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<document id="414B06BD2701F86572528C078B7A6B1F" ID-CLB-Dataset="22106" ID-DOI="10.11646/zootaxa.4778.3.8" ID-GBIF-Dataset="db595c03-4859-4d5b-94f6-f9d73a40db69" ID-ISSN="1175-5326" ID-Zenodo-Dep="3828825" ID-ZooBank="330A0F91-1F61-4DD7-948A-78955F8455D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589541046722" checkinUser="plazi" docAuthor="Rodríguez, William David, Navarrete-Heredia, José L., Arriaga-Varela, Emmanuel &amp; Cuccodoro, Giulio" docDate="2020" docId="03E5ED0AFF9ED56F2B996A19FB60FA55" docLanguage="en" docName="zootaxa.4778.3.8.pdf" docOrigin="Zootaxa 4778 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megarthrus chiapas Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro 2020" docType="treatment" docVersion="7" lastPageNumber="578" masterDocId="FFDC9572FF9FD5682B0E6C20FFE0FFB2" masterDocTitle="Two new species of Megarthrus (Coleoptera, Staphylinidae, Proteininae) from Mexican Cloud Forests" masterLastPageNumber="583" masterPageNumber="571" pageNumber="572" updateTime="1698839835231" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="DE95DEA9B49D156ADC15B05F4059BA23" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E19C086ABCB01BD396DC04E8C5F6939C">
<mods:title id="A9AB85A51AB3BF74150FBA21AEDEA7F1">Two new species of Megarthrus (Coleoptera, Staphylinidae, Proteininae) from Mexican Cloud Forests</mods:title>
</mods:titleInfo>
<mods:name id="CE84DFE0CCF427524B98741282E9CCC4" type="personal">
<mods:role id="5EC5C354F25AB23DACEF7FB003DE3F42">
<mods:roleTerm id="9C770E4E0D9D5F2B7655EDA62328CA76">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5101A8DC80CB30FEC69DA7632CD72226">Rodríguez, William David</mods:namePart>
<mods:affiliation id="E3A50F45E665D0F1D4714F0E88E558A7">Entomología, Centro de Estudios en Zoología, CUCBA, Universidad de Guadalajara, ApartadoPostal 134, 45100 Zapopan, Jalisco, México.</mods:affiliation>
</mods:name>
<mods:name id="BD2F880C7EB30B58AD336D3BD8F9F257" type="personal">
<mods:role id="8A8894836139E27320ED29890620FDAB">
<mods:roleTerm id="9F4A492E8347DB72E2F48D89073EC319">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1095E175EE21D9E972311B93C625142E">Navarrete-Heredia, José L.</mods:namePart>
<mods:affiliation id="1FAC1075B03C56660F2B55698E98FDA0">Entomología, Centro de Estudios en Zoología, CUCBA, Universidad de Guadalajara, ApartadoPostal 134, 45100 Zapopan, Jalisco, México.</mods:affiliation>
</mods:name>
<mods:name id="332E46D82E7BFCD6E14FA8C3DFA8A43B" type="personal">
<mods:role id="ECED64177B85C7EA700A1A324D3260EC">
<mods:roleTerm id="E1B60F8444847EBCBFB5F0FA5C6EC444">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="17824404B3F21245B57198B9EBC9063D">Arriaga-Varela, Emmanuel</mods:namePart>
<mods:affiliation id="B5EF6F566445B88CE057E80AC38F8B29">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 44, Prague, Czech Republic. arriagavarelae @ gmail. com; https: // orcid. org / 0000 - 0001 - 6206 - 9212 &amp; Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Prague, Czech Republic</mods:affiliation>
</mods:name>
<mods:name id="4402E4471D9BCB7E69B8FD85621192F8" type="personal">
<mods:role id="65764541A2BF61E0B945EEB1F34ADC4E">
<mods:roleTerm id="492DFB04431D50D94D5A96DFA937011E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E184C182407EC4F162C0EDFD576996AC">Cuccodoro, Giulio</mods:namePart>
<mods:affiliation id="8E9CD32358B4E9914AA871CD4971A73F">Muséum d’histoire naturelle, CH- 1208, Genève, Switzerland.</mods:affiliation>
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<mods:title id="B21873496C684728523F3C22781A38E3">Zootaxa</mods:title>
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<mods:part id="6708FC37D656BB46CEA63D7FEA19E57D">
<mods:date id="75C9B5B31C60EEAF868A305F1B5B552F">2020</mods:date>
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<mods:number id="1760C40EB3182F390784A605AA03E791">2020-05-15</mods:number>
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<mods:number id="75E4B12E611057323BDBF96584BCF1D7">4778</mods:number>
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<mods:number id="203AEE04A145A7FE790CF0073D49AFF7">3</mods:number>
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<mods:classification id="0991187979EC1949B9E4BD6892286537">journal article</mods:classification>
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<mods:identifier id="E2B8F6E065D89F13194BBB93B3E9CAAA" type="DOI">10.11646/zootaxa.4778.3.8</mods:identifier>
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<treatment id="03E5ED0AFF9ED56F2B996A19FB60FA55" ID-DOI="http://doi.org/10.5281/zenodo.3848008" ID-GBIF-Taxon="164159773" ID-Zenodo-Dep="3848008" LSID="urn:lsid:plazi:treatment:03E5ED0AFF9ED56F2B996A19FB60FA55" httpUri="http://treatment.plazi.org/id/03E5ED0AFF9ED56F2B996A19FB60FA55" lastPageId="7" lastPageNumber="578" pageId="1" pageNumber="572">
<subSubSection id="C3560F97FF9ED5692B996A19FD98F9E1" box="[151,632,1593,1620]" pageId="1" pageNumber="572" type="nomenclature">
<paragraph id="8BF35C1CFF9ED5692B996A19FD98F9E1" blockId="1.[151,885,1593,1684]" box="[151,632,1593,1620]" pageId="1" pageNumber="572">
<heading id="D0BBEB70FF9ED5692B996A19FD98F9E1" bold="true" box="[151,632,1593,1620]" fontSize="11" level="1" pageId="1" pageNumber="572" reason="1">
<emphasis id="B938800EFF9ED5692B996A19FD98F9E1" bold="true" box="[151,632,1593,1620]" pageId="1" pageNumber="572">
<taxonomicName id="4C4C279FFF9ED5692B996A19FDFAF9E1" ID-CoL="8Y69T" authority="Cuccodoro" authorityName="Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro" authorityYear="2020" box="[151,538,1593,1620]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="572" phylum="Arthropoda" rank="species" species="chiapas">
<emphasis id="B938800EFF9ED5692B996A19FE6BF9E6" bold="true" box="[151,395,1593,1620]" italics="true" pageId="1" pageNumber="572">Megarthrus chiapas</emphasis>
Cuccodoro
</taxonomicName>
sp nov.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3560F97FF9ED5692B906A7DFC95F926" pageId="1" pageNumber="572" type="description">
<paragraph id="8BF35C1CFF9ED5692B906A7DFEA5F9C5" blockId="1.[151,885,1593,1684]" box="[158,325,1629,1655]" pageId="1" pageNumber="572">
(
<figureCitation id="13774099FF9ED5692BA86A7DFEF4F9C5" box="[166,276,1629,1655]" captionStart-0="FIGURES 1–3" captionStart-1="FIGURES 4–9" captionStart-2="FIGURES 10–18" captionStart-3="FIGURES 19–24" captionStart-4="FIGURES 25–28" captionStartId-0="2.[151,264,1989,2014]" captionStartId-1="3.[151,264,1927,1952]" captionStartId-2="4.[151,264,1907,1932]" captionStartId-3="5.[151,264,1907,1932]" captionStartId-4="6.[151,264,1907,1932]" captionTargetBox-0="[156,1430,466,1948]" captionTargetBox-1="[177,1417,207,1898]" captionTargetBox-2="[151,1436,181,1866]" captionTargetBox-3="[161,1433,210,1859]" captionTargetBox-4="[151,1436,181,1882]" captionTargetId-0="figure@2.[151,1436,450,1961]" captionTargetId-1="figure@3.[151,1436,201,1902]" captionTargetId-2="figure@4.[151,1436,181,1882]" captionTargetId-3="figure@5.[151,1436,181,1882]" captionTargetId-4="figure@6.[151,1436,181,1882]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURES 1–3. Megarthrus chiapas sp. nov., habitus, male: dorsal (1), ventral (2) and lateral (3) views. Scale bar = 0.5 mm." captionText-1="FIGURES 4–9. Megarthrus chiapas sp. nov., male: aedeagus in ventral (4) and lateral (8) views; mesofemur and mesotro- chanter (5); mesotibia (6); metatibia (7); antenna (9). Scale bars = 0.2 mm: 4–8 (left); 12–18 (right)." captionText-2="FIGURES 10–18. Megarthrus chiapas sp. nov., male: apical segments of abdomen in ventral (10) and lateral (11) views; ab- dominal segments IX and X in lateral (12) and ventral (13) views; abdominal sternite VIII (14), sternite IX (15) and hemitergite IX (18) in ventral view; abdominal tergite VIII in lateral (16) and dorsal (17) views. Scale bars = 0.2 mm: 10–11 (vertical); 12–18 (horizontal)." captionText-3="FIGURES 19–24. Megarthrus chiapas sp. nov., female: genitalia in dorsal (19), posterior (20) and lateral (21) views; abdomi- nal sternite VIII (22) in ventral view; abdominal sternite VIII in lateral (23) and ventral (23) views. Scale bar = 0.2 mm." captionText-4="FIGURES 25–28. Megarthrus chiapas sp. nov. (25, 26): prothorax in dorsal (25) and ventral (26) views, HR = hypomeral ridge. Megarthrus cavianae sp. nov., type locality (27–28): riparian remnants of mountain cloud forest in the Transmexican Volcanic Belt of Xalapa, Veracruz (27); sifted leaf litter in riparian remnants of cloud forests west of Xalapa, Veracruz (28). Scale bar = 0.2 mm: 25–26." figureDoi-0="http://doi.org/10.5281/zenodo.3828827" figureDoi-1="http://doi.org/10.5281/zenodo.3828829" figureDoi-2="http://doi.org/10.5281/zenodo.3828831" figureDoi-3="http://doi.org/10.5281/zenodo.3828833" figureDoi-4="http://doi.org/10.5281/zenodo.3828835" httpUri-0="https://zenodo.org/record/3828827/files/figure.png" httpUri-1="https://zenodo.org/record/3828829/files/figure.png" httpUri-2="https://zenodo.org/record/3828831/files/figure.png" httpUri-3="https://zenodo.org/record/3828833/files/figure.png" httpUri-4="https://zenodo.org/record/3828835/files/figure.png" pageId="1" pageNumber="572">Figs 1-26</figureCitation>
,
<figureCitation id="13774099FF9ED5692A2E6A7DFEDDF9C5" box="[288,317,1629,1655]" captionStart="FIGURE 39" captionStartId="10.[151,250,1537,1562]" captionTargetBox="[151,1436,633,1512]" captionTargetId="figure@10.[151,1436,633,1512]" captionTargetPageId="10" captionText="FIGURE 39. Distribution map of Megarthrus alatorreorum (diamond), M. altivagans (star), M. cavianae sp. nov. (square), M. chiapas sp. nov. (triangle), M. flavosignatus (pentagon) and M. zunilensis (circle)." figureDoi="http://doi.org/10.5281/zenodo.3828841" httpUri="https://zenodo.org/record/3828841/files/figure.png" pageId="1" pageNumber="572">39</figureCitation>
)
</paragraph>
<paragraph id="8BF35C1CFF9ED5692B996A5CFC95F926" blockId="1.[151,885,1593,1684]" box="[151,885,1660,1684]" pageId="1" pageNumber="572">
http://zoobank.org:act:
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</paragraph>
</subSubSection>
<subSubSection id="C3560F97FF9ED5692B996AE2FDF3F8DE" pageId="1" pageNumber="572" type="materials_examined">
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<emphasis id="B938800EFF9ED5692B996AE2FEA7F96E" bold="true" box="[151,327,1729,1756]" pageId="1" pageNumber="572">Type material.</emphasis>
(
<specimenCount id="9D4A9795FF9ED5692A546AE2FE9FF96E" box="[346,383,1730,1756]" pageId="1" pageNumber="572" type="male">7♂</specimenCount>
,
<specimenCount id="9D4A9795FF9ED5692A806AE1FE51F96E" box="[398,433,1729,1756]" pageId="1" pageNumber="572" type="female">5♀</specimenCount>
).
<materialsCitation id="3B245641FF9ED5692AC46AE2FB37F8B3" ID-GBIF-Occurrence="2620268305" collectingDate="1973-08-26" collectingDateMax="1973-08-27" collectingDateMin="1973-08-26" collectionCode="FMNH" collectorName="N Pueblo Nuevo Solistahuacan &amp; A. Newton" country="Mexico" elevation="1830" location="Chiapas" pageId="1" pageNumber="572" specimenCount="1" specimenCount-male="1" stateProvince="Chiapas" typeStatus="holotype">
<emphasis id="B938800EFF9ED5692AC46AE2FDD7F96E" bold="true" box="[458,567,1730,1756]" pageId="1" pageNumber="572">
<typeStatus id="54F7E2BEFF9ED5692AC46AE2FDD7F96E" box="[458,567,1730,1756]" pageId="1" pageNumber="572" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D4A9795FF9ED569294F6AE3FDB9F96F" box="[577,601,1731,1757]" pageId="1" pageNumber="572" type="male">♂</specimenCount>
: labeled
<collectingCountry id="F35B1C8CFF9ED56929CF6AE2FCD4F96E" box="[705,820,1730,1756]" name="Mexico" pageId="1" pageNumber="572">MEXICO</collectingCountry>
:
<collectingRegion id="498892FEFF9ED569284F6AE2FC7EF96E" box="[833,926,1730,1756]" country="Mexico" name="Chiapas" pageId="1" pageNumber="572">Chiapas</collectingRegion>
,
<quantity id="4CB4F1F9FF9ED56928A26AE2FC01F96E" box="[940,993,1730,1756]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="1" pageNumber="572" unit="mi" value="8.0">8 mi</quantity>
[=
<quantity id="4CB4F1F9FF9ED5692F036AE2FB8EF96E" box="[1037,1134,1729,1756]" metricMagnitude="4" metricUnit="m" metricValue="1.29" pageId="1" pageNumber="572" unit="km" value="12.9">12.9 km</quantity>
]
<collectorName id="26B939CAFF9ED5692F706AE3FF09F8B2" pageId="1" pageNumber="572">N Pueblo Nuevo Solistahuacán</collectorName>
, 6000’ [=
<quantity id="4CB4F1F9FF9ED5692A6B6AC6FE22F8B2" box="[357,450,1765,1792]" metricMagnitude="3" metricUnit="m" metricValue="1.83" pageId="1" pageNumber="572" unit="m" value="1830.0">
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</quantity>
],
<date id="FFF27ADCFF9ED5692AD66AC6FD72F8B2" box="[472,658,1766,1792]" pageId="1" pageNumber="572" value="1973-08-26" valueMax="1973-08-27" valueMin="1973-08-26">
<collectingDate id="EFB68334FF9ED5692AD66AC6FD72F8B2" box="[472,658,1766,1792]" pageId="1" pageNumber="572" value="1973-08-26" valueMax="1973-08-27" valueMin="1973-08-26">26-27.VIII.1973</collectingDate>
</date>
,
<collectorName id="26B939CAFF9ED56929AE6AC6FCC2F8B3" box="[672,802,1766,1793]" pageId="1" pageNumber="572">A. Newton</collectorName>
, berlese cloud forest litter, in
<collectionCode id="ED5DC4D9FF9ED5692F8C6AC7FB37F8B3" box="[1154,1239,1767,1793]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="1" pageNumber="572" type="Museum">FMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B245641FF9ED5692FEB6AC6FDEEF8DE" ID-GBIF-Occurrence="2620268304" collectingDate="1991-09-19" collectingDateMax="1973-08-27" collectingDateMin="1973-08-26" collectionCode="FMNH, CZUG, SEMC" collectorName="R. Anderson" country="Mexico" elevation="1500" location="Rayon" pageId="1" pageNumber="572" specimenCount="11" specimenCount-female="5" specimenCount-male="6" stateProvince="Chiapas" typeStatus="paratype">
<emphasis id="B938800EFF9ED5692FEB6AC6FABDF8B2" bold="true" box="[1253,1373,1766,1792]" pageId="1" pageNumber="572">
<typeStatus id="54F7E2BEFF9ED5692FEB6AC6FABDF8B2" box="[1253,1373,1766,1792]" pageId="1" pageNumber="572" type="paratype">Paratypes</typeStatus>
</emphasis>
(11): labaled with the same data as holotype,
<specimenCount id="9D4A9795FF9ED569296D6B2AFD66F896" box="[611,646,1802,1828]" pageId="1" pageNumber="572" type="male">2♂</specimenCount>
and
<specimenCount id="9D4A9795FF9ED56929CF6B2AFD04F896" box="[705,740,1802,1828]" pageId="1" pageNumber="572" type="female">1♀</specimenCount>
in
<collectionCode id="ED5DC4D9FF9ED56928056B2BFCBFF897" box="[779,863,1803,1829]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="1" pageNumber="572" type="Museum">FMNH</collectionCode>
,
<specimenCount id="9D4A9795FF9ED56928626B2AFC78F897" box="[876,920,1802,1829]" pageId="1" pageNumber="572" type="male">1 ♂</specimenCount>
&amp;
<specimenCount id="9D4A9795FF9ED56928CE6B2AFC03F896" box="[960,995,1802,1828]" pageId="1" pageNumber="572" type="female">1♀</specimenCount>
in MHNG and
<specimenCount id="9D4A9795FF9ED5692FAA6B2AFB27F896" box="[1188,1223,1802,1828]" pageId="1" pageNumber="572" type="male">1♂</specimenCount>
in
<collectionCode id="ED5DC4D9FF9ED5692FE06B2AFADEF896" box="[1262,1342,1802,1828]" country="Mexico" httpUri="http://grbio.org/cool/ntgr-f7vv" name="Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia" pageId="1" pageNumber="572">CZUG</collectionCode>
; labeled
<collectingCountry id="F35B1C8CFF9ED5692B996B0EFEEAF8FA" box="[151,266,1838,1864]" name="Mexico" pageId="1" pageNumber="572">MEXICO</collectingCountry>
:
<collectingRegion id="498892FEFF9ED5692A1B6B0EFE92F8FA" box="[277,370,1838,1864]" country="Mexico" name="Chiapas" pageId="1" pageNumber="572">Chiapas</collectingRegion>
,
<quantity id="4CB4F1F9FF9ED5692A706B0EFE2DF8FA" box="[382,461,1838,1864]" metricMagnitude="3" metricUnit="m" metricValue="8.9" pageId="1" pageNumber="572" unit="km" value="8.9">8.9 km</quantity>
E
<location id="8E930AC7FF9ED5692AE06B0EFDDAF8FA" LSID="urn:lsid:plazi:treatment:03E5ED0AFF9ED56F2B996A19FB60FA55:8E930AC7FF9ED5692AE06B0EFDDAF8FA" box="[494,570,1838,1864]" country="Mexico" name="Rayon" pageId="1" pageNumber="572" stateProvince="Chiapas">Rayón</location>
,
<quantity id="4CB4F1F9FF9ED56929486B0DFD76F8FA" box="[582,662,1837,1864]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="572" unit="m" value="1500.0">
<elevation id="0061BB2FFF9ED56929486B0DFD76F8FA" box="[582,662,1837,1864]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="572" unit="m" value="1500.0">1500m</elevation>
</quantity>
,
<date id="FFF27ADCFF9ED56929AD6B0EFCC5F8FA" box="[675,805,1838,1864]" pageId="1" pageNumber="572" value="1991-09-19">
<collectingDate id="EFB68334FF9ED56929AD6B0EFCC5F8FA" box="[675,805,1838,1864]" pageId="1" pageNumber="572" value="1991-09-19">19.IX.1991</collectingDate>
</date>
,
<collectorName id="26B939CAFF9ED569283F6B0FFC21F8FA" box="[817,961,1838,1865]" pageId="1" pageNumber="572">R. Anderson</collectorName>
, 91-109, ex cloud forest litter,
<specimenCount id="9D4A9795FF9ED5692E106B0EFAA1F8FA" box="[1310,1345,1838,1864]" pageId="1" pageNumber="572" type="male">1♂</specimenCount>
and
<specimenCount id="9D4A9795FF9ED5692E776B0EFA7CF8FA" box="[1401,1436,1838,1864]" pageId="1" pageNumber="572" type="female">2♀</specimenCount>
in
<collectionCode id="ED5DC4D9FF9ED5692BBA6B72FEE3F8DE" box="[180,259,1874,1900]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="1" pageNumber="572" type="Museum">SEMC</collectionCode>
,
<specimenCount id="9D4A9795FF9ED5692A006B72FED8F8DF" box="[270,312,1874,1901]" pageId="1" pageNumber="572" type="male">1 ♂</specimenCount>
and
<specimenCount id="9D4A9795FF9ED5692A616B72FE72F8DE" box="[367,402,1874,1900]" pageId="1" pageNumber="572" type="female">1♀</specimenCount>
in MHNG
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3560F97FF9ED56F2BC96B55FC6AFAE5" lastPageId="7" lastPageNumber="578" pageId="1" pageNumber="572" type="description">
<paragraph id="8BF35C1CFF9ED56A2BC96B55FCEBFEAD" blockId="1.[151,1437,1729,2045]" lastBlockId="2.[151,1437,153,431]" lastPageId="2" lastPageNumber="573" pageId="1" pageNumber="572">
<emphasis id="B938800EFF9ED5692BC96B55FEB9F822" bold="true" box="[199,345,1909,1936]" pageId="1" pageNumber="572">Description.</emphasis>
Habitus as in
<figureCitation id="13774099FF9ED5692AF56B56FDBFF822" box="[507,607,1910,1936]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1989,2014]" captionTargetBox="[156,1430,466,1948]" captionTargetId="figure@2.[151,1436,450,1961]" captionTargetPageId="2" captionText="FIGURES 1–3. Megarthrus chiapas sp. nov., habitus, male: dorsal (1), ventral (2) and lateral (3) views. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3828827" httpUri="https://zenodo.org/record/3828827/files/figure.png" pageId="1" pageNumber="572">Figs 1–3</figureCitation>
. Combined length of head, pronotum and elytra =
<quantity id="4CB4F1F9FF9ED5692F946B56FACEF823" box="[1178,1326,1910,1937]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="1" pageNumber="572" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3 - 1.4 mm</quantity>
; maximal pronotal width = 0.9 - 1.0 mm. Body chestnut brown, with lateral margins of pronotum and appendages slightly paler; 1, 2 or 3 yellowish apical antennomeres. Dorsal pubescence denser on head and pronotum than on elytral disc; medial frontal setae directed forward; elytral and pronotal setae almost straight, recumbent, becoming slightly denser along medial groove of pronotum and on anterior portion of elytral disc; pubescence on abdomen uniform, parallel, without macrosetae on sternites IV–VII. Frons, pronotum, humeral area of elytral and anterior portion of prohypomera granulate; frontal granulation conspicuous, with granules about as high as their diameter, or higher; elytral disc and metaventrite coarsely granulo-fossulate.
</paragraph>
<paragraph id="8BF35C1CFF9DD56A2BC96D09FE35FE1D" blockId="2.[151,1437,153,431]" pageId="2" pageNumber="573">
Frons above clypeus forming sharp ridge, the latter finely carinate, evenly; mesal portion of disc moderately convex in lateral view, evenly; anterior margin in dorsal view weakly arcuate in middle and laterally oblique; Ushaped frontal impression shallow. Temples convex, granulate. Occipital ridge indistinct. Antennae (
<figureCitation id="13774099FF9DD56A2FF46D51FADEFE39" box="[1274,1342,369,395]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1927,1952]" captionTargetBox="[177,1417,207,1898]" captionTargetId="figure@3.[151,1436,201,1902]" captionTargetPageId="3" captionText="FIGURES 4–9. Megarthrus chiapas sp. nov., male: aedeagus in ventral (4) and lateral (8) views; mesofemur and mesotro- chanter (5); mesotibia (6); metatibia (7); antenna (9). Scale bars = 0.2 mm: 4–8 (left); 12–18 (right)." figureDoi="http://doi.org/10.5281/zenodo.3828829" httpUri="https://zenodo.org/record/3828829/files/figure.png" pageId="2" pageNumber="573">Fig. 9</figureCitation>
) 1.8-2.0 times longer than pronotum.
</paragraph>
<caption id="DF330C94FF9DD56A2B996BE5FA6EF86F" ID-DOI="http://doi.org/10.5281/zenodo.3828827" ID-Zenodo-Dep="3828827" box="[151,1422,1989,2014]" httpUri="https://zenodo.org/record/3828827/files/figure.png" pageId="2" pageNumber="573" startId="2.[151,264,1989,2014]" targetBox="[156,1430,466,1948]" targetPageId="2">
<paragraph id="8BF35C1CFF9DD56A2B996BE5FA6EF86F" blockId="2.[151,1422,1989,2014]" box="[151,1422,1989,2014]" pageId="2" pageNumber="573">
<emphasis id="B938800EFF9DD56A2B996BE5FEDAF86C" bold="true" box="[151,314,1989,2014]" pageId="2" pageNumber="573">FIGURES 1–3.</emphasis>
<taxonomicName id="4C4C279FFF9DD56A2A4F6BE5FDEEF86F" authority="Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro, 2020" authorityName="Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro" authorityYear="2020" box="[321,526,1989,2013]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="573" phylum="Arthropoda" rank="species" species="chiapas" status="sp. nov.">
<emphasis id="B938800EFF9DD56A2A4F6BE5FDEEF86F" box="[321,526,1989,2013]" italics="true" pageId="2" pageNumber="573">Megarthrus chiapas</emphasis>
</taxonomicName>
<emphasis id="B938800EFF9DD56A291A6BE5FD84F86F" bold="true" box="[532,612,1989,2013]" pageId="2" pageNumber="573">
<taxonomicNameLabel id="A20B3D75FF9DD56A291A6BE5FD84F86F" box="[532,612,1989,2013]" pageId="2" pageNumber="573" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, habitus, male: dorsal (1), ventral (2) and lateral (3) views. Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="DF330C94FF9CD56B2B996BA7FB6DF876" ID-DOI="http://doi.org/10.5281/zenodo.3828829" ID-Zenodo-Dep="3828829" httpUri="https://zenodo.org/record/3828829/files/figure.png" pageId="3" pageNumber="574" startId="3.[151,264,1927,1952]" targetBox="[177,1417,207,1898]" targetPageId="3">
<paragraph id="8BF35C1CFF9CD56B2B996BA7FB6DF876" blockId="3.[151,1436,1927,1988]" pageId="3" pageNumber="574">
<emphasis id="B938800EFF9CD56B2B996BA7FEDDF812" bold="true" box="[151,317,1927,1952]" pageId="3" pageNumber="574">FIGURES 4–9.</emphasis>
<taxonomicName id="4C4C279FFF9CD56B2A4B6BA7FDF4F82D" authority="Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro, 2020" authorityName="Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro" authorityYear="2020" box="[325,532,1927,1951]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="574" phylum="Arthropoda" rank="species" species="chiapas" status="sp. nov.">
<emphasis id="B938800EFF9CD56B2A4B6BA7FDF4F82D" box="[325,532,1927,1951]" italics="true" pageId="3" pageNumber="574">Megarthrus chiapas</emphasis>
</taxonomicName>
<emphasis id="B938800EFF9CD56B29126BA7FD8FF82D" bold="true" box="[540,623,1927,1951]" pageId="3" pageNumber="574">
<taxonomicNameLabel id="A20B3D75FF9CD56B29126BA7FD8FF82D" box="[540,623,1927,1951]" pageId="3" pageNumber="574" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male: aedeagus in ventral (4) and lateral (8) views; mesofemur and mesotro- chanter (5); mesotibia (6); metatibia (7); antenna (9). Scale bars = 0.2 mm: 4–8 (left); 12–18 (right).
</paragraph>
</caption>
<caption id="DF330C94FF9BD56C2B996B53FEBCF84A" ID-DOI="http://doi.org/10.5281/zenodo.3828831" ID-Zenodo-Dep="3828831" httpUri="https://zenodo.org/record/3828831/files/figure.png" pageId="4" pageNumber="575" startId="4.[151,264,1907,1932]" targetBox="[151,1436,181,1866]" targetPageId="4">
<paragraph id="8BF35C1CFF9BD56C2B996B53FEBCF84A" blockId="4.[151,1436,1907,2040]" pageId="4" pageNumber="575">
<emphasis id="B938800EFF9BD56C2B996B53FEB5F83E" bold="true" box="[151,341,1907,1932]" pageId="4" pageNumber="575">FIGURES 10–18.</emphasis>
<taxonomicName id="4C4C279FFF9BD56C2A526B54FDC9F83E" authority="Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro, 2020" authorityName="Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro" authorityYear="2020" box="[348,553,1908,1932]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="575" phylum="Arthropoda" rank="species" species="chiapas" status="sp. nov.">
<emphasis id="B938800EFF9BD56C2A526B54FDC9F83E" box="[348,553,1908,1932]" italics="true" pageId="4" pageNumber="575">Megarthrus chiapas</emphasis>
</taxonomicName>
<emphasis id="B938800EFF9BD56C293E6B54FD61F83E" bold="true" box="[560,641,1908,1932]" pageId="4" pageNumber="575">
<taxonomicNameLabel id="A20B3D75FF9BD56C293E6B54FD61F83E" box="[560,641,1908,1932]" pageId="4" pageNumber="575" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male: apical segments of abdomen in ventral (10) and lateral (11) views; ab- dominal segments IX and X in lateral (12) and ventral (13) views; abdominal sternite VIII (14), sternite IX (15) and hemitergite IX (18) in ventral view; abdominal tergite VIII in lateral (16) and dorsal (17) views. Scale bars = 0.2 mm: 10–11 (vertical); 12–18 (horizontal).
</paragraph>
</caption>
<caption id="DF330C94FF9AD56D2B996B53FAB6F81D" ID-DOI="http://doi.org/10.5281/zenodo.3828833" ID-Zenodo-Dep="3828833" httpUri="https://zenodo.org/record/3828833/files/figure.png" pageId="5" pageNumber="576" startId="5.[151,264,1907,1932]" targetBox="[161,1433,210,1859]" targetPageId="5">
<paragraph id="8BF35C1CFF9AD56D2B996B53FAB6F81D" blockId="5.[151,1436,1907,1968]" pageId="5" pageNumber="576">
<emphasis id="B938800EFF9AD56D2B996B53FEB4F83E" bold="true" box="[151,340,1907,1932]" pageId="5" pageNumber="576">FIGURES 19–24.</emphasis>
<taxonomicName id="4C4C279FFF9AD56D2A546B54FDC7F83E" authority="Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro, 2020" authorityName="Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro" authorityYear="2020" box="[346,551,1908,1932]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="chiapas" status="sp. nov.">
<emphasis id="B938800EFF9AD56D2A546B54FDC7F83E" box="[346,551,1908,1932]" italics="true" pageId="5" pageNumber="576">Megarthrus chiapas</emphasis>
</taxonomicName>
<emphasis id="B938800EFF9AD56D29236B54FD9DF83E" bold="true" box="[557,637,1908,1932]" pageId="5" pageNumber="576">
<taxonomicNameLabel id="A20B3D75FF9AD56D29236B54FD9DF83E" box="[557,637,1908,1932]" pageId="5" pageNumber="576" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: genitalia in dorsal (19), posterior (20) and lateral (21) views; abdomi- nal sternite VIII (22) in ventral view; abdominal sternite VIII in lateral (23) and ventral (23) views. Scale bar = 0.2 mm.
</paragraph>
</caption>
<caption id="DF330C94FF99D56E2B996B53FE51F84A" ID-DOI="http://doi.org/10.5281/zenodo.3828835" ID-Zenodo-Dep="3828835" httpUri="https://zenodo.org/record/3828835/files/figure.png" pageId="6" pageNumber="577" startId="6.[151,264,1907,1932]" targetBox="[151,1436,181,1882]" targetPageId="6">
<paragraph id="8BF35C1CFF99D56E2B996B53FE51F84A" blockId="6.[151,1437,1907,2040]" pageId="6" pageNumber="577">
<emphasis id="B938800EFF99D56E2B996B53FEB6F83E" bold="true" box="[151,342,1907,1932]" pageId="6" pageNumber="577">FIGURES 25–28.</emphasis>
<taxonomicName id="4C4C279FFF99D56E2A516B54FDCEF83E" authority="Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro, 2020" authorityName="Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro" authorityYear="2020" box="[351,558,1908,1932]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="chiapas" status="sp. nov.">
<emphasis id="B938800EFF99D56E2A516B54FDCEF83E" box="[351,558,1908,1932]" italics="true" pageId="6" pageNumber="577">Megarthrus chiapas</emphasis>
</taxonomicName>
<emphasis id="B938800EFF99D56E29386B54FD69F83E" bold="true" box="[566,649,1908,1932]" pageId="6" pageNumber="577">
<taxonomicNameLabel id="A20B3D75FF99D56E29386B54FD69F83E" box="[566,649,1908,1932]" pageId="6" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(25, 26): prothorax in dorsal (25) and ventral (26) views, HR = hypomeral ridge.
<taxonomicName id="4C4C279FFF99D56E2BD46BB8FE55F802" authority="Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro, 2020" authorityName="Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro" authorityYear="2020" box="[218,437,1944,1968]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="cavianae" status="sp. nov.">
<emphasis id="B938800EFF99D56E2BD46BB8FE55F802" box="[218,437,1944,1968]" italics="true" pageId="6" pageNumber="577">Megarthrus cavianae</emphasis>
</taxonomicName>
<emphasis id="B938800EFF99D56E2AB36BB8FDF7F802" bold="true" box="[445,535,1944,1968]" pageId="6" pageNumber="577">
<taxonomicNameLabel id="A20B3D75FF99D56E2AB36BB8FDF2F802" box="[445,530,1944,1968]" pageId="6" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
type locality (27–28): riparian remnants of mountain cloud forest in the Transmexican Volcanic Belt of Xalapa, Veracruz (27); sifted leaf litter in riparian remnants of cloud forests west of Xalapa, Veracruz (28). Scale bar = 0.2 mm: 25–26.
</paragraph>
</caption>
<paragraph id="8BF35C1CFF98D56F2BC96CB9FAF0FE39" blockId="7.[151,1437,152,1511]" pageId="7" pageNumber="578">
Pronotum (
<figureCitation id="13774099FF98D56F2A4D6CB9FE5EFF01" box="[323,446,152,179]" captionStart="FIGURES 25–28" captionStartId="6.[151,264,1907,1932]" captionTargetBox="[151,1436,181,1882]" captionTargetId="figure@6.[151,1436,181,1882]" captionTargetPageId="6" captionText="FIGURES 25–28. Megarthrus chiapas sp. nov. (25, 26): prothorax in dorsal (25) and ventral (26) views, HR = hypomeral ridge. Megarthrus cavianae sp. nov., type locality (27–28): riparian remnants of mountain cloud forest in the Transmexican Volcanic Belt of Xalapa, Veracruz (27); sifted leaf litter in riparian remnants of cloud forests west of Xalapa, Veracruz (28). Scale bar = 0.2 mm: 25–26." figureDoi="http://doi.org/10.5281/zenodo.3828835" httpUri="https://zenodo.org/record/3828835/files/figure.png" pageId="7" pageNumber="578">Figs 25-26</figureCitation>
) with center strongly convex in frontal view; disc deeply depressed near middle of lateral edges, shallowly depressed along anterior and posterior margins; medial groove deep, parallel-sided, slightly arcu- ate in lateral view; hypomera granulate, with ridge extended from anterior margin to laterobasal angle (
<figureCitation id="13774099FF98D56F2E376CC1FA70FF4E" box="[1337,1424,225,252]" captionStart="FIGURES 25–28" captionStartId="6.[151,264,1907,1932]" captionTargetBox="[151,1436,181,1882]" captionTargetId="figure@6.[151,1436,181,1882]" captionTargetPageId="6" captionText="FIGURES 25–28. Megarthrus chiapas sp. nov. (25, 26): prothorax in dorsal (25) and ventral (26) views, HR = hypomeral ridge. Megarthrus cavianae sp. nov., type locality (27–28): riparian remnants of mountain cloud forest in the Transmexican Volcanic Belt of Xalapa, Veracruz (27); sifted leaf litter in riparian remnants of cloud forests west of Xalapa, Veracruz (28). Scale bar = 0.2 mm: 25–26." figureDoi="http://doi.org/10.5281/zenodo.3828835" httpUri="https://zenodo.org/record/3828835/files/figure.png" pageId="7" pageNumber="578">Fig. 26</figureCitation>
). Proventrite (
<figureCitation id="13774099FF98D56F2A2C6D25FE93FE92" box="[290,371,261,288]" captionStart="FIGURES 25–28" captionStartId="6.[151,264,1907,1932]" captionTargetBox="[151,1436,181,1882]" captionTargetId="figure@6.[151,1436,181,1882]" captionTargetPageId="6" captionText="FIGURES 25–28. Megarthrus chiapas sp. nov. (25, 26): prothorax in dorsal (25) and ventral (26) views, HR = hypomeral ridge. Megarthrus cavianae sp. nov., type locality (27–28): riparian remnants of mountain cloud forest in the Transmexican Volcanic Belt of Xalapa, Veracruz (27); sifted leaf litter in riparian remnants of cloud forests west of Xalapa, Veracruz (28). Scale bar = 0.2 mm: 25–26." figureDoi="http://doi.org/10.5281/zenodo.3828835" httpUri="https://zenodo.org/record/3828835/files/figure.png" pageId="7" pageNumber="578">Fig. 26</figureCitation>
) lacking medial ridge. Scutellum with anterior margin rounded; posterior margin laterally mod- erately arcuate toward strongly arcuate apex. Elytra gradually widened posteriorly (
<figureCitation id="13774099FF98D56F2F386D09FB96FEF1" box="[1078,1142,297,323]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1989,2014]" captionTargetBox="[156,1430,466,1948]" captionTargetId="figure@2.[151,1436,450,1961]" captionTargetPageId="2" captionText="FIGURES 1–3. Megarthrus chiapas sp. nov., habitus, male: dorsal (1), ventral (2) and lateral (3) views. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3828827" httpUri="https://zenodo.org/record/3828827/files/figure.png" pageId="7" pageNumber="578">Fig. 1</figureCitation>
); disc with humeral callus strongly convex, two moderate swellings along anterior and posterior portions of suture, and moderately depressed posteriorly along lateral edge; the latter finely carinate, not denticulate, slightly arcuate in dorsal view.
</paragraph>
<paragraph id="8BF35C1CFF98D56F2BC96DB5FAEEFD35" blockId="7.[151,1437,152,1511]" pageId="7" pageNumber="578">
Male: Frontoclypeal area not modified. Protarsomere 1 lacking ventral patch of modified adhesive setae. Mesofemora (
<figureCitation id="13774099FF98D56F2A1F6D99FEB5FE61" box="[273,341,440,467]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1927,1952]" captionTargetBox="[177,1417,207,1898]" captionTargetId="figure@3.[151,1436,201,1902]" captionTargetPageId="3" captionText="FIGURES 4–9. Megarthrus chiapas sp. nov., male: aedeagus in ventral (4) and lateral (8) views; mesofemur and mesotro- chanter (5); mesotibia (6); metatibia (7); antenna (9). Scale bars = 0.2 mm: 4–8 (left); 12–18 (right)." figureDoi="http://doi.org/10.5281/zenodo.3828829" httpUri="https://zenodo.org/record/3828829/files/figure.png" pageId="7" pageNumber="578">Fig. 5</figureCitation>
) as long as metafemora. Mesotibiae (
<figureCitation id="13774099FF98D56F28066D99FCADFE66" box="[776,845,441,468]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1927,1952]" captionTargetBox="[177,1417,207,1898]" captionTargetId="figure@3.[151,1436,201,1902]" captionTargetPageId="3" captionText="FIGURES 4–9. Megarthrus chiapas sp. nov., male: aedeagus in ventral (4) and lateral (8) views; mesofemur and mesotro- chanter (5); mesotibia (6); metatibia (7); antenna (9). Scale bars = 0.2 mm: 4–8 (left); 12–18 (right)." figureDoi="http://doi.org/10.5281/zenodo.3828829" httpUri="https://zenodo.org/record/3828829/files/figure.png" pageId="7" pageNumber="578">Fig. 6</figureCitation>
) shorter than metatibiae (
<figureCitation id="13774099FF98D56F2F7B6D99FB5CFE61" box="[1141,1212,441,467]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1927,1952]" captionTargetBox="[177,1417,207,1898]" captionTargetId="figure@3.[151,1436,201,1902]" captionTargetPageId="3" captionText="FIGURES 4–9. Megarthrus chiapas sp. nov., male: aedeagus in ventral (4) and lateral (8) views; mesofemur and mesotro- chanter (5); mesotibia (6); metatibia (7); antenna (9). Scale bars = 0.2 mm: 4–8 (left); 12–18 (right)." figureDoi="http://doi.org/10.5281/zenodo.3828829" httpUri="https://zenodo.org/record/3828829/files/figure.png" pageId="7" pageNumber="578">Fig. 7</figureCitation>
), the latter straight. Metatarsomeres 1 about 1.3 times as long as combined length of metatarsomeres 2-4. Peg-like setae present only on mesotrochanters (
<figureCitation id="13774099FF98D56F2A506E21FE41FDA9" box="[350,417,512,539]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1927,1952]" captionTargetBox="[177,1417,207,1898]" captionTargetId="figure@3.[151,1436,201,1902]" captionTargetPageId="3" captionText="FIGURES 4–9. Megarthrus chiapas sp. nov., male: aedeagus in ventral (4) and lateral (8) views; mesofemur and mesotro- chanter (5); mesotibia (6); metatibia (7); antenna (9). Scale bars = 0.2 mm: 4–8 (left); 12–18 (right)." figureDoi="http://doi.org/10.5281/zenodo.3828829" httpUri="https://zenodo.org/record/3828829/files/figure.png" pageId="7" pageNumber="578">Fig. 5</figureCitation>
; arranged in a row), on distal two thirds of mesotibiae (
<figureCitation id="13774099FF98D56F2F046E21FBAEFDA9" box="[1034,1102,513,539]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1927,1952]" captionTargetBox="[177,1417,207,1898]" captionTargetId="figure@3.[151,1436,201,1902]" captionTargetPageId="3" captionText="FIGURES 4–9. Megarthrus chiapas sp. nov., male: aedeagus in ventral (4) and lateral (8) views; mesofemur and mesotro- chanter (5); mesotibia (6); metatibia (7); antenna (9). Scale bars = 0.2 mm: 4–8 (left); 12–18 (right)." figureDoi="http://doi.org/10.5281/zenodo.3828829" httpUri="https://zenodo.org/record/3828829/files/figure.png" pageId="7" pageNumber="578">Fig. 6</figureCitation>
; arranged in 1-2 rows) and on distal half of metatibiae (
<figureCitation id="13774099FF98D56F2AA56E05FE0FFD8D" box="[427,495,549,575]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1927,1952]" captionTargetBox="[177,1417,207,1898]" captionTargetId="figure@3.[151,1436,201,1902]" captionTargetPageId="3" captionText="FIGURES 4–9. Megarthrus chiapas sp. nov., male: aedeagus in ventral (4) and lateral (8) views; mesofemur and mesotro- chanter (5); mesotibia (6); metatibia (7); antenna (9). Scale bars = 0.2 mm: 4–8 (left); 12–18 (right)." figureDoi="http://doi.org/10.5281/zenodo.3828829" httpUri="https://zenodo.org/record/3828829/files/figure.png" pageId="7" pageNumber="578">Fig. 7</figureCitation>
; grouped in a field). Apex of abdomen as in
<figureCitation id="13774099FF98D56F28D96E05FBB1FD8D" box="[983,1105,549,575]" captionStart="FIGURES 10–18" captionStartId="4.[151,264,1907,1932]" captionTargetBox="[151,1436,181,1866]" captionTargetId="figure@4.[151,1436,181,1882]" captionTargetPageId="4" captionText="FIGURES 10–18. Megarthrus chiapas sp. nov., male: apical segments of abdomen in ventral (10) and lateral (11) views; ab- dominal segments IX and X in lateral (12) and ventral (13) views; abdominal sternite VIII (14), sternite IX (15) and hemitergite IX (18) in ventral view; abdominal tergite VIII in lateral (16) and dorsal (17) views. Scale bars = 0.2 mm: 10–11 (vertical); 12–18 (horizontal)." figureDoi="http://doi.org/10.5281/zenodo.3828831" httpUri="https://zenodo.org/record/3828831/files/figure.png" pageId="7" pageNumber="578">Figs 10-11</figureCitation>
; tergite VIII as in
<figureCitation id="13774099FF98D56F2E196E05FA77FD8D" box="[1303,1431,549,575]" captionStart="FIGURES 10–18" captionStartId="4.[151,264,1907,1932]" captionTargetBox="[151,1436,181,1866]" captionTargetId="figure@4.[151,1436,181,1882]" captionTargetPageId="4" captionText="FIGURES 10–18. Megarthrus chiapas sp. nov., male: apical segments of abdomen in ventral (10) and lateral (11) views; ab- dominal segments IX and X in lateral (12) and ventral (13) views; abdominal sternite VIII (14), sternite IX (15) and hemitergite IX (18) in ventral view; abdominal tergite VIII in lateral (16) and dorsal (17) views. Scale bars = 0.2 mm: 10–11 (vertical); 12–18 (horizontal)." figureDoi="http://doi.org/10.5281/zenodo.3828831" httpUri="https://zenodo.org/record/3828831/files/figure.png" pageId="7" pageNumber="578">Figs. 16-17</figureCitation>
, with apex mucronate in dorsal view; sternite VIII as in
<figureCitation id="13774099FF98D56F280B6E69FCB8FDD1" box="[773,856,585,611]" captionStart="FIGURES 10–18" captionStartId="4.[151,264,1907,1932]" captionTargetBox="[151,1436,181,1866]" captionTargetId="figure@4.[151,1436,181,1882]" captionTargetPageId="4" captionText="FIGURES 10–18. Megarthrus chiapas sp. nov., male: apical segments of abdomen in ventral (10) and lateral (11) views; ab- dominal segments IX and X in lateral (12) and ventral (13) views; abdominal sternite VIII (14), sternite IX (15) and hemitergite IX (18) in ventral view; abdominal tergite VIII in lateral (16) and dorsal (17) views. Scale bars = 0.2 mm: 10–11 (vertical); 12–18 (horizontal)." figureDoi="http://doi.org/10.5281/zenodo.3828831" httpUri="https://zenodo.org/record/3828831/files/figure.png" pageId="7" pageNumber="578">Fig. 14</figureCitation>
; segments IX-X as in
<figureCitation id="13774099FF98D56F2F5C6E69FB2DFDD1" box="[1106,1229,585,611]" captionStart="FIGURES 10–18" captionStartId="4.[151,264,1907,1932]" captionTargetBox="[151,1436,181,1866]" captionTargetId="figure@4.[151,1436,181,1882]" captionTargetPageId="4" captionText="FIGURES 10–18. Megarthrus chiapas sp. nov., male: apical segments of abdomen in ventral (10) and lateral (11) views; ab- dominal segments IX and X in lateral (12) and ventral (13) views; abdominal sternite VIII (14), sternite IX (15) and hemitergite IX (18) in ventral view; abdominal tergite VIII in lateral (16) and dorsal (17) views. Scale bars = 0.2 mm: 10–11 (vertical); 12–18 (horizontal)." figureDoi="http://doi.org/10.5281/zenodo.3828831" httpUri="https://zenodo.org/record/3828831/files/figure.png" pageId="7" pageNumber="578">Figs 12-13</figureCitation>
, with hemitergites IX as in
<figureCitation id="13774099FF98D56F2BF96E4DFEA6FD35" box="[247,326,621,647]" captionStart="FIGURES 10–18" captionStartId="4.[151,264,1907,1932]" captionTargetBox="[151,1436,181,1866]" captionTargetId="figure@4.[151,1436,181,1882]" captionTargetPageId="4" captionText="FIGURES 10–18. Megarthrus chiapas sp. nov., male: apical segments of abdomen in ventral (10) and lateral (11) views; ab- dominal segments IX and X in lateral (12) and ventral (13) views; abdominal sternite VIII (14), sternite IX (15) and hemitergite IX (18) in ventral view; abdominal tergite VIII in lateral (16) and dorsal (17) views. Scale bars = 0.2 mm: 10–11 (vertical); 12–18 (horizontal)." figureDoi="http://doi.org/10.5281/zenodo.3828831" httpUri="https://zenodo.org/record/3828831/files/figure.png" pageId="7" pageNumber="578">Fig. 18</figureCitation>
and sternite IX as in
<figureCitation id="13774099FF98D56F29386E4DFD61FD35" box="[566,641,620,647]" captionStart="FIGURES 10–18" captionStartId="4.[151,264,1907,1932]" captionTargetBox="[151,1436,181,1866]" captionTargetId="figure@4.[151,1436,181,1882]" captionTargetPageId="4" captionText="FIGURES 10–18. Megarthrus chiapas sp. nov., male: apical segments of abdomen in ventral (10) and lateral (11) views; ab- dominal segments IX and X in lateral (12) and ventral (13) views; abdominal sternite VIII (14), sternite IX (15) and hemitergite IX (18) in ventral view; abdominal tergite VIII in lateral (16) and dorsal (17) views. Scale bars = 0.2 mm: 10–11 (vertical); 12–18 (horizontal)." figureDoi="http://doi.org/10.5281/zenodo.3828831" httpUri="https://zenodo.org/record/3828831/files/figure.png" pageId="7" pageNumber="578">Fig 15</figureCitation>
. Aedeagus as in
<figureCitation id="13774099FF98D56F28356E4DFC26FD35" box="[827,966,621,647]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1927,1952]" captionTargetBox="[177,1417,207,1898]" captionTargetId="figure@3.[151,1436,201,1902]" captionTargetPageId="3" captionText="FIGURES 4–9. Megarthrus chiapas sp. nov., male: aedeagus in ventral (4) and lateral (8) views; mesofemur and mesotro- chanter (5); mesotibia (6); metatibia (7); antenna (9). Scale bars = 0.2 mm: 4–8 (left); 12–18 (right)." figureDoi="http://doi.org/10.5281/zenodo.3828829" httpUri="https://zenodo.org/record/3828829/files/figure.png" pageId="7" pageNumber="578">Figs 4 and 8</figureCitation>
, with dorsal valve triangular.
</paragraph>
<paragraph id="8BF35C1CFF98D56F2BC96EB1FEA8FD41" blockId="7.[151,1437,152,1511]" pageId="7" pageNumber="578">
<emphasis id="B938800EFF98D56F2BC96EB1FEFBFD19" box="[199,283,657,683]" italics="true" pageId="7" pageNumber="578">Female</emphasis>
: Abdominal tergite VIII (
<figureCitation id="13774099FF98D56F29476EB1FD28FD19" box="[585,712,657,683]" captionStart="FIGURES 19–24" captionStartId="5.[151,264,1907,1932]" captionTargetBox="[161,1433,210,1859]" captionTargetId="figure@5.[151,1436,181,1882]" captionTargetPageId="5" captionText="FIGURES 19–24. Megarthrus chiapas sp. nov., female: genitalia in dorsal (19), posterior (20) and lateral (21) views; abdomi- nal sternite VIII (22) in ventral view; abdominal sternite VIII in lateral (23) and ventral (23) views. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3828833" httpUri="https://zenodo.org/record/3828833/files/figure.png" pageId="7" pageNumber="578">Figs 23-24</figureCitation>
) forming a medioapical projection, with dorsal outline evenly convex in lateral view. Sternite VIII as in
<figureCitation id="13774099FF98D56F29606E95FD21FD7D" box="[622,705,693,719]" captionStart="FIGURES 19–24" captionStartId="5.[151,264,1907,1932]" captionTargetBox="[161,1433,210,1859]" captionTargetId="figure@5.[151,1436,181,1882]" captionTargetPageId="5" captionText="FIGURES 19–24. Megarthrus chiapas sp. nov., female: genitalia in dorsal (19), posterior (20) and lateral (21) views; abdomi- nal sternite VIII (22) in ventral view; abdominal sternite VIII in lateral (23) and ventral (23) views. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3828833" httpUri="https://zenodo.org/record/3828833/files/figure.png" pageId="7" pageNumber="578">Fig. 22</figureCitation>
. Genital segments as in
<figureCitation id="13774099FF98D56F28DC6E95FBAEFD7D" box="[978,1102,693,719]" captionStart="FIGURES 19–24" captionStartId="5.[151,264,1907,1932]" captionTargetBox="[161,1433,210,1859]" captionTargetId="figure@5.[151,1436,181,1882]" captionTargetPageId="5" captionText="FIGURES 19–24. Megarthrus chiapas sp. nov., female: genitalia in dorsal (19), posterior (20) and lateral (21) views; abdomi- nal sternite VIII (22) in ventral view; abdominal sternite VIII in lateral (23) and ventral (23) views. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3828833" httpUri="https://zenodo.org/record/3828833/files/figure.png" pageId="7" pageNumber="578">Figs 19-21</figureCitation>
; gonocoxal plate without me- diodorsal ridge.
</paragraph>
<paragraph id="8BF35C1CFF98D56F2BC96EDCFA6FFB11" blockId="7.[151,1437,152,1511]" pageId="7" pageNumber="578">
<emphasis id="B938800EFF98D56F2BC96EDCFD88FCA5" bold="true" box="[199,616,764,791]" pageId="7" pageNumber="578">Comparisons and diagnostic notes.</emphasis>
<taxonomicName id="4C4C279FFF98D56F29616EDDFC34FCA5" authority="Cuccodoro" authorityName="Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro" authorityYear="2020" box="[623,980,765,791]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="chiapas" status="sp. nov.">
<emphasis id="B938800EFF98D56F29616EDDFCB2FCA5" box="[623,850,765,791]" italics="true" pageId="7" pageNumber="578">Megarthrus chiapas</emphasis>
Cuccodoro
</taxonomicName>
<emphasis id="B938800EFF98D56F28D56EDDFBD5FCA5" bold="true" box="[987,1077,765,791]" pageId="7" pageNumber="578">
<taxonomicNameLabel id="A20B3D75FF98D56F28D56EDDFBD5FCA5" box="[987,1077,765,791]" pageId="7" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is easily distinguished from the Mexican
<taxonomicName id="4C4C279FFF98D56F2BF16F02FE79FC89" authorityName="Bernhauer" authorityYear="1929" box="[255,409,801,827]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="altivagans">
<emphasis id="B938800EFF98D56F2BF16F02FE79FC89" box="[255,409,801,827]" italics="true" pageId="7" pageNumber="578">M. altivagans</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4C279FFF98D56F2AC36F02FD68FC89" authorityName="Rodriguez &amp; Navarrete-Heredia" authorityYear="2015" box="[461,648,801,827]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="alatorreorum">
<emphasis id="B938800EFF98D56F2AC36F02FD68FC89" box="[461,648,801,827]" italics="true" pageId="7" pageNumber="578">M. alatorreorum</emphasis>
</taxonomicName>
by the presence of a prohypomeral ridge (
<figureCitation id="13774099FF98D56F2F526F01FB4CFC8E" box="[1116,1196,801,828]" captionStart="FIGURES 25–28" captionStartId="6.[151,264,1907,1932]" captionTargetBox="[151,1436,181,1882]" captionTargetId="figure@6.[151,1436,181,1882]" captionTargetPageId="6" captionText="FIGURES 25–28. Megarthrus chiapas sp. nov. (25, 26): prothorax in dorsal (25) and ventral (26) views, HR = hypomeral ridge. Megarthrus cavianae sp. nov., type locality (27–28): riparian remnants of mountain cloud forest in the Transmexican Volcanic Belt of Xalapa, Veracruz (27); sifted leaf litter in riparian remnants of cloud forests west of Xalapa, Veracruz (28). Scale bar = 0.2 mm: 25–26." figureDoi="http://doi.org/10.5281/zenodo.3828835" httpUri="https://zenodo.org/record/3828835/files/figure.png" pageId="7" pageNumber="578">Fig. 26</figureCitation>
) and the medial frontal setae directed forward. It shares these features with
<taxonomicName id="4C4C279FFF98D56F280E6F66FBBBFCED" authority="Bierig (1940)" authorityName="Bierig" authorityYear="1940" box="[768,1115,837,863]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="flavosignatus">
<emphasis id="B938800EFF98D56F280E6F66FC5AFCED" box="[768,954,837,863]" italics="true" pageId="7" pageNumber="578">M. flavosignatus</emphasis>
<bibRefCitation id="EFDD21EDFF98D56F28CF6F65FBBBFCED" author="Bierig, A." box="[961,1115,837,863]" pageId="7" pageNumber="578" pagination="373 - 380" refId="ref4326" refString="Bierig, A. (1940) Proteinini (Col. Staph.) Costaricenses. Revista de Entomologia, 11, 373 - 380." type="journal article" year="1940">Bierig (1940)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C4C279FFF98D56F2F696F66FA77FCED" authority="Sharp (1887)" authorityName="Sharp" authorityYear="1887" box="[1127,1431,837,863]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="zunilensis">
<emphasis id="B938800EFF98D56F2F696F66FB1BFCED" box="[1127,1275,837,863]" italics="true" pageId="7" pageNumber="578">M. zunilensis</emphasis>
<bibRefCitation id="EFDD21EDFF98D56F2E0C6F65FA77FCED" author="Sharp, D." box="[1282,1431,837,863]" pageId="7" pageNumber="578" pagination="673 - 824" refId="ref4996" refString="Sharp, D. (1887) Staphylinidae. In: Biologia Centrali-Americana. Insecta, Coleoptera. 1 (2). Taylor &amp; Francis, London, pp. 673 - 824." type="book chapter" year="1887">Sharp (1887)</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C4C279FFF98D56F2BCB6F4AFC8BFC31" authority="Rodriguez, Navarrete-Heredia &amp; Arriaga-Varela" authorityName="Rodriguez, Navarrete-Heredia &amp; Arriaga-Varela" authorityYear="2020" box="[197,875,873,899]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="cavianae" status="sp. nov.">
<emphasis id="B938800EFF98D56F2BCB6F4AFEAEFC31" box="[197,334,874,899]" italics="true" pageId="7" pageNumber="578">M. cavianae</emphasis>
Rodríguez, Navarrete-Heredia &amp; Arriaga-Varela
</taxonomicName>
<emphasis id="B938800EFF98D56F287F6F49FC29FC31" bold="true" box="[881,969,873,899]" pageId="7" pageNumber="578">
<taxonomicNameLabel id="A20B3D75FF98D56F287F6F49FC29FC31" box="[881,969,873,899]" pageId="7" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, which are also the only other members of the genus to have the posterior margin of the male abdominal sternite VIII forming two projecting pointed processes (
<figureCitation id="13774099FF98D56F2B916F91FE8EFC79" box="[159,366,945,971]" captionStart="FIGURES 10–18" captionStartId="4.[151,264,1907,1932]" captionTargetBox="[151,1436,181,1866]" captionTargetId="figure@4.[151,1436,181,1882]" captionTargetPageId="4" captionText="FIGURES 10–18. Megarthrus chiapas sp. nov., male: apical segments of abdomen in ventral (10) and lateral (11) views; ab- dominal segments IX and X in lateral (12) and ventral (13) views; abdominal sternite VIII (14), sternite IX (15) and hemitergite IX (18) in ventral view; abdominal tergite VIII in lateral (16) and dorsal (17) views. Scale bars = 0.2 mm: 10–11 (vertical); 12–18 (horizontal)." figureDoi="http://doi.org/10.5281/zenodo.3828831" httpUri="https://zenodo.org/record/3828831/files/figure.png" pageId="7" pageNumber="578">Figs 10, 11 and 14</figureCitation>
). Within these four species, the males of
<taxonomicName id="4C4C279FFF98D56F283F6F92FC0BFC79" authorityName="Bierig" authorityYear="1940" box="[817,1003,945,971]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="flavosignatus">
<emphasis id="B938800EFF98D56F283F6F92FC0BFC79" box="[817,1003,945,971]" italics="true" pageId="7" pageNumber="578">M. flavosignatus</emphasis>
</taxonomicName>
can be easily distinguished by the peglike setae on the mesotibiae arranged in a row (
<bibRefCitation id="EFDD21EDFF98D56F29A26FF5FC8BFC5D" author="Cuccodoro, G." box="[684,875,981,1007]" pageId="7" pageNumber="578" pagination="107 - 147" refId="ref4399" refString="Cuccodoro, G. (2011) Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae). Revue Suisse de Zoologie, 118 (1), 107 - 147. https: // doi. org / 10.5962 / bhl. part. 117802" type="journal article" year="2011">Cuccodoro 2011</bibRefCitation>
: Fig. 60) instead of grouped in a field (
<figureCitation id="13774099FF98D56F2E2B6FF5FA8CFC5D" box="[1317,1388,981,1007]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1927,1952]" captionTargetBox="[177,1417,207,1898]" captionTargetId="figure@3.[151,1436,201,1902]" captionTargetPageId="3" captionText="FIGURES 4–9. Megarthrus chiapas sp. nov., male: aedeagus in ventral (4) and lateral (8) views; mesofemur and mesotro- chanter (5); mesotibia (6); metatibia (7); antenna (9). Scale bars = 0.2 mm: 4–8 (left); 12–18 (right)." figureDoi="http://doi.org/10.5281/zenodo.3828829" httpUri="https://zenodo.org/record/3828829/files/figure.png" pageId="7" pageNumber="578">Figs 6</figureCitation>
and
<figureCitation id="13774099FF98D56F2B996FD9FF56FBA1" box="[151,182,1017,1043]" captionStart="FIGURES 32–38" captionStartId="9.[151,264,1914,1939]" captionTargetBox="[163,1421,187,1881]" captionTargetId="figure@9.[151,1436,181,1889]" captionTargetPageId="9" captionText="FIGURES 32–38. Megarthrus cavianae sp. nov., male: aedeagus in ventral (32) and lateral (37) views; mesofemur and mes- otrochanter (33); mesotibia (34); metatibia (35); antenna (36); abdominal tergite VIII in dorsal view (38). Scale bars = 0.2 mm: 32–35, 37, 38 (horizontal); 36 (vertical)." figureDoi="http://doi.org/10.5281/zenodo.3828839" httpUri="https://zenodo.org/record/3828839/files/figure.png" pageId="7" pageNumber="578">34</figureCitation>
;
<bibRefCitation id="EFDD21EDFF98D56F2BCD6FD9FE60FBA1" author="Cuccodoro, G." box="[195,384,1017,1043]" pageId="7" pageNumber="578" pagination="107 - 147" refId="ref4399" refString="Cuccodoro, G. (2011) Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae). Revue Suisse de Zoologie, 118 (1), 107 - 147. https: // doi. org / 10.5962 / bhl. part. 117802" type="journal article" year="2011">Cuccodoro 2011</bibRefCitation>
: Fig. 174), and by the metatibiae bearing peg-like setae only in distal third (
<bibRefCitation id="EFDD21EDFF98D56F2FD76FD9FA78FBA1" author="Cuccodoro, G." box="[1241,1432,1017,1043]" pageId="7" pageNumber="578" pagination="107 - 147" refId="ref4399" refString="Cuccodoro, G. (2011) Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae). Revue Suisse de Zoologie, 118 (1), 107 - 147. https: // doi. org / 10.5962 / bhl. part. 117802" type="journal article" year="2011">Cuccodoro 2011</bibRefCitation>
: Fig. 59) intead of distal half (
<figureCitation id="13774099FF98D56F2AD5683DFDC1FB85" box="[475,545,1053,1079]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1927,1952]" captionTargetBox="[177,1417,207,1898]" captionTargetId="figure@3.[151,1436,201,1902]" captionTargetPageId="3" captionText="FIGURES 4–9. Megarthrus chiapas sp. nov., male: aedeagus in ventral (4) and lateral (8) views; mesofemur and mesotro- chanter (5); mesotibia (6); metatibia (7); antenna (9). Scale bars = 0.2 mm: 4–8 (left); 12–18 (right)." figureDoi="http://doi.org/10.5281/zenodo.3828829" httpUri="https://zenodo.org/record/3828829/files/figure.png" pageId="7" pageNumber="578">Figs 7</figureCitation>
and
<figureCitation id="13774099FF98D56F2958683CFD95FB85" box="[598,629,1052,1079]" captionStart="FIGURES 32–38" captionStartId="9.[151,264,1914,1939]" captionTargetBox="[163,1421,187,1881]" captionTargetId="figure@9.[151,1436,181,1889]" captionTargetPageId="9" captionText="FIGURES 32–38. Megarthrus cavianae sp. nov., male: aedeagus in ventral (32) and lateral (37) views; mesofemur and mes- otrochanter (33); mesotibia (34); metatibia (35); antenna (36); abdominal tergite VIII in dorsal view (38). Scale bars = 0.2 mm: 32–35, 37, 38 (horizontal); 36 (vertical)." figureDoi="http://doi.org/10.5281/zenodo.3828839" httpUri="https://zenodo.org/record/3828839/files/figure.png" pageId="7" pageNumber="578">35</figureCitation>
;
<bibRefCitation id="EFDD21EDFF98D56F298E683DFCDDFB85" author="Cuccodoro, G." box="[640,829,1053,1079]" pageId="7" pageNumber="578" pagination="107 - 147" refId="ref4399" refString="Cuccodoro, G. (2011) Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae). Revue Suisse de Zoologie, 118 (1), 107 - 147. https: // doi. org / 10.5962 / bhl. part. 117802" type="journal article" year="2011">Cuccodoro 2011</bibRefCitation>
: Fig. 173). The males of three remaining species differ from each other by the shape of the tip of the abdominal tergite VIII, which is broadly rounded to slightly concave in
<taxonomicName id="4C4C279FFF98D56F2B996846FECAFBCD" authorityName="Sharp" authorityYear="1887" box="[151,298,1125,1151]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="zunilensis">
<emphasis id="B938800EFF98D56F2B996846FECAFBCD" box="[151,298,1125,1151]" italics="true" pageId="7" pageNumber="578">M. zunilensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFDD21EDFF98D56F2A376845FE17FBCD" author="Cuccodoro, G." box="[313,503,1125,1151]" pageId="7" pageNumber="578" pagination="107 - 147" refId="ref4399" refString="Cuccodoro, G. (2011) Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae). Revue Suisse de Zoologie, 118 (1), 107 - 147. https: // doi. org / 10.5962 / bhl. part. 117802" type="journal article" year="2011">Cuccodoro 2011</bibRefCitation>
: Fig. 176), mucronate in
<taxonomicName id="4C4C279FFF98D56F28016846FC6BFBCD" authorityName="Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro" authorityYear="2020" box="[783,907,1125,1151]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="chiapas">
<emphasis id="B938800EFF98D56F28016846FC6BFBCD" box="[783,907,1125,1151]" italics="true" pageId="7" pageNumber="578">M. chiapas</emphasis>
</taxonomicName>
(
<figureCitation id="13774099FF98D56F28946845FC0CFBCD" box="[922,1004,1125,1151]" captionStart="FIGURES 10–18" captionStartId="4.[151,264,1907,1932]" captionTargetBox="[151,1436,181,1866]" captionTargetId="figure@4.[151,1436,181,1882]" captionTargetPageId="4" captionText="FIGURES 10–18. Megarthrus chiapas sp. nov., male: apical segments of abdomen in ventral (10) and lateral (11) views; ab- dominal segments IX and X in lateral (12) and ventral (13) views; abdominal sternite VIII (14), sternite IX (15) and hemitergite IX (18) in ventral view; abdominal tergite VIII in lateral (16) and dorsal (17) views. Scale bars = 0.2 mm: 10–11 (vertical); 12–18 (horizontal)." figureDoi="http://doi.org/10.5281/zenodo.3828831" httpUri="https://zenodo.org/record/3828831/files/figure.png" pageId="7" pageNumber="578">Fig. 17</figureCitation>
), and sinuate in
<taxonomicName id="4C4C279FFF98D56F2FAD6846FACDFBCD" authorityName="Rodriguez, Navarrete-Heredia &amp; Arriaga-Varela" authorityYear="2020" box="[1187,1325,1126,1151]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="cavianae">
<emphasis id="B938800EFF98D56F2FAD6846FACDFBCD" box="[1187,1325,1126,1151]" italics="true" pageId="7" pageNumber="578">M. cavianae</emphasis>
</taxonomicName>
(
<figureCitation id="13774099FF98D56F2E326845FA70FBCD" box="[1340,1424,1125,1151]" captionStart="FIGURES 32–38" captionStartId="9.[151,264,1914,1939]" captionTargetBox="[163,1421,187,1881]" captionTargetId="figure@9.[151,1436,181,1889]" captionTargetPageId="9" captionText="FIGURES 32–38. Megarthrus cavianae sp. nov., male: aedeagus in ventral (32) and lateral (37) views; mesofemur and mes- otrochanter (33); mesotibia (34); metatibia (35); antenna (36); abdominal tergite VIII in dorsal view (38). Scale bars = 0.2 mm: 32–35, 37, 38 (horizontal); 36 (vertical)." figureDoi="http://doi.org/10.5281/zenodo.3828839" httpUri="https://zenodo.org/record/3828839/files/figure.png" pageId="7" pageNumber="578">Fig. 38</figureCitation>
). The shape of the aedeagus, notably that of the apical portion of the ventral wall, is diagnostic for the four species.
</paragraph>
<paragraph id="8BF35C1CFF98D56F2BC9688DFC6AFAE5" blockId="7.[151,1437,152,1511]" pageId="7" pageNumber="578">
The female of
<taxonomicName id="4C4C279FFF98D56F2A62688EFDC6FB75" authorityName="Bierig" authorityYear="1940" box="[364,550,1197,1223]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="flavosignatus">
<emphasis id="B938800EFF98D56F2A62688EFDC6FB75" box="[364,550,1197,1223]" italics="true" pageId="7" pageNumber="578">M. flavosignatus</emphasis>
</taxonomicName>
is still unknown. However the mediodorsal margin of the valvifers is contiguous in apical half in
<taxonomicName id="4C4C279FFF98D56F2A7668F2FDEAFB59" authorityName="Sharp" authorityYear="1887" box="[376,522,1233,1259]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="zunilensis">
<emphasis id="B938800EFF98D56F2A7668F2FDEAFB59" box="[376,522,1233,1259]" italics="true" pageId="7" pageNumber="578">M. zunilensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFDD21EDFF98D56F291768F1FD37FB59" author="Cuccodoro, G." box="[537,727,1233,1259]" pageId="7" pageNumber="578" pagination="107 - 147" refId="ref4399" refString="Cuccodoro, G. (2011) Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae). Revue Suisse de Zoologie, 118 (1), 107 - 147. https: // doi. org / 10.5962 / bhl. part. 117802" type="journal article" year="2011">Cuccodoro 2011</bibRefCitation>
: Fig. 180), while they are not contiguous in
<taxonomicName id="4C4C279FFF98D56F2FB168F2FAA9FB59" authorityName="Rodriguez, Navarrete-Heredia &amp; Arriaga-Varela" authorityYear="2020" box="[1215,1353,1234,1259]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="cavianae">
<emphasis id="B938800EFF98D56F2FB168F2FAA9FB59" box="[1215,1353,1234,1259]" italics="true" pageId="7" pageNumber="578">M. cavianae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4C279FFF98D56F2E7068F2FF0EFABD" authorityName="Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="chiapas">
<emphasis id="B938800EFF98D56F2E7068F2FF0EFABD" italics="true" pageId="7" pageNumber="578">M. chiapas</emphasis>
</taxonomicName>
(
<figureCitation id="13774099FF98D56F2BF068D5FEB1FABD" box="[254,337,1269,1295]" captionStart="FIGURES 19–24" captionStartId="5.[151,264,1907,1932]" captionTargetBox="[161,1433,210,1859]" captionTargetId="figure@5.[151,1436,181,1882]" captionTargetPageId="5" captionText="FIGURES 19–24. Megarthrus chiapas sp. nov., female: genitalia in dorsal (19), posterior (20) and lateral (21) views; abdomi- nal sternite VIII (22) in ventral view; abdominal sternite VIII in lateral (23) and ventral (23) views. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3828833" httpUri="https://zenodo.org/record/3828833/files/figure.png" pageId="7" pageNumber="578">Fig. 19</figureCitation>
), and the posterior apical margin of the abdominal stenite VIII is evenly narrowed to subangulate apex in
<taxonomicName id="4C4C279FFF98D56F2BFE693AFE8DFA81" authorityName="Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro" authorityYear="2020" box="[240,365,1305,1331]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="chiapas">
<emphasis id="B938800EFF98D56F2BFE693AFE8DFA81" box="[240,365,1305,1331]" italics="true" pageId="7" pageNumber="578">M. chiapas</emphasis>
</taxonomicName>
(
<figureCitation id="13774099FF98D56F2A736939FE31FA81" box="[381,465,1305,1331]" captionStart="FIGURES 19–24" captionStartId="5.[151,264,1907,1932]" captionTargetBox="[161,1433,210,1859]" captionTargetId="figure@5.[151,1436,181,1882]" captionTargetPageId="5" captionText="FIGURES 19–24. Megarthrus chiapas sp. nov., female: genitalia in dorsal (19), posterior (20) and lateral (21) views; abdomi- nal sternite VIII (22) in ventral view; abdominal sternite VIII in lateral (23) and ventral (23) views. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3828833" httpUri="https://zenodo.org/record/3828833/files/figure.png" pageId="7" pageNumber="578">Fig. 22</figureCitation>
), while it is slightly sinuate to rounded at apex in
<taxonomicName id="4C4C279FFF98D56F2F0D693AFB76FA81" authorityName="Sharp" authorityYear="1887" box="[1027,1174,1305,1331]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="zunilensis">
<emphasis id="B938800EFF98D56F2F0D693AFB76FA81" box="[1027,1174,1305,1331]" italics="true" pageId="7" pageNumber="578">M. zunilensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFDD21EDFF98D56F2FA86939FA85FA81" author="Cuccodoro, G." box="[1190,1381,1305,1331]" pageId="7" pageNumber="578" pagination="107 - 147" refId="ref4399" refString="Cuccodoro, G. (2011) Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae). Revue Suisse de Zoologie, 118 (1), 107 - 147. https: // doi. org / 10.5962 / bhl. part. 117802" type="journal article" year="2011">Cuccodoro 2011</bibRefCitation>
: Fig. 180), and deeply sinuate to rounded apex in
<taxonomicName id="4C4C279FFF98D56F2981691EFCF9FAE5" authorityName="Rodriguez, Navarrete-Heredia &amp; Arriaga-Varela" authorityYear="2020" box="[655,793,1342,1367]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="cavianae">
<emphasis id="B938800EFF98D56F2981691EFCF9FAE5" box="[655,793,1342,1367]" italics="true" pageId="7" pageNumber="578">M. cavianae</emphasis>
</taxonomicName>
(
<figureCitation id="13774099FF98D56F2827691DFC9DFAE5" box="[809,893,1341,1367]" captionStart="FIGURES 29–31" captionStartId="8.[151,264,1938,1963]" captionTargetBox="[197,1398,417,1905]" captionTargetId="figure@8.[189,1398,417,1913]" captionTargetPageId="8" captionText="FIGURES 29–31. Megarthrus cavianae sp. nov., female: habitus in dorsal view (29); abdominal sternite VIII in ventral views (30); apical segments of abdomen in lateral view (31). Scale bars; vertical = 0.5 mm (29); 25–26. horizontal = 0.2 mm (30, 31)." figureDoi="http://doi.org/10.5281/zenodo.3828837" httpUri="https://zenodo.org/record/3828837/files/figure.png" pageId="7" pageNumber="578">Fig. 30</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3560F97FF98D56F2BC96940FD92FA71" pageId="7" pageNumber="578" type="distribution">
<paragraph id="8BF35C1CFF98D56F2BC96940FD92FA71" blockId="7.[151,1437,152,1511]" pageId="7" pageNumber="578">
<emphasis id="B938800EFF98D56F2BC96940FDADFAC9" bold="true" box="[199,589,1376,1403]" pageId="7" pageNumber="578">Distribution and natural history.</emphasis>
<taxonomicName id="4C4C279FFF98D56F295D6941FC57FAC9" authority="Cuccodoro" authorityName="Rodríguez &amp; Navarrete-Heredia &amp; Arriaga-Varela &amp; Cuccodoro" authorityYear="2020" box="[595,951,1377,1403]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="chiapas">
<emphasis id="B938800EFF98D56F295D6941FCD5FAC9" box="[595,821,1377,1403]" italics="true" pageId="7" pageNumber="578">Megarthrus chiapas</emphasis>
Cuccodoro
</taxonomicName>
<emphasis id="B938800EFF98D56F28B06941FBF0FAC9" bold="true" box="[958,1040,1377,1403]" pageId="7" pageNumber="578">sp nov.</emphasis>
in known only from the northwestern part of the Sierra Madre de
<collectingRegion id="498892FEFF98D56F2AF869A5FDB0FA2D" box="[502,592,1413,1439]" country="Mexico" name="Chiapas" pageId="7" pageNumber="578">Chiapas</collectingRegion>
(
<figureCitation id="13774099FF98D56F296E69A5FD53FA2D" box="[608,691,1413,1439]" captionStart="FIGURE 39" captionStartId="10.[151,250,1537,1562]" captionTargetBox="[151,1436,633,1512]" captionTargetId="figure@10.[151,1436,633,1512]" captionTargetPageId="10" captionText="FIGURE 39. Distribution map of Megarthrus alatorreorum (diamond), M. altivagans (star), M. cavianae sp. nov. (square), M. chiapas sp. nov. (triangle), M. flavosignatus (pentagon) and M. zunilensis (circle)." figureDoi="http://doi.org/10.5281/zenodo.3828841" httpUri="https://zenodo.org/record/3828841/files/figure.png" pageId="7" pageNumber="578">Fig. 39</figureCitation>
), where specimens were collected at elevations ranging from
<quantity id="4CB4F1F9FF98D56F2E6A69A4FEE9FA71" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="7" pageNumber="578" unit="m" value="1650.0" valueMax="1800.0" valueMin="1500.0">
<date id="FFF27ADCFF98D56F2E6A69A4FF0CFA71" pageId="7" pageNumber="578" value="1500" valueMax="1800">1500 to 1800</date>
m
</quantity>
sieving cloud forest’s leaf litter.
</paragraph>
</subSubSection>
<subSubSection id="C3560F97FF98D56F2BC969EDFB60FA55" box="[199,1152,1485,1511]" pageId="7" pageNumber="578" type="etymology">
<paragraph id="8BF35C1CFF98D56F2BC969EDFB60FA55" blockId="7.[151,1437,152,1511]" box="[199,1152,1485,1511]" pageId="7" pageNumber="578">
<emphasis id="B938800EFF98D56F2BC969EDFEAEFA55" bold="true" box="[199,334,1485,1511]" pageId="7" pageNumber="578">Etymology.</emphasis>
The specific name is a noun in apposition derived from the
<typeStatus id="54F7E2BEFF98D56F28E169EDFBFFFA55" box="[1007,1055,1485,1511]" pageId="7" pageNumber="578">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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