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<document id="13C7A99D12AE630B6FA4BAF57F31E4F8" ID-CLB-Dataset="276843" ID-DOI="10.5852/ejt.2023.901.2317" ID-GBIF-Dataset="f82db014-3609-4e5e-a3d9-c90c9de565e3" ID-ISSN="2118-9773" ID-Zenodo-Dep="10088606" ID-ZooBank="FB29CFDF-0064-4C6B-9C8A-3673B9437837" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1699006741891" checkinUser="plazi" docAuthor="Dhadwal, Tarun &amp; Bharti, Himender" docDate="2023" docId="60418785FFA5A332FDCCFDE9687FFC3F" docLanguage="en" docName="EJT.2023.901.1-51.pdf" docOrigin="European Journal of Taxonomy 901 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2317/10057" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Camponotus keihitoi Forel 1913" docType="treatment" docVersion="5" lastPageNumber="23" masterDocId="9C78FFFDFFB5A324FF98FFBE6C3DFFB8" masterDocTitle="Two new species of the genus Camponotus Mayr, 1861 (Hymenoptera: Formicidae) with five new records from India" masterLastPageNumber="51" masterPageNumber="1" pageNumber="17" updateTime="1699514390746" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6A9C2143BE671D549A2788E0AC6FC9DE">Two new species of the genus Camponotus Mayr, 1861 (Hymenoptera: Formicidae) with five new records from India</mods:title>
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<mods:namePart id="BC8105A8D2E48BCD42E6AE85CB15FD01">Dhadwal, Tarun</mods:namePart>
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<mods:affiliation id="10C123E90477A91F4F9CE89974706AC6">Department of Zoology and Environmental Sciences, Punjabi University, Patiala, Punjab, India.</mods:affiliation>
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<mods:namePart id="7A9E63779A4FDB8B0C41D1EEF1B8ACC2">Bharti, Himender</mods:namePart>
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<mods:affiliation id="D0B3EA91D61E832AF47716FD4971F136">Department of Zoology and Environmental Sciences, Punjabi University, Patiala, Punjab, India.</mods:affiliation>
<mods:nameIdentifier id="D0D106B94F8FD8DB6F7101AB7A5B0F16" type="email">himenderbharti@gmail.com</mods:nameIdentifier>
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<mods:title id="3FAA50E6DA2D98DC6CD0F76E8113DA76">European Journal of Taxonomy</mods:title>
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<mods:date id="EFF54F4D79E1F29035145B6E8BC57923">2023</mods:date>
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<mods:number id="FA055028D418D37472B6DD77D32CDA75">2023-11-03</mods:number>
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<mods:number id="5201BE68BA7E4C20E4CD73A7C3FF7383">901</mods:number>
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<treatment id="60418785FFA5A332FDCCFDE9687FFC3F" ID-GBIF-Taxon="217349322" LSID="urn:lsid:plazi:treatment:60418785FFA5A332FDCCFDE9687FFC3F" httpUri="http://treatment.plazi.org/id/60418785FFA5A332FDCCFDE9687FFC3F" lastPageId="22" lastPageNumber="23" pageId="16" pageNumber="17">
<subSubSection id="A0F26518FFA5A334FDCCFDE96FE2FDCA" box="[596,991,599,626]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="E8573693FFA5A334FDCCFDE96FE2FDCA" blockId="16.[596,991,599,662]" box="[596,991,599,626]" pageId="16" pageNumber="17">
<heading id="B31F81FFFFA5A334FDCCFDE96FE2FDCA" box="[596,991,599,626]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="17" reason="2">
<taxonomicName id="2FE84D10FFA5A334FDCCFDE96FE2FDCA" ID-CoL="QDWF" authority="Forel, 1913" authorityName="Forel" authorityYear="1913" box="[596,991,599,626]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="keihitoi">
<emphasis id="DA9CEA81FFA5A334FDCCFDE96F6DFDCA" bold="true" box="[596,848,599,626]" italics="true" pageId="16" pageNumber="17">Camponotus keihitoi</emphasis>
Forel, 1913
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A0F26518FFA5A334FD40FDC26F61FD2E" box="[728,860,636,662]" pageId="16" pageNumber="17" type="description">
<paragraph id="E8573693FFA5A334FD40FDC26F61FD2E" blockId="16.[596,991,599,662]" box="[728,860,636,662]" pageId="16" pageNumber="17">
<figureCitation id="70D32A16FFA5A334FD40FDC26F61FD2E" box="[728,860,636,662]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="17.[189,232,1963,1989]" captionStartId-1="18.[189,232,1963,1989]" captionStartId-2="19.[189,232,1963,1989]" captionStartId-3="20.[189,232,1963,1989]" captionTargetBox-0="[471,1117,265,1915]" captionTargetBox-1="[415,1172,265,1915]" captionTargetBox-2="[421,1166,265,1920]" captionTargetBox-3="[459,1129,265,1920]" captionTargetId-0="figure-11@17.[471,1117,265,1915]" captionTargetId-1="figure-20@18.[415,1172,265,1915]" captionTargetId-2="figure-12@19.[421,1166,265,1920]" captionTargetId-3="figure-20@20.[459,1129,265,1920]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionTargetPageId-3="20" captionText-0="Fig. 10. Camponotus keihitoiForel, 1913, major worker (PUAC T82). A. Head in full face view. B. Body in profile view. C. Body in dorsal view." captionText-1="Fig. 11. Camponotus keihitoi Forel, 1913, minor worker (PUAC T85). A. Head in full face view. B. Body in profile view. C. Body in dorsal view." captionText-2="Fig. 12. Camponotus keihitoi Forel, 1913, gyne (PUAC T89). A. Head in full face view. B. Body in profile view. C. Body in dorsal view." captionText-3="Fig. 13. Camponotus keihitoi Forel, 1913, ♂ (PUAC T90). A. Head in full face view. B. Body in profile view. C. Body in dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.10088626" figureDoi-1="http://doi.org/10.5281/zenodo.10088628" figureDoi-2="http://doi.org/10.5281/zenodo.10088630" figureDoi-3="http://doi.org/10.5281/zenodo.10088632" httpUri-0="https://zenodo.org/record/10088626/files/figure.png" httpUri-1="https://zenodo.org/record/10088628/files/figure.png" httpUri-2="https://zenodo.org/record/10088630/files/figure.png" httpUri-3="https://zenodo.org/record/10088632/files/figure.png" pageId="16" pageNumber="17">Figs 10–13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A0F26518FFA5A334FF25FD006932FD45" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="E8573693FFA5A334FF25FD006F3BFD60" blockId="16.[189,1295,702,765]" box="[189,774,702,728]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="C8F811BDFFA5A334FF25FD006F3BFD60" box="[189,774,702,728]" pageId="16" pageNumber="17">
<treatmentCitation id="69491082FFA5A334FF25FD006F3BFD60" author="Forel A." box="[189,774,702,728]" page="663" pageId="16" pageNumber="17" year="1913">
<taxonomicName id="2FE84D10FFA5A334FF25FD006F3BFD60" ID-CoL="QDWF" authority="Forel, 1913 b: 663" authorityName="Forel" authorityPageNumber="663" authorityYear="1913" box="[189,774,702,728]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="variety" species="fallax" variety="keihitoi">
<emphasis id="DA9CEA81FFA5A334FF25FD006DA1FD60" box="[189,412,702,728]" italics="true" pageId="16" pageNumber="17">Camponotus fallax</emphasis>
var.
<emphasis id="DA9CEA81FFA5A334FE4DFD006E10FD60" box="[469,557,702,728]" italics="true" pageId="16" pageNumber="17">keihitoi</emphasis>
<bibRefCitation id="8C794B62FFA5A334FDADFD006F3BFD60" author="Forel A." box="[565,774,702,728]" pageId="16" pageNumber="17" pagination="659 - 673" refId="ref15093" refString="Forel A. 1913 b. Quelques fourmis des Indes, du Japon et d'Afrique. Revue suisse de Zoologie 21: 659 - 673. https: // doi. org / 10.5962 / bhl. part. 37159" type="journal article" year="1913">Forel, 1913b: 663</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="E8573693FFA5A334FF25FD5C6932FD45" blockId="16.[189,1295,702,765]" box="[189,1295,738,765]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="C8F811BDFFA5A334FF25FD5C6932FD45" box="[189,1295,738,765]" pageId="16" pageNumber="17">
<treatmentCitation id="69491082FFA5A334FF25FD5C6EF1FD44" author="Teranishi C." box="[189,716,738,765]" page="137" pageId="16" pageNumber="17" year="1915">
<taxonomicName id="2FE84D10FFA5A334FF25FD5C6EF1FD44" authority="Teranishi, 1915: 137" authorityName="Teranishi" authorityPageNumber="137" authorityYear="1915" box="[189,716,738,765]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tokyoensis">
<emphasis id="DA9CEA81FFA5A334FF25FD5C6DEEFD44" box="[189,467,738,764]" italics="true" pageId="16" pageNumber="17">Camponotus tokyoensis</emphasis>
<bibRefCitation id="8C794B62FFA5A334FE42FD5C6EF1FD44" author="Teranishi C." box="[474,716,738,765]" pageId="16" pageNumber="17" pagination="137 - 138" refId="ref15579" refString="Teranishi C. 1915 A new species of Formicidae from Japan. Entomological Magazine. Kyoto 1: 137 - 138." type="journal article" year="1915">Teranishi, 1915: 137</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(synonymized by
<treatmentCitation id="69491082FFA5A334FC3DFD5C693EFD45" author="Terayama M. &amp; Satoh T." box="[933,1283,738,765]" page="532" pageId="16" pageNumber="17" year="1990">
<bibRefCitation id="8C794B62FFA5A334FC3DFD5C693EFD45" author="Terayama M. &amp; Satoh T." box="[933,1283,738,765]" pageId="16" pageNumber="17" pagination="532" refId="ref15601" refString="Terayama M. &amp; Satoh T. 1990. Taxonomic notes on two Japanese species of Formicidae (Hymenoptera). Japanese Journal of Entomology 58: 532." type="journal article" year="1990">Terayama &amp; Satoh 1990: 532</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A0F26518FFA5A334FF25FC9A6D04FB99" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="E8573693FFA5A334FF25FC9A6D0BFC86" blockId="16.[189,310,804,830]" box="[189,310,804,830]" pageId="16" pageNumber="17">
<heading id="B31F81FFFFA5A334FF25FC9A6D0BFC86" bold="true" box="[189,310,804,830]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="DA9CEA81FFA5A334FF25FC9A6D0BFC86" bold="true" box="[189,310,804,830]" pageId="16" pageNumber="17">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="E8573693FFA5A334FF25FCEA6D04FB99" blockId="16.[188,1398,851,1057]" pageId="16" pageNumber="17">
<taxonomicName id="2FE84D10FFA5A334FF25FCEA6E6AFCD6" authority="Forel, 1913" authorityName="Forel" authorityYear="1913" box="[189,599,851,878]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="keihitoi">
<emphasis id="DA9CEA81FFA5A334FF25FCEA6D81FCD5" box="[189,444,851,878]" italics="true" pageId="16" pageNumber="17">Camponotus keihitoi</emphasis>
Forel, 1913
</taxonomicName>
is allied to
<taxonomicName id="2FE84D10FFA5A334FD6CFCEA686CFCD5" authority="Forel, 1886" authorityName="Forel" authorityYear="1886" box="[756,1105,851,878]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="quarinotatus">
<emphasis id="DA9CEA81FFA5A334FD6CFCEA6F8BFCD5" box="[756,950,852,878]" italics="true" pageId="16" pageNumber="17">C. quarinotatus</emphasis>
<bibRefCitation id="8C794B62FFA5A334FC5AFCED686CFCD5" author="Forel A." box="[962,1105,851,877]" pageId="16" pageNumber="17" pagination="131 - 215" refId="ref14826" refString="Forel A. 1886. Etudes myrmecologiques en 1886. Annales de la Societe entomologique de Belgique 30: 131 - 215." type="journal article" year="1886">Forel, 1886</bibRefCitation>
</taxonomicName>
, however, both species can be fairly distinguished by the following combination of characteristics: in
<taxonomicName id="2FE84D10FFA5A334FB01FCC66926FC29" authorityName="Forel" authorityYear="1913" box="[1177,1307,887,914]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="keihitoi">
<emphasis id="DA9CEA81FFA5A334FB01FCC66926FC29" box="[1177,1307,887,914]" italics="true" pageId="16" pageNumber="17">C. keihitoi</emphasis>
</taxonomicName>
(major worker), the metanotal depression is distinct; pilosity is absent on the mesosoma and the petiole; and the anterior clypeal margin is transverse. While in
<taxonomicName id="2FE84D10FFA5A334FCA0FC7E6FC4FC61" authorityName="Forel" authorityYear="1886" box="[824,1017,960,986]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="quarinotatus">
<emphasis id="DA9CEA81FFA5A334FCA0FC7E6FC4FC61" box="[824,1017,960,986]" italics="true" pageId="16" pageNumber="17">C. quarinotatus</emphasis>
</taxonomicName>
(major worker), the metanotal depression is indistinct; the mesosoma and the petiole are pilose; the clypeal margin is convex anteriorly.
</paragraph>
</subSubSection>
<subSubSection id="A0F26518FFA5A334FF25FBF76904FB0F" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="E8573693FFA5A334FF25FBF76D97FBDB" blockId="16.[189,426,1097,1123]" box="[189,426,1097,1123]" pageId="16" pageNumber="17">
<heading id="B31F81FFFFA5A334FF25FBF76D97FBDB" bold="true" box="[189,426,1097,1123]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="DA9CEA81FFA5A334FF25FBF76D97FBDB" bold="true" box="[189,426,1097,1123]" pageId="16" pageNumber="17">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="E8573693FFA5A334FF25FBC76904FB0F" blockId="16.[189,1399,1143,1208]" pageId="16" pageNumber="17">
<materialsCitation id="58803CCEFFA5A334FF25FBC76908FB0F" ID-GBIF-Occurrence="4436650304" collectingDate="2019-07-04" collectionCode="PUAC" collectorName="T. Dhadwal" country="India" elevation="3600" latitude="30.728" location="Nanda Devi National Park" longLatPrecision="6" longitude="79.6053" municipality="Flower Valley" pageId="16" pageNumber="17" specimenCode="PUAC T81, PUAC T82, PUAC T83, PUAC T84, PUAC T85, PUAC T86, PUAC T87, PUAC T88, PUAC T89, PUAC T90, PUAC T91, PUAC T92, PUAC T93, PUAC T94, PUAC T95, PUAC T96, PUAC T97, PUAC T98" specimenCount="18" specimenCount-female="2" specimenCount-male="2" specimenCount-worker="14" stateProvince="Uttarakhand">
<collectingCountry id="90FF7603FFA5A334FF25FBC76D33FB2B" box="[189,270,1145,1171]" name="India" pageId="16" pageNumber="17">INDIA</collectingCountry>
•
<specimenCount id="FEEEFD1AFFA5A334FEBCFBC76D94FB2B" box="[292,425,1145,1171]" count="14" pageId="16" pageNumber="17" type="worker">14 workers</specimenCount>
,
<specimenCount id="FEEEFD1AFFA5A334FE2BFBC76DC9FB2C" box="[435,500,1145,1172]" count="2" pageId="16" pageNumber="17" type="female">2 ♀♀</specimenCount>
, 2 ♂♂;
<collectingRegion id="2A2CF871FFA5A334FDD7FBC76EDEFB2B" box="[591,739,1145,1171]" country="India" name="Uttarakhand" pageId="16" pageNumber="17">Uttarakhand</collectingRegion>
,
<collectingMunicipality id="0833ACE9FFA5A334FD75FBC76FADFB2B" box="[749,912,1145,1171]" pageId="16" pageNumber="17">Flower Valley</collectingMunicipality>
(
<location id="ED376048FFA5A334FC06FBC768EAFB2B" LSID="urn:lsid:plazi:treatment:60418785FFA5A332FDCCFDE9687FFC3F:ED376048FFA5A334FC06FBC768EAFB2B" box="[926,1239,1145,1171]" country="India" latitude="30.728" longLatPrecision="6" longitude="79.6053" municipality="Flower Valley" name="Nanda Devi National Park" pageId="16" pageNumber="17" stateProvince="Uttarakhand">Nanda Devi National Park</location>
);
<geoCoordinate id="8DDC5054FFA5A334FB71FBC7694FFB2C" box="[1257,1394,1145,1172]" degrees="30.7280" direction="north" orientation="latitude" pageId="16" pageNumber="17" precision="5" value="30.728">30.7280° N</geoCoordinate>
,
<geoCoordinate id="8DDC5054FFA5A334FF25FB236D79FB00" box="[189,324,1181,1208]" degrees="79.6053" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="5" value="79.6053">79.6053° E</geoCoordinate>
; elev.
<quantity id="2F109B76FFA5A334FE16FB236DD7FB00" box="[398,490,1181,1208]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="16" pageNumber="17" unit="m" value="3600.0">
<elevation id="63C5D1A0FFA5A334FE16FB236DD7FB00" box="[398,490,1181,1208]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="16" pageNumber="17" unit="m" value="3600.0">3600 m</elevation>
</quantity>
;
<date id="9C561053FFA5A334FE6FFB236E41FB0F" box="[503,636,1181,1207]" pageId="16" pageNumber="17" value="2019-07-04">
<collectingDate id="8C12E9BBFFA5A334FE6FFB236E41FB0F" box="[503,636,1181,1207]" pageId="16" pageNumber="17" value="2019-07-04">4 Jul. 2019</collectingDate>
</date>
;
<collectionCode id="8EF9AE56FFA5A334FD1FFB206EA7FB00" box="[647,666,1182,1208]" name="Tavera, Department of Geology and Geophysics" pageId="16" pageNumber="17">T</collectionCode>
.
<collectorName id="451D5345FFA5A334FD3EFB236F2DFB0F" box="[678,784,1181,1207]" pageId="16" pageNumber="17">Dhadwal</collectorName>
leg.; hand picking method;
<collectionCode id="8EF9AE56FFA5A334FBC1FB23689AFB0F" box="[1113,1191,1181,1207]" pageId="16" pageNumber="17">PUAC</collectionCode>
<specimenCode id="B84E9EE8FFA5A334FB36FB2368E0FB0F" box="[1198,1245,1181,1207]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="16" pageNumber="17">T81</specimenCode>
to
<specimenCode id="B84E9EE8FFA5A334FA9AFB236908FB0F" box="[1282,1333,1181,1207]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="16" pageNumber="17">T98</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A0F26518FFA5A332FF25FB616F02FE13" lastPageId="22" lastPageNumber="23" pageId="16" pageNumber="17" type="description">
<paragraph id="E8573693FFA5A334FF25FB616D48FB41" blockId="16.[189,373,1247,1273]" box="[189,373,1247,1273]" pageId="16" pageNumber="17">
<heading id="B31F81FFFFA5A334FF25FB616D48FB41" bold="true" box="[189,373,1247,1273]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="DA9CEA81FFA5A334FF25FB616D48FB41" bold="true" box="[189,373,1247,1273]" pageId="16" pageNumber="17">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="E8573693FFA5A334FF72FAB06DCEFA91" blockId="16.[189,1399,1294,1429]" box="[234,499,1294,1321]" pageId="16" pageNumber="17">
<emphasis id="DA9CEA81FFA5A334FF72FAB06DA1FA91" bold="true" box="[234,412,1294,1321]" pageId="16" pageNumber="17">Major worker</emphasis>
(n = 7)
</paragraph>
<paragraph id="E8573693FFA5A334FF25FA8D6E37FA2C" blockId="16.[189,1399,1294,1429]" pageId="16" pageNumber="17">
HL 1.56–1.70; HW 1.26–1.44; EL 0.42–0.45; SL 1.47–1.59; PW 0.96–1.05; WL 2.07–2.19; MTL 1.35– 1.68; HTL 1.59–1.84; PL 0.51–0.57;
<collectingCountry id="90FF7603FFA5A334FDE3FAE96E9DFAC9" box="[635,672,1367,1393]" name="Philippines" pageId="16" pageNumber="17">PH</collectingCountry>
0.57–0.75; GL 1.81–2.64; TL 5.95–7.10; CI 80–84; SI 110– 116; REL 26–27; PrI 72–76.
</paragraph>
<paragraph id="E8573693FFA5A334FF72FA036DCEFA6F" blockId="16.[189,1399,1469,1603]" box="[234,499,1469,1495]" pageId="16" pageNumber="17">
<emphasis id="DA9CEA81FFA5A334FF72FA036DA1FA6F" bold="true" box="[234,412,1469,1495]" pageId="16" pageNumber="17">Minor worker</emphasis>
(n = 7)
</paragraph>
<paragraph id="E8573693FFA5A334FF25FA5F6E36F9FB" blockId="16.[189,1399,1469,1603]" pageId="16" pageNumber="17">
HL 1.35–1.38; HW 1.14–1.20; EL 0.36–0.39; SL 1.38–1.44; PW 0.91–0.93; WL 1.95–2.01; MTL 1.17– 1.23; HTL 1.51–1.56; PL 0.45–0.54;
<collectingCountry id="90FF7603FFA5A334FDE2F9BB6EA2F9A7" box="[634,671,1541,1567]" name="Philippines" pageId="16" pageNumber="17">PH</collectingCountry>
0.54–0.57; GL 1.71–2.04; TL 5.46–5.97; CI 84–86; SI 120– 121; REL 26–28; PrI 77–79.
</paragraph>
<paragraph id="E8573693FFA5A334FF72F9D56DB8F93D" blockId="16.[189,1399,1643,1742]" box="[234,389,1643,1670]" pageId="16" pageNumber="17">
<emphasis id="DA9CEA81FFA5A334FF72F9D56D10F93D" bold="true" box="[234,301,1643,1669]" pageId="16" pageNumber="17">Gyne</emphasis>
(n = 2)
</paragraph>
<paragraph id="E8573693FFA5A334FF25F92E695EF975" blockId="16.[189,1399,1643,1742]" pageId="16" pageNumber="17">
HL 1.59–1.72; HW 1.47–1.51; EL 0.49–0.57; SL 1.59–1.63; WL 2.54–2.66; MTL 1.41–1.43; HTL 1.80– 1.96; PL 0.57–0.65;
<collectingCountry id="90FF7603FFA5A334FE35F90A6DEFF976" box="[429,466,1716,1742]" name="Philippines" pageId="16" pageNumber="17">PH</collectingCountry>
0.82–0.90; GL 2.74–2.82; TL 7.44–7.85; CI 87–92; SI 107–108; REL 30–33.
</paragraph>
<paragraph id="E8573693FFA5A334FF72F94B6DBCF8B7" blockId="16.[189,1398,1781,1916]" box="[234,385,1781,1808]" pageId="16" pageNumber="17">
<emphasis id="DA9CEA81FFA5A334FF72F94B6D17F8B7" bold="true" box="[234,298,1781,1807]" pageId="16" pageNumber="17">Male</emphasis>
(n = 2)
</paragraph>
<paragraph id="E8573693FFA5A334FF25F8A46D73F8C3" blockId="16.[189,1398,1781,1916]" pageId="16" pageNumber="17">
HL 1.02–1.11; HW 1.08–1.10; EL 0.39–0.42; SL 1.41–1.43; WL 2.01–2.13; MTL 1.33–1.35; HTL 1.72–1.74; PL 0.51–0.53;
<collectingCountry id="90FF7603FFA5A334FDAEF8806E61F8E0" box="[566,604,1854,1880]" name="Philippines" pageId="16" pageNumber="17">PH</collectingCountry>
0.45–0.48; GL 2.37–2.54; TL 5.91–6.31; CI 99–105; SI 130–131; REL 37–38.
</paragraph>
<caption id="BC97661BFFA4A335FF25F8156EB1F850" ID-DOI="http://doi.org/10.5281/zenodo.10088626" ID-Zenodo-Dep="10088626" httpUri="https://zenodo.org/record/10088626/files/figure.png" pageId="17" pageNumber="18" startId="17.[189,232,1963,1989]" targetBox="[471,1117,265,1915]" targetPageId="17">
<paragraph id="E8573693FFA4A335FF25F8156EB1F850" blockId="17.[187,1399,1963,2025]" pageId="17" pageNumber="18">
<emphasis id="DA9CEA81FFA4A335FF25F8156D25F87D" bold="true" box="[189,280,1963,1989]" pageId="17" pageNumber="18">Fig. 10.</emphasis>
<taxonomicName id="2FE84D10FFA4A335FE86F8126EA6F87D" authority="Forel, 1913" authorityName="Forel" authorityYear="1913" box="[286,667,1963,1990]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="keihitoi">
<emphasis id="DA9CEA81FFA4A335FE86F8126E2DF87D" box="[286,528,1963,1990]" italics="true" pageId="17" pageNumber="18">Camponotus keihitoi</emphasis>
Forel, 1913
</taxonomicName>
, major worker (PUAC T82).
<emphasis id="DA9CEA81FFA4A335FC73F815683DF87D" bold="true" box="[1003,1024,1963,1989]" pageId="17" pageNumber="18">A</emphasis>
. Head in full face view.
<emphasis id="DA9CEA81FFA4A335FA8DF8156914F87D" bold="true" box="[1301,1321,1963,1989]" pageId="17" pageNumber="18">B</emphasis>
. Body in profile view.
<emphasis id="DA9CEA81FFA4A335FEEDF8706DB7F850" bold="true" box="[373,394,1998,2024]" pageId="17" pageNumber="18">C</emphasis>
. Body in dorsal view.
</paragraph>
</caption>
<caption id="BC97661BFFA7A336FF25F8156EB1F850" ID-DOI="http://doi.org/10.5281/zenodo.10088628" ID-Zenodo-Dep="10088628" httpUri="https://zenodo.org/record/10088628/files/figure.png" pageId="18" pageNumber="19" startId="18.[189,232,1963,1989]" targetBox="[415,1172,265,1915]" targetPageId="18">
<paragraph id="E8573693FFA7A336FF25F8156EB1F850" blockId="18.[187,1399,1963,2025]" pageId="18" pageNumber="19">
<emphasis id="DA9CEA81FFA7A336FF25F8156D2AF87D" bold="true" box="[189,279,1963,1989]" pageId="18" pageNumber="19">Fig. 11.</emphasis>
<taxonomicName id="2FE84D10FFA7A336FE84F8126EA4F87D" authority="Forel, 1913" authorityName="Forel" authorityYear="1913" box="[284,665,1963,1990]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="keihitoi">
<emphasis id="DA9CEA81FFA7A336FE84F8126E33F87D" box="[284,526,1963,1990]" italics="true" pageId="18" pageNumber="19">Camponotus keihitoi</emphasis>
Forel, 1913
</taxonomicName>
, minor worker (PUAC T85).
<emphasis id="DA9CEA81FFA7A336FC73F815683DF87D" bold="true" box="[1003,1024,1963,1989]" pageId="18" pageNumber="19">A</emphasis>
. Head in full face view.
<emphasis id="DA9CEA81FFA7A336FA8DF8156914F87D" bold="true" box="[1301,1321,1963,1989]" pageId="18" pageNumber="19">B</emphasis>
. Body in profile view.
<emphasis id="DA9CEA81FFA7A336FEEDF8706DB7F850" bold="true" box="[373,394,1998,2024]" pageId="18" pageNumber="19">C</emphasis>
. Body in dorsal view.
</paragraph>
</caption>
<caption id="BC97661BFFA6A337FF25F8156E53F850" ID-DOI="http://doi.org/10.5281/zenodo.10088630" ID-Zenodo-Dep="10088630" httpUri="https://zenodo.org/record/10088630/files/figure.png" pageId="19" pageNumber="20" startId="19.[189,232,1963,1989]" targetBox="[421,1166,265,1920]" targetPageId="19">
<paragraph id="E8573693FFA6A337FF25F8156E53F850" blockId="19.[189,1399,1963,2025]" pageId="19" pageNumber="20">
<emphasis id="DA9CEA81FFA6A337FF25F8156D20F87D" bold="true" box="[189,285,1963,1989]" pageId="19" pageNumber="20">Fig. 12.</emphasis>
<taxonomicName id="2FE84D10FFA6A337FEBEF8126E89F87D" authority="Forel, 1913" authorityName="Forel" authorityYear="1913" box="[294,692,1963,1990]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="keihitoi">
<emphasis id="DA9CEA81FFA6A337FEBEF8126E21F87D" box="[294,540,1963,1990]" italics="true" pageId="19" pageNumber="20">Camponotus keihitoi</emphasis>
Forel, 1913
</taxonomicName>
, gyne (PUAC T89).
<emphasis id="DA9CEA81FFA6A337FC29F8156FF9F87D" bold="true" box="[945,964,1963,1989]" pageId="19" pageNumber="20">A</emphasis>
. Head in full face view.
<emphasis id="DA9CEA81FFA6A337FB6CF8156938F87D" bold="true" box="[1268,1285,1963,1989]" pageId="19" pageNumber="20">B</emphasis>
. Body in profile view.
<emphasis id="DA9CEA81FFA6A337FECFF8706D51F850" bold="true" box="[343,364,1998,2024]" pageId="19" pageNumber="20">C</emphasis>
. Body in dorsal view.
</paragraph>
</caption>
<caption id="BC97661BFFA1A330FF25F8156E27F850" ID-DOI="http://doi.org/10.5281/zenodo.10088632" ID-Zenodo-Dep="10088632" httpUri="https://zenodo.org/record/10088632/files/figure.png" pageId="20" pageNumber="21" startId="20.[189,232,1963,1989]" targetBox="[459,1129,265,1920]" targetPageId="20">
<paragraph id="E8573693FFA1A330FF25F8156E27F850" blockId="20.[189,1399,1961,2025]" pageId="20" pageNumber="21">
<emphasis id="DA9CEA81FFA1A330FF25F8156D27F87D" bold="true" box="[189,282,1963,1989]" pageId="20" pageNumber="21">Fig. 13.</emphasis>
<taxonomicName id="2FE84D10FFA1A330FEB9F8126E9EF87D" authority="Forel, 1913" authorityName="Forel" authorityYear="1913" box="[289,675,1963,1990]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="keihitoi">
<emphasis id="DA9CEA81FFA1A330FEB9F8126E29F87D" box="[289,532,1963,1990]" italics="true" pageId="20" pageNumber="21">Camponotus keihitoi</emphasis>
Forel, 1913
</taxonomicName>
, ♂ (PUAC T90).
<emphasis id="DA9CEA81FFA1A330FCEAF8156FBAF87D" bold="true" box="[882,903,1963,1989]" pageId="20" pageNumber="21">A</emphasis>
. Head in full face view.
<emphasis id="DA9CEA81FFA1A330FB3CF8156885F87D" bold="true" box="[1188,1208,1963,1989]" pageId="20" pageNumber="21">B</emphasis>
. Body in profile view.
<emphasis id="DA9CEA81FFA1A330FE9BF8706D25F850" bold="true" box="[259,280,1998,2024]" pageId="20" pageNumber="21">C</emphasis>
. Body in dorsal view.
</paragraph>
</caption>
<paragraph id="E8573693FFA0A331FF25FEB66D73FE9A" blockId="21.[189,334,264,290]" box="[189,334,264,290]" pageId="21" pageNumber="22">
<heading id="B31F81FFFFA0A331FF25FEB66D73FE9A" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="DA9CEA81FFA0A331FF25FEB66D73FE9A" bold="true" box="[189,334,264,290]" pageId="21" pageNumber="22">Description</emphasis>
</heading>
</paragraph>
<paragraph id="E8573693FFA0A331FF72FE896E2FFEE9" blockId="21.[187,1400,311,725]" box="[234,530,311,337]" pageId="21" pageNumber="22">
<emphasis id="DA9CEA81FFA0A331FF72FE896DA1FEE9" bold="true" box="[234,412,311,337]" pageId="21" pageNumber="22">Major worker</emphasis>
(
<figureCitation id="70D32A16FFA0A331FE34FE896E39FEE9" box="[428,516,311,337]" captionStart="Fig" captionStartId="17.[189,232,1963,1989]" captionTargetBox="[471,1117,265,1915]" captionTargetId="figure-11@17.[471,1117,265,1915]" captionTargetPageId="17" captionText="Fig. 10. Camponotus keihitoiForel, 1913, major worker (PUAC T82). A. Head in full face view. B. Body in profile view. C. Body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10088626" httpUri="https://zenodo.org/record/10088626/files/figure.png" pageId="21" pageNumber="22">Fig. 10</figureCitation>
):
</paragraph>
<paragraph id="E8573693FFA0A331FF25FEE56E0EFD6C" blockId="21.[187,1400,311,725]" pageId="21" pageNumber="22">
<smallCapsWord id="EEB1A04FFFA0A331FF25FEE56D20FECD" baselines="366,366" box="[189,285,347,373]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Habitus" pageId="21" pageNumber="22">HABITUS</smallCapsWord>
. In full-face view, head as long as broad or slightly longer than broad (CI 80–84), posterior head margin convex, occipital corners broadly rounded and lateral sides convex; clypeus broad, anterior margin transverse; mandibles subtriangular and masticatory margin with 5 teeth; antennae slender with 12 segments, scape long (SI 107–108), exceeds posterior head margin by ¼ of its length; eyes moderate in size placed laterally above the mid-length of the head. In dorsal view, mesosoma anteriorly broad (PrI 72–76) and narrow posteriorly; pronotum broader than long; pro-mesonotal suture and metanotal groove distinct; propodeum slightly impressed behind the metanotal groove; propodeum laterally compressed behind the mesonotum; propodeal declivity steep; mesosoma does not form a single convexity, convexity interrupted at propodeum being truncate; propodeal declivity concave; propodeal spiracle oval in shape, placed below the propodeal declivity; petiole anteriorly convex and posteriorly flat; tibiae cylindrical; gaster elongate and subglobose.
</paragraph>
<paragraph id="E8573693FFA0A331FF25FD426921FC81" blockId="21.[189,1399,764,825]" pageId="21" pageNumber="22">
<smallCapsWord id="EEB1A04FFFA0A331FF25FD426D05FCAE" baselines="783,783" box="[189,312,764,790]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Sculpture" pageId="21" pageNumber="22">SCULPTURE</smallCapsWord>
. Head, mesosoma, petiole and gaster all glossy; rest of body is reticulate-rugulose; clypeus with a median keel and scattered punctures; mandibles longitudinally rugulose with sparse pits.
</paragraph>
<paragraph id="E8573693FFA0A331FF25FCDF6F83FC79" blockId="21.[189,1399,864,961]" pageId="21" pageNumber="22">
<smallCapsWord id="EEB1A04FFFA0A331FF25FCDF6D23FCC3" baselines="884,884" box="[189,286,865,891]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Pilosity" pageId="21" pageNumber="22">PILOSITY</smallCapsWord>
<smallCapsWord id="EEB1A04FFFA0A331FEBEFCD86D6EFCC1" baselines="884" box="[294,339,870,889]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="21" pageNumber="22">AND</smallCapsWord>
<smallCapsWord id="EEB1A04FFFA0A331FEC3FCD86DDDFCC1" baselines="884" box="[347,480,870,889]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="pubescence" pageId="21" pageNumber="22">PUBESCENCE</smallCapsWord>
. Body coated in appressed hairs; pale yellow erect or sub-erect hairs plentiful on clypeus and mandibles; gaster covered in long hairs and hind tibia of the legs with dispersed setae underneath as well as 3–4 suberect setae close to the apical spurs.
</paragraph>
<paragraph id="E8573693FFA0A331FF25FC566902FBBA" blockId="21.[189,1343,1000,1027]" box="[189,1343,1000,1027]" pageId="21" pageNumber="22">
<smallCapsWord id="EEB1A04FFFA0A331FF25FC566D6AFBBA" baselines="1019,1020" box="[189,343,1000,1026]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colouration" pageId="21" pageNumber="22">COLOURATION</smallCapsWord>
. Body blackish; mandibles reddish brown and antennae and appendages dark brown.
</paragraph>
<paragraph id="E8573693FFA0A331FF72FB976E35FBFC" blockId="21.[188,1399,1065,1232]" box="[234,520,1065,1092]" pageId="21" pageNumber="22">
<emphasis id="DA9CEA81FFA0A331FF72FB976DA1FBFC" bold="true" box="[234,412,1065,1092]" pageId="21" pageNumber="22">Minor worker</emphasis>
(
<figureCitation id="70D32A16FFA0A331FE34FB976E3DFBFC" box="[428,512,1065,1092]" captionStart="Fig" captionStartId="18.[189,232,1963,1989]" captionTargetBox="[415,1172,265,1915]" captionTargetId="figure-20@18.[415,1172,265,1915]" captionTargetPageId="18" captionText="Fig. 11. Camponotus keihitoi Forel, 1913, minor worker (PUAC T85). A. Head in full face view. B. Body in profile view. C. Body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10088628" httpUri="https://zenodo.org/record/10088628/files/figure.png" pageId="21" pageNumber="22">Fig. 11</figureCitation>
)
</paragraph>
<paragraph id="E8573693FFA0A331FF25FBF36D32FB68" blockId="21.[188,1399,1065,1232]" pageId="21" pageNumber="22">All characteristics are the same as of major worker except: in minor worker head is comparatively small (CI 84–86) and oval with a convex posterior margin and sub-parallel lateral margins; the anterior clypeal margin is slightly convex; scape long (SI 120–121), surpassing posterior margin of head by half of its length.</paragraph>
<paragraph id="E8573693FFA0A331FF72FB466DA6FAAA" blockId="21.[187,1399,1272,1474]" box="[234,411,1272,1298]" pageId="21" pageNumber="22">
<emphasis id="DA9CEA81FFA0A331FF72FB466D10FAAA" bold="true" box="[234,301,1272,1298]" pageId="21" pageNumber="22">Gyne</emphasis>
(
<figureCitation id="70D32A16FFA0A331FEA5FB466DAFFAAA" box="[317,402,1272,1298]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[421,1166,265,1920]" captionTargetId="figure-12@19.[421,1166,265,1920]" captionTargetPageId="19" captionText="Fig. 12. Camponotus keihitoi Forel, 1913, gyne (PUAC T89). A. Head in full face view. B. Body in profile view. C. Body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10088630" httpUri="https://zenodo.org/record/10088630/files/figure.png" pageId="21" pageNumber="22">Fig. 12</figureCitation>
)
</paragraph>
<paragraph id="E8573693FFA0A331FF25FAA56F9DFA7A" blockId="21.[187,1399,1272,1474]" pageId="21" pageNumber="22">Similar to the major worker with few modifications depicting the caste and the following differences: head narrower (CI 87–92) with subparallel lateral margins and convex posterior margin; cephalic dorsum with 3 prominent ocelli; mandibles with 5 teeth; scape of antennae surpassing posterior margin of head by 1/₆ of its length (SI 107–108); scutum and scutellum minutely reticulated; dorsal surface of petiole transverse; propodeal declivity almost straight slightly convex.</paragraph>
<paragraph id="E8573693FFA0A331FF72FA576DA5F9BB" blockId="21.[188,1401,1513,1962]" box="[234,408,1513,1539]" pageId="21" pageNumber="22">
<emphasis id="DA9CEA81FFA0A331FF72FA576D17F9BB" bold="true" box="[234,298,1513,1539]" pageId="21" pageNumber="22">Male</emphasis>
(
<figureCitation id="70D32A16FFA0A331FEA2FA576DB2F9BB" box="[314,399,1513,1539]" captionStart="Fig" captionStartId="20.[189,232,1963,1989]" captionTargetBox="[459,1129,265,1920]" captionTargetId="figure-20@20.[459,1129,265,1920]" captionTargetPageId="20" captionText="Fig. 13. Camponotus keihitoi Forel, 1913, ♂ (PUAC T90). A. Head in full face view. B. Body in profile view. C. Body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10088632" httpUri="https://zenodo.org/record/10088632/files/figure.png" pageId="21" pageNumber="22">Fig. 13</figureCitation>
)
</paragraph>
<paragraph id="E8573693FFA0A331FF25F9B36926F812" blockId="21.[188,1401,1513,1962]" pageId="21" pageNumber="22">
<smallCapsWord id="EEB1A04FFFA0A331FF25F9B36D20F99F" baselines="1568,1568" box="[189,285,1549,1575]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Habitus" pageId="21" pageNumber="22">HABITUS</smallCapsWord>
. In full-face view, head as long as broad (CI 99–105), posterior margin of the head slightly concave; cephalic dorsum with 3 prominent ocelli; clypeus carinate in the middle; mandibles slender, curved strap like apical tooth acute, remainder without any teeth or denticles, when closed their tips overlap; eyes subglobose, convex, large and bulging, breaking lateral cephalic head outline; antennae 13-segmented and filiform, scape long (SI 130–131), surpassing posterior margin of head by about half of their length. Mesosoma enlarged, pronotum transverse, narrow and convex; scutum large, rounded anteriorly and transverse posteriorly; dorsally without notauli; parapsidal lines prominent and diverging anteriorly; scutellum pentagonal in shape; mesepimeron with a posterodorsal (epimeral) lobe that covers mesothoracic spiracle and forms a seemingly isolated plate; jugal lobe of hind wing present; petiole triangular, dorsal margin convex; propodeal declivity smoothly rounded; propodeal spiracle round. Pygostyles tubular, projecting outward; parameres elongated; cuspi small bent toward digiti, shorter than digiti; digiti long with short peg-like teeth bent toward parameres; penis valves projecting.
</paragraph>
<paragraph id="E8573693FFA3A332FF25FEB76861FED1" blockId="22.[189,1398,265,362]" pageId="22" pageNumber="23">
<smallCapsWord id="EEB1A04FFFA3A332FF25FEB76D23FE9B" baselines="284,284" box="[189,286,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Pilosity" pageId="22" pageNumber="23">PILOSITY</smallCapsWord>
<smallCapsWord id="EEB1A04FFFA3A332FEB2FEB16D6AFE9A" baselines="285" box="[298,343,271,290]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="22" pageNumber="23">AND</smallCapsWord>
<smallCapsWord id="EEB1A04FFFA3A332FEFBFEB06DD5FE99" baselines="284" box="[355,488,270,289]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="pubescence" pageId="22" pageNumber="23">PUBESCENCE</smallCapsWord>
. Clypeus and posterior margin of head with a few thin setae, scutum and scutellum with sparse erect short setae; gaster with adpressed short hairs; pygostyles and distal part of parameres setose; hind tibia without a row of spiny bristles on ventral margins.
</paragraph>
<paragraph id="E8573693FFA3A332FF25FE2F6F02FE13" blockId="22.[189,831,401,427]" box="[189,831,401,427]" pageId="22" pageNumber="23">
<smallCapsWord id="EEB1A04FFFA3A332FF25FE2F6D6AFE13" baselines="420,420" box="[189,343,401,427]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colouration" pageId="22" pageNumber="23">COLOURATION</smallCapsWord>
. Colour and sculpture as of worker caste.
</paragraph>
</subSubSection>
<subSubSection id="A0F26518FFA3A332FF25FE6C6E0EFDA3" pageId="22" pageNumber="23" type="distribution">
<paragraph id="E8573693FFA3A332FF25FE6C6D90FE54" blockId="22.[189,429,466,492]" box="[189,429,466,492]" pageId="22" pageNumber="23">
<heading id="B31F81FFFFA3A332FF25FE6C6D90FE54" bold="true" box="[189,429,466,492]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="DA9CEA81FFA3A332FF25FE6C6D90FE54" bold="true" box="[189,429,466,492]" pageId="22" pageNumber="23">Global distribution</emphasis>
</heading>
</paragraph>
<paragraph id="E8573693FFA3A332FF25FDBF6E0EFDA3" blockId="22.[189,563,513,539]" box="[189,563,513,539]" pageId="22" pageNumber="23">
<collectingCountry id="90FF7603FFA3A332FF25FDBF6D3EFDA3" box="[189,259,513,539]" name="China" pageId="22" pageNumber="23">China</collectingCountry>
and
<collectingCountry id="90FF7603FFA3A332FEA4FDBF6D42FDA3" box="[316,383,513,539]" name="Japan" pageId="22" pageNumber="23">Japan</collectingCountry>
(
<typeStatus id="37538831FFA3A332FE16FDBF6DFFFDA3" box="[398,450,513,539]" pageId="22" pageNumber="23">type</typeStatus>
locality).
</paragraph>
</subSubSection>
<subSubSection id="A0F26518FFA3A332FF25FDFC6CCAFD4D" pageId="22" pageNumber="23" type="biology_ecology">
<paragraph id="E8573693FFA3A332FF25FDFC6D20FDE4" blockId="22.[189,285,578,604]" box="[189,285,578,604]" pageId="22" pageNumber="23">
<heading id="B31F81FFFFA3A332FF25FDFC6D20FDE4" bold="true" box="[189,285,578,604]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="DA9CEA81FFA3A332FF25FDFC6D20FDE4" bold="true" box="[189,285,578,604]" pageId="22" pageNumber="23">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="E8573693FFA3A332FF25FDCF6CCAFD4D" blockId="22.[188,1399,625,757]" pageId="22" pageNumber="23">
During the field survey, the species was documented in
<collectingRegion id="2A2CF871FFA3A332FCF1FDCF6FC3FD33" box="[873,1022,625,651]" country="India" name="Uttarakhand" pageId="22" pageNumber="23">Uttarakhand</collectingRegion>
. The nest was found under the boulder. Some of the workers were also collected moving on the grass. The area is mostly surrounded by short grass and flowers with an average daily temperature of 22°C. The habitat is mostly open grassland
<typeStatus id="37538831FFA3A332FF25FD656CCEFD4D" box="[189,243,731,757]" pageId="22" pageNumber="23">type</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="A0F26518FFA3A332FF25FCA2687FFC3F" pageId="22" pageNumber="23" type="discussion">
<paragraph id="E8573693FFA3A332FF25FCA26D01FC8E" blockId="22.[189,316,796,822]" box="[189,316,796,822]" pageId="22" pageNumber="23">
<heading id="B31F81FFFFA3A332FF25FCA26D01FC8E" bold="true" box="[189,316,796,822]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="DA9CEA81FFA3A332FF25FCA26D01FC8E" bold="true" box="[189,316,796,822]" pageId="22" pageNumber="23">Relevance</emphasis>
</heading>
</paragraph>
<paragraph id="E8573693FFA3A332FF25FCF5687FFC3F" blockId="22.[188,1399,842,904]" pageId="22" pageNumber="23">
<taxonomicName id="2FE84D10FFA3A332FF25FCF56E01FCDC" authority="Forel, 1913" authorityName="Forel" authorityYear="1913" box="[189,572,842,869]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="keihitoi">
<emphasis id="DA9CEA81FFA3A332FF25FCF56D92FCDC" box="[189,431,842,869]" italics="true" pageId="22" pageNumber="23">Camponotus keihitoi</emphasis>
Forel, 1913
</taxonomicName>
represents a new record for
<collectingCountry id="90FF7603FFA3A332FC1BFCF46FFEFCDC" box="[899,963,842,868]" name="India" pageId="22" pageNumber="23">India</collectingCountry>
. Previously, this species was reported from
<collectingCountry id="90FF7603FFA3A332FF64FCD36D7FFC3F" box="[252,322,877,903]" name="China" pageId="22" pageNumber="23">China</collectingCountry>
and
<collectingCountry id="90FF7603FFA3A332FEE3FCD06DFFFC30" box="[379,450,878,904]" name="Japan" pageId="22" pageNumber="23">Japan</collectingCountry>
. The male of the species is described for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>
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