treatments-xml/data/03/FD/14/03FD141FFF8FFFD4FCAF2809FC21F9E1.xml

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<mods:title>A New Species Of Isoperla Banks (Plecoptera: Perlodidae) From The Southern Appalachians, With Notes On The I. Montana Group</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Verdone, Chris J.</mods:namePart>
<mods:affiliation>Department of Bioagricultural Sciences and Pest Management, Colorado State University, Fort Collins, Colorado, 80523, U. S. A. E-mail: verdonec @ gmail. com E-mail: boris. kondratieff @ colostate. edu</mods:affiliation>
<mods:nameIdentifier type="email">verdonec@gmail.comE-mail</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kondratieff, Boris C.</mods:namePart>
<mods:affiliation>Department of Bioagricultural Sciences and Pest Management, Colorado State University, Fort Collins, Colorado, 80523, U. S. A. E-mail: verdonec @ gmail. com E-mail: boris. kondratieff @ colostate. edu</mods:affiliation>
<mods:nameIdentifier type="email">verdonec@gmail.comE-mail</mods:nameIdentifier>
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<mods:date>2017</mods:date>
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<mods:number>13</mods:number>
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<taxonomicName authority="Verdone &amp; Kondratieff" authorityName="Verdone &amp; Kondratieff" authorityYear="2017" box="[860,1346,299,324]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="dewalti" status="sp. n.">
<emphasis bold="true" box="[860,1056,299,323]" italics="true" pageId="2" pageNumber="113">Isoperla dewalti</emphasis>
Verdone &amp; Kondratieff
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http://lsid.speciesfile.org/urn:lsid: 
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.speciesfile.org: TaxonName:501137
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(
<figureCitation box="[1081,1198,401,425]" captionStart-0="Fig" captionStart-1="Figs" captionStart-2="Figs" captionStart-3="Figs" captionStart-4="Figs" captionStart-5="Figs" captionStart-6="Figs" captionStartId-0="1.[189,226,968,992]" captionStartId-1="3.[189,238,1284,1308]" captionStartId-2="4.[189,238,1284,1308]" captionStartId-3="5.[189,238,1284,1308]" captionStartId-4="6.[189,238,1360,1384]" captionStartId-5="7.[189,238,1284,1308]" captionStartId-6="8.[189,238,1360,1384]" captionTargetBox-0="[248,1383,254,921]" captionTargetBox-1="[324,1307,218,1201]" captionTargetBox-2="[324,1307,218,1201]" captionTargetBox-3="[324,1307,218,1201]" captionTargetBox-4="[286,1344,218,1276]" captionTargetBox-5="[324,1307,218,1201]" captionTargetBox-6="[286,1344,218,1276]" captionTargetId-0="figure-409@1.[248,1383,254,921]" captionTargetId-1="figure-282@3.[324,1307,218,1201]" captionTargetId-2="figure-268@4.[324,1307,218,1201]" captionTargetId-3="figure-275@5.[324,1307,218,1201]" captionTargetId-4="figure-242@6.[286,1344,218,1276]" captionTargetId-5="figure-266@7.[324,1307,218,1201]" captionTargetId-6="figure-278@8.[286,1344,218,1276]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="8" captionText-0="Fig. 1. Adult habitus of Isoperla dewalti sp. n., Helton Creek Grayson Co., Virginia, 2X." captionText-1="Figs. 2–4. Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia, 5X. Fig. 2. Head and pronotum, dorsal. Fig. 3. Male sterna 7–9, ventral. Fig. 4. Male paraprocts, dorsal." captionText-2="Figs. 5–7. Aedeagus of Isoperla dewalti sp. n., lateral, 5x. A. (vestigial) posteroventral membranous aedeagal tube, B. Posteroventral lobe, C. Posterior sclerotized arrowhead shaped aedeagal spine plate with upturned distal stout spine, D. Paired posterolateral lobes, E. Posteromedial rectangular patch of dense, stout, rust colored spines, F. Dorsal lobe, G. Dense short sharp spinulae, H. Anterior lobe. Fig. 5. Helton Creek, Grayson Co., Virginia. Fig. 6. Tributary to Le Conte Creek, Sevier Co., Tennessee. Fig. 7. Helton Creek, Grayson Co., Virginia." captionText-3="Figs. 8–10. Aedeagus of Isoperla dewalti sp. n., posterior, 5x. B. Posteroventral lobe, C. Posterior sclerotized arrowhead shaped aedeagal spine plate with upturned distal stout spine, D. Paired posterolateral lobes, E. Posteromedial rectangular patch of dense, stout, rust colored spines, F. Dorsal lobe, G. Dense short sharp spinulae, Fig. 8. Helton Creek, Grayson Co., Virginia. Fig. 9. Tributary to Le Conte Creek, Sevier Co., Tennessee. Fig. 10. Helton Creek, Grayson Co., Virginia." captionText-4="Figs. 11–14. Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia. Fig. 11. Posterior sclerotized arrowhead shaped aedeagal spine plate with upturned distal stout spine and broad flat sclerotized plates with numerous small apical finger-like projections, 1,800X. Fig. 12. Short sharp spinulae, 700X. Fig. 13. Shelflike scales with apical hair-like projections, 1,200X. Fig. 14. Aedeagus, lateral, lateral area with dense short sharp spinulae (arrow) 65X." captionText-5="Figs. 15–17. Female subgenital plate of Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia, 5X. Fig. 15. Broadly rounded Fig. 16. Broadly triangular, truncate with shallow posteromedian emargination. Fig. 17. Broadly triangular." captionText-6="Figs. 18–20. Isoperla dewalti sp. n., Tributary to Le Conte Creek, Sevier Co., Tennessee. Fig. 18. Ovum, 400X. Fig. 19. Ovum collar, 1,800X. Fig. 20. Ovum follicle cell impressions and micropyles, 800X." figureDoi-0="http://doi.org/10.5281/zenodo.4757787" figureDoi-1="http://doi.org/10.5281/zenodo.4757791" figureDoi-2="http://doi.org/10.5281/zenodo.4757793" figureDoi-3="http://doi.org/10.5281/zenodo.4757797" figureDoi-4="http://doi.org/10.5281/zenodo.4757799" figureDoi-5="http://doi.org/10.5281/zenodo.4757803" figureDoi-6="http://doi.org/10.5281/zenodo.4757807" httpUri-0="https://zenodo.org/record/4757787/files/figure.png" httpUri-1="https://zenodo.org/record/4757791/files/figure.png" httpUri-2="https://zenodo.org/record/4757793/files/figure.png" httpUri-3="https://zenodo.org/record/4757797/files/figure.png" httpUri-4="https://zenodo.org/record/4757799/files/figure.png" httpUri-5="https://zenodo.org/record/4757803/files/figure.png" httpUri-6="https://zenodo.org/record/4757807/files/figure.png" pageId="2" pageNumber="113">Figs. 1–20</figureCitation>
)
</paragraph>
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<emphasis bold="true" box="[835,1074,472,496]" pageId="2" pageNumber="113">Material examined.</emphasis>
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<emphasis bold="true" pageId="2" pageNumber="113">
<collectingCountry box="[1223,1301,472,496]" name="United States of America" pageId="2" pageNumber="113">U.S.A.</collectingCountry>
<emphasis bold="true" box="[1311,1324,475,496]" italics="true" pageId="2" pageNumber="113">–</emphasis>
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: 
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,
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, 
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, 
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, 
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, 
<date box="[1235,1399,545,569]" pageId="2" pageNumber="113" value="2017-05-13">
<collectingDate box="[1235,1399,545,569]" pageId="2" pageNumber="113" value="2017-05-13">13 May 2017</collectingDate>
</date>
, 
<collectorName pageId="2" pageNumber="113">C. Verdone</collectorName>
, 
<collectorName box="[952,1145,581,605]" pageId="2" pageNumber="113">B.C. Kondratieff</collectorName>
(
<collectionCode box="[1162,1232,581,605]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="2" pageNumber="113" type="Other type of research institution/biorepository">INHS</collectionCode>
, Insect Collection 
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)
</materialsCitation>
. 
<emphasis box="[950,1056,617,640]" italics="true" pageId="2" pageNumber="113">
<typeStatus box="[950,1056,617,640]" pageId="2" pageNumber="113">Paratypes</typeStatus>
</emphasis>
: 
<emphasis bold="true" box="[1081,1443,616,641]" pageId="2" pageNumber="113">
<collectingRegion box="[1081,1279,616,641]" country="United States of America" name="North Carolina" pageId="2" pageNumber="113">North Carolina</collectingRegion>
: Avery Co.,
</emphasis>
Shantee Creek, Adams Apple Dr., SW of Foscoe, 
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, 
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, 
<date box="[1090,1236,689,713]" pageId="2" pageNumber="113" value="2010-05-13">13 May 2010</date>
, B.C. Kondratieff, 
<collectionCode box="[835,856,725,748]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="113">R</collectionCode>
.E. Zuellig, D. 
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. Lenat, 
<collectionCode box="[1134,1155,725,748]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="113">R</collectionCode>
.F. Kirchner, 
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(
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). 
<emphasis bold="true" box="[835,973,760,784]" pageId="2" pageNumber="113">Macon Co.,</emphasis>
Tellico Creek, Tellico Rd. ~ 
<quantity box="[1299,1378,760,784]" metricMagnitude="2" metricUnit="m" metricValue="8.04672" pageId="2" pageNumber="113" unit="mi" value="0.5">0.5 mi.</quantity>
W of Sugar Cove Rd., 
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, 
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, 
<date pageId="2" pageNumber="113" value="2017-05-15">15 May 2017</date>
, C. Verdone, B.C. Kondratieff, 
<specimenCount box="[1301,1321,832,856]" pageId="2" pageNumber="113" type="male">♂</specimenCount>
(
<collectionCode box="[1345,1428,833,857]" pageId="2" pageNumber="113">CSUIC</collectionCode>
). 
<emphasis bold="true" box="[835,966,868,892]" pageId="2" pageNumber="113">Swain Co.,</emphasis>
Beech Flats Prong, GSMNP, Hwy 441, 
<geoCoordinate degrees="35.60227" direction="north" orientation="latitude" pageId="2" pageNumber="113" precision="1" value="35.60227">N 35.60227</geoCoordinate>
, 
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, 
<date box="[1092,1239,904,929]" pageId="2" pageNumber="113" value="2017-05-16">16 May 2017</date>
, C. Verdone, B.C. Kondratieff, 
<specimenCount box="[987,1007,940,964]" pageId="2" pageNumber="113" type="male">♂</specimenCount>
(
<collectionCode box="[1023,1110,940,964]" pageId="2" pageNumber="113">CSUIC</collectionCode>
). 
<emphasis bold="true" box="[1129,1292,940,964]" pageId="2" pageNumber="113">Watauga Co.,</emphasis>
Boone Fork, CR 1561, W of Boone Fork, [
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, 
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], 
<date box="[835,984,1012,1037]" pageId="2" pageNumber="113" value="2005-05-02">2 May 2005</date>
, B.C. Kondratieff, 
<collectionCode box="[1224,1245,1013,1036]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="113">R</collectionCode>
.E. Zuellig, D. 
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. Lenat, 
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.F. Kirchner, 
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, 
<specimenCount box="[1143,1177,1048,1072]" pageId="2" pageNumber="113" type="female">2♀</specimenCount>
(
<collectionCode box="[1198,1285,1048,1072]" pageId="2" pageNumber="113">CSUIC</collectionCode>
). 
<emphasis bold="true" pageId="2" pageNumber="113">
<collectingRegion box="[1309,1438,1048,1072]" country="United States of America" name="Tennessee" pageId="2" pageNumber="113">Tennessee</collectingRegion>
: Sevier Co.,
</emphasis>
trib. to Le Conte Creek, GSMNP, Twin Creeks Uplands Research Lab, 
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, 
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, 
<date box="[947,1093,1156,1181]" pageId="2" pageNumber="113" value="2017-05-16">16 May 2017</date>
, C. Verdone, B.C. Kondratieff, 
<specimenCount box="[835,868,1192,1216]" pageId="2" pageNumber="113" type="male">2♂</specimenCount>
, 
<specimenCount box="[889,923,1192,1216]" pageId="2" pageNumber="113" type="female">2♀</specimenCount>
(
<collectionCode box="[945,1032,1192,1216]" pageId="2" pageNumber="113">CSUIC</collectionCode>
). 
<emphasis bold="true" box="[1057,1347,1192,1216]" pageId="2" pageNumber="113">
<collectingRegion box="[1057,1159,1192,1216]" country="United States of America" name="Virginia" pageId="2" pageNumber="113">Virginia</collectingRegion>
, Franklin Co.,
</emphasis>
Grassy Fork, [Co. Rd. 619, 2 mi. SE Snow Creek Rd.], [
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, 
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], 
<date box="[1169,1361,1264,1289]" pageId="2" pageNumber="113" value="2012-05-21">21 May 2012</date>
, B.C. Kondratieff, 
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.F. Kirchner, 
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(
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). 
<emphasis bold="true" box="[1287,1443,1300,1324]" pageId="2" pageNumber="113">Grayson Co.,</emphasis>
Same location as 
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, 
<date box="[1149,1295,1336,1361]" pageId="2" pageNumber="113" value="2017-05-12">12 May 2017</date>
, C. Verdone, B.C. Kondratieff, J. Richards, 
<collectionCode box="[1194,1215,1373,1396]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="113">R</collectionCode>
. Agbalog, 
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, 
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(
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); Same location as 
<typeStatus box="[1145,1252,1408,1432]" pageId="2" pageNumber="113">holotype</typeStatus>
, 
<date box="[1262,1408,1408,1433]" pageId="2" pageNumber="113" value="2017-05-13">13 May 2017</date>
, C. Verdone, B.C. Kondratieff, 
<specimenCount box="[1161,1195,1444,1468]" pageId="2" pageNumber="113" type="male">6♂</specimenCount>
, 
<specimenCount box="[1210,1257,1444,1468]" pageId="2" pageNumber="113" type="female">71♀</specimenCount>
(
<collectionCode box="[1273,1359,1444,1468]" pageId="2" pageNumber="113">CSUIC</collectionCode>
); Same location as 
<typeStatus box="[968,1073,1480,1504]" pageId="2" pageNumber="113">holotype</typeStatus>
, 
<date box="[1086,1236,1480,1505]" pageId="2" pageNumber="113" value="2017-05-17">17 May 2017</date>
, C. Verdone, B.C. Kondratieff, 
<specimenCount box="[985,1005,1516,1540]" pageId="2" pageNumber="113" type="female">♀</specimenCount>
(
<collectionCode box="[1020,1090,1516,1540]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="2" pageNumber="113" type="Other type of research institution/biorepository">INHS</collectionCode>
, Insect Collection 659584).
</paragraph>
</subSubSection>
<subSubSection box="[835,1358,1552,1576]" pageId="2" pageNumber="113" type="distribution">
<paragraph blockId="2.[835,1444,472,1828]" box="[835,1358,1552,1576]" pageId="2" pageNumber="113">
<emphasis bold="true" box="[835,994,1552,1576]" pageId="2" pageNumber="113">Distribution.</emphasis>
<collectingCountry box="[1001,1077,1552,1576]" name="United States of America" pageId="2" pageNumber="113">U.S.A.</collectingCountry>
– NC, TN, VA (
<figureCitation box="[1264,1343,1552,1576]" captionStart="Fig" captionStartId="10.[189,226,1826,1850]" captionTargetBox="[249,1383,284,1716]" captionTargetId="figure-61@10.[249,1383,283,1750]" captionTargetPageId="10" captionText="Fig. 22. Distribution map of examined material and physiographic provinces." figureDoi="http://doi.org/10.5281/zenodo.4757813" httpUri="https://zenodo.org/record/4757813/files/figure.png" pageId="2" pageNumber="113">Fig. 22</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="113" type="etymology">
<paragraph blockId="2.[835,1444,472,1828]" pageId="2" pageNumber="113">
<emphasis bold="true" box="[835,973,1588,1612]" pageId="2" pageNumber="113">Etymology.</emphasis>
The new species is named in honor of Dr. R. Edward DeWalt, 
<collectingRegion box="[1149,1231,1624,1648]" country="United States of America" name="Illinois" pageId="2" pageNumber="113">Illinois</collectingRegion>
Natural History Survey, Champaign, 
<collectingRegion box="[1085,1169,1660,1684]" country="United States of America" name="Illinois" pageId="2" pageNumber="113">Illinois</collectingRegion>
, for his dedication and numerous contributions to the study of stoneflies. The proposed common name is the “DeWalt Stripetail”.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="117" pageId="2" pageNumber="113" type="description">
<paragraph blockId="2.[835,1444,472,1828]" lastBlockId="5.[189,797,1536,1884]" lastPageId="5" lastPageNumber="116" pageId="2" pageNumber="113">
<emphasis bold="true" box="[835,904,1804,1828]" pageId="2" pageNumber="113">Male.</emphasis>
Macropterous; forewing length 
<quantity box="[1283,1443,1804,1828]" metricMagnitude="-2" metricUnit="m" metricValue="1.095" metricValueMax="1.13" metricValueMin="1.06" pageId="2" pageNumber="113" unit="mm" value="10.95" valueMax="11.3" valueMin="10.6">10.6–11.3 mm</quantity>
(n=5). Body length 9.6–10.0 mm (n=5). Head and pronotum pale yellow with contrasting pigmentation. General body color pale yellow with light brown to brown markings. (
<figureCitation box="[617,689,1536,1560]" captionStart="Fig" captionStartId="1.[189,226,968,992]" captionTargetBox="[248,1383,254,921]" captionTargetId="figure-409@1.[248,1383,254,921]" captionTargetPageId="1" captionText="Fig. 1. Adult habitus of Isoperla dewalti sp. n., Helton Creek Grayson Co., Virginia, 2X." figureDoi="http://doi.org/10.5281/zenodo.4757787" httpUri="https://zenodo.org/record/4757787/files/figure.png" pageId="3" pageNumber="114">Fig. 1</figureCitation>
). Dorsal background of head pale yellow with medium brown bands connecting ocelli; bands widest anteriorly; bands truncate at anterior ocellus; triangular dusky spot anterior to anterior ocellus (
<figureCitation box="[197,267,1716,1740]" captionStart="Figs" captionStartId="3.[189,238,1284,1308]" captionTargetBox="[324,1307,218,1201]" captionTargetId="figure-282@3.[324,1307,218,1201]" captionTargetPageId="3" captionText="Figs. 2–4. Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia, 5X. Fig. 2. Head and pronotum, dorsal. Fig. 3. Male sterna 7–9, ventral. Fig. 4. Male paraprocts, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4757791" httpUri="https://zenodo.org/record/4757791/files/figure.png" pageId="3" pageNumber="114">Fig. 2</figureCitation>
). Antennal scape pale yellow with a diffuse diagonal brown stripe; flagellum segments 1–7 pale; remaining segments medium brown (
<figureCitation box="[669,784,1788,1812]" captionStart-0="Fig" captionStart-1="Figs" captionStartId-0="1.[189,226,968,992]" captionStartId-1="3.[189,238,1284,1308]" captionTargetBox-0="[248,1383,254,921]" captionTargetBox-1="[324,1307,218,1201]" captionTargetId-0="figure-409@1.[248,1383,254,921]" captionTargetId-1="figure-282@3.[324,1307,218,1201]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionText-0="Fig. 1. Adult habitus of Isoperla dewalti sp. n., Helton Creek Grayson Co., Virginia, 2X." captionText-1="Figs. 2–4. Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia, 5X. Fig. 2. Head and pronotum, dorsal. Fig. 3. Male sterna 7–9, ventral. Fig. 4. Male paraprocts, dorsal." figureDoi-0="http://doi.org/10.5281/zenodo.4757787" figureDoi-1="http://doi.org/10.5281/zenodo.4757791" httpUri-0="https://zenodo.org/record/4757787/files/figure.png" httpUri-1="https://zenodo.org/record/4757791/files/figure.png" pageId="3" pageNumber="114">Figs. 1–2</figureCitation>
). Pronotum with a wide medial pale stripe; rugosities irregular, raised, medium brown; anterolateral margins medium brown; posterolateral margins pale (
<figureCitation box="[901,971,1464,1488]" captionStart="Figs" captionStartId="3.[189,238,1284,1308]" captionTargetBox="[324,1307,218,1201]" captionTargetId="figure-282@3.[324,1307,218,1201]" captionTargetPageId="3" captionText="Figs. 2–4. Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia, 5X. Fig. 2. Head and pronotum, dorsal. Fig. 3. Male sterna 7–9, ventral. Fig. 4. Male paraprocts, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4757791" httpUri="https://zenodo.org/record/4757791/files/figure.png" pageId="3" pageNumber="114">Fig. 2</figureCitation>
). Femora light brown dorsally, covered with uniformly spaced, short, brown setae; femora pale ventrally; tibia light brown, covered with uniformly spaced, short brown setae; tarsi medium brown (
<figureCitation box="[933,1003,1608,1632]" captionStart="Fig" captionStartId="1.[189,226,968,992]" captionTargetBox="[248,1383,254,921]" captionTargetId="figure-409@1.[248,1383,254,921]" captionTargetPageId="1" captionText="Fig. 1. Adult habitus of Isoperla dewalti sp. n., Helton Creek Grayson Co., Virginia, 2X." figureDoi="http://doi.org/10.5281/zenodo.4757787" httpUri="https://zenodo.org/record/4757787/files/figure.png" pageId="3" pageNumber="114">Fig. 1</figureCitation>
). Meso- and metanota light brown (
<figureCitation box="[843,917,1644,1668]" captionStart="Fig" captionStartId="1.[189,226,968,992]" captionTargetBox="[248,1383,254,921]" captionTargetId="figure-409@1.[248,1383,254,921]" captionTargetPageId="1" captionText="Fig. 1. Adult habitus of Isoperla dewalti sp. n., Helton Creek Grayson Co., Virginia, 2X." figureDoi="http://doi.org/10.5281/zenodo.4757787" httpUri="https://zenodo.org/record/4757787/files/figure.png" pageId="3" pageNumber="114">Fig. 1</figureCitation>
). Wings hyaline, veins of forewing light basally, brown distally (
<figureCitation box="[1126,1193,1680,1704]" captionStart="Fig" captionStartId="1.[189,226,968,992]" captionTargetBox="[248,1383,254,921]" captionTargetId="figure-409@1.[248,1383,254,921]" captionTargetPageId="1" captionText="Fig. 1. Adult habitus of Isoperla dewalti sp. n., Helton Creek Grayson Co., Virginia, 2X." figureDoi="http://doi.org/10.5281/zenodo.4757787" httpUri="https://zenodo.org/record/4757787/files/figure.png" pageId="3" pageNumber="114">Fig. 1</figureCitation>
). Terga light brown, posterior ⅓ medium brown; terga 1–8 with uniformly spaced brown setae. Sterna pale yellow to light brown; sternum 8 with a light brown recessed vesicle, 0.65X as wide as long; evenly rounded, extending to the posterior margin of sternum 8 (
<figureCitation box="[197,281,1572,1596]" captionStart="Figs" captionStartId="3.[189,238,1284,1308]" captionTargetBox="[324,1307,218,1201]" captionTargetId="figure-282@3.[324,1307,218,1201]" captionTargetPageId="3" captionText="Figs. 2–4. Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia, 5X. Fig. 2. Head and pronotum, dorsal. Fig. 3. Male sterna 7–9, ventral. Fig. 4. Male paraprocts, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4757791" httpUri="https://zenodo.org/record/4757791/files/figure.png" pageId="4" pageNumber="115">Fig. 3</figureCitation>
). Paraprocts short, triangular, slightly incurved, moderately sclerotized; trichoid sensilla scattered over dorsal surface, absent on apex; apex heavily sclerotized; tips acutely pointed (
<figureCitation box="[710,785,1680,1704]" captionStart="Figs" captionStartId="3.[189,238,1284,1308]" captionTargetBox="[324,1307,218,1201]" captionTargetId="figure-282@3.[324,1307,218,1201]" captionTargetPageId="3" captionText="Figs. 2–4. Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia, 5X. Fig. 2. Head and pronotum, dorsal. Fig. 3. Male sterna 7–9, ventral. Fig. 4. Male paraprocts, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4757791" httpUri="https://zenodo.org/record/4757791/files/figure.png" pageId="4" pageNumber="115">Fig. 4</figureCitation>
). Cerci brown, setose; cercal segments with a single, long, stout, ventral hair at posterior margin. Aedeagus with a long basal stalk, a highly reduced (vestigial) posteroventral membranous aedeagal tube, a posteroventral lobe, paired posterolateral lobes, a dorsal lobe, and an anterior lobe (
<figureCitation captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="4.[189,238,1284,1308]" captionStartId-1="5.[189,238,1284,1308]" captionTargetBox-0="[324,1307,218,1201]" captionTargetBox-1="[324,1307,218,1201]" captionTargetId-0="figure-268@4.[324,1307,218,1201]" captionTargetId-1="figure-275@5.[324,1307,218,1201]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Figs. 5–7. Aedeagus of Isoperla dewalti sp. n., lateral, 5x. A. (vestigial) posteroventral membranous aedeagal tube, B. Posteroventral lobe, C. Posterior sclerotized arrowhead shaped aedeagal spine plate with upturned distal stout spine, D. Paired posterolateral lobes, E. Posteromedial rectangular patch of dense, stout, rust colored spines, F. Dorsal lobe, G. Dense short sharp spinulae, H. Anterior lobe. Fig. 5. Helton Creek, Grayson Co., Virginia. Fig. 6. Tributary to Le Conte Creek, Sevier Co., Tennessee. Fig. 7. Helton Creek, Grayson Co., Virginia." captionText-1="Figs. 8–10. Aedeagus of Isoperla dewalti sp. n., posterior, 5x. B. Posteroventral lobe, C. Posterior sclerotized arrowhead shaped aedeagal spine plate with upturned distal stout spine, D. Paired posterolateral lobes, E. Posteromedial rectangular patch of dense, stout, rust colored spines, F. Dorsal lobe, G. Dense short sharp spinulae, Fig. 8. Helton Creek, Grayson Co., Virginia. Fig. 9. Tributary to Le Conte Creek, Sevier Co., Tennessee. Fig. 10. Helton Creek, Grayson Co., Virginia." figureDoi-0="http://doi.org/10.5281/zenodo.4757793" figureDoi-1="http://doi.org/10.5281/zenodo.4757797" httpUri-0="https://zenodo.org/record/4757793/files/figure.png" httpUri-1="https://zenodo.org/record/4757797/files/figure.png" pageId="4" pageNumber="115">Figs. 5– 10</figureCitation>
); posteroventral membranous aedeagal tube highly reduced (vestigial), appearing only occasionally as a small lobe at the base of the basal stalk (
<figureCitation box="[919,1000,1716,1740]" captionStart="Figs" captionStartId="4.[189,238,1284,1308]" captionTargetBox="[324,1307,218,1201]" captionTargetId="figure-268@4.[324,1307,218,1201]" captionTargetPageId="4" captionText="Figs. 5–7. Aedeagus of Isoperla dewalti sp. n., lateral, 5x. A. (vestigial) posteroventral membranous aedeagal tube, B. Posteroventral lobe, C. Posterior sclerotized arrowhead shaped aedeagal spine plate with upturned distal stout spine, D. Paired posterolateral lobes, E. Posteromedial rectangular patch of dense, stout, rust colored spines, F. Dorsal lobe, G. Dense short sharp spinulae, H. Anterior lobe. Fig. 5. Helton Creek, Grayson Co., Virginia. Fig. 6. Tributary to Le Conte Creek, Sevier Co., Tennessee. Fig. 7. Helton Creek, Grayson Co., Virginia." figureDoi="http://doi.org/10.5281/zenodo.4757793" httpUri="https://zenodo.org/record/4757793/files/figure.png" pageId="4" pageNumber="115">Fig. 5</figureCitation>
); aedeagus with a posteromedial rectangular patch of dense, stout, rust colored spines, and a single posterior sclerotized arrowhead shaped aedeagal spine plate with upturned distal stout spine (
<figureCitation box="[979,1098,1860,1884]" captionStart="Figs" captionStartId="5.[189,238,1284,1308]" captionTargetBox="[324,1307,218,1201]" captionTargetId="figure-275@5.[324,1307,218,1201]" captionTargetPageId="5" captionText="Figs. 8–10. Aedeagus of Isoperla dewalti sp. n., posterior, 5x. B. Posteroventral lobe, C. Posterior sclerotized arrowhead shaped aedeagal spine plate with upturned distal stout spine, D. Paired posterolateral lobes, E. Posteromedial rectangular patch of dense, stout, rust colored spines, F. Dorsal lobe, G. Dense short sharp spinulae, Fig. 8. Helton Creek, Grayson Co., Virginia. Fig. 9. Tributary to Le Conte Creek, Sevier Co., Tennessee. Fig. 10. Helton Creek, Grayson Co., Virginia." figureDoi="http://doi.org/10.5281/zenodo.4757797" httpUri="https://zenodo.org/record/4757797/files/figure.png" pageId="4" pageNumber="115">Figs. 8–10</figureCitation>
); apex of structure upturned with approximately 10 large stout rust colored spines (
<figureCitation box="[290,380,1572,1596]" captionStart="Figs" captionStartId="6.[189,238,1360,1384]" captionTargetBox="[286,1344,218,1276]" captionTargetId="figure-242@6.[286,1344,218,1276]" captionTargetPageId="6" captionText="Figs. 11–14. Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia. Fig. 11. Posterior sclerotized arrowhead shaped aedeagal spine plate with upturned distal stout spine and broad flat sclerotized plates with numerous small apical finger-like projections, 1,800X. Fig. 12. Short sharp spinulae, 700X. Fig. 13. Shelflike scales with apical hair-like projections, 1,200X. Fig. 14. Aedeagus, lateral, lateral area with dense short sharp spinulae (arrow) 65X." figureDoi="http://doi.org/10.5281/zenodo.4757799" httpUri="https://zenodo.org/record/4757799/files/figure.png" pageId="5" pageNumber="116">Fig. 11</figureCitation>
); base of plate with broad flat sclerotized plates with numerous small apical finger-like projections (
<figureCitation box="[479,567,1644,1668]" captionStart="Figs" captionStartId="6.[189,238,1360,1384]" captionTargetBox="[286,1344,218,1276]" captionTargetId="figure-242@6.[286,1344,218,1276]" captionTargetPageId="6" captionText="Figs. 11–14. Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia. Fig. 11. Posterior sclerotized arrowhead shaped aedeagal spine plate with upturned distal stout spine and broad flat sclerotized plates with numerous small apical finger-like projections, 1,800X. Fig. 12. Short sharp spinulae, 700X. Fig. 13. Shelflike scales with apical hair-like projections, 1,200X. Fig. 14. Aedeagus, lateral, lateral area with dense short sharp spinulae (arrow) 65X." figureDoi="http://doi.org/10.5281/zenodo.4757799" httpUri="https://zenodo.org/record/4757799/files/figure.png" pageId="5" pageNumber="116">Fig. 11</figureCitation>
); basal stalk with short sharp spinulae (
<figureCitation box="[455,539,1680,1704]" captionStart="Figs" captionStartId="6.[189,238,1360,1384]" captionTargetBox="[286,1344,218,1276]" captionTargetId="figure-242@6.[286,1344,218,1276]" captionTargetPageId="6" captionText="Figs. 11–14. Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia. Fig. 11. Posterior sclerotized arrowhead shaped aedeagal spine plate with upturned distal stout spine and broad flat sclerotized plates with numerous small apical finger-like projections, 1,800X. Fig. 12. Short sharp spinulae, 700X. Fig. 13. Shelflike scales with apical hair-like projections, 1,200X. Fig. 14. Aedeagus, lateral, lateral area with dense short sharp spinulae (arrow) 65X." figureDoi="http://doi.org/10.5281/zenodo.4757799" httpUri="https://zenodo.org/record/4757799/files/figure.png" pageId="5" pageNumber="116">Fig. 12</figureCitation>
); posterolateral lobes with sparse scale like plates medially; posteromedial area with shallow shelf-like scales with apical hair-like projections (
<figureCitation box="[571,649,1788,1812]" captionStart="Figs" captionStartId="6.[189,238,1360,1384]" captionTargetBox="[286,1344,218,1276]" captionTargetId="figure-242@6.[286,1344,218,1276]" captionTargetPageId="6" captionText="Figs. 11–14. Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia. Fig. 11. Posterior sclerotized arrowhead shaped aedeagal spine plate with upturned distal stout spine and broad flat sclerotized plates with numerous small apical finger-like projections, 1,800X. Fig. 12. Short sharp spinulae, 700X. Fig. 13. Shelflike scales with apical hair-like projections, 1,200X. Fig. 14. Aedeagus, lateral, lateral area with dense short sharp spinulae (arrow) 65X." figureDoi="http://doi.org/10.5281/zenodo.4757799" httpUri="https://zenodo.org/record/4757799/files/figure.png" pageId="5" pageNumber="116">Fig. 13</figureCitation>
); lateral area with dense short sharp spinulae (
<figureCitation box="[581,663,1824,1848]" captionStart="Figs" captionStartId="6.[189,238,1360,1384]" captionTargetBox="[286,1344,218,1276]" captionTargetId="figure-242@6.[286,1344,218,1276]" captionTargetPageId="6" captionText="Figs. 11–14. Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia. Fig. 11. Posterior sclerotized arrowhead shaped aedeagal spine plate with upturned distal stout spine and broad flat sclerotized plates with numerous small apical finger-like projections, 1,800X. Fig. 12. Short sharp spinulae, 700X. Fig. 13. Shelflike scales with apical hair-like projections, 1,200X. Fig. 14. Aedeagus, lateral, lateral area with dense short sharp spinulae (arrow) 65X." figureDoi="http://doi.org/10.5281/zenodo.4757799" httpUri="https://zenodo.org/record/4757799/files/figure.png" pageId="5" pageNumber="116">Fig. 14</figureCitation>
).
</paragraph>
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<paragraph blockId="3.[189,1442,1284,1344]" pageId="3" pageNumber="114">
Figs. 2–4. 
<taxonomicName authority="Verdone &amp; Kondratieff, 2017" authorityName="Verdone &amp; Kondratieff" authorityYear="2017" box="[303,471,1284,1308]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="species" species="dewalti" status="sp. n.">
<emphasis box="[303,471,1284,1308]" italics="true" pageId="3" pageNumber="114">Isoperla dewalti</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[477,540,1285,1308]" pageId="3" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
, Helton Creek, Grayson Co., Virginia, 5X. Fig. 2. Head and pronotum, dorsal. Fig. 3. Male sterna 7–9, ventral. Fig. 4. Male paraprocts, dorsal.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4757793" ID-Zenodo-Dep="4757793" httpUri="https://zenodo.org/record/4757793/files/figure.png" pageId="4" pageNumber="115" startId="4.[189,238,1284,1308]" targetBox="[324,1307,218,1201]" targetPageId="4">
<paragraph blockId="4.[189,1443,1284,1488]" pageId="4" pageNumber="115">
Figs. 5–7. Aedeagus of 
<taxonomicName authority="Verdone &amp; Kondratieff, 2017" authorityName="Verdone &amp; Kondratieff" authorityYear="2017" box="[460,628,1284,1308]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="dewalti" status="sp. n.">
<emphasis box="[460,628,1284,1308]" italics="true" pageId="4" pageNumber="115">Isoperla dewalti</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[636,699,1285,1308]" pageId="4" pageNumber="115" rank="species">sp. n.</taxonomicNameLabel>
, lateral, 5x. A. (vestigial) posteroventral membranous aedeagal tube, B. Posteroventral lobe, C. Posterior sclerotized arrowhead shaped aedeagal spine plate with upturned distal stout spine, D. Paired posterolateral lobes, E. Posteromedial rectangular patch of dense, stout, rust colored spines, F. Dorsal lobe, G. Dense short sharp spinulae, H. Anterior lobe. Fig. 5. Helton Creek, Grayson Co., Virginia. Fig. 6. Tributary to Le Conte Creek, Sevier Co., Tennessee. Fig. 7. Helton Creek, Grayson Co., Virginia.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4757797" ID-Zenodo-Dep="4757797" httpUri="https://zenodo.org/record/4757797/files/figure.png" pageId="5" pageNumber="116" startId="5.[189,238,1284,1308]" targetBox="[324,1307,218,1201]" targetPageId="5">
<paragraph blockId="5.[189,1443,1284,1452]" pageId="5" pageNumber="116">
Figs. 8–10. Aedeagus of 
<taxonomicName authority="Verdone &amp; Kondratieff, 2017" authorityName="Verdone &amp; Kondratieff" authorityYear="2017" box="[476,646,1284,1308]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="116" phylum="Arthropoda" rank="species" species="dewalti" status="sp. n.">
<emphasis box="[476,646,1284,1308]" italics="true" pageId="5" pageNumber="116">Isoperla dewalti</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[654,719,1285,1308]" pageId="5" pageNumber="116" rank="species">sp. n.</taxonomicNameLabel>
, posterior, 5x. B. Posteroventral lobe, C. Posterior sclerotized arrowhead shaped aedeagal spine plate with upturned distal stout spine, D. Paired posterolateral lobes, E. Posteromedial rectangular patch of dense, stout, rust colored spines, F. Dorsal lobe, G. Dense short sharp spinulae, Fig. 8. Helton Creek, Grayson Co., Virginia. Fig. 9. Tributary to Le Conte Creek, Sevier Co., Tennessee. Fig. 10. Helton Creek, Grayson Co., Virginia.
</paragraph>
</caption>
<paragraph blockId="5.[189,797,1536,1884]" lastBlockId="5.[835,1443,1536,1884]" pageId="5" pageNumber="116">
<emphasis bold="true" box="[189,284,1860,1884]" pageId="5" pageNumber="116">Female.</emphasis>
Macropterous; forewing wing length 
<quantity box="[737,934,1536,1884]" metricMagnitude="-2" metricUnit="m" metricValue="1.1949999999999998" metricValueMax="1.26" metricValueMin="1.13" pageId="5" pageNumber="116" unit="mm" value="11.95" valueMax="12.6" valueMin="11.3">11.3– 12.6 mm</quantity>
(n=5). Body length 
<quantity box="[1158,1304,1536,1560]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.09" metricValueMin="0.9099999999999999" pageId="5" pageNumber="116" unit="mm" value="10.0" valueMax="10.9" valueMin="9.1">9.1–10.9 mm</quantity>
(n=5). Body coloration and morphology similar to male. Sternum 8 with variable subgenital plate extending approximately ⅓–½ over sternum 9; subgenital plate broadly rounded (
<figureCitation box="[1167,1264,1680,1704]" captionStart="Figs" captionStartId="7.[189,238,1284,1308]" captionTargetBox="[324,1307,218,1201]" captionTargetId="figure-266@7.[324,1307,218,1201]" captionTargetPageId="7" captionText="Figs. 15–17. Female subgenital plate of Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia, 5X. Fig. 15. Broadly rounded Fig. 16. Broadly triangular, truncate with shallow posteromedian emargination. Fig. 17. Broadly triangular." figureDoi="http://doi.org/10.5281/zenodo.4757803" httpUri="https://zenodo.org/record/4757803/files/figure.png" pageId="5" pageNumber="116">Fig. 15</figureCitation>
), or broadly triangular, truncate with shallow posteromedian emargination (
<figureCitation box="[1012,1097,1752,1776]" captionStart="Figs" captionStartId="7.[189,238,1284,1308]" captionTargetBox="[324,1307,218,1201]" captionTargetId="figure-266@7.[324,1307,218,1201]" captionTargetPageId="7" captionText="Figs. 15–17. Female subgenital plate of Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia, 5X. Fig. 15. Broadly rounded Fig. 16. Broadly triangular, truncate with shallow posteromedian emargination. Fig. 17. Broadly triangular." figureDoi="http://doi.org/10.5281/zenodo.4757803" httpUri="https://zenodo.org/record/4757803/files/figure.png" pageId="5" pageNumber="116">Fig. 16</figureCitation>
), or broadly triangular (
<figureCitation captionStart="Figs" captionStartId="7.[189,238,1284,1308]" captionTargetBox="[324,1307,218,1201]" captionTargetId="figure-266@7.[324,1307,218,1201]" captionTargetPageId="7" captionText="Figs. 15–17. Female subgenital plate of Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia, 5X. Fig. 15. Broadly rounded Fig. 16. Broadly triangular, truncate with shallow posteromedian emargination. Fig. 17. Broadly triangular." figureDoi="http://doi.org/10.5281/zenodo.4757803" httpUri="https://zenodo.org/record/4757803/files/figure.png" pageId="5" pageNumber="116">Fig. 17</figureCitation>
).
</paragraph>
<paragraph blockId="5.[835,1443,1536,1884]" lastBlockId="6.[835,1443,1612,1888]" lastPageId="6" lastPageNumber="117" pageId="5" pageNumber="116">
<emphasis bold="true" box="[835,918,1823,1847]" pageId="5" pageNumber="116">Ovum.</emphasis>
Length 226.3–234.4 μm (n=2); width 184.9– 192.5 μm (n=2). General shape oblong; cross section triangular, distinct cross shaped ridges dividing anterior and posterior poles; anterior end concave (
<figureCitation box="[197,280,1684,1708]" captionStart="Figs" captionStartId="8.[189,238,1360,1384]" captionTargetBox="[286,1344,218,1276]" captionTargetId="figure-278@8.[286,1344,218,1276]" captionTargetPageId="8" captionText="Figs. 18–20. Isoperla dewalti sp. n., Tributary to Le Conte Creek, Sevier Co., Tennessee. Fig. 18. Ovum, 400X. Fig. 19. Ovum collar, 1,800X. Fig. 20. Ovum follicle cell impressions and micropyles, 800X." figureDoi="http://doi.org/10.5281/zenodo.4757807" httpUri="https://zenodo.org/record/4757807/files/figure.png" pageId="6" pageNumber="117">Fig. 18</figureCitation>
). Color pale brown. Collar well-developed, flared apically with elevated longitudinal ridges; base not offset from egg body (
<figureCitation box="[565,647,1756,1780]" captionStart="Figs" captionStartId="8.[189,238,1360,1384]" captionTargetBox="[286,1344,218,1276]" captionTargetId="figure-278@8.[286,1344,218,1276]" captionTargetPageId="8" captionText="Figs. 18–20. Isoperla dewalti sp. n., Tributary to Le Conte Creek, Sevier Co., Tennessee. Fig. 18. Ovum, 400X. Fig. 19. Ovum collar, 1,800X. Fig. 20. Ovum follicle cell impressions and micropyles, 800X." figureDoi="http://doi.org/10.5281/zenodo.4757807" httpUri="https://zenodo.org/record/4757807/files/figure.png" pageId="6" pageNumber="117">Fig. 19</figureCitation>
). Hexagonal follicle cell impressions well-developed with thickened ridges and with numerous small mounds between reticulations (
<figureCitation box="[470,615,1864,1888]" captionStart="Figs" captionStartId="8.[189,238,1360,1384]" captionTargetBox="[286,1344,218,1276]" captionTargetId="figure-278@8.[286,1344,218,1276]" captionTargetPageId="8" captionText="Figs. 18–20. Isoperla dewalti sp. n., Tributary to Le Conte Creek, Sevier Co., Tennessee. Fig. 18. Ovum, 400X. Fig. 19. Ovum collar, 1,800X. Fig. 20. Ovum follicle cell impressions and micropyles, 800X." figureDoi="http://doi.org/10.5281/zenodo.4757807" httpUri="https://zenodo.org/record/4757807/files/figure.png" pageId="6" pageNumber="117">Figs. 18, 20</figureCitation>
). Eclosion line absent, micropyles arranged singularly or in threes, positioned on top of cross ridges near anterior ⅓ of egg (
<figureCitation box="[892,1025,1684,1708]" captionStart="Figs" captionStartId="8.[189,238,1360,1384]" captionTargetBox="[286,1344,218,1276]" captionTargetId="figure-278@8.[286,1344,218,1276]" captionTargetPageId="8" captionText="Figs. 18–20. Isoperla dewalti sp. n., Tributary to Le Conte Creek, Sevier Co., Tennessee. Fig. 18. Ovum, 400X. Fig. 19. Ovum collar, 1,800X. Fig. 20. Ovum follicle cell impressions and micropyles, 800X." figureDoi="http://doi.org/10.5281/zenodo.4757807" httpUri="https://zenodo.org/record/4757807/files/figure.png" pageId="6" pageNumber="117">Figs. 18, 20</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4757799" ID-Zenodo-Dep="4757799" httpUri="https://zenodo.org/record/4757799/files/figure.png" pageId="6" pageNumber="117" startId="6.[189,238,1360,1384]" targetBox="[286,1344,218,1276]" targetPageId="6">
<paragraph blockId="6.[189,1444,1360,1528]" pageId="6" pageNumber="117">
Figs. 11–14. 
<taxonomicName authority="Verdone &amp; Kondratieff, 2017" authorityName="Verdone &amp; Kondratieff" authorityYear="2017" box="[344,519,1360,1384]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="117" phylum="Arthropoda" rank="species" species="dewalti" status="sp. n.">
<emphasis box="[344,519,1360,1384]" italics="true" pageId="6" pageNumber="117">Isoperla dewalti</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[532,601,1361,1384]" pageId="6" pageNumber="117" rank="species">sp. n.</taxonomicNameLabel>
, Helton Creek, Grayson Co., Virginia. Fig. 11. Posterior sclerotized arrowhead shaped aedeagal spine plate with upturned distal stout spine and broad flat sclerotized plates with numerous small apical finger-like projections, 1,800X. Fig. 12. Short sharp spinulae, 700X. Fig. 13. Shelflike scales with apical hair-like projections, 1,200X. Fig. 14. Aedeagus, lateral, lateral area with dense short sharp spinulae (arrow) 65X.
</paragraph>
</caption>
<paragraph blockId="6.[835,1443,1612,1888]" box="[835,1042,1720,1744]" pageId="6" pageNumber="117">
<emphasis bold="true" box="[835,910,1720,1744]" pageId="6" pageNumber="117">Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="120" pageId="6" pageNumber="117" type="diagnosis">
<paragraph blockId="6.[835,1443,1612,1888]" lastBlockId="7.[189,798,1464,1848]" lastPageId="7" lastPageNumber="118" pageId="6" pageNumber="117">
<emphasis bold="true" box="[835,964,1755,1779]" pageId="6" pageNumber="117">Diagnosis.</emphasis>
<taxonomicName authorityName="Verdone &amp; Kondratieff" authorityYear="2017" box="[971,1139,1755,1779]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="117" phylum="Arthropoda" rank="species" species="dewalti">
<emphasis box="[971,1139,1755,1779]" italics="true" pageId="6" pageNumber="117">Isoperla dewalti</emphasis>
</taxonomicName>
is placed in the 
<taxonomicName baseAuthorityName="Banks" baseAuthorityYear="1989" box="[1327,1443,1756,1779]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="117" phylum="Arthropoda" rank="species" species="montana">
<emphasis box="[1327,1443,1756,1779]" italics="true" pageId="6" pageNumber="117">I. montana</emphasis>
</taxonomicName>
Group based on the presence of a single posterior sclerotized arrowhead shaped aedeagal spine plate with an upturned distal stout spine, dark brown bars connecting the ocelli, and–albeit vestigial–a posteroventral membranous aedeagal tube. The new species is easily distinguished from 
<taxonomicName baseAuthorityName="Banks" baseAuthorityYear="1989" box="[679,797,1537,1560]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="montana">
<emphasis box="[679,797,1537,1560]" italics="true" pageId="7" pageNumber="118">I. montana</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Szczytko &amp; Kondratieff" authorityYear="2015" box="[251,361,1572,1596]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="nelsoni">
<emphasis box="[251,361,1572,1596]" italics="true" pageId="7" pageNumber="118">I. nelsoni</emphasis>
</taxonomicName>
by the lack of a well-developed posteroventral membranous aedeagal tube.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4757803" ID-Zenodo-Dep="4757803" httpUri="https://zenodo.org/record/4757803/files/figure.png" pageId="7" pageNumber="118" startId="7.[189,238,1284,1308]" targetBox="[324,1307,218,1201]" targetPageId="7">
<paragraph blockId="7.[189,1443,1284,1380]" pageId="7" pageNumber="118">
Figs. 15–17. Female subgenital plate of 
<taxonomicName authority="Verdone &amp; Kondratieff, 2017" authorityName="Verdone &amp; Kondratieff" authorityYear="2017" box="[653,822,1284,1308]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="dewalti" status="sp. n.">
<emphasis box="[653,822,1284,1308]" italics="true" pageId="7" pageNumber="118">Isoperla dewalti</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[830,894,1285,1308]" pageId="7" pageNumber="118" rank="species">sp. n.</taxonomicNameLabel>
, Helton Creek, Grayson Co., Virginia, 5X. Fig. 15. Broadly rounded Fig. 16. Broadly triangular, truncate with shallow posteromedian emargination. Fig. 17. Broadly triangular.
</paragraph>
</caption>
<paragraph blockId="7.[189,798,1464,1848]" lastBlockId="8.[189,798,1504,1924]" lastPageId="8" lastPageNumber="119" pageId="7" pageNumber="118">
Several 
<typeStatus box="[321,426,1644,1668]" pageId="7" pageNumber="118">paratype</typeStatus>
specimens of 
<taxonomicName authorityName="Verdone &amp; Kondratieff" authorityYear="2017" box="[597,698,1644,1668]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="dewalti">
<emphasis box="[597,698,1644,1668]" italics="true" pageId="7" pageNumber="118">I. dewalti</emphasis>
</taxonomicName>
that are deposited in the C. P. Gillette Museum of Arthropod Diversity had previously been identified as 
<taxonomicName baseAuthorityName="Banks" baseAuthorityYear="1989" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="montana">
<emphasis italics="true" pageId="7" pageNumber="118">I. montana</emphasis>
</taxonomicName>
. The two species are superficially similar and it was likely thought that the posteroventral membranous aedeagal tube did not evert. However, it is the experience of the authors that when a welldeveloped aedeagal tube is present it is the first structure to evert. Additionally, differences in the posteromedial patch of spines distinguish the two species. In 
<taxonomicName authorityName="Verdone &amp; Kondratieff" authorityYear="2017" box="[979,1088,1608,1632]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="dewalti">
<emphasis box="[979,1088,1608,1632]" italics="true" pageId="7" pageNumber="118">I. dewalti</emphasis>
</taxonomicName>
the posteromedial patch of spines is broad, appearing as a uniform rectangular patch (
<figureCitation box="[927,1058,1680,1704]" captionStart="Figs" captionStartId="5.[189,238,1284,1308]" captionTargetBox="[324,1307,218,1201]" captionTargetId="figure-275@5.[324,1307,218,1201]" captionTargetPageId="5" captionText="Figs. 8–10. Aedeagus of Isoperla dewalti sp. n., posterior, 5x. B. Posteroventral lobe, C. Posterior sclerotized arrowhead shaped aedeagal spine plate with upturned distal stout spine, D. Paired posterolateral lobes, E. Posteromedial rectangular patch of dense, stout, rust colored spines, F. Dorsal lobe, G. Dense short sharp spinulae, Fig. 8. Helton Creek, Grayson Co., Virginia. Fig. 9. Tributary to Le Conte Creek, Sevier Co., Tennessee. Fig. 10. Helton Creek, Grayson Co., Virginia." figureDoi="http://doi.org/10.5281/zenodo.4757797" httpUri="https://zenodo.org/record/4757797/files/figure.png" pageId="7" pageNumber="118">Figs. 8–10</figureCitation>
). Whereas, in 
<taxonomicName baseAuthorityName="Banks" baseAuthorityYear="1989" box="[1259,1388,1681,1704]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="montana">
<emphasis box="[1259,1388,1681,1704]" italics="true" pageId="7" pageNumber="118">I. montana</emphasis>
</taxonomicName>
the posteromedial patch of spines is narrow, appearing as a thin ridge (figs. 33.15–33.16, 
<bibRefCitation author="Szczytko, S. W. &amp; B. C. Kondratieff" pageId="7" pageNumber="118" pagination="1 - 289" refId="ref8269" refString="Szczytko, S. W. &amp; B. C. Kondratieff. 2015. A review of the eastern Nearctic Isoperlinae (Plecoptera: Perlodidae) with the description of twenty-two new species. Monographs of Illiesia, Number 1: 1 - 289. http: // illiesia. speciesfile. org / papers / Monographiae-of-Illiesia. pdf" type="book chapter" year="2015">Szczytko &amp; Kondratieff 2015</bibRefCitation>
). Furthermore, 
<taxonomicName baseAuthorityName="Banks" baseAuthorityYear="1989" box="[1299,1443,1789,1812]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="montana">
<emphasis box="[1299,1443,1789,1812]" italics="true" pageId="7" pageNumber="118">I. montana</emphasis>
</taxonomicName>
possesses a dorsal rounded mound of dense, long, stout, rust colored spines above the posteromedial patch of spines (figs. 33.15–33.16, 
<bibRefCitation author="Szczytko, S. W. &amp; B. C. Kondratieff" pageId="8" pageNumber="119" pagination="1 - 289" refId="ref8269" refString="Szczytko, S. W. &amp; B. C. Kondratieff. 2015. A review of the eastern Nearctic Isoperlinae (Plecoptera: Perlodidae) with the description of twenty-two new species. Monographs of Illiesia, Number 1: 1 - 289. http: // illiesia. speciesfile. org / papers / Monographiae-of-Illiesia. pdf" type="book chapter" year="2015">Szczytko &amp; Kondratieff 2015</bibRefCitation>
), which the new species lacks.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4757807" ID-Zenodo-Dep="4757807" httpUri="https://zenodo.org/record/4757807/files/figure.png" pageId="8" pageNumber="119" startId="8.[189,238,1360,1384]" targetBox="[286,1344,218,1276]" targetPageId="8">
<paragraph blockId="8.[189,1444,1360,1420]" pageId="8" pageNumber="119">
Figs. 18–20. 
<taxonomicName authority="Verdone &amp; Kondratieff, 2017" authorityName="Verdone &amp; Kondratieff" authorityYear="2017" box="[332,501,1360,1384]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="dewalti" status="sp. n.">
<emphasis box="[332,501,1360,1384]" italics="true" pageId="8" pageNumber="119">Isoperla dewalti</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[508,571,1361,1384]" pageId="8" pageNumber="119" rank="species">sp. n.</taxonomicNameLabel>
, Tributary to Le Conte Creek, Sevier Co., Tennessee. Fig. 18. Ovum, 400X. Fig. 19. Ovum collar, 1,800X. Fig. 20. Ovum follicle cell impressions and micropyles, 800X.
</paragraph>
</caption>
<paragraph blockId="8.[189,798,1504,1924]" lastBlockId="9.[189,798,1145,1925]" lastPageId="9" lastPageNumber="120" pageId="8" pageNumber="119">
Females of the new species are most similar to 
<taxonomicName baseAuthorityName="Banks" baseAuthorityYear="1989" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="montana">
<emphasis italics="true" pageId="8" pageNumber="119">I. montana</emphasis>
</taxonomicName>
and 
<taxonomicName authority="Szczytko &amp; Kondratieff, 2015" authorityName="Szczytko &amp; Kondratieff" authorityYear="2015" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="kirchneri">
<emphasis box="[357,484,1648,1672]" italics="true" pageId="8" pageNumber="119">I. kirchneri</emphasis>
<bibRefCitation author="Szczytko, S. W. &amp; B. C. Kondratieff" pageId="8" pageNumber="119" pagination="1 - 289" refId="ref8269" refString="Szczytko, S. W. &amp; B. C. Kondratieff. 2015. A review of the eastern Nearctic Isoperlinae (Plecoptera: Perlodidae) with the description of twenty-two new species. Monographs of Illiesia, Number 1: 1 - 289. http: // illiesia. speciesfile. org / papers / Monographiae-of-Illiesia. pdf" type="book chapter" year="2015">Szczytko &amp; Kondratieff, 2015</bibRefCitation>
</taxonomicName>
. Females of the new species have a subgenital plate that extends approximately ⅓–½ over sternum 9 and appears broadly rounded (
<figureCitation box="[574,654,1756,1780]" captionStart="Figs" captionStartId="7.[189,238,1284,1308]" captionTargetBox="[324,1307,218,1201]" captionTargetId="figure-266@7.[324,1307,218,1201]" captionTargetPageId="7" captionText="Figs. 15–17. Female subgenital plate of Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia, 5X. Fig. 15. Broadly rounded Fig. 16. Broadly triangular, truncate with shallow posteromedian emargination. Fig. 17. Broadly triangular." figureDoi="http://doi.org/10.5281/zenodo.4757803" httpUri="https://zenodo.org/record/4757803/files/figure.png" pageId="8" pageNumber="119">Fig. 15</figureCitation>
), or broadly triangular, truncate with shallow posteromedian emargination (
<figureCitation box="[367,451,1828,1852]" captionStart="Figs" captionStartId="7.[189,238,1284,1308]" captionTargetBox="[324,1307,218,1201]" captionTargetId="figure-266@7.[324,1307,218,1201]" captionTargetPageId="7" captionText="Figs. 15–17. Female subgenital plate of Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia, 5X. Fig. 15. Broadly rounded Fig. 16. Broadly triangular, truncate with shallow posteromedian emargination. Fig. 17. Broadly triangular." figureDoi="http://doi.org/10.5281/zenodo.4757803" httpUri="https://zenodo.org/record/4757803/files/figure.png" pageId="8" pageNumber="119">Fig. 16</figureCitation>
), or broadly triangular (
<figureCitation captionStart="Figs" captionStartId="7.[189,238,1284,1308]" captionTargetBox="[324,1307,218,1201]" captionTargetId="figure-266@7.[324,1307,218,1201]" captionTargetPageId="7" captionText="Figs. 15–17. Female subgenital plate of Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia, 5X. Fig. 15. Broadly rounded Fig. 16. Broadly triangular, truncate with shallow posteromedian emargination. Fig. 17. Broadly triangular." figureDoi="http://doi.org/10.5281/zenodo.4757803" httpUri="https://zenodo.org/record/4757803/files/figure.png" pageId="8" pageNumber="119">Fig. 17</figureCitation>
), and have a dorsal head pattern with brown bands connecting the ocelli that are truncate at the anterior ocellus with a triangular dusky spot anterior to the anterior ocellus (
<figureCitation box="[1207,1274,1540,1564]" captionStart="Figs" captionStartId="3.[189,238,1284,1308]" captionTargetBox="[324,1307,218,1201]" captionTargetId="figure-282@3.[324,1307,218,1201]" captionTargetPageId="3" captionText="Figs. 2–4. Isoperla dewalti sp. n., Helton Creek, Grayson Co., Virginia, 5X. Fig. 2. Head and pronotum, dorsal. Fig. 3. Male sterna 7–9, ventral. Fig. 4. Male paraprocts, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4757791" httpUri="https://zenodo.org/record/4757791/files/figure.png" pageId="8" pageNumber="119">Fig. 2</figureCitation>
). Females of 
<taxonomicName baseAuthorityName="Banks" baseAuthorityYear="1989" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="montana">
<emphasis italics="true" pageId="8" pageNumber="119">I. montana</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Szczytko &amp; Kondratieff" authorityYear="2015" box="[989,1108,1576,1600]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="kirchneri">
<emphasis box="[989,1108,1576,1600]" italics="true" pageId="8" pageNumber="119">I. kirchneri</emphasis>
</taxonomicName>
also have a subgenital plate that is variable, appearing broadly triangular usually with a shallow posteromedian emargination or pointed (figs. 25.5, 33.7, 
<bibRefCitation author="Szczytko, S. W. &amp; B. C. Kondratieff" pageId="8" pageNumber="119" pagination="1 - 289" refId="ref8269" refString="Szczytko, S. W. &amp; B. C. Kondratieff. 2015. A review of the eastern Nearctic Isoperlinae (Plecoptera: Perlodidae) with the description of twenty-two new species. Monographs of Illiesia, Number 1: 1 - 289. http: // illiesia. speciesfile. org / papers / Monographiae-of-Illiesia. pdf" type="book chapter" year="2015">Szczytko &amp; Kondratieff 2015</bibRefCitation>
) and have dorsal head patterns with brown bands connecting the ocelli. In 
<taxonomicName baseAuthorityName="Banks" baseAuthorityYear="1989" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="montana">
<emphasis italics="true" pageId="8" pageNumber="119">I. montana</emphasis>
</taxonomicName>
the bands extend anteriorly to near the frons (occasionally pale brown or faint), and without a medium brown anterior patch near the frons (fig. 33.1, 
<bibRefCitation author="Szczytko, S. W. &amp; B. C. Kondratieff" box="[1016,1356,1900,1924]" pageId="8" pageNumber="119" pagination="1 - 289" refId="ref8269" refString="Szczytko, S. W. &amp; B. C. Kondratieff. 2015. A review of the eastern Nearctic Isoperlinae (Plecoptera: Perlodidae) with the description of twenty-two new species. Monographs of Illiesia, Number 1: 1 - 289. http: // illiesia. speciesfile. org / papers / Monographiae-of-Illiesia. pdf" type="book chapter" year="2015">Szczytko &amp; Kondratieff 2015</bibRefCitation>
). In 
<emphasis box="[1406,1422,1901,1924]" italics="true" pageId="8" pageNumber="119">I.</emphasis>
<taxonomicName authorityName="Szczytko &amp; Kondratieff" authorityYear="2015" box="[189,286,1145,1169]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="kirchneri">
<emphasis box="[189,286,1145,1169]" italics="true" pageId="9" pageNumber="120">kirchneri</emphasis>
</taxonomicName>
the bands sometimes extend anteriorly from the anterior ocellus, with a well-defined to diffuse spot on frons (fig. 25.1, 
<bibRefCitation author="Szczytko, S. W. &amp; B. C. Kondratieff" pageId="9" pageNumber="120" pagination="1 - 289" refId="ref8269" refString="Szczytko, S. W. &amp; B. C. Kondratieff. 2015. A review of the eastern Nearctic Isoperlinae (Plecoptera: Perlodidae) with the description of twenty-two new species. Monographs of Illiesia, Number 1: 1 - 289. http: // illiesia. speciesfile. org / papers / Monographiae-of-Illiesia. pdf" type="book chapter" year="2015">Szczytko &amp; Kondratieff 2015</bibRefCitation>
). The variation in subgenital plate morphology and dorsal head pattern exhibited by these species make the separation of isolated females very difficult.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4757809" ID-Zenodo-Dep="4757809" box="[189,951,1037,1062]" httpUri="https://zenodo.org/record/4757809/files/figure.png" pageId="9" pageNumber="120" startId="9.[189,226,1037,1061]" targetBox="[324,1307,254,991]" targetPageId="9">
<paragraph blockId="9.[189,951,1037,1062]" box="[189,951,1037,1062]" pageId="9" pageNumber="120">Fig. 21. Type locality. Helton Creek, Grayson Co., Virginia, USA.</paragraph>
</caption>
<paragraph blockId="9.[189,798,1145,1925]" pageId="9" pageNumber="120">
Ova of the new species are similar to 
<taxonomicName authorityName="Szczytko &amp; Kondratieff" authorityYear="2015" box="[671,790,1397,1421]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="kirchneri">
<emphasis box="[671,790,1397,1421]" italics="true" pageId="9" pageNumber="120">I. kirchneri</emphasis>
</taxonomicName>
, 
<emphasis box="[189,428,1433,1457]" italics="true" pageId="9" pageNumber="120">
<taxonomicName baseAuthorityName="Banks" baseAuthorityYear="1989" box="[189,309,1434,1457]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="montana">I. montana</taxonomicName>
, 
<taxonomicName authorityName="Szczytko &amp; Kondratieff" authorityYear="2015" box="[321,423,1433,1457]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="nelsoni">I. nelsoni</taxonomicName>
,
</emphasis>
and 
<taxonomicName authority="(Banks, 1902)" baseAuthorityName="Banks" baseAuthorityYear="1902" box="[488,757,1433,1457]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="signata">
<emphasis box="[488,589,1434,1457]" italics="true" pageId="9" pageNumber="120">I. signata</emphasis>
(Banks, 1902)
</taxonomicName>
, all of which are oblong and triangular in cross section with distinct cross shaped ridges dividing anterior and posterior poles. Ova of the new species are distinguished from other similar ova by the interreticulate follicle cell impressions, which are developed as mounds in the new species (
<figureCitation captionStart="Figs" captionStartId="8.[189,238,1360,1384]" captionTargetBox="[286,1344,218,1276]" captionTargetId="figure-278@8.[286,1344,218,1276]" captionTargetPageId="8" captionText="Figs. 18–20. Isoperla dewalti sp. n., Tributary to Le Conte Creek, Sevier Co., Tennessee. Fig. 18. Ovum, 400X. Fig. 19. Ovum collar, 1,800X. Fig. 20. Ovum follicle cell impressions and micropyles, 800X." figureDoi="http://doi.org/10.5281/zenodo.4757807" httpUri="https://zenodo.org/record/4757807/files/figure.png" pageId="9" pageNumber="120">Figs. 18, 20</figureCitation>
) and punctations in the other similar species (figs. 25.19–25.22, 33.25–33.28, 36.9–36.11, 50.20–50.23, 
<bibRefCitation author="Szczytko, S. W. &amp; B. C. Kondratieff" box="[189,527,1757,1781]" pageId="9" pageNumber="120" pagination="1 - 289" refId="ref8269" refString="Szczytko, S. W. &amp; B. C. Kondratieff. 2015. A review of the eastern Nearctic Isoperlinae (Plecoptera: Perlodidae) with the description of twenty-two new species. Monographs of Illiesia, Number 1: 1 - 289. http: // illiesia. speciesfile. org / papers / Monographiae-of-Illiesia. pdf" type="book chapter" year="2015">Szczytko &amp; Kondratieff 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="120" type="biology_ecology">
<paragraph blockId="9.[189,798,1145,1925]" lastBlockId="9.[835,1443,1145,1565]" pageId="9" pageNumber="120">
<emphasis bold="true" box="[189,391,1792,1817]" pageId="9" pageNumber="120">Biological notes.</emphasis>
There is no information about the biology or life cycle of 
<taxonomicName authorityName="Verdone &amp; Kondratieff" authorityYear="2017" box="[456,556,1829,1853]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="dewalti">
<emphasis box="[456,556,1829,1853]" italics="true" pageId="9" pageNumber="120">I. dewalti</emphasis>
</taxonomicName>
. Based on the above records, the emergence period appears to be in early to mid-May, and as with most eastern 
<taxonomicName box="[680,764,1901,1925]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis box="[680,764,1901,1925]" italics="true" pageId="9" pageNumber="120">Isoperla</emphasis>
</taxonomicName>
, a univoltine life cycle is presumed. The 
<typeStatus box="[1289,1341,1146,1170]" pageId="9" pageNumber="120">type</typeStatus>
locality, Helton Creek, has a unique stonefly fauna, species with generally northern distributions (e.g. 
<taxonomicName authority="Ricker, 1936" authorityName="Ricker" authorityYear="1936" class="Insecta" family="Chloroperlidae" genus="Alloperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="concolor">
<emphasis italics="true" pageId="9" pageNumber="120">Alloperla concolor</emphasis>
Ricker, 1936
</taxonomicName>
) and species with generally southern distributions (e.g. 
<taxonomicName authority="Kirchner &amp; Kondratieff, 2005" authorityName="Kirchner &amp; Kondratieff" authorityYear="2005" class="Insecta" family="Chloroperlidae" genus="Haploperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="120" phylum="Arthropoda" rank="species" species="parkeri">
<emphasis box="[1222,1443,1289,1313]" italics="true" pageId="9" pageNumber="120">Haploperla parkeri</emphasis>
Kirchner &amp; Kondratieff, 2005
</taxonomicName>
) have been documented from this location. From the limited records available it is inferred that the new species also has a generally southern distribution, occurring in 1 
<superScript attach="none" box="[878,890,1470,1484]" fontSize="6" pageId="9" pageNumber="120">st</superScript>
–3 
<superScript attach="left" box="[917,933,1470,1484]" fontSize="6" pageId="9" pageNumber="120">rd</superScript>
order streams in the Piedmont Plateau and the southern Blue Ridge physiographic provinces (
<figureCitation box="[843,925,1541,1565]" captionStart="Fig" captionStartId="10.[189,226,1826,1850]" captionTargetBox="[249,1383,284,1716]" captionTargetId="figure-61@10.[249,1383,283,1750]" captionTargetPageId="10" captionText="Fig. 22. Distribution map of examined material and physiographic provinces." figureDoi="http://doi.org/10.5281/zenodo.4757813" httpUri="https://zenodo.org/record/4757813/files/figure.png" pageId="9" pageNumber="120">Fig. 22</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>