182 lines
20 KiB
XML
182 lines
20 KiB
XML
<document id="372AA6BDA73FA84563FB11EFEC57C6C4" ID-DOI="10.1206/0003-0090(2005)293<0001:CNHAEO>2.0.CO;2" ID-ISSN="0003-0090" ID-Zenodo-Dep="5362748" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630237563515" checkinUser="felipe" docAuthor="OPITZ, WESTON" docDate="2005" docId="03E787FE99591139FD72FACEFB25FECD" docLanguage="en" docName="B293.pdf" docOrigin="Bulletin of the American Museum of Natural History 2005 (293)" docSource="http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0090(2005)293%3C0001%3ACNHAEO%3E2.0.CO%3B2" docStyle="DocumentStyle:2B34016C1EBFE58B07368F9517618743.3:BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleId="2B34016C1EBFE58B07368F9517618743" docStyleName="BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleVersion="3" docTitle="Aphelocerus cheliferous OPITZ 2005, new species" docType="treatment" docVersion="3" lastPageNumber="90" masterDocId="FFDEFF8699011160FFAFFF8EFFB3FFDE" masterDocTitle="Classification, Natural History, And Evolution Of The Genus Aphelocerus Kirsch (Coleoptera: Cleridae: Clerinae)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="89" updateTime="1699194784552" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="97B35807A1695A353B390EA05A497664">Classification, Natural History, And Evolution Of The Genus Aphelocerus Kirsch (Coleoptera: Cleridae: Clerinae)</mods:title>
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<mods:namePart id="537C764C04096F0F81B4BEB5A8518AA3">OPITZ, WESTON</mods:namePart>
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<mods:title id="8AF36C337F7B1AAEE1814F3CFAE3CB6C">Bulletin of the American Museum of Natural History</mods:title>
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<mods:part id="68B7EE3B1D54DBAC64C1A2895FE0DD86">
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<mods:date id="0879927F71518E5AA691B63ACFEF2873">2005</mods:date>
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<mods:number id="743DEBD5E31EFDE41EE07742ECEF2490">2005-05-11</mods:number>
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<mods:number id="A8899D78C74AA8CDCEFCF873CD09A771">2005</mods:number>
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<mods:number id="87A18A97FAE36E71BF0F03239991BB68">293</mods:number>
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<mods:classification id="EC110555EC3153C9EA26C92BDCBCE347">journal article</mods:classification>
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<mods:identifier id="DE5B8D1F2CF5A771B1BE231DD8935DE4" type="DOI">10.1206/0003-0090(2005)293<0001:CNHAEO>2.0.CO;2</mods:identifier>
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<mods:identifier id="3166E93CEFA09AD40C538EC881E547A2" type="ISSN">0003-0090</mods:identifier>
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<mods:identifier id="68F3E352F04646CC1A73A806A4389D13" type="Zenodo-Dep">5362748</mods:identifier>
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<treatment id="03E787FE99591139FD72FACEFB25FECD" LSID="urn:lsid:plazi:treatment:03E787FE99591139FD72FACEFB25FECD" httpUri="http://treatment.plazi.org/id/03E787FE99591139FD72FACEFB25FECD" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89">
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<subSubSection id="C354656399591138FD72FACEFB00FAAB" pageId="88" pageNumber="89" type="nomenclature">
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<paragraph id="8BF136E899591138FD72FACEFB00FAAB" blockId="88.[726,1203,1344,1397]" pageId="88" pageNumber="89">
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<heading id="D0B9818499591138FD72FACEFB00FAAB" centered="true" fontSize="9" level="2" pageId="88" pageNumber="89" reason="2">
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<emphasis id="B93AEAFA99591138FD72FACEFBB9FA86" bold="true" box="[733,1034,1344,1368]" pageId="88" pageNumber="89">
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<taxonomicName id="4C4E4D6B99591138FD72FACEFBB0FA86" authority="OPITZ, 2005" authorityName="OPITZ" authorityYear="2005" box="[733,1027,1344,1368]" class="Insecta" family="Cleridae" genus="Aphelocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="cheliferous" status="sp. nov.">
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<emphasis id="B93AEAFA99591138FD72FACEFBB0FA86" bold="true" box="[733,1027,1344,1368]" italics="true" pageId="88" pageNumber="89">Aphelocerus cheliferous</emphasis>
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</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A209578199591138FBBCFACEFB11FA89" box="[1043,1186,1344,1367]" pageId="88" pageNumber="89" rank="species">new species</taxonomicNameLabel>
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<figureCitation id="13752A6D99591138FD79FAD0FCE5FAAB" box="[726,854,1374,1397]" captionStart="Figs" captionStartId="45.[149,197,1162,1183]" captionTargetBox="[123,1227,188,1140]" captionTargetId="figure-16@45.[123,1227,187,1141]" captionTargetPageId="45" captionText="Figs. 85–120. Antennae, pygidia, and sixth visible abdominal sterna. 85–90. Antennae. 85. Aphelocerus affaniatis. 86. A. dispilis. 87. A. formicoides. 88. A. cheliferous. 89. A. panus. 90. A. calvus. 91– 120. Pygidium and/or sixth visible abdominal sternum. 91–94. A. ciliaris. 91–92. male (91, pygidium; 92, sixth visible abdominal sternum). 93–94. female (93, pygidium; 94, sixth visible dominal sternum). 95. A. coactus, female pygdium. 96. A. domus, female pygidium. 97–100. A. coalitus, male (97, pygidium; 98, sixth visible abdominal sternum), 99–100. Female (99, pygidium; 100, sixth visible abdominal sternum). 101. A. leucomelas, male pygidium. 102–104. A. coactus. 102–103. male (102, pygidium; 103, sixth visible abdominal sternum). 104, female, pygidium. 105–106. A. acutus, pygidea (105, male; 106, female). 107–109. A. lividus. 107, male (pygidium). 108–109, female (108, pygidium; 109, sixth visible abdominal sternum). 110–113. A. sagittarius. 110–111, male (110, pygidium; 111, sixth visible abdominal sternum). 113, female, pygidium. 114–115. A. humerus, female (114, pygidium; 115, sixth visible abdominal sternum). 116–117. A. naevius, female (116, pygidium; 117, sixth visible abdominal sternum). 118–119. A. triangulus, female (118, pygidium; 119, sixth visible abdominal sternum). 120. A. arenatus, male, pygidium." figureDoi="http://doi.org/10.5281/zenodo.5362776" httpUri="https://zenodo.org/record/5362776/files/figure.png" pageId="88" pageNumber="89">Figures 88</figureCitation>
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,
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<figureCitation id="13752A6D99591138FCCAFAD0FC63FAAB" box="[869,976,1374,1397]" captionStart="Figs" captionStartId="51.[149,197,1352,1373]" captionTargetBox="[131,1217,188,1330]" captionTargetId="figure-16@51.[131,1218,187,1331]" captionTargetPageId="51" captionText="Figs. 139–153. Tegmina or phallus. 139. Aphelocerus immarginatus, tegmen. 140. A. vietus, tegmen. 141. A. patulus, tegmen. 142. A. yungas, tegmen. 143. A. naevius, tegmen. 144a. A. catie, tegmen. 144b. A. affaniatis, tegmen. 144c. A. echinatus, tegmen. 145. A. scutellaris. 146–147. A. cheliferous (146, tegmen; 147, phallus). 148. A. panus, tegmen. 149. A. formicoides, tegmen. 150. A. acanthus, tegmen. 151. A. bufustis, tegmen. 152. A. batesi, tegmen. 153a. A. calvus, tegmen. 153b. A. capillus, tegmen." figureDoi="http://doi.org/10.5281/zenodo.5362780" httpUri="https://zenodo.org/record/5362780/files/figure.png" pageId="88" pageNumber="89">146, 147</figureCitation>
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,
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<figureCitation id="13752A6D99591138FC70FAD0FBBEFAAB" box="[991,1037,1374,1397]" captionStart="Figs" captionStartId="62.[150,198,1596,1617]" captionTargetBox="[198,1154,188,1573]" captionTargetId="figure-9@62.[197,1155,187,1575]" captionTargetPageId="62" captionText="Figs. 184–192. Internal reproductive organs. 184–186. Aphelocerus inconstans. 184–185, male (184, accessory glands, testis, and ejaculatory duct; 185, base of medial accessory gland; 186, base of lateral accessory gland). 187–189. A. myrmecoides. 187–188, male (187, accessory glands, testis, and ejaculatory duct; 188, medial accessory gland). 189. Female. 190. A. leucomelas, male. 191. A. discapillus, male. 192. A. cheliferous male." figureDoi="http://doi.org/10.5281/zenodo.5362786" httpUri="https://zenodo.org/record/5362786/files/figure.png" pageId="88" pageNumber="89">192</figureCitation>
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, 228; map 19
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C354656399591139FD76FA1FFDBBFF2B" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89" type="materials_examined">
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<paragraph id="8BF136E899591138FD76FA1FFB1AF94D" blockId="88.[700,1229,1425,1741]" pageId="88" pageNumber="89">
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<materialsCitation id="3B263CB599591138FD76FA1FFB1AF94D" collectingDate="1983-05-20" collectionCode="LACM" collectorName="C. & L. O'Brien & Marshall" country="Mexico" location="Los Tuxtlas" pageId="88" pageNumber="89" specimenCount="2" typeStatus="holotype">
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<typeStatus id="54F5884A99591138FD76FA1FFCFCFA79" box="[729,847,1425,1448]" pageId="88" pageNumber="89" type="holotype">HOLOTYPE</typeStatus>
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: Male.
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<collectingCountry id="F359767899591138FC1CFA1FFBA3FA76" box="[947,1040,1425,1448]" name="Mexico" pageId="88" pageNumber="89">Mexico</collectingCountry>
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. Ver. 2509,
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<location id="8E91603399591138FB0EFA1FFCA7FA1B" LSID="urn:lsid:plazi:treatment:03E787FE99591139FD72FACEFB25FECD:8E91603399591138FB0EFA1FFCA7FA1B" country="Mexico" name="Los Tuxtlas" pageId="88" pageNumber="89">Los Tuxtlas</location>
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, Biol. Sta. UNAM,
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<date id="FFF0102899591138FBABFA20FB37FA1B" box="[1028,1156,1454,1477]" pageId="88" pageNumber="89" value="1983-05-20">
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<collectingDate id="EFB4E9C099591138FBABFA20FB37FA1B" box="[1028,1156,1454,1477]" pageId="88" pageNumber="89" value="1983-05-20">20V1983</collectingDate>
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</date>
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,
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<collectorName id="26BB533E99591138FB3DFA20FC8DFA3D" pageId="88" pageNumber="89">C. & L. O’Brien</collectorName>
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&
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<collectorName id="26BB533E99591138FCC7FA42FC7CFA3D" box="[872,975,1484,1507]" pageId="88" pageNumber="89">Marshall</collectorName>
|
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(
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<collectionCode id="ED5FAE2D99591138FC4EFA42FB8FFA3D" box="[993,1084,1484,1507]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="88" pageNumber="89" type="Museum">LACM</collectionCode>
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||
). (Specimen point mounted, sex label affixed to paper point, white, hand printed; support card, white; locality label, white, machine printed;
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<collectionCode id="ED5FAE2D99591138FD13F9CFFCA0F986" box="[700,787,1601,1624]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="88" pageNumber="89" type="Museum">LACM</collectionCode>
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repository label, white, machine printed;
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<typeStatus id="54F5884A99591138FCB0F9D0FC36F9AB" box="[799,901,1630,1653]" pageId="88" pageNumber="89" type="holotype">holotype</typeStatus>
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label, red, machine printed; plastic vial with abdomen and aedeagus).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BF136E899591139FD76F917FDBBFF2B" blockId="88.[700,1229,1425,1741]" lastBlockId="89.[123,652,193,1742]" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89">
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||
<materialsCitation id="3B263CB599591139FD76F917FF44FF06" collectingDate="1983-05-20" collectionCode="JNRC, WOPC" collectorName="C. & L. O'Brien & Marshall" country="Mexico" lastPageId="89" lastPageNumber="90" location="Los Tuxtlas" pageId="88" pageNumber="89" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54F5884A99591138FD76F917FCE4F971" box="[729,855,1689,1712]" pageId="88" pageNumber="89" type="paratype">PARATYPES</typeStatus>
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: Fifteen specimens from the same locality as the holotype: (
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<collectionCode id="ED5FAE2D99591138FBFAF938FB12F913" box="[1109,1185,1718,1741]" pageId="88" pageNumber="89">JNRC</collectionCode>
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, 4;
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<collectionCode id="ED5FAE2D99581139FFD4FF4FFF61FF06" box="[123,210,193,216]" pageId="89" pageNumber="90">WOPC</collectionCode>
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, 1)
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</materialsCitation>
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;
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<materialsCitation id="3B263CB599581139FEA9FF4FFD35FF06" box="[262,646,193,216]" collectingDate="1983-05-21" collectionCode="JNRC, WOPC" country="Mexico" pageId="89" pageNumber="90" specimenCount="1" typeStatus="paratype">
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||
<date id="FFF0102899581139FEA9FF4FFE31FF06" box="[262,386,193,216]" pageId="89" pageNumber="90" value="1983-05-21">
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||
<collectingDate id="EFB4E9C099581139FEA9FF4FFE31FF06" box="[262,386,193,216]" pageId="89" pageNumber="90" value="1983-05-21">21V1983</collectingDate>
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||
</date>
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||
(
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||
<collectionCode id="ED5FAE2D99581139FE3CFF4FFE6CFF06" box="[403,479,193,216]" pageId="89" pageNumber="90">JNRC</collectionCode>
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, 5;
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<collectionCode id="ED5FAE2D99581139FDA7FF4FFDECFF06" box="[520,607,193,216]" pageId="89" pageNumber="90">WOPC</collectionCode>
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, 2)
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</materialsCitation>
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||
;
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<materialsCitation id="3B263CB599581139FFD4FF50FDB6FF2B" box="[123,517,222,245]" collectingDate="1983-05-19" collectionCode="JNRC, WOPC" country="Mexico" pageId="89" pageNumber="90" specimenCount="1" typeStatus="paratype">
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||
<date id="FFF0102899581139FFD4FF50FF4BFF2B" box="[123,248,222,245]" pageId="89" pageNumber="90" value="1983-05-19">
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||
<collectingDate id="EFB4E9C099581139FFD4FF50FF4BFF2B" box="[123,248,222,245]" pageId="89" pageNumber="90" value="1983-05-19">19V1983</collectingDate>
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||
</date>
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||
(
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||
<collectionCode id="ED5FAE2D99581139FEA5FF50FEE5FF2B" box="[266,342,222,245]" pageId="89" pageNumber="90">JNRC</collectionCode>
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||
, 1;
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||
<collectionCode id="ED5FAE2D99581139FE2BFF50FE68FF2B" box="[388,475,222,245]" pageId="89" pageNumber="90">WOPC</collectionCode>
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||
, 2)
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</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C354656399581139FF37FF72FE9AFE23" pageId="89" pageNumber="90" type="diagnosis">
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||
<paragraph id="8BF136E899581139FF37FF72FE9AFE23" blockId="89.[123,652,193,1742]" pageId="89" pageNumber="90">
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DIAGNOSIS: The antennal apex of these beetles is testaceous, which is also a characteristic of some other species across speciesgroup lines. Among the members of this species complex,
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<taxonomicName id="4C4E4D6B99581139FEE2FEFFFE7DFE56" authorityName="OPITZ" authorityYear="2005" box="[333,462,369,392]" class="Insecta" family="Cleridae" genus="Aphelocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="cheliferous">
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||
<emphasis id="B93AEAFA99581139FEE2FEFFFE7DFE56" box="[333,462,369,392]" italics="true" pageId="89" pageNumber="90">cheliferous</emphasis>
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</taxonomicName>
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specimens may be identified by having the elytral surface polished, the outer margin of the elytral apex obtuse, and the side margins of the pronotum feebly convex.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C354656399581139FF37FD8AFE3DF986" pageId="89" pageNumber="90" type="description">
|
||
<paragraph id="8BF136E899581139FF37FD8AFE41FA1B" blockId="89.[123,652,193,1742]" pageId="89" pageNumber="90">
|
||
DESCRIPTION:
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<emphasis id="B93AEAFA99581139FEEEFD8AFEC3FDC5" box="[321,368,516,539]" italics="true" pageId="89" pageNumber="90">Size</emphasis>
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||
: Length 5.0–5.0 mm; width 2.0–
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<quantity id="4CB69B0D99581139FEABFDAFFEDFFDE6" box="[260,364,545,568]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="89" pageNumber="90" unit="mm" value="2.5">2.5 mm</quantity>
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||
.
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<emphasis id="B93AEAFA99581139FE2EFDAFFDB7FDE6" box="[385,516,545,568]" italics="true" pageId="89" pageNumber="90">Integument</emphasis>
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||
: Cranium, pronotum, venter, and legs cyanescent; elytra piceous.
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<emphasis id="B93AEAFA99581139FF4AFDD2FEFDFDAD" box="[229,334,604,627]" italics="true" pageId="89" pageNumber="90">Vestiture</emphasis>
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||
: Integument densely vest ed with prominent light and dark setae, light setae particularly prominent along lower edge of pronotum and along the epipleuron.
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<emphasis id="B93AEAFA99581139FFD4FD5FFF09FD36" box="[123,186,721,744]" italics="true" pageId="89" pageNumber="90">Head</emphasis>
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||
: Genae and epicranium expanded; interocular depression and frontal umbo well developed; frons, epicranium, and gena finely punctate; eyes subspherical, boldly convex; width across eyes feebly narrower than width of pronotum (31:33); antenna (figs. 88, 228) elongate, nearly as long as length of pronotum (35:37).
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<emphasis id="B93AEAFA99581139FECAFC11FE0AFC68" box="[357,441,927,950]" italics="true" pageId="89" pageNumber="90">Thorax</emphasis>
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||
: Pronotum elongate (33:37), considerably narrower than width of elytra across humeri (33:40); lateral margins feebly arcuate; anterior transverse depression faintly indicated; elytra strongly convex, disc with nine longitudinal rows of fine punctations, and shallow longitudinal ridges, depth at humerus 18, greatest depth in posterior half 28; legs prominent, metafemur extended to elytral apex.
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<emphasis id="B93AEAFA99581139FDBAFB29FD30FB60" box="[533,643,1191,1214]" italics="true" pageId="89" pageNumber="90">Abdomen</emphasis>
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||
: Posterior margin of pygdium evenly arcuate in both sexes.
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<emphasis id="B93AEAFA99581139FE8BFB6FFF01FAC8" italics="true" pageId="89" pageNumber="90">Male internal reproductive organs</emphasis>
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(fig. 192): Lateral accessory gland narrow and convoluted, coiled; medial gland longer and vastly more robust than lateral gland.
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<emphasis id="B93AEAFA99581139FF60FAD9FECCFAB0" box="[207,383,1367,1390]" italics="true" pageId="89" pageNumber="90">Male genitalia</emphasis>
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(fig. 146): Parameres well developed; ventral and dorsal sinuses equal in length; phallic plate (fig. 147) set with row of marginal denticles.
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</paragraph>
|
||
<paragraph id="8BF136E899581139FF37FA42FDCCF9C3" blockId="89.[123,652,193,1742]" pageId="89" pageNumber="90">VARIATION: The shallow longitudinal elytral ridges vary in expression as does the extent of testaceousness of the antennal apex.</paragraph>
|
||
<paragraph id="8BF136E899581139FF37F9AAFE3DF986" blockId="89.[123,652,193,1742]" pageId="89" pageNumber="90">
|
||
NATURAL HISTORY: Specimens were collected in May at
|
||
<specimenCount id="9D48FD6199581139FEE6F9CFFE3DF986" box="[329,398,1601,1624]" pageId="89" pageNumber="90" type="generic">
|
||
<quantity id="4CB69B0D99581139FEE6F9CFFE39F986" box="[329,394,1601,1624]" metricMagnitude="1" metricUnit="m" metricValue="7.6" pageId="89" pageNumber="90" unit="m" value="76.0">76 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C354656399581139FF37F9D0FEE4F96E" pageId="89" pageNumber="90" type="distribution">
|
||
<paragraph id="8BF136E899581139FF37F9D0FEE4F96E" blockId="89.[123,652,193,1742]" pageId="89" pageNumber="90">
|
||
DISTRIBUTION (map 19): This species is known from the slopes of Volvan San Andres Tuxtla, in
|
||
<collectingCountry id="F359767899581139FF59F917FEE0F96E" box="[246,339,1689,1712]" name="Mexico" pageId="89" pageNumber="90">Mexico</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C354656399581139FF37F938FB25FECD" pageId="89" pageNumber="90" type="etymology">
|
||
<paragraph id="8BF136E899581139FF37F938FB25FECD" blockId="89.[123,652,193,1742]" lastBlockId="89.[699,1227,193,275]" pageId="89" pageNumber="90">
|
||
ETYMOLOGY: The trivial name
|
||
<taxonomicName id="4C4E4D6B99581139FDA5F939FD38F910" authorityName="OPITZ" authorityYear="2005" box="[522,651,1719,1742]" class="Insecta" family="Cleridae" genus="Aphelocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="cheliferous">
|
||
<emphasis id="B93AEAFA99581139FDA5F939FD38F910" box="[522,651,1719,1742]" italics="true" pageId="89" pageNumber="90">cheliferous</emphasis>
|
||
</taxonomicName>
|
||
comes from the Latin noun
|
||
<emphasis id="B93AEAFA99581139FBA9FF4FFBF6FF06" box="[1030,1093,193,216]" italics="true" pageId="89" pageNumber="90">chela</emphasis>
|
||
(claw) and
|
||
<emphasis id="B93AEAFA99581139FD6BFF51FCBEFF28" box="[708,781,223,246]" italics="true" pageId="89" pageNumber="90">ferous</emphasis>
|
||
(carry). I refer to the presence of marginal denticles on the phallic plates.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |