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<mods:title id="8DB32CF0164635BCBC34FB2E45D44257">A revision of the Pediacus Shuckard 1839 (Coleoptera: Cucujidae) of Asia and Australasia</mods:title>
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<taxonomicName id="4C4E4D06A935FFAC0AEE15A1CCD3B5BC" ID-CoL="6JYC" authority="Shuckard" authorityName="Shuckard" box="[151,389,221,247]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis id="B93AEA97A935FFAC0AEE15A1CC52B5BC" bold="true" box="[151,260,221,247]" italics="true" pageId="2" pageNumber="34">Pediacus</emphasis>
Shuckard
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<taxonomicName id="4C4E4D06A935FFAC0AEE1458CC98B471" ID-CoL="6JYC" authority="Shuckard, 1839: 150" authorityName="Shuckard" authorityPageNumber="150" authorityYear="1839" box="[151,462,292,314]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B93AEA97A935FFAC0AEE1458CDA5B471" box="[151,243,292,314]" italics="true" pageId="2" pageNumber="34">Pediacus</emphasis>
<treatmentCitation id="0AEF1094A935FFAC0A831458CC98B471" author="Shuckard" box="[250,462,292,314]" page="150" pageId="2" pageNumber="34" year="1839">
<bibRefCitation id="EFDF4B74A935FFAC0A831458CC98B471" author="Shuckard, W. E." bookContentInfo="240 pp." box="[250,462,292,314]" firstAuthor="Shuckard" journalOrPublisher="Bailliere, London" page="150" pageId="2" pageNumber="34" refId="ref16203" refString="Shuckard, W. E. (1839) Elements of British Entomology. Part 1. Bailliere, London, 240 pp." title="Elements of British Entomology. Part 1" type="book" year="1839">Shuckard, 1839: 150</bibRefCitation>
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, 185.
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<typeStatus id="54F58827A935FFAC0AEE143FCD9FB413" box="[151,201,323,344]" pageId="2" pageNumber="34">Type</typeStatus>
species.
<taxonomicName id="4C4E4D06A935FFAC0B5C143FCF87B413" ID-CoL="328YW" authority="Fabricius, 1792: 96" authorityName="Fabricius" authorityPageNumber="96" authorityYear="1792" box="[293,721,322,344]" class="Insecta" family="Cucujidae" genus="Cucujus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="dermestoides">
<emphasis id="B93AEA97A935FFAC0B5C143FCF52B413" box="[293,516,322,344]" italics="true" pageId="2" pageNumber="34">Cucujus dermestoides</emphasis>
<bibRefCitation id="EFDF4B74A935FFAC0873143ECF87B413" author="Fabricius, J. C." bookContentInfo="Vol. 1, i - xx, 330 pp., Vol. 2, 538 pp" box="[522,721,322,344]" firstAuthor="Fabricius" journalOrPublisher="C. G. Proft, Copenhagen" page="96" pageId="2" pageNumber="34" refId="ref15454" refString="Fabricius, J. C. (1792) Entomologia Systematica Emendata et Aucta 1. C. G. Proft, Copenhagen, Vol. 1, i - xx, 330 pp., Vol. 2, 538 pp." title="Entomologia Systematica Emendata et Aucta 1" type="book" year="1792">Fabricius, 1792: 96</bibRefCitation>
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, by monotypy.
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<emphasis id="B93AEA97A935FFAC0AEE14F9CC47B4D5" bold="true" box="[151,273,389,414]" pageId="2" pageNumber="34">Diagnosis.</emphasis>
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<emphasis id="B93AEA97A935FFAC0B6314F9CCD7B4D6" box="[282,385,389,413]" italics="true" pageId="2" pageNumber="34">Pediacus</emphasis>
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species may be distinguished from the other two genera known from Asia and Australasia,
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<emphasis id="B93AEA97A935FFAC0AEE14D5CDA5B48B" box="[151,243,425,448]" italics="true" pageId="2" pageNumber="34">Cucujus</emphasis>
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and
<taxonomicName id="4C4E4D06A935FFAC0B5314D5CCD2B48A" box="[298,388,425,449]" class="Insecta" family="Cucujidae" genus="Platisus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B93AEA97A935FFAC0B5314D5CCD2B48A" box="[298,388,425,449]" italics="true" pageId="2" pageNumber="34">Platisus</emphasis>
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, by the following characters: size small (length
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), temples absent or reduced to a small tooth-like projection, antennae with a well-developed three-segmented club.
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<emphasis id="B93AEA97A935FFAC0ABE148DCC0FB741" bold="true" box="[199,345,497,522]" pageId="2" pageNumber="34">Description.</emphasis>
As for
<bibRefCitation id="EFDF4B74A935FFAC0BCA148DCF37B742" author="Thomas, M. C." box="[435,609,497,522]" firstAuthor="Thomas" journalOrPublisher="Insecta Mundi" pageId="2" pageNumber="34" pagination="157 - 177" part="17" refId="ref16298" refString="Thomas, M. C. (2004 [&quot; 2003 &quot;]) A revision of Pediacus Shuckard (Coleoptera: Cucujidae) for America north of Mexico, with notes on other species. Insecta Mundi, 17 (3 - 4), 157 - 177." title="A revision of Pediacus Shuckard (Coleoptera: Cucujidae) for America north of Mexico, with notes on other species" type="journal article" year="2004">Thomas (2004)</bibRefCitation>
but with the following changes: pronotum with anterior angle obtuse to acute; elytra broadly rounded to moderately costate or carinate sublaterally.
</paragraph>
</subSubSection>
<subSubSection id="C354650EA935FFAC0ABE1745C9A1B605" pageId="2" pageNumber="34" type="biology_ecology">
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<emphasis id="B93AEA97A935FFAC0ABE1745CC71B719" bold="true" box="[199,295,569,594]" pageId="2" pageNumber="34">Biology.</emphasis>
There is little information available about the biology of
<taxonomicName id="4C4E4D06A935FFAC09C41745C972B71A" box="[957,1060,569,593]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B93AEA97A935FFAC09C41745C972B71A" box="[957,1060,569,593]" italics="true" pageId="2" pageNumber="34">Pediacus</emphasis>
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. As for other cucujids,
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<emphasis id="B93AEA97A935FFAC0F4C1745C8CDB71A" box="[1333,1435,569,593]" italics="true" pageId="2" pageNumber="34">Pediacus</emphasis>
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are apparently found under bark. In his revision of the North American fauna,
<bibRefCitation id="EFDF4B74A935FFAC0E6A1721C997B73D" author="Thomas, M. C." box="[1043,1217,605,630]" firstAuthor="Thomas" journalOrPublisher="Insecta Mundi" pageId="2" pageNumber="34" pagination="157 - 177" part="17" refId="ref16298" refString="Thomas, M. C. (2004 [&quot; 2003 &quot;]) A revision of Pediacus Shuckard (Coleoptera: Cucujidae) for America north of Mexico, with notes on other species. Insecta Mundi, 17 (3 - 4), 157 - 177." title="A revision of Pediacus Shuckard (Coleoptera: Cucujidae) for America north of Mexico, with notes on other species" type="journal article" year="2004">Thomas (2004)</bibRefCitation>
noted that they are predominantly found under bark of dead conifers. There is minimal collection data available among the specimens examined in this revision. This includes: dead oak bark; large cut tree-
<taxonomicName id="4C4E4D06A935FFAC098917DAC918B7F6" box="[1008,1102,678,701]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="34" phylum="Tracheophyta" rank="genus">
<emphasis id="B93AEA97A935FFAC098917DAC918B7F6" box="[1008,1102,678,701]" italics="true" pageId="2" pageNumber="34">Quercus</emphasis>
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sp.; on spruce; small cut sapling; light trap; insect flight trap and sifted moss and bark. Several specimens were collected from high altitudes, ranging up to
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, and for one species,
<taxonomicName id="4C4E4D06A935FFAC08911792CED3B64E" box="[744,901,750,773]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="montivagus">
<emphasis id="B93AEA97A935FFAC08911792CED3B64E" box="[744,901,750,773]" italics="true" pageId="2" pageNumber="34">P. montivagus</emphasis>
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, ca.
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.
</paragraph>
<paragraph id="8BF13685A935FFAC0ABE166DC9A1B605" blockId="2.[151,1436,389,846]" pageId="2" pageNumber="34">
Little is known about
<taxonomicName id="4C4E4D06A935FFAC0BC5166DCF75B662" box="[444,547,785,809]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B93AEA97A935FFAC0BC5166DCF75B662" box="[444,547,785,809]" italics="true" pageId="2" pageNumber="34">Pediacus</emphasis>
</taxonomicName>
larvae. They are likely to be predacious, as is apparently the case for the larvae of
<taxonomicName id="4C4E4D06A935FFAC0ACC164ACC47B606" box="[181,273,822,845]" class="Insecta" family="Cucujidae" genus="Cucujus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B93AEA97A935FFAC0ACC164ACC47B606" box="[181,273,822,845]" italics="true" pageId="2" pageNumber="34">Cucujus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4E4D06A935FFAC0B3E1649CE65B605" authority="Marris &amp; Klimaszewski" authorityName="Marris &amp; Klimaszewski" box="[327,819,821,846]" class="Insecta" family="Cucujidae" genus="Platisus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="zelandicus">
<emphasis id="B93AEA97A935FFAC0B3E1649CF49B606" box="[327,543,821,845]" italics="true" pageId="2" pageNumber="34">Platisus zelandicus</emphasis>
Marris &amp; Klimaszewski
</taxonomicName>
(
<bibRefCitation id="EFDF4B74A935FFAC09381649C97EB605" author="Smith, D. B. &amp; Sears, M. K." box="[833,1064,821,846]" firstAuthor="Smith" journalOrPublisher="Canadian Entomologist" pageId="2" pageNumber="34" pagination="173 - 175" part="114" refId="ref16227" refString="Smith, D. B. &amp; Sears, M. K. (1982) Mandibular structure and feeding habits of three morphologically similar coleopterous larvae: Cucujus clavipes (Cucujidae), Dendroides canadensis (Pyrochroidae), and Pytho depressus (Salpingidae). Canadian Entomologist, 114, 173 - 175. http: // dx. doi. org / 10.4039 / ent 114173 - 2" title="Mandibular structure and feeding habits of three morphologically similar coleopterous larvae: Cucujus clavipes (Cucujidae), Dendroides canadensis (Pyrochroidae), and Pytho depressus (Salpingidae)" type="journal article" year="1982">Smith &amp; Sears 1982</bibRefCitation>
;
<bibRefCitation id="EFDF4B74A935FFAC0E4D164AC9BCB605" author="Watt, J. C. &amp; Marris, J. W. M. &amp; Klimaszewski, J." box="[1076,1258,821,846]" etAl="et al." firstAuthor="Watt" journalOrPublisher="Journal of the Royal Society of New Zealand" pageId="2" pageNumber="34" pagination="327 - 339" part="31" refId="ref16490" refString="Watt, J. C., Marris, J. W. M. &amp; Klimaszewski, J. (2001) A new species of Platisus (Coleoptera: Cucujidae) from New Zealand, described from the adult and larva. Journal of the Royal Society of New Zealand, 31, 327 - 339. http: // dx. doi. org / 10.1080 / 03014223.2001.9517657" title="A new species of Platisus (Coleoptera: Cucujidae) from New Zealand, described from the adult and larva" type="journal article" year="2001">
Watt
<emphasis id="B93AEA97A935FFAC0E16164AC9FEB606" box="[1135,1192,821,845]" italics="true" pageId="2" pageNumber="34">et al.</emphasis>
2001
</bibRefCitation>
).
</paragraph>
</subSubSection>
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<heading id="D0B981E9A935FFAC0AEE16DDCF8AB6F0" bold="true" box="[151,732,929,955]" fontSize="11" level="1" pageId="2" pageNumber="34" reason="1">
<emphasis id="B93AEA97A935FFAC0AEE16DDCF8AB6F0" bold="true" box="[151,732,929,955]" pageId="2" pageNumber="34">
Key to
<taxonomicName id="4C4E4D06A935FFAC0A8916DDCC0BB6F0" box="[240,349,929,955]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B93AEA97A935FFAC0A8916DDCC0BB6F0" bold="true" box="[240,349,929,955]" italics="true" pageId="2" pageNumber="34">Pediacus</emphasis>
</taxonomicName>
adults of Asia and Australasia
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C354650EA935FFAD0AEE169BC8CDB469" lastPageId="3" lastPageNumber="35" pageId="2" pageNumber="34" type="key">
<keyStep id="30BA2E20A935FFAC0AEE169BC8CAB151" pageId="2" pageNumber="34">
<paragraph id="8BF13685A935FFAC0AEE169BC8CAB6B6" blockId="2.[151,1437,999,2018]" box="[151,1436,999,1021]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE169BC8CAB6B6" box="[151,1436,999,1021]" pageId="2" pageNumber="34">
1. Pronotum strongly explanate with broad glabrous lateral margins (
<figureCitation id="13752A00A935FFAC0924169BCECAB6B6" box="[861,924,999,1021]" captionStart="FIGURE 1" captionStartId="4.[151,255,1802,1826]" captionTargetBox="[151,1391,194,1765]" captionTargetId="figure@4.[151,1436,194,1781]" captionTargetPageId="4" captionText="FIGURE 1. Pediacus ater Grouvelle (A, C, E, H) specimen from Borneo; (B, D, F, G) specimen from Sumatra. (A, B) Head and pronotum; (C, D) dorsal habitus; (E, F) antenna; (G, H) aedeagus." httpUri="https://zenodo.org/record/252770/files/figure.png" pageId="2" pageNumber="34">Figs. 1</figureCitation>
A, B), pronotal disc broadly planar
<emphasis id="B93AEA97A935FFAC0E81169BC864B6B7" box="[1272,1330,999,1020]" italics="true" pageId="2" pageNumber="34">P. at e r</emphasis>
Grouvelle
</keyLead>
</paragraph>
<paragraph id="8BF13685A935FFAC0AEE1179C8CAB151" blockId="2.[151,1437,999,2018]" box="[151,1436,1028,1050]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE1179C8CAB151" box="[151,1436,1028,1050]" pageId="2" pageNumber="34">- Pronotum not strongly explanate and lacking broad glabrous lateral margins, pronotal disc convex to impressed.......... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BA2E20A935FFAC0AEE115DC8CDB11E" pageId="2" pageNumber="34">
<paragraph id="8BF13685A935FFAC0AEE115DC8CDB17C" blockId="2.[151,1437,999,2018]" box="[151,1435,1057,1079]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE115DC8CDB17C" box="[151,1435,1057,1079]" pageId="2" pageNumber="34">
2(1) Head with small but distinct postocular denticle (
<figureCitation id="13752A00A935FFAC08C1115DCFAFB17C" box="[696,761,1057,1079]" captionStart="FIGURE 7" captionStartId="16.[151,255,1946,1970]" captionTargetBox="[159,1435,251,1828]" captionTargetId="figure@16.[151,1436,241,1828]" captionTargetPageId="16" captionText="FIGURE 7. Pediacus species: (A D) P. leei sp. nov.; (E H) P. montivagus Champion. (A, F) Head and pronotum; (C, E) antenna; (B, H) dorsal habitus; (D, G) aedeagus." httpUri="https://zenodo.org/record/252776/files/figure.png" pageId="2" pageNumber="34">Figs. 7</figureCitation>
F, 9B)................................................. 3
</keyLead>
</paragraph>
<paragraph id="8BF13685A935FFAC0AEE1143C8CDB11E" blockId="2.[151,1437,999,2018]" box="[151,1435,1087,1109]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE1143C8CDB11E" box="[151,1435,1087,1109]" pageId="2" pageNumber="34">- Head lacking a distinct postocular denticle................................................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BA2E20A935FFAC0AEE1120C8CAB181" pageId="2" pageNumber="34">
<paragraph id="8BF13685A935FFAC0AEE1120C8CAB1DB" blockId="2.[151,1437,999,2018]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE1120C8CAB1DB" pageId="2" pageNumber="34">
3(2) Pronotum usually with disc distinctly darker than surrounding areas, anterior denticle small and broadly obtuse (
<figureCitation id="13752A00A935FFAC0F751120C813B139" box="[1292,1349,1116,1138]" captionStart="FIGURE 9" captionStartId="21.[151,255,1974,1998]" captionTargetBox="[151,1433,363,1948]" captionTargetId="figure@21.[151,1436,363,1951]" captionTargetPageId="21" captionText="FIGURE 9. Pediacus species: (A D) P. sinensis sp. nov.; (E H) P. smirnovi Nikitsky and Belov. (A, F) Head and pronotum; (B, E) antenna; (C, H) dorsal habitus; (D, G) aedeagus." httpUri="https://zenodo.org/record/252778/files/figure.png" pageId="2" pageNumber="34">Fig. 9</figureCitation>
B); male genitalia with short, globular parameres (
<figureCitation id="13752A00A935FFAC08121105CFF2B1C4" box="[619,676,1145,1167]" captionStart="FIGURE 9" captionStartId="21.[151,255,1974,1998]" captionTargetBox="[151,1433,363,1948]" captionTargetId="figure@21.[151,1436,363,1951]" captionTargetPageId="21" captionText="FIGURE 9. Pediacus species: (A D) P. sinensis sp. nov.; (E H) P. smirnovi Nikitsky and Belov. (A, F) Head and pronotum; (B, E) antenna; (C, H) dorsal habitus; (D, G) aedeagus." httpUri="https://zenodo.org/record/252778/files/figure.png" pageId="2" pageNumber="34">Fig. 9</figureCitation>
D).......................................
<taxonomicName id="4C4E4D06A935FFAC0EE61106C81FB1C4" box="[1183,1353,1146,1167]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="sinensis" status="sp. nov.">
<emphasis id="B93AEA97A935FFAC0EE61106C81FB1C4" box="[1183,1353,1146,1167]" italics="true" pageId="2" pageNumber="34">Pediacus sinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957ECA935FFAC0F361106C8CAB1DB" box="[1359,1436,1146,1168]" pageId="2" pageNumber="34" rank="species">
<emphasis id="B93AEA97A935FFAC0F361106C8CAB1DB" bold="true" box="[1359,1436,1146,1168]" pageId="2" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BF13685A935FFAC0AEE11EBC8CAB181" blockId="2.[151,1437,999,2018]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE11EBC8CAB181" pageId="2" pageNumber="34">
- Pronotum colouration uniform, anterior denticle more or less obsolete (
<figureCitation id="13752A00A935FFAC09F511EBCE90B1E7" box="[908,966,1175,1197]" captionStart="FIGURE 7" captionStartId="16.[151,255,1946,1970]" captionTargetBox="[159,1435,251,1828]" captionTargetId="figure@16.[151,1436,241,1828]" captionTargetPageId="16" captionText="FIGURE 7. Pediacus species: (A D) P. leei sp. nov.; (E H) P. montivagus Champion. (A, F) Head and pronotum; (C, E) antenna; (B, H) dorsal habitus; (D, G) aedeagus." httpUri="https://zenodo.org/record/252776/files/figure.png" pageId="2" pageNumber="34">Fig. 7</figureCitation>
F); male genitalia with elongate parameres (
<figureCitation id="13752A00A935FFAC0F1211EBCDBBB181" captionStart="FIGURE 7" captionStartId="16.[151,255,1946,1970]" captionTargetBox="[159,1435,251,1828]" captionTargetId="figure@16.[151,1436,241,1828]" captionTargetPageId="16" captionText="FIGURE 7. Pediacus species: (A D) P. leei sp. nov.; (E H) P. montivagus Champion. (A, F) Head and pronotum; (C, E) antenna; (B, H) dorsal habitus; (D, G) aedeagus." httpUri="https://zenodo.org/record/252776/files/figure.png" pageId="2" pageNumber="34">Fig. 7</figureCitation>
G).............................................................................
<taxonomicName id="4C4E4D06A935FFAC0ED511C9C8CAB181" ID-CoL="7692J" authority="Champion" authorityName="Champion" box="[1196,1436,1204,1226]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="montivagus">
<emphasis id="B93AEA97A935FFAC0ED511C9C864B181" box="[1196,1330,1205,1226]" italics="true" pageId="2" pageNumber="34">P. montivagus</emphasis>
Champion
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BA2E20A935FFAC0AEE11ADC8CDB04E" pageId="2" pageNumber="34">
<paragraph id="8BF13685A935FFAC0AEE11ADC8CDB1AC" blockId="2.[151,1437,999,2018]" box="[151,1435,1233,1255]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE11ADC8CDB1AC" box="[151,1435,1233,1255]" pageId="2" pageNumber="34">4(2) Elytra broad (&lt;1.95× longer than wide).................................................................... 5</keyLead>
</paragraph>
<paragraph id="8BF13685A935FFAC0AEE1193C8CDB04E" blockId="2.[151,1437,999,2018]" box="[151,1435,1263,1285]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE1193C8CDB04E" box="[151,1435,1263,1285]" pageId="2" pageNumber="34">- Elytra elongate (&gt;2.00× longer than wide).................................................................. 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BA2E20A935FFAC0AEE1070C8CDB074" pageId="2" pageNumber="34">
<paragraph id="8BF13685A935FFAC0AEE1070C8CAB069" blockId="2.[151,1437,999,2018]" box="[151,1436,1292,1314]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE1070C8CAB069" box="[151,1436,1292,1314]" pageId="2" pageNumber="34">
5(4) Elytra with sublateral margins of disc carinate (
<figureCitation id="13752A00A935FFAC08DE1070CFB6B069" box="[679,736,1292,1314]" captionStart="FIGURE 3" captionStartId="8.[151,250,1807,1829]" captionTargetBox="[153,1434,201,1765]" captionTargetId="figure@8.[151,1436,195,1783]" captionTargetPageId="8" captionText="FIGURE 3. Pediacus species: (A D) Pediacus bhutanicus Sen Gupta, holotype; P. carinatus sp. nov. (E, H, I) specimen from Sumatra; (F, G) specimen from Thailand. (A, E, F) Head and pronotum; (B, I) dorsal habitus; (C, G) antenna; (D, H) aedeagus." httpUri="https://zenodo.org/record/252772/files/figure.png" pageId="2" pageNumber="34">
Fig.
<date id="FFF01045A935FFAC08AD1070CFB6B069" box="[724,736,1292,1314]" pageId="2" pageNumber="34">3</date>
</figureCitation>
I)..................................
<taxonomicName id="4C4E4D06A935FFAC0EE81071C81FB069" box="[1169,1353,1293,1314]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="carinatus" status="sp. nov.">
<emphasis id="B93AEA97A935FFAC0EE81071C81FB069" box="[1169,1353,1293,1314]" italics="true" pageId="2" pageNumber="34">Pediacus carinatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957ECA935FFAC0F361070C8CAB069" box="[1359,1436,1292,1314]" pageId="2" pageNumber="34" rank="species">
<emphasis id="B93AEA97A935FFAC0F361070C8CAB069" bold="true" box="[1359,1436,1292,1314]" pageId="2" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BF13685A935FFAC0AEE1056C8CDB074" blockId="2.[151,1437,999,2018]" box="[151,1435,1321,1343]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE1056C8CDB074" box="[151,1435,1321,1343]" pageId="2" pageNumber="34">- Elytra with sublateral margins of disc costate to broadly curved................................................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BA2E20A935FFAC0AEE103BC8C0B031" pageId="2" pageNumber="34">
<paragraph id="8BF13685A935FFAC0AEE103BC8CDB016" blockId="2.[151,1437,999,2018]" box="[151,1435,1351,1373]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE103BC8CDB016" box="[151,1435,1351,1373]" pageId="2" pageNumber="34">
6(5) Pronotum anterior angles rounded (
<figureCitation id="13752A00A935FFAC084C103BCF3BB016" box="[565,621,1351,1373]" captionStart="FIGURE 5" captionStartId="12.[151,255,1972,1996]" captionTargetBox="[164,1436,365,1948]" captionTargetId="figure@12.[151,1436,361,1948]" captionTargetPageId="12" captionText="FIGURE 5. Pediacus species: (A D) P. f us c us Erichson; (E H) P. japonicoides sp. nov. (A, E) Head and pronotum; (B, E) antenna; (C, H) dorsal habitus; (D, G) aedeagus." httpUri="https://zenodo.org/record/252774/files/figure.png" pageId="2" pageNumber="34">Fig. 5</figureCitation>
A), disc of pronotum convex.............................
<emphasis id="B93AEA97A935FFAC0E94103BC86BB017" box="[1261,1341,1351,1372]" italics="true" pageId="2" pageNumber="34">P. f us c us</emphasis>
Erichson
</keyLead>
</paragraph>
<paragraph id="8BF13685A935FFAC0AEE1019C8C0B031" blockId="2.[151,1437,999,2018]" box="[151,1430,1380,1402]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE1019C8C0B031" box="[151,1430,1380,1402]" pageId="2" pageNumber="34">- Pronotum anterior angles with an acute projecting denticle, disc of pronotum weakly to moderately impressed.......... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BA2E20A935FFAC0AEE10FDC8CAB0FE" pageId="2" pageNumber="34">
<paragraph id="8BF13685A935FFAC0AEE10FDC8CAB0D3" blockId="2.[151,1437,999,2018]" box="[151,1436,1409,1432]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE10FDC8CAB0D3" box="[151,1436,1409,1432]" pageId="2" pageNumber="34">
7(6) Pronotum with sides convergent anteriorly, anteromedial denticle distinctly produced (
<figureCitation id="13752A00A935FFAC0E6710FDC900B0DC" box="[1054,1110,1409,1431]" captionStart="FIGURE 8" captionStartId="19.[151,255,1759,1783]" captionTargetBox="[151,1415,187,1723]" captionTargetId="figure@19.[151,1415,179,1729]" captionTargetPageId="19" captionText="FIGURE 8. Pediacus species: (A D) P. pendelburyi sp. nov.; (E G) P. rufipes Grouvelle, holotype. (A, E) Head and pronotum; (B, G) dorsal habitus; (C) aedeagus; (D, F) antenna." httpUri="https://zenodo.org/record/252777/files/figure.png" pageId="2" pageNumber="34">Fig. 8</figureCitation>
A)
<taxonomicName id="4C4E4D06A935FFAC0E0010FEC81FB0DC" box="[1145,1353,1410,1431]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="pendleburyi" status="sp. nov.">
<emphasis id="B93AEA97A935FFAC0E0010FEC81FB0DC" box="[1145,1353,1410,1431]" italics="true" pageId="2" pageNumber="34">Pediacus pendleburyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957ECA935FFAC0F3610FEC8CAB0D3" box="[1359,1436,1410,1432]" pageId="2" pageNumber="34" rank="species">
<emphasis id="B93AEA97A935FFAC0F3610FEC8CAB0D3" bold="true" box="[1359,1436,1410,1432]" pageId="2" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BF13685A935FFAC0AEE10E3C8CAB0FE" blockId="2.[151,1437,999,2018]" box="[151,1436,1439,1461]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE10E3C8CAB0FE" box="[151,1436,1439,1461]" pageId="2" pageNumber="34">
- Pronotum with sides subparallel anteriorly, anteromedial denticle weak to obsolete (
<figureCitation id="13752A00A935FFAC0E7F10E3C968B0FE" box="[1030,1086,1439,1461]" captionStart="FIGURE 3" captionStartId="8.[151,250,1807,1829]" captionTargetBox="[153,1434,201,1765]" captionTargetId="figure@8.[151,1436,195,1783]" captionTargetPageId="8" captionText="FIGURE 3. Pediacus species: (A D) Pediacus bhutanicus Sen Gupta, holotype; P. carinatus sp. nov. (E, H, I) specimen from Sumatra; (F, G) specimen from Thailand. (A, E, F) Head and pronotum; (B, I) dorsal habitus; (C, G) antenna; (D, H) aedeagus." httpUri="https://zenodo.org/record/252772/files/figure.png" pageId="2" pageNumber="34">Fig. 3</figureCitation>
A)......
<taxonomicName id="4C4E4D06A935FFAC0EC910E3C8CAB0FE" ID-CoL="7692G" authority="Sen Gupta" authorityName="Sen Gupta" box="[1200,1436,1439,1461]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="bhutanicus">
<emphasis id="B93AEA97A935FFAC0EC910E3C867B0FF" box="[1200,1329,1439,1460]" italics="true" pageId="2" pageNumber="34">P. bhutanicus</emphasis>
Sen Gupta
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BA2E20A935FFAC0AEE10C0C8C3B346" pageId="2" pageNumber="34">
<paragraph id="8BF13685A935FFAC0AEE10C0C8CBB0A4" blockId="2.[151,1437,999,2018]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE10C0C8CBB0A4" pageId="2" pageNumber="34">
8(4) Eyes large, coarsely faceted, with facets distinctly convex (
<figureCitation id="13752A00A935FFAC096710C0CE36B099" box="[798,864,1468,1490]" captionStart="FIGURE 2" captionStartId="6.[151,250,1807,1829]" captionTargetBox="[151,1405,195,1782]" captionTargetId="figure@6.[151,1436,195,1783]" captionTargetPageId="6" captionText="FIGURE 2. Pediacus australis sp. nov. (A, D, E, H) specimen from Philippines, Mindanao; (B, C, F, G) specimen from Australia. (A, B) Head and pronotum; (C, D) antenna; (E, F) dorsal habitus; (G, H) aedeagus." httpUri="https://zenodo.org/record/252771/files/figure.png" pageId="2" pageNumber="34">Figs. 2</figureCitation>
A, B); body colour pale orange-brown (
<figureCitation id="13752A00A935FFAC0EA510C0C848B099" box="[1244,1310,1468,1490]" captionStart="FIGURE 2" captionStartId="6.[151,250,1807,1829]" captionTargetBox="[151,1405,195,1782]" captionTargetId="figure@6.[151,1436,195,1783]" captionTargetPageId="6" captionText="FIGURE 2. Pediacus australis sp. nov. (A, D, E, H) specimen from Philippines, Mindanao; (B, C, F, G) specimen from Australia. (A, B) Head and pronotum; (C, D) antenna; (E, F) dorsal habitus; (G, H) aedeagus." httpUri="https://zenodo.org/record/252771/files/figure.png" pageId="2" pageNumber="34">Figs. 2</figureCitation>
E, F).....................................................................................
<taxonomicName id="4C4E4D06A935FFAC0EE310A6C81AB0A4" box="[1178,1356,1498,1519]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="australis" status="sp. nov.">
<emphasis id="B93AEA97A935FFAC0EE310A6C81AB0A4" box="[1178,1356,1498,1519]" italics="true" pageId="2" pageNumber="34">Pediacus australis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957ECA935FFAC0F2B10A6C8CBB0A4" box="[1362,1437,1498,1519]" pageId="2" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BF13685A935FFAC0AEE108BC8C3B346" blockId="2.[151,1437,999,2018]" box="[151,1429,1527,1549]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE108BC8C3B346" box="[151,1429,1527,1549]" pageId="2" pageNumber="34">- Eyes variable in size, finely faceted, with facets faintly convex; body colour variable.............................. 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BA2E20A935FFAC0AEE1368C8CDB30C" pageId="2" pageNumber="34">
<paragraph id="8BF13685A935FFAC0AEE1368C8CDB361" blockId="2.[151,1437,999,2018]" box="[151,1435,1556,1578]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE1368C8CDB361" box="[151,1435,1556,1578]" pageId="2" pageNumber="34">9(8) Pronotum anterior angles with a distinctly produced, acute to narrowly obtuse denticle............................. 10</keyLead>
</paragraph>
<paragraph id="8BF13685A935FFAC0AEE134EC8CDB30C" blockId="2.[151,1437,999,2018]" box="[151,1435,1585,1607]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE134EC8CDB30C" box="[151,1435,1585,1607]" pageId="2" pageNumber="34">- Pronotum anterior angles broadly obtuse, blunt or rounded, lacking a distinct denticle.............................. 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BA2E20A935FFAC0AEE1333C8CDB3C9" pageId="2" pageNumber="34">
<paragraph id="8BF13685A935FFAC0AEE1333C8CDB32E" blockId="2.[151,1437,999,2018]" box="[151,1435,1615,1637]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE1333C8CDB32E" box="[151,1435,1615,1637]" pageId="2" pageNumber="34">
10(9) Pronotum with sides convergent anteriorly (
<figureCitation id="13752A00A935FFAC08FF1333CF9FB32E" box="[646,713,1615,1637]" captionStart="FIGURE 4" captionStartId="10.[151,255,1914,1938]" captionTargetBox="[159,1436,303,1884]" captionTargetId="figure@10.[151,1436,303,1890]" captionTargetPageId="10" captionText="FIGURE 4. Pediacus species: (A D) Pediacus elongatus Sen Gupta, holotype; (E H) P. fujianensis sp. nov. (A, F) Head and pronotum; (B, E) antenna; (C, H) dorsal habitus; (D, G) aedeagus." httpUri="https://zenodo.org/record/252773/files/figure.png" pageId="2" pageNumber="34">Figs. 4</figureCitation>
A, 10A)................................................... 11
</keyLead>
</paragraph>
<paragraph id="8BF13685A935FFAC0AEE1311C8CDB3C9" blockId="2.[151,1437,999,2018]" box="[151,1435,1644,1666]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE1311C8CDB3C9" box="[151,1435,1644,1666]" pageId="2" pageNumber="34">- Pronotum with sides parallel in anterior half............................................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BA2E20A935FFAC0AEE13F5C8CAB3F6" pageId="2" pageNumber="34">
<paragraph id="8BF13685A935FFAC0AEE13F5C8CAB3EB" blockId="2.[151,1437,999,2018]" box="[151,1436,1673,1696]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE13F5C8CAB3EB" box="[151,1436,1673,1696]" pageId="2" pageNumber="34">
11(10) Antennae relatively broad, antennomere III distinctly broader than IV (
<figureCitation id="13752A00A935FFAC09F013F5CE98B3D4" box="[905,974,1673,1695]" captionStart="FIGURE 10" captionStartId="23.[151,255,1806,1830]" captionTargetBox="[173,1416,205,1782]" captionTargetId="figure@23.[162,1424,195,1783]" captionTargetPageId="23" captionText="FIGURE 10. Pediacus species: (A C) P. thomasi sp. nov.; (D H) P. taiwanensis sp. nov. (A, E) Head and pronotum; (C, D) antenna; (B, H) dorsal habitus; (F, G) variation of aedeagus." httpUri="https://zenodo.org/record/252779/files/figure.png" pageId="2" pageNumber="34">Fig. 10</figureCitation>
C)...............
<taxonomicName id="4C4E4D06A935FFAC0EE613F6C81FB3D4" box="[1183,1353,1674,1695]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="thomasi" status="sp. nov.">
<emphasis id="B93AEA97A935FFAC0EE613F6C81FB3D4" box="[1183,1353,1674,1695]" italics="true" pageId="2" pageNumber="34">Pediacus thomasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957ECA935FFAC0F3613F6C8CAB3EB" box="[1359,1436,1674,1696]" pageId="2" pageNumber="34" rank="species">
<emphasis id="B93AEA97A935FFAC0F3613F6C8CAB3EB" bold="true" box="[1359,1436,1674,1696]" pageId="2" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BF13685A935FFAC0AEE13DBC8CAB3F6" blockId="2.[151,1437,999,2018]" box="[151,1436,1703,1725]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE13DBC8CAB3F6" box="[151,1436,1703,1725]" pageId="2" pageNumber="34">
- Antennae relatively slender, antennomere III faintly broader than IV (
<figureCitation id="13752A00A935FFAC090513DBCEE0B3F6" box="[892,950,1703,1725]" captionStart="FIGURE 4" captionStartId="10.[151,255,1914,1938]" captionTargetBox="[159,1436,303,1884]" captionTargetId="figure@10.[151,1436,303,1890]" captionTargetPageId="10" captionText="FIGURE 4. Pediacus species: (A D) Pediacus elongatus Sen Gupta, holotype; (E H) P. fujianensis sp. nov. (A, F) Head and pronotum; (B, E) antenna; (C, H) dorsal habitus; (D, G) aedeagus." httpUri="https://zenodo.org/record/252773/files/figure.png" pageId="2" pageNumber="34">Fig. 4</figureCitation>
B)...................
<taxonomicName id="4C4E4D06A935FFAC0EC513DBC8CAB3F6" ID-CoL="75X2H" authority="Sen Gupta" authorityName="Sen Gupta" box="[1212,1436,1703,1725]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B93AEA97A935FFAC0EC513DBC867B3F7" box="[1212,1329,1703,1724]" italics="true" pageId="2" pageNumber="34">P. elongatus</emphasis>
Sen Gupta
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BA2E20A935FFAC0AEE13B8C8CDB3BC" pageId="2" pageNumber="34">
<paragraph id="8BF13685A935FFAC0AEE13B8C8CAB391" blockId="2.[151,1437,999,2018]" box="[151,1436,1732,1754]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE13B8C8CAB391" box="[151,1436,1732,1754]" pageId="2" pageNumber="34">
12(10) Pronotum anterior angles with a small but distinct denticle (
<figureCitation id="13752A00A935FFAC096713B8CE01B391" box="[798,855,1732,1754]" captionStart="FIGURE 8" captionStartId="19.[151,255,1759,1783]" captionTargetBox="[151,1415,187,1723]" captionTargetId="figure@19.[151,1415,179,1729]" captionTargetPageId="19" captionText="FIGURE 8. Pediacus species: (A D) P. pendelburyi sp. nov.; (E G) P. rufipes Grouvelle, holotype. (A, E) Head and pronotum; (B, G) dorsal habitus; (C) aedeagus; (D, F) antenna." httpUri="https://zenodo.org/record/252777/files/figure.png" pageId="2" pageNumber="34">Fig. 8</figureCitation>
E), disc deeply impressed anteriorly...
<taxonomicName id="4C4E4D06A935FFAC0EA513B9C8CAB391" ID-CoL="75WQK" authority="Grouvelle" authorityName="Grouvelle" box="[1244,1436,1732,1754]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="B93AEA97A935FFAC0EA513B9C860B391" box="[1244,1334,1733,1754]" italics="true" pageId="2" pageNumber="34">P. rufipes</emphasis>
Grouvelle
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF13685A935FFAC0AEE139EC8CDB3BC" blockId="2.[151,1437,999,2018]" box="[151,1435,1761,1783]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE139EC8CDB3BC" box="[151,1435,1761,1783]" pageId="2" pageNumber="34">- Pronotum anterior angles with a strongly produced acute to narrowly obtuse denticle, disc not deeply impressed anteriorly. 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BA2E20A935FFAC0AEE1383C8CAB226" pageId="2" pageNumber="34">
<paragraph id="8BF13685A935FFAC0AEE1383C8CAB279" blockId="2.[151,1437,999,2018]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE1383C8CAB279" pageId="2" pageNumber="34">
13(12). Eyes large and strongly projecting (ocular index &lt;0.70), relatively coarsely faceted; pronotum with weakly projecting obtuse denticles (
<figureCitation id="13752A00A935FFAC0B3A1260CCDEB279" box="[323,392,1820,1842]" captionStart="FIGURE 10" captionStartId="23.[151,255,1806,1830]" captionTargetBox="[173,1416,205,1782]" captionTargetId="figure@23.[162,1424,195,1783]" captionTargetPageId="23" captionText="FIGURE 10. Pediacus species: (A C) P. thomasi sp. nov.; (D H) P. taiwanensis sp. nov. (A, E) Head and pronotum; (C, D) antenna; (B, H) dorsal habitus; (F, G) variation of aedeagus." httpUri="https://zenodo.org/record/252779/files/figure.png" pageId="2" pageNumber="34">Fig. 10</figureCitation>
E)............................................................
<taxonomicName id="4C4E4D06A935FFAC0E031261C81FB279" box="[1146,1353,1821,1842]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="taiwanensis" status="sp. nov.">
<emphasis id="B93AEA97A935FFAC0E031261C81FB279" box="[1146,1353,1821,1842]" italics="true" pageId="2" pageNumber="34">Pediacus taiwanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957ECA935FFAC0F361260C8CAB279" box="[1359,1436,1820,1842]" pageId="2" pageNumber="34" rank="species">
<emphasis id="B93AEA97A935FFAC0F361260C8CAB279" bold="true" box="[1359,1436,1820,1842]" pageId="2" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BF13685A935FFAC0AEE1246C8CAB226" blockId="2.[151,1437,999,2018]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE1246C8CAB226" pageId="2" pageNumber="34">
- Eyes smaller and less strongly projecting (ocular index&gt;0.70), finely faceted; pronotum anterior angles with strongly projecting acute denticles (
<figureCitation id="13752A00A935FFAC0BD9122BCC8FB226" box="[416,473,1879,1901]" captionStart="FIGURE 7" captionStartId="16.[151,255,1946,1970]" captionTargetBox="[159,1435,251,1828]" captionTargetId="figure@16.[151,1436,241,1828]" captionTargetPageId="16" captionText="FIGURE 7. Pediacus species: (A D) P. leei sp. nov.; (E H) P. montivagus Champion. (A, F) Head and pronotum; (C, E) antenna; (B, H) dorsal habitus; (D, G) aedeagus." httpUri="https://zenodo.org/record/252776/files/figure.png" pageId="2" pageNumber="34">Fig. 7</figureCitation>
A)...........................................................
<taxonomicName id="4C4E4D06A935FFAC0EB0122BC81FB227" box="[1225,1353,1879,1900]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="leei" status="sp. nov.">
<emphasis id="B93AEA97A935FFAC0EB0122BC81FB227" box="[1225,1353,1879,1900]" italics="true" pageId="2" pageNumber="34">Pediacus leei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957ECA935FFAC0F36122BC8CAB226" box="[1359,1436,1879,1901]" pageId="2" pageNumber="34" rank="species">
<emphasis id="B93AEA97A935FFAC0F36122BC8CAB226" bold="true" box="[1359,1436,1879,1901]" pageId="2" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BA2E20A935FFAC0AEE1208C8CDB2EC" pageId="2" pageNumber="34">
<paragraph id="8BF13685A935FFAC0AEE1208C8CAB2C1" blockId="2.[151,1437,999,2018]" box="[151,1436,1908,1930]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE1208C8CAB2C1" box="[151,1436,1908,1930]" pageId="2" pageNumber="34">
14(9) Elytra strongly elongate (&gt;2.20× longer than wide) (
<figureCitation id="13752A00A935FFAC08AF1208CE59B2C1" box="[726,783,1908,1930]" captionStart="FIGURE 4" captionStartId="10.[151,255,1914,1938]" captionTargetBox="[159,1436,303,1884]" captionTargetId="figure@10.[151,1436,303,1890]" captionTargetPageId="10" captionText="FIGURE 4. Pediacus species: (A D) Pediacus elongatus Sen Gupta, holotype; (E H) P. fujianensis sp. nov. (A, F) Head and pronotum; (B, E) antenna; (C, H) dorsal habitus; (D, G) aedeagus." httpUri="https://zenodo.org/record/252773/files/figure.png" pageId="2" pageNumber="34">Fig. 4</figureCitation>
H)............................
<taxonomicName id="4C4E4D06A935FFAC0EFA1209C81FB2C1" box="[1155,1353,1909,1930]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="fujianensis" status="sp. nov.">
<emphasis id="B93AEA97A935FFAC0EFA1209C81FB2C1" box="[1155,1353,1909,1930]" italics="true" pageId="2" pageNumber="34">Pediacus fujianensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957ECA935FFAC0F361208C8CAB2C1" box="[1359,1436,1908,1930]" pageId="2" pageNumber="34" rank="species">
<emphasis id="B93AEA97A935FFAC0F361208C8CAB2C1" bold="true" box="[1359,1436,1908,1930]" pageId="2" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BF13685A935FFAC0AEE12EEC8CDB2EC" blockId="2.[151,1437,999,2018]" box="[151,1435,1937,1959]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE12EEC8CDB2EC" box="[151,1435,1937,1959]" pageId="2" pageNumber="34">- Elytra less strongly elongate (&lt;2.20× longer than wide)...................................................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BA2E20A935FFAC0AEE12D3C8CDB2A9" pageId="2" pageNumber="34">
<paragraph id="8BF13685A935FFAC0AEE12D3C8CDB28E" blockId="2.[151,1437,999,2018]" box="[151,1435,1967,1989]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE12D3C8CDB28E" box="[151,1435,1967,1989]" pageId="2" pageNumber="34">
15(14) Eyes small, weakly projecting (ocular index&gt;0.80) (
<figureCitation id="13752A00A935FFAC08A012D3CE44B28E" box="[729,786,1967,1989]" captionStart="FIGURE 6" captionStartId="14.[151,255,1914,1938]" captionTargetBox="[162,1421,309,1885]" captionTargetId="figure@14.[151,1436,303,1890]" captionTargetPageId="14" captionText="FIGURE 6. Pediacus species: (A D) P. japonicus Reitter; (E G) P. kurosawai Sasaji, holotype. (A, E) Head and pronotum; (B, G) dorsal habitus; (C) aedeagus; (D, F) antenna." httpUri="https://zenodo.org/record/252775/files/figure.png" pageId="2" pageNumber="34">Fig. 6</figureCitation>
E)...................................
<taxonomicName id="4C4E4D06A935FFAC0E9B12D3C8CDB28E" ID-CoL="75WQJ" authority="Sasaji" authorityName="Sasaji" box="[1250,1435,1967,1989]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="kurosawai">
<emphasis id="B93AEA97A935FFAC0E9B12D3C80AB28F" box="[1250,1372,1967,1988]" italics="true" pageId="2" pageNumber="34">P. kurosawai</emphasis>
Sasaji
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF13685A935FFAC0AEE12B1C8CDB2A9" blockId="2.[151,1437,999,2018]" box="[151,1435,1996,2018]" pageId="2" pageNumber="34">
<keyLead id="30BF95B0A935FFAC0AEE12B1C8CDB2A9" box="[151,1435,1996,2018]" pageId="2" pageNumber="34">- Eyes larger, more strongly projecting (ocular index &lt;0.78).................................................... 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BA2E20A934FFAD0AEE15EBC8CAB5AC" pageId="3" pageNumber="35">
<paragraph id="8BF13685A934FFAD0AEE15EBC8CAB581" blockId="3.[151,1436,151,290]" pageId="3" pageNumber="35">
<keyLead id="30BF95B0A934FFAD0AEE15EBC8CAB581" pageId="3" pageNumber="35">
16(15) Pronotum with disc deeply impressed, setation short and inconspicuous, punctures sparse and weakly impressed medially (
<figureCitation id="13752A00A934FFAD0F0915EBCDBBB581" captionStart="FIGURE 9" captionStartId="21.[151,255,1974,1998]" captionTargetBox="[151,1433,363,1948]" captionTargetId="figure@21.[151,1436,363,1951]" captionTargetPageId="21" captionText="FIGURE 9. Pediacus species: (A D) P. sinensis sp. nov.; (E H) P. smirnovi Nikitsky and Belov. (A, F) Head and pronotum; (B, E) antenna; (C, H) dorsal habitus; (D, G) aedeagus." httpUri="https://zenodo.org/record/252778/files/figure.png" pageId="3" pageNumber="35">Fig. 9</figureCitation>
F)........................................................................
<taxonomicName id="4C4E4D06A934FFAD0E0915C9C8CAB581" authority="Nikitsky and Belov" authorityName="Nikitsky and Belov" box="[1136,1436,180,202]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="smi" subSpecies="rnovi">
<emphasis id="B93AEA97A934FFAD0E0915C9C98FB581" box="[1136,1241,181,202]" italics="true" pageId="3" pageNumber="35">P. smi rnovi</emphasis>
Nikitsky and Belov
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF13685A934FFAD0AEE15AEC8CAB5AC" blockId="3.[151,1436,151,290]" box="[151,1436,209,231]" pageId="3" pageNumber="35">
<keyLead id="30BF95B0A934FFAD0AEE15AEC8CAB5AC" box="[151,1436,209,231]" pageId="3" pageNumber="35">- Pronotum with disc moderately impressed, setation conspicuous, punctures dense and more strongly impressed medially.. 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BA2E20A934FFAD0AEE1593C8CDB469" pageId="3" pageNumber="35">
<paragraph id="8BF13685A934FFAD0AEE1593C8CAB44E" blockId="3.[151,1436,151,290]" box="[151,1436,239,261]" pageId="3" pageNumber="35">
<keyLead id="30BF95B0A934FFAD0AEE1593C8CAB44E" box="[151,1436,239,261]" pageId="3" pageNumber="35">
17(16) Pronotum colouration uniform, anterior angles broadly obtuse (
<figureCitation id="13752A00A934FFAD09391593CE2EB44E" box="[832,888,239,261]" captionStart="FIGURE 5" captionStartId="12.[151,255,1972,1996]" captionTargetBox="[164,1436,365,1948]" captionTargetId="figure@12.[151,1436,361,1948]" captionTargetPageId="12" captionText="FIGURE 5. Pediacus species: (A D) P. f us c us Erichson; (E H) P. japonicoides sp. nov. (A, E) Head and pronotum; (B, E) antenna; (C, H) dorsal habitus; (D, G) aedeagus." httpUri="https://zenodo.org/record/252774/files/figure.png" pageId="3" pageNumber="35">Fig. 5</figureCitation>
F)..................
<taxonomicName id="4C4E4D06A934FFAD0E081593C81FB44F" box="[1137,1353,239,260]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="japonicoides" status="sp. nov.">
<emphasis id="B93AEA97A934FFAD0E081593C81FB44F" box="[1137,1353,239,260]" italics="true" pageId="3" pageNumber="35">Pediacus japonicoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957ECA934FFAD0F361593C8CAB44E" box="[1359,1436,239,261]" pageId="3" pageNumber="35" rank="species">
<emphasis id="B93AEA97A934FFAD0F361593C8CAB44E" bold="true" box="[1359,1436,239,261]" pageId="3" pageNumber="35">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BF13685A934FFAD0AEE1471C8CDB469" blockId="3.[151,1436,151,290]" box="[151,1435,268,290]" pageId="3" pageNumber="35">
<keyLead id="30BF95B0A934FFAD0AEE1471C8CDB469" box="[151,1435,268,290]" pageId="3" pageNumber="35">
- Pronotum usually with disc distinctly darker than surrounding areas; anterior angles rounded (
<figureCitation id="13752A00A934FFAD0E141470C9F3B469" box="[1133,1189,268,290]" captionStart="FIGURE 6" captionStartId="14.[151,255,1914,1938]" captionTargetBox="[162,1421,309,1885]" captionTargetId="figure@14.[151,1436,303,1890]" captionTargetPageId="14" captionText="FIGURE 6. Pediacus species: (A D) P. japonicus Reitter; (E G) P. kurosawai Sasaji, holotype. (A, E) Head and pronotum; (B, G) dorsal habitus; (C) aedeagus; (D, F) antenna." httpUri="https://zenodo.org/record/252775/files/figure.png" pageId="3" pageNumber="35">Fig. 6</figureCitation>
A)..
<taxonomicName id="4C4E4D06A934FFAD0E991471C8CDB469" ID-CoL="6TX2J" authority="Reitter" authorityName="Reitter" box="[1248,1435,268,290]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B93AEA97A934FFAD0E991471C802B469" box="[1248,1364,269,290]" italics="true" pageId="3" pageNumber="35">P. japonicus</emphasis>
Reitter
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>