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<document id="08AC144BF748C224C8FDAC9380D6C137" ID-CLB-Dataset="46694" ID-DOI="10.11646/zootaxa.3754.1.2" ID-GBIF-Dataset="f435fbaf-802a-4202-92ff-6d69aa405739" ID-ISSN="1175-5326" ID-Zenodo-Dep="252769" ID-ZooBank="D873973D-A4F5-455F-83DD-4B3E564DB630" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460367436938" checkinUser="plazi" docAuthor="Marris, John W. M. &amp; Ślipiński, Adam" docDate="2014" docId="03E78793A934FFAD0AEE140EC833B226" docLanguage="en" docName="zt03754p058.pdf" docOrigin="Zootaxa 3754 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pediacus ater Grouvelle" docType="treatment" docVersion="11" lastPageNumber="35" masterDocId="FFDEFFEBA937FFAE0A79157CCD56B54B" masterDocTitle="A revision of the Pediacus Shuckard 1839 (Coleoptera: Cucujidae) of Asia and Australasia" masterLastPageNumber="58" masterPageNumber="32" pageNumber="35" updateTime="1698295345156" updateUser="ExternalLinkService">
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<mods:title id="921E70119CFE0A8B5DB5A71CDB4DC395">A revision of the Pediacus Shuckard 1839 (Coleoptera: Cucujidae) of Asia and Australasia</mods:title>
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<mods:namePart id="D28FF900F1F9911871FCCB5E9B753C60">Marris, John W. M.</mods:namePart>
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<mods:namePart id="4F6BAB4161988BC41DD01670E023608C">Ślipiński, Adam</mods:namePart>
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<mods:date id="A9A20C776D72465FC25B638568401B4F">2014</mods:date>
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<treatment id="03E78793A934FFAD0AEE140EC833B226" ID-DOI="http://doi.org/10.5281/zenodo.6139870" ID-GBIF-Taxon="119411055" ID-Zenodo-Dep="6139870" LSID="urn:lsid:plazi:treatment:03E78793A934FFAD0AEE140EC833B226" httpUri="http://treatment.plazi.org/id/03E78793A934FFAD0AEE140EC833B226" lastPageNumber="35" pageId="3" pageNumber="35">
<subSubSection id="C354650EA934FFAD0AEE140ECCEBB4C7" box="[151,445,370,396]" pageId="3" pageNumber="35" type="nomenclature">
<paragraph id="8BF13685A934FFAD0AEE140ECCEBB4C7" blockId="3.[151,445,370,430]" box="[151,445,370,396]" pageId="3" pageNumber="35">
<heading id="D0B981E9A934FFAD0AEE140ECCEBB4C7" bold="true" box="[151,445,370,396]" fontSize="11" level="1" pageId="3" pageNumber="35" reason="1">
<taxonomicName id="4C4E4D06A934FFAD0AEE140ECCEBB4C7" ID-CoL="768QG" authority="Grouvelle" authorityName="Grouvelle" box="[151,445,370,396]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="ater">
<emphasis id="B93AEA97A934FFAD0AEE140ECCEBB4C7" bold="true" box="[151,445,370,396]" pageId="3" pageNumber="35">
<emphasis id="B93AEA97A934FFAD0AEE140ECC6CB4C7" bold="true" box="[151,314,370,396]" italics="true" pageId="3" pageNumber="35">Pediacus ater</emphasis>
Grouvelle
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C354650EA934FFAD0AEE14E9CC77B4E5" box="[151,289,405,430]" pageId="3" pageNumber="35" type="description">
<paragraph id="8BF13685A934FFAD0AEE14E9CC77B4E5" blockId="3.[151,445,370,430]" box="[151,289,405,430]" pageId="3" pageNumber="35">
<figureCitation id="13752A00A934FFAD0AEE14E9CD88B4E5" box="[151,222,405,430]" captionStart="FIGURE 1" captionStartId="4.[151,255,1802,1826]" captionTargetBox="[151,1391,194,1765]" captionTargetId="figure@4.[151,1436,194,1781]" captionTargetPageId="4" captionText="FIGURE 1. Pediacus ater Grouvelle (A, C, E, H) specimen from Borneo; (B, D, F, G) specimen from Sumatra. (A, B) Head and pronotum; (C, D) dorsal habitus; (E, F) antenna; (G, H) aedeagus." httpUri="https://zenodo.org/record/252770/files/figure.png" pageId="3" pageNumber="35">Figs. 1</figureCitation>
AH.
</paragraph>
</subSubSection>
<subSubSection id="C354650EA934FFAD0AEE14A1CF5EB4B8" box="[151,520,477,499]" pageId="3" pageNumber="35" type="reference_group">
<paragraph id="8BF13685A934FFAD0AEE14A1CF5EB4B8" blockId="3.[151,520,477,499]" box="[151,520,477,499]" pageId="3" pageNumber="35">
<treatmentCitationGroup id="AB5E11ABA934FFAD0AEE14A1CF5EB4B8" box="[151,520,477,499]" pageId="3" pageNumber="35">
<taxonomicName id="4C4E4D06A934FFAD0AEE14A1CF52B4B8" ID-CoL="768QG" authority="Grouvelle, 1897: 396" authorityName="Grouvelle" authorityPageNumber="396" authorityYear="1897" box="[151,516,477,499]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="ater">
<emphasis id="B93AEA97A934FFAD0AEE14A1CC75B4B8" box="[151,291,477,499]" italics="true" pageId="3" pageNumber="35">Pediacus ater</emphasis>
<treatmentCitation id="0AEF1094A934FFAD0B5014A1CF52B4B8" author="Grouvelle" box="[297,516,477,499]" page="396" pageId="3" pageNumber="35" year="1897">
<bibRefCitation id="EFDF4B74A934FFAD0B5014A1CF52B4B8" author="Grouvelle, A." box="[297,516,477,499]" firstAuthor="Grouvelle" journalOrPublisher="Annali del Museo Civico di Storia Naturale. Giacomo Doria. (Series 2)" page="396" pageId="3" pageNumber="35" pagination="342 - 398" part="28" refId="ref15497" refString="Grouvelle, A. (1897) Clavicornes nouveaux des indes orientales et pays voisins. Annali del Museo Civico di Storia Naturale. Giacomo Doria. (Series 2), 28, 342 - 398." title="Clavicornes nouveaux des indes orientales et pays voisins" type="journal article" year="1897">Grouvelle, 1897: 396</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
</subSubSection>
<subSubSection id="C354650EA934FFAD0AEE1763C9AFB716" pageId="3" pageNumber="35" type="diagnosis">
<paragraph id="8BF13685A934FFAD0AEE1763C9AFB716" blockId="3.[151,1436,543,1901]" pageId="3" pageNumber="35">
<emphasis id="B93AEA97A934FFAD0AEE1763CC47B773" bold="true" box="[151,273,543,568]" pageId="3" pageNumber="35">Diagnosis.</emphasis>
Recognisable by the combination of the following characters: body form broad, surface highly glossy, pronotum strongly explanate with glabrous lateral margins, elytra with epipleura strongly explanate.
</paragraph>
</subSubSection>
<subSubSection id="C354650EA934FFAD0ABE171BCCEEB1D6" pageId="3" pageNumber="35" type="description">
<paragraph id="8BF13685A934FFAD0ABE171BCCEEB1D6" blockId="3.[151,1436,543,1901]" pageId="3" pageNumber="35">
<emphasis id="B93AEA97A934FFAD0ABE171BCCF6B7CB" bold="true" box="[199,416,615,640]" pageId="3" pageNumber="35">Description. Body</emphasis>
length
<quantity id="4CB69B60A934FFAD0B801714CFD5B734" box="[505,643,616,640]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.7" metricValueMin="3.1" pageId="3" pageNumber="35" unit="mm" value="3.9" valueMax="4.7" valueMin="3.1">3.14.7 mm</quantity>
(n=9), broad, 2.55× longer than wide; colour variable, ranging from black-brown to orange-brown, disc of pronotum occasionally darker, with head, antennae, lateral margins of pronotum and legs paler; pubescence short, pale, appressed and inconspicuous; surface highly glossy.
<emphasis id="B93AEA97A934FFAD0F2517D3C8CDB783" bold="true" box="[1372,1435,687,712]" pageId="3" pageNumber="35">Head</emphasis>
strongly transverse, 1.86× wider than long; puncturation moderately impressed, broad and sparse medially, increasing in size and density laterally with punctures separated by distinctly less than their diameter; setae very short and appressed, inconspicuous; microsculpture absent, surface glossy; eyes large, very strongly projecting, ocular index 0.69, finely faceted, postocular denticle absent; antennae as in
<figureCitation id="13752A00A934FFAD0E7B1643C918B613" box="[1026,1102,831,856]" captionStart="FIGURE 1" captionStartId="4.[151,255,1802,1826]" captionTargetBox="[151,1391,194,1765]" captionTargetId="figure@4.[151,1436,194,1781]" captionTargetPageId="4" captionText="FIGURE 1. Pediacus ater Grouvelle (A, C, E, H) specimen from Borneo; (B, D, F, G) specimen from Sumatra. (A, B) Head and pronotum; (C, D) dorsal habitus; (E, F) antenna; (G, H) aedeagus." httpUri="https://zenodo.org/record/252770/files/figure.png" pageId="3" pageNumber="35">Figs. 1</figureCitation>
E, F, antennomere VII larger than VI and VIII.
<emphasis id="B93AEA97A934FFAD0B1E1618CCB6B636" bold="true" box="[359,480,868,893]" pageId="3" pageNumber="35">Pronotum</emphasis>
strongly transverse, 1.40× wider than long, widest at about midpoint, sides broadly curved; lateral margins strongly explanate, denticles ranging from broadly rounded to narrowly obtuse, anterior denticle almost obsolete in some specimens; disc planar to faintly impressed; puncturation as for head, with a narrow, impunctate median strip most prominent basally; setation and microsculpture as for head, lateral margins glabrous.
<emphasis id="B93AEA97A934FFAD0B741688CC0FB146" bold="true" box="[269,345,1012,1037]" pageId="3" pageNumber="35">Elytra</emphasis>
broadly elongate, 1.70× longer than wide, sides subparallel; epipleura strongly explanate, reflexed; puncturation moderately dense, moderately impressed basally, becoming weakly impressed laterally and posteriorly; setation short and inconspicuous; microsculpture absent, surface glossy.
<emphasis id="B93AEA97A934FFAD0E341140C9EDB11E" bold="true" box="[1101,1211,1084,1109]" pageId="3" pageNumber="35">Genitalia</emphasis>
for male as in
<figureCitation id="13752A00A934FFAD0F1C1140CDF7B133" captionStart="FIGURE 1" captionStartId="4.[151,255,1802,1826]" captionTargetBox="[151,1391,194,1765]" captionTargetId="figure@4.[151,1436,194,1781]" captionTargetPageId="4" captionText="FIGURE 1. Pediacus ater Grouvelle (A, C, E, H) specimen from Borneo; (B, D, F, G) specimen from Sumatra. (A, B) Head and pronotum; (C, D) dorsal habitus; (E, F) antenna; (G, H) aedeagus." httpUri="https://zenodo.org/record/252770/files/figure.png" pageId="3" pageNumber="35">Figs. 1</figureCitation>
G, H, parameres elongate, tapering apically; internal sac with strong tooth-like projections basally and a sclerotized apical section.
</paragraph>
</subSubSection>
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<emphasis id="B93AEA97A934FFAD0ABE11DBCC22B18B" bold="true" box="[199,372,1191,1216]" pageId="3" pageNumber="35">
<typeStatus id="54F58827A934FFAD0ABE11DBCC54B18B" box="[199,258,1191,1216]" pageId="3" pageNumber="35">Type</typeStatus>
material.
</emphasis>
<materialsCitation id="3B263CD8A934FFAD0B0311D4C95AB18B" ID-GBIF-Occurrence="3923183335" box="[378,1036,1191,1216]" pageId="3" pageNumber="35" typeStatus="syntype">
<typeStatus id="54F58827A934FFAD0B0311D4CC80B18B" box="[378,470,1192,1216]" pageId="3" pageNumber="35" type="syntype">Syntype</typeStatus>
from Sumatra, Si-Rambe, E. Modigliani (
<collectionCode id="ED5FAE40A934FFAD09D311D4C955B18B" LSID="urn:lsid:biocol.org:col:34988" box="[938,1027,1192,1216]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="35">MNHN</collectionCode>
)
</materialsCitation>
was examined and photographed in 1997 by
<collectingCountry id="F3597615A934FFAD0A8011B0CC76B1AF" box="[249,288,1228,1252]" name="American Samoa" pageId="3" pageNumber="35">AS</collectingCountry>
.
</paragraph>
<paragraph id="8BF13685A934FFAD0ABE1193C8C2B24B" blockId="3.[151,1436,543,1901]" pageId="3" pageNumber="35">
<emphasis id="B93AEA97A934FFAD0ABE1193CF65B043" bold="true" box="[199,563,1263,1288]" pageId="3" pageNumber="35">Other Specimens Examined.</emphasis>
<materialsCitation id="3B263CD8A934FFAD083E1193C9EEB067" ID-GBIF-Occurrence="3923183314" collectingDate="1921-02-19" collectingDateMax="1921-02-23" collectingDateMin="1921-02-19" collectionCode="NHMB" collectorName="Bocak &amp; Bocakova" country="Indonesia" elevation="1675" location="Sumatra" pageId="3" pageNumber="35" specimenCount="10" stateProvince="Sumatra">
<emphasis id="B93AEA97A934FFAD083E1193CE22B043" bold="true" box="[583,884,1263,1288]" pageId="3" pageNumber="35">
<collectingCountry id="F3597615A934FFAD083E1193CFBCB043" box="[583,746,1263,1288]" name="Indonesia" pageId="3" pageNumber="35">INDONESIA</collectingCountry>
:
<collectingRegion id="498AF867A934FFAD097B1193CE39B043" box="[770,879,1263,1288]" pageId="3" pageNumber="35">Sumatra</collectingRegion>
:
</emphasis>
SUMATRA (N. [North]), BRASTAGI [
<emphasis id="B93AEA97A934FFAD0F0F118CC8C3B04C" box="[1398,1429,1264,1287]" italics="true" pageId="3" pageNumber="35">sic</emphasis>
, =Berasatagi] G. Sibayak,
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<date id="FFF01045A934FFAD082D1068CE55B067" box="[596,771,1300,1324]" pageId="3" pageNumber="35" value="1921-02-19" valueMax="1921-02-23" valueMin="1921-02-19">
<collectingDate id="EFB4E9ADA934FFAD082D1068CE55B067" box="[596,771,1300,1324]" pageId="3" pageNumber="35" value="1921-02-19" valueMax="1921-02-23" valueMin="1921-02-19">19.23.II. 1921</collectingDate>
</date>
<collectorName id="26BB5353A934FFAD09721068CEBFB066" box="[779,1001,1300,1325]" pageId="3" pageNumber="35">Bocák &amp; Bocáková</collectorName>
lgt. (
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,
<collectionCode id="ED5FAE40A934FFAD0E2A1068C9FAB067" box="[1107,1196,1300,1324]" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="3" pageNumber="35">NHMB</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B263CD8A934FFAD0EB91068C9EDB01B" ID-GBIF-Occurrence="3923183334" collectingDate="1987-05-16" collectionCode="MHNG" collectorName="A. Smetana" elevation="1500" location="Mt. Kinabalu Nat. Pk. HQ at Liwagu Rv." pageId="3" pageNumber="35" specimenCount="2" stateProvince="BORNEO SABAH">
<emphasis id="B93AEA97A934FFAD0EB91068C876B066" bold="true" box="[1216,1312,1300,1325]" pageId="3" pageNumber="35">Borneo:</emphasis>
<collectingRegion id="498AF867A934FFAD0F501068CDA0B01B" pageId="3" pageNumber="35">BORNEO SABAH</collectingRegion>
<location id="8E91605EA934FFAD0A841045CF9BB01B" LSID="urn:lsid:plazi:treatment:03E78793A934FFAD0AEE140EC833B226:8E91605EA934FFAD0A841045CF9BB01B" box="[253,717,1335,1360]" name="Mt. Kinabalu Nat. Pk. HQ at Liwagu Rv." pageId="3" pageNumber="35" stateProvince="BORNEO SABAH">Mt. Kinabalu Nat. Pk. HQ at Liwagu Rv.</location>
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<elevation id="0063D1B6A934FFAD08AD104BCE74B01B" box="[724,802,1335,1360]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="35" unit="m" value="1500.0">1500m</elevation>
</quantity>
<date id="FFF01045A934FFAD0950104BCECAB01B" box="[809,924,1335,1360]" pageId="3" pageNumber="35" value="1987-05-16">
<collectingDate id="EFB4E9ADA934FFAD0950104BCECAB01B" box="[809,924,1335,1360]" pageId="3" pageNumber="35" value="1987-05-16">16.V.1987</collectingDate>
</date>
<collectorName id="26BB5353A934FFAD09DD1044C971B01B" box="[932,1063,1336,1360]" pageId="3" pageNumber="35">A. Smetana</collectorName>
(
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,
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);
</materialsCitation>
<materialsCitation id="3B263CD8A934FFAD0EBB1044C8CEB03F" ID-GBIF-Occurrence="3923183364" pageId="3" pageNumber="35">
BORNEO
<collectionCode id="ED5FAE40A934FFAD0F441044C8CAB01B" box="[1341,1436,1336,1360]" pageId="3" pageNumber="35">SABAH</collectionCode>
Mt. Kinabalu N. P. Headquarters
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<date id="FFF01045A934FFAD08111020CF8BB03F" box="[616,733,1372,1396]" pageId="3" pageNumber="35" value="1987-04-29">IV2987</date>
, Large cut tree-Quercus sp. D.E. Bright collector (1,
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;
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N.[North] BORNEO, BETTOTAN. Nr. SANDAKAN.
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[Following text on reverse] C. B. K. &amp; H. M. P., F.M.S Museums (3,
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)
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;
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BORNEO: Sabah km. 62, Lahad Datu To Danum Valley F. S.
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, Under bark D.E. Bright Collector (1,
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;
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Mt Tibang
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., O. Borneo Mjöberg (1,
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); Pajau River, O. Borneo Mjöberg (1,
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);
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<materialsCitation id="3B263CD8A934FFAD0B831090C8CAB34F" ID-GBIF-Occurrence="3923183349" box="[506,1436,1516,1541]" pageId="3" pageNumber="35">
BRITISH N. BORNEO Tenompok
<date id="FFF01045A934FFAD09E41090C943B34E" box="[925,1045,1516,1541]" pageId="3" pageNumber="35" value="1959-02-13">13.II.1959</date>
, T.C. Maa Collector (2,
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),
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<materialsCitation id="3B263CD8A934FFAD0AEE136CC8CAB363" ID-GBIF-Occurrence="3923183347" box="[151,1436,1551,1576]" collectingDate="1988-08-24" collectionCode="MHNG" collectorName="A. Smetana" elevation="485" location="Mt. Kinabalu Nat. Pk. Poring Hot Spgs" pageId="3" pageNumber="35" specimenCount="1" stateProvince="BORNEO SABAH">
<collectingRegion id="498AF867A934FFAD0AEE136CCC25B363" box="[151,371,1552,1576]" pageId="3" pageNumber="35">BORNEO SABAH</collectingRegion>
<location id="8E91605EA934FFAD0B04136DCE1AB363" LSID="urn:lsid:plazi:treatment:03E78793A934FFAD0AEE140EC833B226:8E91605EA934FFAD0B04136DCE1AB363" box="[381,844,1551,1576]" name="Mt. Kinabalu Nat. Pk. Poring Hot Spgs" pageId="3" pageNumber="35" stateProvince="BORNEO SABAH">Mt. Kinabalu Nat. Pk. Poring Hot Spgs</location>
.
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<date id="FFF01045A934FFAD09DD136CC94FB363" box="[932,1049,1552,1576]" pageId="3" pageNumber="35" value="1988-08-24">
<collectingDate id="EFB4E9ADA934FFAD09DD136CC94FB363" box="[932,1049,1552,1576]" pageId="3" pageNumber="35" value="1988-08-24">24.VIII.88</collectingDate>
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<collectorName id="26BB5353A934FFAD0E5D136CC9FFB363" box="[1060,1193,1552,1576]" pageId="3" pageNumber="35">A. Smetana</collectorName>
[B142] (
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,
<collectionCode id="ED5FAE40A934FFAD0F4F136CC8C7B363" box="[1334,1425,1552,1576]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="3" pageNumber="35">MHNG</collectionCode>
).
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<materialsCitation id="3B263CD8A934FFAD0AEE1348CCF8B33B" ID-GBIF-Occurrence="3923183322" pageId="3" pageNumber="35">
<emphasis id="B93AEA97A934FFAD0AEE1348CC60B306" bold="true" box="[151,310,1588,1613]" pageId="3" pageNumber="35">
<collectingCountry id="F3597615A934FFAD0AEE1348CC67B306" box="[151,305,1588,1613]" name="Malaysia" pageId="3" pageNumber="35">MALAYSIA</collectingCountry>
:
</emphasis>
PAHANG, F. M. S. [
<collectingCountry id="F3597615A934FFAD08461348CE0EB307" box="[575,856,1588,1613]" name="Malaysia" pageId="3" pageNumber="35">Federated Malay States</collectingCountry>
] Lubok Tamang
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.
<date id="FFF01045A934FFAD0EEC1348C9AAB307" box="[1173,1276,1588,1613]" pageId="3" pageNumber="35">7th June</date>
, 1923 H. M. Pendlebury. (1,
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);
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<collectingCountry id="F3597615A934FFAD0BCF1324CF1AB33B" box="[438,588,1624,1648]" name="Malaysia" pageId="3" pageNumber="35">MALAYSIA</collectingCountry>
: Pahang Tioman Island Umg. Kampung Tekek 15.
<date id="FFF01045A934FFAD0EF7132BC852B33B" box="[1166,1284,1623,1648]" pageId="3" pageNumber="35" value="1993-07-24">24.7.1993</date>
lg. Schuh (1, CRS);
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: Selangor Fraser Hill,
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<date id="FFF01045A934FFAD08B71300CE68B3DF" box="[718,830,1660,1684]" pageId="3" pageNumber="35" value="1993-08-14">14.8.1993</date>
lg. Schuh (1, CRS).
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<emphasis id="B93AEA97A934FFAD0E551300C9BEB3DE" bold="true" box="[1068,1256,1660,1685]" pageId="3" pageNumber="35">PHILIPPINES:</emphasis>
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,
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<elevation id="0063D1B6A934FFAD0AEE13E3CDB3B3F3" box="[151,229,1695,1720]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="35" unit="m" value="1600.0">1600m</elevation>
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,
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<quantity id="4CB69B60A934FFAD0B0B13E3CCE4B3F3" box="[370,434,1695,1720]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="35" unit="km" value="30.0">30km</quantity>
W of Maramag
</locationDeviation>
,
<date id="FFF01045A934FFAD080713DCCE17B3F3" box="[638,833,1696,1720]" pageId="3" pageNumber="35" value="1990-12-28" valueMax="1990-12-30" valueMin="1990-12-28">
<collectingDate id="EFB4E9ADA934FFAD080713DCCE17B3F3" box="[638,833,1696,1720]" pageId="3" pageNumber="35" value="1990-12-28" valueMax="1990-12-30" valueMin="1990-12-28">28.30.Dec 1990</collectingDate>
</date>
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lgt. (
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,
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).
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<emphasis id="B93AEA97A934FFAD0E2313E3C9ACB3F3" bold="true" box="[1114,1274,1695,1720]" pageId="3" pageNumber="35">
<collectingCountry id="F3597615A934FFAD0E2313E3C9A3B3F3" box="[1114,1269,1695,1720]" name="Thailand" pageId="3" pageNumber="35">THAILAND</collectingCountry>
:
</emphasis>
<collectingCountry id="F3597615A934FFAD0F7D13DCC8CEB3F3" box="[1284,1432,1696,1720]" name="Thailand" pageId="3" pageNumber="35">THAILAND</collectingCountry>
, Chiang MaiP [Chiang Mai Province] DoiSuthep, St. LouisChr. Camp, V.28.1999, CW&amp;LB. O'Brien (4,
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);
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NW
<collectingCountry id="F3597615A934FFAD0AB5139BCC64B24B" box="[204,306,1767,1792]" name="Thailand" pageId="3" pageNumber="35">Thailand</collectingCountry>
, 1991
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,
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,
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19.23.4.,
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leg. (
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,
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).
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</paragraph>
</subSubSection>
<subSubSection id="C354650EA934FFAD0ABE1270C973B26E" box="[199,1061,1804,1829]" pageId="3" pageNumber="35" type="distribution">
<paragraph id="8BF13685A934FFAD0ABE1270C973B26E" blockId="3.[151,1436,543,1901]" box="[199,1061,1804,1829]" pageId="3" pageNumber="35">
<emphasis id="B93AEA97A934FFAD0ABE1270CC36B26E" bold="true" box="[199,352,1804,1829]" pageId="3" pageNumber="35">Distribution.</emphasis>
<collectingCountry id="F3597615A934FFAD0B1E1270CC82B26E" box="[359,468,1804,1829]" name="Indonesia" pageId="3" pageNumber="35">Indonesia</collectingCountry>
(Sumatra, Borneo),
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,
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,
<collectingCountry id="F3597615A934FFAD09C31270C977B26E" box="[954,1057,1804,1829]" name="Thailand" pageId="3" pageNumber="35">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C354650EA934FFAD0ABE1253C833B226" pageId="3" pageNumber="35" type="discussion">
<paragraph id="8BF13685A934FFAD0ABE1253C833B226" blockId="3.[151,1436,543,1901]" pageId="3" pageNumber="35">
<emphasis id="B93AEA97A934FFAD0ABE1253CC1AB203" bold="true" box="[199,332,1839,1864]" pageId="3" pageNumber="35">Discussion.</emphasis>
This species shows considerable colour variation, ranging from unicolorous black-brown to orange-brown. In some specimens the disc of the pronotum is significantly darker than the rest of the dorsum.
</paragraph>
</subSubSection>
</treatment>
</document>