageNumber="1000">
|
||
<taxonomicNameLabel box="[640,695,2053,2074]" pageId="6" pageNumber="1000" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, left, antiaxial view. Scale bar 50 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7159902" ID-Zenodo-Dep="7159902" box="[119,1274,1895,1917]" httpUri="https://zenodo.org/record/7159902/files/figure.png" pageId="7" pageNumber="1001" startId="7.[119,186,1896,1917]" targetBox="[103,1494,287,1840]" targetPageId="7">
|
||
<paragraph blockId="7.[119,1274,1895,1917]" box="[119,1274,1895,1917]" pageId="7" pageNumber="1001">
|
||
<emphasis bold="true" box="[119,204,1896,1917]" pageId="7" pageNumber="1001">Figure 4</emphasis>
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[216,364,1895,1917]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="1001" phylum="Arthropoda" rank="species" species="becki" status="n. sp.">
|
||
<emphasis box="[216,364,1895,1917]" italics="true" pageId="7" pageNumber="1001">Leobodes becki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[370,425,1896,1917]" pageId="7" pageNumber="1001">
|
||
<taxonomicNameLabel box="[370,425,1896,1917]" pageId="7" pageNumber="1001" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, adult, SEM micrographs: a, b – dorsal view; c – ventral view; d – ventrolateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7159906" ID-Zenodo-Dep="7159906" httpUri="https://zenodo.org/record/7159906/files/figure.png" pageId="8" pageNumber="1002" startId="8.[119,186,1887,1908]" targetBox="[103,1494,278,1830]" targetPageId="8">
|
||
<paragraph blockId="8.[118,1505,1886,1931]" pageId="8" pageNumber="1002">
|
||
<emphasis bold="true" box="[119,204,1887,1908]" pageId="8" pageNumber="1002">Figure 5</emphasis>
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[216,364,1886,1908]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="1002" phylum="Arthropoda" rank="species" species="becki" status="n. sp.">
|
||
<emphasis box="[216,364,1886,1908]" italics="true" pageId="8" pageNumber="1002">Leobodes becki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[371,425,1887,1908]" pageId="8" pageNumber="1002">
|
||
<taxonomicNameLabel box="[371,425,1887,1908]" pageId="8" pageNumber="1002" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, adult, SEM micrographs: a – dorsoanterior view; b – lateral view; c – ventroanterior view; d – subcapitulum, ventrolateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7159912" ID-Zenodo-Dep="7159912" httpUri="https://zenodo.org/record/7159912/files/figure.png" pageId="9" pageNumber="1003" startId="9.[119,186,1704,1725]" targetBox="[103,1494,221,1647]" targetPageId="9">
|
||
<paragraph blockId="9.[118,1487,1704,1771]" pageId="9" pageNumber="1003">
|
||
<emphasis bold="true" box="[119,205,1704,1725]" pageId="9" pageNumber="1003">Figure 6</emphasis>
|
||
<taxonomicName box="[219,359,1704,1726]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="1003" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis box="[219,311,1704,1726]" italics="true" pageId="9" pageNumber="1003">Leobodes</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
, adult, microscope images: a –
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[668,744,1704,1726]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="1003" phylum="Arthropoda" rank="species" species="becki" status="n. sp.">
|
||
<emphasis box="[668,744,1704,1726]" italics="true" pageId="9" pageNumber="1003">L. becki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[751,806,1704,1725]" pageId="9" pageNumber="1003">
|
||
<taxonomicNameLabel box="[751,806,1704,1725]" pageId="9" pageNumber="1003" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, dorsal view; b –
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[982,1058,1704,1726]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="1003" phylum="Arthropoda" rank="species" species="becki" status="n. sp.">
|
||
<emphasis box="[982,1058,1704,1726]" italics="true" pageId="9" pageNumber="1003">L. becki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1065,1120,1704,1725]" pageId="9" pageNumber="1003">
|
||
<taxonomicNameLabel box="[1065,1120,1704,1725]" pageId="9" pageNumber="1003" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, lateral view; c –
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[1296,1414,1704,1726]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="1003" phylum="Arthropoda" rank="species" species="schalleri" status="n. sp.">
|
||
<emphasis box="[1296,1414,1704,1726]" italics="true" pageId="9" pageNumber="1003">L. schalleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1420,1475,1704,1725]" pageId="9" pageNumber="1003">
|
||
<taxonomicNameLabel box="[1420,1475,1704,1725]" pageId="9" pageNumber="1003" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, dorsal view; d –
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[282,398,1726,1748]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="1003" phylum="Arthropoda" rank="species" species="schalleri" status="n. sp.">
|
||
<emphasis box="[282,398,1726,1748]" italics="true" pageId="9" pageNumber="1003">L. schalleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[398,453,1727,1748]" pageId="9" pageNumber="1003">
|
||
<taxonomicNameLabel box="[398,453,1727,1748]" pageId="9" pageNumber="1003" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, lateral view; e –
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[628,772,1726,1748]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="1003" phylum="Arthropoda" rank="species" species="schawalleri" status="n. sp.">
|
||
<emphasis box="[628,772,1726,1748]" italics="true" pageId="9" pageNumber="1003">L. schawalleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[773,828,1727,1748]" pageId="9" pageNumber="1003">
|
||
<taxonomicNameLabel box="[773,828,1727,1748]" pageId="9" pageNumber="1003" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, dorsal view; f –
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[999,1143,1726,1748]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="1003" phylum="Arthropoda" rank="species" species="schawalleri" status="n. sp.">
|
||
<emphasis box="[999,1143,1726,1748]" italics="true" pageId="9" pageNumber="1003">L. schawalleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1143,1198,1727,1748]" pageId="9" pageNumber="1003">
|
||
<taxonomicNameLabel box="[1143,1198,1727,1748]" pageId="9" pageNumber="1003" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, lateral view. Magnification:
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="9.[118,1487,1704,1771]" box="[118,256,1749,1771]" pageId="9" pageNumber="1003">400 (10 × 40).</paragraph>
|
||
<paragraph blockId="9.[491,1543,1889,2016]" lastBlockId="10.[112,1428,547,643]" lastPageId="10" lastPageNumber="1004" pageId="9" pageNumber="1003">
|
||
seta of medium length, distinctly shorter than rostral and lamellar setae (versus long, not shorter than rostral and lamellar setae) directed posterolaterad (versus erect) in dorsal view. Also,
|
||
<taxonomicName authorityName="Chen and Wang" authorityYear="2007" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="1003" phylum="Arthropoda" rank="species" species="praeconcavus">
|
||
<emphasis italics="true" pageId="9" pageNumber="1003">L. praeconcavus</emphasis>
|
||
</taxonomicName>
|
||
has smooth (versus barbed) notogastral setae (except
|
||
<emphasis box="[1221,1233,1959,1982]" italics="true" pageId="9" pageNumber="1003">c</emphasis>
|
||
) of medium length,
|
||
<emphasis box="[1454,1481,1958,1982]" italics="true" pageId="9" pageNumber="1003">lm</emphasis>
|
||
reaching insertion
|
||
<emphasis box="[689,710,1992,2016]" italics="true" pageId="9" pageNumber="1003">lp</emphasis>
|
||
(versus short,
|
||
<emphasis box="[867,894,1992,2016]" italics="true" pageId="9" pageNumber="1003">lm</emphasis>
|
||
not reaching insertion
|
||
<emphasis box="[1139,1160,1992,2016]" italics="true" pageId="9" pageNumber="1003">lp</emphasis>
|
||
);
|
||
<taxonomicName authorityName="Aoki" authorityYear="2000" box="[1182,1275,1993,2016]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="1003" phylum="Arthropoda" rank="species" species="yinae">
|
||
<emphasis box="[1182,1275,1993,2016]" italics="true" pageId="9" pageNumber="1003">L. yinae</emphasis>
|
||
</taxonomicName>
|
||
has bifurcate (versus Note: Roman letters refer to normal setae (except ɛ = famulus); Greek letters refer to solenidia. Single quotation mark (′)
|
||
</paragraph>
|
||
<caption box="[119,1175,247,270]" pageId="10" pageNumber="1004" startId="10.[119,175,248,269]">
|
||
<paragraph blockId="10.[119,1175,247,270]" box="[119,1175,247,270]" pageId="10" pageNumber="1004">
|
||
<emphasis bold="true" box="[119,193,248,269]" pageId="10" pageNumber="1004">Table 1</emphasis>
|
||
Leg setation and solenidia of adult
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[541,689,247,269]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="1004" phylum="Arthropoda" rank="species" species="becki" status="n. sp.">
|
||
<emphasis box="[541,689,247,269]" italics="true" pageId="10" pageNumber="1004">Leobodes becki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[695,749,248,269]" pageId="10" pageNumber="1004">
|
||
<taxonomicNameLabel box="[695,749,248,269]" pageId="10" pageNumber="1004" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[761,876,247,269]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="1004" phylum="Arthropoda" rank="species" species="schalleri" status="n. sp.">
|
||
<emphasis box="[761,876,247,269]" italics="true" pageId="10" pageNumber="1004">L. schalleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[876,930,248,269]" pageId="10" pageNumber="1004">
|
||
<taxonomicNameLabel box="[876,930,248,269]" pageId="10" pageNumber="1004" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName authority="Ermilov & Martens, 2021" authorityName="Ermilov & Martens" authorityYear="2021" box="[978,1121,247,269]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="1004" phylum="Arthropoda" rank="species" species="schawalleri" status="n. sp.">
|
||
<emphasis box="[978,1121,247,269]" italics="true" pageId="10" pageNumber="1004">L. schawalleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1121,1175,248,269]" pageId="10" pageNumber="1004" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="10" pageNumber="1004">
|
||
<table box="[112,1408,343,537]" gridcols="10" gridrows="5" pageId="10" pageNumber="1004">
|
||
<tr box="[112,1408,343,369]" gridrow="0" pageId="10" pageNumber="1004" rowspan-4="1" rowspan-6="1" rowspan-8="1" rowspan-9="1">
|
||
<th box="[112,159,343,369]" gridcol="0" gridrow="0" pageId="10" pageNumber="1004">Leg</th>
|
||
<th box="[192,242,343,369]" gridcol="1" gridrow="0" pageId="10" pageNumber="1004">
|
||
<emphasis box="[192,221,343,368]" italics="true" pageId="10" pageNumber="1004">Tr</emphasis>
|
||
</th>
|
||
<th box="[304,420,343,369]" gridcol="2" gridrow="0" pageId="10" pageNumber="1004">
|
||
<emphasis box="[304,336,343,368]" italics="true" pageId="10" pageNumber="1004">Fe</emphasis>
|
||
</th>
|
||
<th box="[479,537,343,369]" gridcol="3" gridrow="0" pageId="10" pageNumber="1004">
|
||
<emphasis box="[479,515,343,369]" italics="true" pageId="10" pageNumber="1004">Ge</emphasis>
|
||
</th>
|
||
<th box="[622,718,343,369]" gridcol="5" gridrow="0" pageId="10" pageNumber="1004">
|
||
<emphasis box="[622,648,343,368]" italics="true" pageId="10" pageNumber="1004">Ti</emphasis>
|
||
</th>
|
||
<th box="[849,1221,343,369]" gridcol="7" gridrow="0" pageId="10" pageNumber="1004">
|
||
<emphasis box="[849,881,343,368]" italics="true" pageId="10" pageNumber="1004">Ta</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[112,1408,382,414]" gridrow="1" pageId="10" pageNumber="1004">
|
||
<th box="[112,159,382,414]" gridcol="0" gridrow="1" pageId="10" pageNumber="1004">I</th>
|
||
<td box="[192,242,382,414]" gridcol="1" gridrow="1" pageId="10" pageNumber="1004">
|
||
<emphasis box="[192,212,383,411]" italics="true" pageId="10" pageNumber="1004">v′</emphasis>
|
||
</td>
|
||
<td box="[304,420,382,414]" gridcol="2" gridrow="1" pageId="10" pageNumber="1004">
|
||
<emphasis box="[304,420,383,411]" italics="true" pageId="10" pageNumber="1004">d, (l), bv″</emphasis>
|
||
</td>
|
||
<td box="[479,537,382,414]" gridcol="3" gridrow="1" pageId="10" pageNumber="1004">
|
||
<emphasis box="[479,537,383,411]" italics="true" pageId="10" pageNumber="1004">l′, v′,</emphasis>
|
||
</td>
|
||
<td box="[551,567,382,414]" gridcol="4" gridrow="1" pageId="10" pageNumber="1004">σ</td>
|
||
<td box="[622,718,382,414]" gridcol="5" gridrow="1" pageId="10" pageNumber="1004">
|
||
<emphasis box="[622,650,382,408]" italics="true" pageId="10" pageNumber="1004">(l)</emphasis>
|
||
,
|
||
<emphasis box="[671,705,383,409]" italics="true" pageId="10" pageNumber="1004">(v)</emphasis>
|
||
,
|
||
</td>
|
||
<td box="[731,801,382,414]" gridcol="6" gridrow="1" pageId="10" pageNumber="1004">φ1, φ2</td>
|
||
<td box="[849,1221,382,414]" gridcol="7" gridrow="1" pageId="10" pageNumber="1004">
|
||
<emphasis box="[849,886,383,409]" italics="true" pageId="10" pageNumber="1004">(ft)</emphasis>
|
||
,
|
||
<emphasis box="[907,949,383,409]" italics="true" pageId="10" pageNumber="1004">(tc)</emphasis>
|
||
,
|
||
<emphasis box="[969,1007,383,409]" italics="true" pageId="10" pageNumber="1004">(it)</emphasis>
|
||
,
|
||
<emphasis box="[1027,1063,383,409]" italics="true" pageId="10" pageNumber="1004">(p)</emphasis>
|
||
,
|
||
<emphasis box="[1084,1119,383,409]" italics="true" pageId="10" pageNumber="1004">(u)</emphasis>
|
||
,
|
||
<emphasis box="[1140,1176,383,409]" italics="true" pageId="10" pageNumber="1004">(a)</emphasis>
|
||
,
|
||
<emphasis box="[1196,1208,383,408]" italics="true" pageId="10" pageNumber="1004">s</emphasis>
|
||
,
|
||
</td>
|
||
<td box="[1229,1291,382,414]" gridcol="8" gridrow="1" pageId="10" pageNumber="1004">
|
||
<emphasis box="[1229,1278,383,409]" italics="true" pageId="10" pageNumber="1004">(pv)</emphasis>
|
||
,
|
||
</td>
|
||
<td box="[1304,1408,382,414]" gridcol="9" gridrow="1" pageId="10" pageNumber="1004">ɛ, ω1, ω2</td>
|
||
</tr>
|
||
<tr box="[112,1408,427,458]" gridrow="2" pageId="10" pageNumber="1004" rowspan-6="1">
|
||
<th box="[112,159,427,458]" gridcol="0" gridrow="2" pageId="10" pageNumber="1004">II</th>
|
||
<td box="[192,242,427,458]" gridcol="1" gridrow="2" pageId="10" pageNumber="1004">
|
||
<emphasis box="[192,212,427,455]" italics="true" pageId="10" pageNumber="1004">v′</emphasis>
|
||
</td>
|
||
<td box="[304,420,427,458]" gridcol="2" gridrow="2" pageId="10" pageNumber="1004">
|
||
<emphasis box="[304,420,427,455]" italics="true" pageId="10" pageNumber="1004">d, (l), bv″</emphasis>
|
||
</td>
|
||
<td box="[479,537,427,458]" gridcol="3" gridrow="2" pageId="10" pageNumber="1004">
|
||
<emphasis box="[479,537,427,455]" italics="true" pageId="10" pageNumber="1004">l′, v′,</emphasis>
|
||
</td>
|
||
<td box="[551,567,427,458]" gridcol="4" gridrow="2" pageId="10" pageNumber="1004">σ</td>
|
||
<td box="[622,718,427,458]" gridcol="5" gridrow="2" pageId="10" pageNumber="1004">
|
||
<emphasis box="[622,655,429,455]" italics="true" pageId="10" pageNumber="1004">(v)</emphasis>
|
||
, φ
|
||
</td>
|
||
<td box="[849,1221,427,458]" gridcol="7" gridrow="2" pageId="10" pageNumber="1004">
|
||
<emphasis box="[849,886,427,453]" italics="true" pageId="10" pageNumber="1004">(ft)</emphasis>
|
||
,
|
||
<emphasis box="[907,949,427,453]" italics="true" pageId="10" pageNumber="1004">(tc)</emphasis>
|
||
,
|
||
<emphasis box="[969,1007,427,453]" italics="true" pageId="10" pageNumber="1004">(it)</emphasis>
|
||
,
|
||
<emphasis box="[1027,1063,427,453]" italics="true" pageId="10" pageNumber="1004">(p)</emphasis>
|
||
,
|
||
<emphasis box="[1084,1119,427,453]" italics="true" pageId="10" pageNumber="1004">(u)</emphasis>
|
||
,
|
||
<emphasis box="[1140,1176,427,453]" italics="true" pageId="10" pageNumber="1004">(a)</emphasis>
|
||
,
|
||
<emphasis box="[1196,1208,428,453]" italics="true" pageId="10" pageNumber="1004">s</emphasis>
|
||
,
|
||
</td>
|
||
<td box="[1229,1291,427,458]" gridcol="8" gridrow="2" pageId="10" pageNumber="1004">
|
||
<emphasis box="[1229,1278,427,453]" italics="true" pageId="10" pageNumber="1004">(pv)</emphasis>
|
||
,
|
||
</td>
|
||
<td box="[1304,1408,427,458]" gridcol="9" gridrow="2" pageId="10" pageNumber="1004">ω1, ω2</td>
|
||
</tr>
|
||
<tr box="[112,1408,469,497]" gridrow="3" pageId="10" pageNumber="1004" rowspan-4="1" rowspan-6="1" rowspan-9="1">
|
||
<th box="[112,159,469,497]" gridcol="0" gridrow="3" pageId="10" pageNumber="1004">III</th>
|
||
<td box="[192,242,469,497]" gridcol="1" gridrow="3" pageId="10" pageNumber="1004">
|
||
<emphasis box="[192,242,469,497]" italics="true" pageId="10" pageNumber="1004">l′, v′</emphasis>
|
||
</td>
|
||
<td box="[304,420,469,497]" gridcol="2" gridrow="3" pageId="10" pageNumber="1004">
|
||
<emphasis box="[304,399,469,497]" italics="true" pageId="10" pageNumber="1004">d, l′, ev′</emphasis>
|
||
</td>
|
||
<td box="[479,537,469,497]" gridcol="3" gridrow="3" pageId="10" pageNumber="1004">σ</td>
|
||
<td box="[622,718,469,497]" gridcol="5" gridrow="3" pageId="10" pageNumber="1004">
|
||
<emphasis box="[622,655,471,497]" italics="true" pageId="10" pageNumber="1004">(v)</emphasis>
|
||
, φ
|
||
</td>
|
||
<td box="[849,1221,469,497]" gridcol="7" gridrow="3" pageId="10" pageNumber="1004">
|
||
<emphasis box="[849,886,471,497]" italics="true" pageId="10" pageNumber="1004">(ft)</emphasis>
|
||
,
|
||
<emphasis box="[907,949,471,497]" italics="true" pageId="10" pageNumber="1004">(tc)</emphasis>
|
||
,
|
||
<emphasis box="[969,1007,471,497]" italics="true" pageId="10" pageNumber="1004">(it)</emphasis>
|
||
,
|
||
<emphasis box="[1027,1063,471,497]" italics="true" pageId="10" pageNumber="1004">(p)</emphasis>
|
||
,
|
||
<emphasis box="[1084,1119,471,497]" italics="true" pageId="10" pageNumber="1004">(u)</emphasis>
|
||
,
|
||
<emphasis box="[1140,1176,471,497]" italics="true" pageId="10" pageNumber="1004">(a)</emphasis>
|
||
,
|
||
<emphasis box="[1196,1208,471,496]" italics="true" pageId="10" pageNumber="1004">s</emphasis>
|
||
,
|
||
</td>
|
||
<td box="[1229,1291,469,497]" gridcol="8" gridrow="3" pageId="10" pageNumber="1004">
|
||
<emphasis box="[1229,1278,471,497]" italics="true" pageId="10" pageNumber="1004">(pv)</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr box="[112,1408,509,537]" gridrow="4" pageId="10" pageNumber="1004" rowspan-4="1" rowspan-6="1" rowspan-8="1" rowspan-9="1">
|
||
<th box="[112,159,509,537]" gridcol="0" gridrow="4" pageId="10" pageNumber="1004">IV</th>
|
||
<td box="[192,242,509,537]" gridcol="1" gridrow="4" pageId="10" pageNumber="1004">
|
||
<emphasis box="[192,212,509,537]" italics="true" pageId="10" pageNumber="1004">v′</emphasis>
|
||
</td>
|
||
<td box="[304,420,509,537]" gridcol="2" gridrow="4" pageId="10" pageNumber="1004">
|
||
<emphasis box="[304,369,509,537]" italics="true" pageId="10" pageNumber="1004">d, ev′</emphasis>
|
||
</td>
|
||
<td box="[479,537,509,537]" gridcol="3" gridrow="4" pageId="10" pageNumber="1004">
|
||
<emphasis box="[479,494,510,536]" italics="true" pageId="10" pageNumber="1004">d</emphasis>
|
||
</td>
|
||
<td box="[622,718,509,537]" gridcol="5" gridrow="4" pageId="10" pageNumber="1004">
|
||
<emphasis box="[622,655,510,536]" italics="true" pageId="10" pageNumber="1004">(v)</emphasis>
|
||
, φ
|
||
</td>
|
||
<td box="[849,1221,509,537]" gridcol="7" gridrow="4" pageId="10" pageNumber="1004">ft″, (tc), (p), (u), (a), s, (pv)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph blockId="10.[112,1428,547,643]" pageId="10" pageNumber="1004">marks setae on the anterior and double quotation mark (″) setae on the posterior side of a given leg segment; parentheses refer to a pair of setae.</paragraph>
|
||
<paragraph blockId="10.[491,1285,714,738]" box="[491,1285,714,738]" pageId="10" pageNumber="1004">
|
||
simple) notogastral seta
|
||
<emphasis box="[750,762,715,738]" italics="true" pageId="10" pageNumber="1004">c</emphasis>
|
||
and foveolate (versus not foveolate) notogaster.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |