treatments-xml/data/03/DE/3E/03DE3E2AE905FFFCDACCF900C977F8ED.xml
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<document id="0DBBA73E376CA246C46605B206E93466" ID-CLB-Dataset="7489" ID-DOI="10.11646/zootaxa.4949.2.11" ID-GBIF-Dataset="e874c6d1-463e-48b3-8d44-2c45ef752a2b" ID-ISSN="1175-5326" ID-Zenodo-Dep="4636238" ID-ZooBank="C792D613-C435-4BC8-8EF7-150427B0D5C4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616665745697" checkinUser="plazi" docAuthor="Narayanan, S. Prasanth, Sathrumithra, S., Anuja, R., Christopher, G., Thomas, A. P. &amp; Julka, J. M." docDate="2021" docId="03DE3E2AE905FFFCDACCF900C977F8ED" docLanguage="en" docName="zootaxa.4949.2.11.pdf" docOrigin="Zootaxa 4949 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Moniligaster keralensis Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="388" masterDocId="FFE74652E900FFFBDA5BFFA2CA01FFA1" masterDocTitle="Three new species and four new species records of earthworms of the genus Moniligaster Perrier, 1872 (Clitellata: Moniligastridae) from Kerala region of the Western Ghats Biodiversity Hotspot, India" masterLastPageNumber="397" masterPageNumber="381" pageNumber="386" updateTime="1698934973357" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="94B7A97393472177EE411743F4A02B23" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="4863F435888550372913F2FB279C9C50">Three new species and four new species records of earthworms of the genus Moniligaster Perrier, 1872 (Clitellata: Moniligastridae) from Kerala region of the Western Ghats Biodiversity Hotspot, India</mods:title>
</mods:titleInfo>
<mods:name id="0BD140CB08EC9C2E6D26884BECF65CAC" type="personal">
<mods:role id="D7179FAE852CAC1D75E8883D0A5641F8">
<mods:roleTerm id="4D45DCB5C2650FE679C41545EC1FECCD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6CCD50446BC4CE3A300C0801C5F1636D">Narayanan, S. Prasanth</mods:namePart>
<mods:affiliation id="10532DBFFF600FF3620E9355B5D9ADCF">Advanced Centre of Environmental Studies and Sustainable Development, Mahatma Gandhi University, Priyadarsini Hills, Kottayam - 686560, Kerala, India</mods:affiliation>
</mods:name>
<mods:name id="A1EBCD27A9E9E1A6F68FD8763F0521C8" type="personal">
<mods:role id="F38ECBFD89E65C2D8D16C503087AAB20">
<mods:roleTerm id="FECD9F8A54AF0D4B41C1F78D218327F2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C5137932C070C9B1DD73D13BCAD905A0">Sathrumithra, S.</mods:namePart>
<mods:affiliation id="A8F92B7A67DC29F8899E4D5DE8ADA72A">Advanced Centre of Environmental Studies and Sustainable Development, Mahatma Gandhi University, Priyadarsini Hills, Kottayam - 686560, Kerala, India</mods:affiliation>
</mods:name>
<mods:name id="607B14D41560D46D1A08B213CF2DC1CC" type="personal">
<mods:role id="0F10F36084F25DAF2230AF78101AC916">
<mods:roleTerm id="2E00EB40B4DBFA7E57460CE14D83B7F2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A869F71FCF11754632E2961CDAD60051">Anuja, R.</mods:namePart>
<mods:affiliation id="F7D9710E8DF8DA4365D494CB22C12F5E">School of Environmental Sciences, Mahatma Gandhi University, Priyadarsini Hills, Kottayam - 686560, Kerala, India</mods:affiliation>
</mods:name>
<mods:name id="1D6A5FDDC0B0153E6DCDC4495D98E082" type="personal">
<mods:role id="0AC7BB883C394C438BB658A82149ED94">
<mods:roleTerm id="6D8CE96C181D1AE36BF263BFB90366F7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D6D847488E60F1C80CE49D530022C28F">Christopher, G.</mods:namePart>
<mods:affiliation id="E81BBA5737E4C6B7D36F55D9B100DCC2">Advanced Centre of Environmental Studies and Sustainable Development, Mahatma Gandhi University, Priyadarsini Hills, Kottayam - 686560, Kerala, India</mods:affiliation>
</mods:name>
<mods:name id="855EDDF4A796F14D48542557B5C0DAFC" type="personal">
<mods:role id="0CA49B9797F78912AC9FBA550E30BB59">
<mods:roleTerm id="3917AC27F7E6D9A26E7C07064B5A617E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A81BC0E624B98A158B38ECF676673CBB">Thomas, A. P.</mods:namePart>
<mods:affiliation id="ED8ED876890BE4E0164071F475D737F9">Advanced Centre of Environmental Studies and Sustainable Development, Mahatma Gandhi University, Priyadarsini Hills, Kottayam - 686560, Kerala, India</mods:affiliation>
</mods:name>
<mods:name id="C3B392BBA8D245D870577B12A949801D" type="personal">
<mods:role id="2AAB2C9C5C951417356104D3814CD1DA">
<mods:roleTerm id="3A682AE69EE2D5DF8A80D2014DF06C17">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="017DA1CCD85CDCC8415294B15C137869">Julka, J. M.</mods:namePart>
<mods:affiliation id="0A4374D407FAC0D13895A76FBEA8B750">School of Biological and Environmental Sciences, Faculty of Basic Sciences, Shoolini University of Biotechnology and Management Sciences, Solan- 173212, Himachal Pradesh, India</mods:affiliation>
</mods:name>
<mods:typeOfResource id="072EE1237E4E611E7D8986A2CE6E821B">text</mods:typeOfResource>
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<mods:titleInfo id="433709426C8C6BECBC441CC42BC845E9">
<mods:title id="4E2CE6C728A775FF0C4ADD4F4E8D204E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="45C24A0862DFA97116BE57B20A80D3AE">
<mods:date id="F09EA6FE54CBC8774977BC24F51F193B">2021</mods:date>
<mods:detail id="DB3B27744A7CD5DF61DD4D4FCF53BA85" type="pubDate">
<mods:number id="DD65935FFBE1E19EF4428C20509656BA">2021-03-25</mods:number>
</mods:detail>
<mods:detail id="122274A87D20DAA2644D15FE194C662B" type="volume">
<mods:number id="6C2458B067CDC6F0CF1CB9A036219C46">4949</mods:number>
</mods:detail>
<mods:detail id="88025811488113721318B13E5B56DC6C" type="issue">
<mods:number id="7CC9DB55AD52C4F3F49562FB243B526E">2</mods:number>
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<mods:start id="36C52D09D05BA6CFBB01F214ECF2191D">381</mods:start>
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<mods:classification id="2BE3AE4AD5213C22030B77C9C649C86E">journal article</mods:classification>
<mods:identifier id="BB58E46CC8DF85721429AB2F81A9CD43" type="CLB-Dataset">7489</mods:identifier>
<mods:identifier id="33E08C539B1C462396410A4E6DA78902" type="DOI">10.11646/zootaxa.4949.2.11</mods:identifier>
<mods:identifier id="FFDC48263E14E968FB76CC4CE8F18AB7" type="GBIF-Dataset">e874c6d1-463e-48b3-8d44-2c45ef752a2b</mods:identifier>
<mods:identifier id="95B50CD09ED024FD7A8CA7634B5BECD8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0DEA52F19924DD01918F6C23CB73CA9F" type="Zenodo-Dep">4636238</mods:identifier>
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<treatment id="03DE3E2AE905FFFCDACCF900C977F8ED" ID-DOI="http://doi.org/10.5281/zenodo.4664601" ID-GBIF-Taxon="180876798" ID-Zenodo-Dep="4664601" LSID="urn:lsid:plazi:treatment:03DE3E2AE905FFFCDACCF900C977F8ED" httpUri="http://treatment.plazi.org/id/03DE3E2AE905FFFCDACCF900C977F8ED" lastPageId="7" lastPageNumber="388" pageId="5" pageNumber="386">
<subSubSection id="C36DDCB7E905FFFEDACCF900C920F91D" box="[151,801,1698,1725]" pageId="5" pageNumber="386" type="nomenclature">
<paragraph id="8BC88F3CE905FFFEDACCF900C920F91D" blockId="5.[151,801,1698,1760]" box="[151,801,1698,1725]" pageId="5" pageNumber="386">
<heading id="D0803850E905FFFEDACCF900C920F91D" bold="true" box="[151,801,1698,1725]" fontSize="11" level="1" pageId="5" pageNumber="386" reason="1">
<emphasis id="B903532EE905FFFEDACCF900C920F91D" bold="true" box="[151,801,1698,1725]" pageId="5" pageNumber="386">
<taxonomicName id="4C77F4BFE905FFFEDACCF900C8B7F91D" authority="Narayanan &amp; Julka" authorityName="Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka" authorityYear="2021" box="[151,694,1698,1725]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Moniligastrida" pageId="5" pageNumber="386" phylum="Annelida" rank="species" species="keralensis" status="sp. nov.">
<emphasis id="B903532EE905FFFEDACCF900CBB2F91C" bold="true" box="[151,435,1698,1725]" italics="true" pageId="5" pageNumber="386">Moniligaster keralensis</emphasis>
Narayanan &amp; Julka
</taxonomicName>
,
<taxonomicNameLabel id="A230EE55E905FFFED898F900C920F91D" box="[707,801,1698,1724]" pageId="5" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36DDCB7E905FFFEDACCF964CB09F941" box="[151,264,1734,1760]" pageId="5" pageNumber="386" type="description">
<paragraph id="8BC88F3CE905FFFEDACCF964CB09F941" blockId="5.[151,801,1698,1760]" box="[151,264,1734,1760]" pageId="5" pageNumber="386">
(
<figureCitation id="134C93B9E905FFFEDAC4F964CAFEF941" box="[159,255,1734,1760]" captionStart="FIGURE 3" captionStartId="6.[151,250,1777,1801]" captionTargetBox="[169,1422,783,1746]" captionTargetId="figure-269@6.[160,1428,777,1754]" captionTargetPageId="6" captionText="FIGURE 3. Moniligaster keralensis Narayanan &amp; Julka sp. nov. A. Ventral view. B. Prostate of right hand side, dorsal view. C. Prostatic capsule of right hand side, dorsal view. D. Spermatheca of left hand side. Abbreviations. M.H.L.—Mass of hairpin loops; Oe.—Oesophagus; Pr.C.—Prostatic capsule; Pr.—Prostate; Pr.D.—Prostatic duct; Sep.—Septum; Sp.A.—Spermathecal ampulla; Sp.Atr.G.—Spermathecal atrial gland; Sp.D.—Spermathecal duct; S.M.A.—Secondary male aperture; Sp.P.—Spermathecal pore; T.S.—Testis sac; Vd—Vas deferens." figureDoi="http://doi.org/10.5281/zenodo.4636246" httpUri="https://zenodo.org/record/4636246/files/figure.png" pageId="5" pageNumber="386">Figure 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36DDCB7E905FFFDDACCF8ACCE40FD50" lastPageId="6" lastPageNumber="387" pageId="5" pageNumber="386" type="materials_examined">
<paragraph id="8BC88F3CE905FFFEDACCF8ACCBF3F835" blockId="5.[151,1436,1806,2013]" pageId="5" pageNumber="386">
<emphasis id="B903532EE905FFFEDACCF8ACCB45F888" bold="true" box="[151,324,1806,1833]" pageId="5" pageNumber="386">Type material.</emphasis>
<materialsCitation id="3B1F8561E905FFFEDB10F8ACC947F8D1" ID-GBIF-Occurrence="3068455313" collectingDate="2015-05-03" collectionCode="WGRC, IR" collectorName="S. Sathrumithra &amp; J. K. Joseph &amp; A. Sasi" country="India" county="Pathanamthitta District" latitude="9.204945" location="Clitellate" longLatPrecision="1" longitude="77.010086" municipality="Kurichikanam in Konni Reserve Forest" pageId="5" pageNumber="386" specimenCount="1" stateProvince="Kerala" typeStatus="holotype">
<emphasis id="B903532EE905FFFEDB10F8ACCBB3F889" box="[331,434,1806,1832]" italics="true" pageId="5" pageNumber="386">
<typeStatus id="54CC319EE905FFFEDB10F8ACCBB3F889" box="[331,434,1806,1832]" pageId="5" pageNumber="386" type="holotype">Holotype</typeStatus>
</emphasis>
:
<location id="8EA8D9E7E905FFFEDB9BF8ACC826F889" LSID="urn:lsid:plazi:treatment:03DE3E2AE905FFFCDACCF900C977F8ED:8EA8D9E7E905FFFEDB9BF8ACC826F889" box="[448,551,1806,1832]" country="India" county="Pathanamthitta District" latitude="9.204945" longLatPrecision="1" longitude="77.010086" municipality="Kurichikanam in Konni Reserve Forest" name="Clitellate" pageId="5" pageNumber="386" stateProvince="Kerala">Clitellate</location>
(ZSI/
<collectionCode id="ED6617F9E905FFFED833F8ACC8BEF889" box="[616,703,1806,1832]" pageId="5" pageNumber="386">WGRC</collectionCode>
/
<collectionCode id="ED6617F9E905FFFED89CF8ACC8E2F889" box="[711,739,1806,1832]" pageId="5" pageNumber="386">IR</collectionCode>
/INV/8839), evergreen forest,
<collectingMunicipality id="6BAC1546E905FFFEDE69F8ACCADCF8EC" pageId="5" pageNumber="386">Kurichikanam in Konni Reserve Forest</collectingMunicipality>
(
<geoCoordinate id="EE43E9FBE905FFFEDAAAF890CB7FF8ED" box="[241,382,1842,1868]" degrees="9" direction="north" minutes="12" orientation="latitude" pageId="5" pageNumber="386" precision="1" seconds="17.8" value="9.204945">9°12'17.8&quot;N</geoCoordinate>
,
<geoCoordinate id="EE43E9FBE905FFFEDBD4F890C827F8EC" box="[399,550,1842,1869]" degrees="77" direction="east" minutes="00" orientation="longitude" pageId="5" pageNumber="386" precision="1" seconds="36.3" value="77.010086">77°00'36.3&quot;E</geoCoordinate>
) (
<quantity id="4C8F22D9E905FFFED819F890C88CF8ED" box="[578,653,1842,1868]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="386" unit="km" value="10.0">10 km</quantity>
from Kokkathodu),
<collectingCounty id="62A9F7B0E905FFFED9DBF890CE8BF8ED" box="[896,1162,1842,1868]" pageId="5" pageNumber="386">Pathanamthitta District</collectingCounty>
,
<collectingRegion id="49B341DEE905FFFEDEC2F890CEE7F8ED" box="[1177,1254,1842,1868]" country="India" name="Kerala" pageId="5" pageNumber="386">Kerala</collectingRegion>
,
<collectingCountry id="F360CFACE905FFFEDEADF890CF33F8ED" box="[1270,1330,1842,1868]" name="India" pageId="5" pageNumber="386">India</collectingCountry>
,
<date id="FFC9A9FCE905FFFEDF19F890CAD3F8D0" pageId="5" pageNumber="386" value="2015-05-03">
<collectingDate id="EF8D5014E905FFFEDF19F890CAD3F8D0" pageId="5" pageNumber="386" value="2015-05-03">
03 May
<quantity id="4C8F22D9E905FFFEDACCF8F4CAD3F8D0" box="[151,210,1878,1905]" metricMagnitude="6" metricUnit="m" metricValue="2.015534" pageId="5" pageNumber="386" unit="m" value="2015534.0">2015</quantity>
</collectingDate>
</date>
, 534 m a.s.l.,
<collectorName id="2682EAEAE905FFFEDB2FF8F4C828F8D1" box="[372,553,1878,1904]" pageId="5" pageNumber="386">S. Sathrumithra</collectorName>
,
<collectorName id="2682EAEAE905FFFED86DF8F5C8B6F8D1" box="[566,695,1878,1905]" pageId="5" pageNumber="386">J.K. Joseph</collectorName>
and
<collectorName id="2682EAEAE905FFFED8ABF8F4C947F8D1" box="[752,838,1878,1904]" pageId="5" pageNumber="386">A. Sasi</collectorName>
</materialsCitation>
.
<materialsCitation id="3B1F8561E905FFFED908F8F5CBEFF835" ID-GBIF-Occurrence="3068455302" collectingDate="2015-05-03" collectionCode="WGRC, IR" collectorName="S. Sathrumithra &amp; J. K. Joseph &amp; A. Sasi" country="India" county="Pathanamthitta District" latitude="9.204945" location="Clitellate" longLatPrecision="1" longitude="77.010086" municipality="Kurichikanam in Konni Reserve Forest" pageId="5" pageNumber="386" specimenCount="1" stateProvince="Kerala" typeStatus="paratype">
<emphasis id="B903532EE905FFFED908F8F5C9C4F8D1" box="[851,965,1879,1904]" italics="true" pageId="5" pageNumber="386">
<typeStatus id="54CC319EE905FFFED908F8F5C9C4F8D1" box="[851,965,1879,1904]" pageId="5" pageNumber="386" type="paratype">Paratypes</typeStatus>
</emphasis>
: 4 clitellates (ZSI/
<collectionCode id="ED6617F9E905FFFEDEC1F8F4CEF1F8D1" box="[1178,1264,1878,1904]" pageId="5" pageNumber="386">WGRC</collectionCode>
/
<collectionCode id="ED6617F9E905FFFEDEA2F8F4CF14F8D1" box="[1273,1301,1878,1904]" pageId="5" pageNumber="386">IR</collectionCode>
/INV/8840), collection data as for holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BC88F3CE905FFFDDA9CF83CCE40FD50" blockId="5.[151,1436,1806,2013]" lastBlockId="6.[151,1437,150,753]" lastPageId="6" lastPageNumber="387" pageId="5" pageNumber="386">
<emphasis id="B903532EE905FFFEDA9CF83CCB82F818" bold="true" box="[199,387,1950,1977]" pageId="5" pageNumber="386">Other material.</emphasis>
<specimenCount id="9D7144B5E905FFFEDBD7F83CC827F819" box="[396,550,1950,1976]" pageId="5" pageNumber="386" type="generic">41 specimens</specimenCount>
altogether. 3 clitellates (
<collectionCode id="ED6617F9E905FFFED91BF83CC9AFF819" box="[832,942,1950,1976]" pageId="5" pageNumber="386">ACESSD</collectionCode>
/EW/729),
<taxonomicName id="4C77F4BFE905FFFEDE77F83CCF33F819" box="[1068,1330,1950,1976]" class="Liliopsida" family="Zingiberaceae" genus="Elettaria" kingdom="Plantae" order="Zingiberales" pageId="5" pageNumber="386" phylum="Tracheophyta" rank="species" species="cardamomum">
<emphasis id="B903532EE905FFFEDE77F83CCF33F819" box="[1068,1330,1950,1976]" italics="true" pageId="5" pageNumber="386">Elettaria cardamomum</emphasis>
</taxonomicName>
and
<emphasis id="B903532EE905FFFEDF37F83DCB09F87D" italics="true" pageId="5" pageNumber="386">Piper nigrum</emphasis>
plantation, Chathurangappara in Cardamom Hill Reserve (
<geoCoordinate id="EE43E9FBE905FFFED9F2F860CE3CF87C" box="[937,1085,1986,2013]" degrees="9" direction="north" minutes="53" orientation="latitude" pageId="5" pageNumber="386" precision="1" seconds="46.8" value="9.896333">9°5346.8”N</geoCoordinate>
,
<geoCoordinate id="EE43E9FBE905FFFEDE11F860CED8F87C" box="[1098,1241,1986,2013]" degrees="77" direction="east" minutes="13" orientation="longitude" pageId="5" pageNumber="386" precision="1" seconds="5.3" value="77.21814">77°13 5.3”E</geoCoordinate>
), Idukki District, Kerala,
<collectingCountry id="F360CFACE906FFFDDAA9FF35CB2FFF10" box="[242,302,151,177]" name="India" pageId="6" pageNumber="387">India</collectingCountry>
,
<date id="FFC9A9FCE906FFFDDB60FF35CBFAFF10" box="[315,507,151,177]" pageId="6" pageNumber="387" value="2011-01-12">12 January 2011</date>
, 1105 m a.s.l., S.P. Narayanan; 1 clitellate and 7 aclitellates (
<collectionCode id="ED6617F9E906FFFDDEE1FF35CF29FF10" box="[1210,1320,151,177]" pageId="6" pageNumber="387">ACESSD</collectionCode>
/EW/730), evergreen forest, Aluvamkudy (
<geoCoordinate id="EE43E9FBE906FFFDDBA1FF19C87BFF74" box="[506,634,187,213]" degrees="9" direction="north" minutes="16" orientation="latitude" pageId="6" pageNumber="387" precision="1" seconds="7.8" value="9.268833">9°16'7.8&quot;N</geoCoordinate>
,
<geoCoordinate id="EE43E9FBE906FFFDD8DDFF19C91EFF77" box="[646,799,187,214]" degrees="77" direction="east" minutes="00" orientation="longitude" pageId="6" pageNumber="387" precision="1" seconds="16.8" value="77.00467">77°00'16.8&quot;E</geoCoordinate>
), Pathanamthitta District,
<collectingRegion id="49B341DEE906FFFDDE1FFF19CE90FF74" box="[1092,1169,187,213]" country="India" name="Kerala" pageId="6" pageNumber="387">Kerala</collectingRegion>
,
<collectingCountry id="F360CFACE906FFFDDEC7FF19CED9FF74" box="[1180,1240,187,213]" name="India" pageId="6" pageNumber="387">India</collectingCountry>
,
<date id="FFC9A9FCE906FFFDDEBFFF19CF99FF74" box="[1252,1432,187,213]" pageId="6" pageNumber="387" value="2013-08-21">
21 August
<quantity id="4C8F22D9E906FFFDDF06FF19CF99FF74" box="[1373,1432,187,213]" metricMagnitude="6" metricUnit="m" metricValue="2.0135129999999997" pageId="6" pageNumber="387" unit="m" value="2013513.0">2013</quantity>
</date>
, 513 m a.s.l., S.P. Narayanan, S. Sathrumithra, D. Kuriakose and A. Sasi; 1 clitellate and 6 aclitellates (
<collectionCode id="ED6617F9E906FFFDDF7CFF7DCF92FF58" box="[1319,1427,223,249]" pageId="6" pageNumber="387">ACESSD</collectionCode>
/ EW/731), riparian stretch, Mundomoozhy (
<geoCoordinate id="EE43E9FBE906FFFDD8D9FEA1C911FEBC" box="[642,784,259,285]" degrees="9" direction="north" minutes="14" orientation="latitude" pageId="6" pageNumber="387" precision="1" seconds="43.8" value="9.245501">9°14'43.8&quot;N</geoCoordinate>
,
<geoCoordinate id="EE43E9FBE906FFFDD947FEA0C9B7FEBC" box="[796,950,258,285]" degrees="76" direction="east" minutes="55" orientation="longitude" pageId="6" pageNumber="387" precision="1" seconds="20.1" value="76.92225">76°55'20.1&quot;E</geoCoordinate>
), Pathanamthitta District,
<collectingRegion id="49B341DEE906FFFDDE85FEA1CF2AFEBC" box="[1246,1323,259,285]" country="India" name="Kerala" pageId="6" pageNumber="387">Kerala</collectingRegion>
,
<collectingCountry id="F360CFACE906FFFDDF63FEA1CF75FEBC" box="[1336,1396,259,285]" name="India" pageId="6" pageNumber="387">India</collectingCountry>
,
<date id="FFC9A9FCE906FFFDDFDBFEA1CB2CFEE0" pageId="6" pageNumber="387" value="2013-08-21">21 August 2013</date>
, 68 m a.s.l., S.P. Narayanan, S. Sathrumithra, D. Kuriakose and A. Sasi; 4 aclitellates (
<collectionCode id="ED6617F9E906FFFDDF7CFE85CF92FEE0" box="[1319,1427,295,321]" pageId="6" pageNumber="387">ACESSD</collectionCode>
/ EW/732), evergreen forest, Kottavasal in Achankovil Reserve Forest (
<geoCoordinate id="EE43E9FBE906FFFDD990FEE9CE4AFEC4" box="[971,1099,331,357]" degrees="9" direction="north" minutes="4" orientation="latitude" pageId="6" pageNumber="387" precision="1" seconds="17.4" value="9.0715">9°4'17.4&quot;N</geoCoordinate>
,
<geoCoordinate id="EE43E9FBE906FFFDDE00FEE8CEF5FEC4" box="[1115,1268,330,357]" degrees="77" direction="east" minutes="12" orientation="longitude" pageId="6" pageNumber="387" precision="1" seconds="24.5" value="77.2068">77°12'24.5&quot;E</geoCoordinate>
), Kollam District,
<collectingRegion id="49B341DEE906FFFDDA8FFECDCB20FE28" box="[212,289,367,393]" country="India" name="Kerala" pageId="6" pageNumber="387">Kerala</collectingRegion>
,
<collectingCountry id="F360CFACE906FFFDDB74FECDCB6AFE28" box="[303,363,367,393]" name="India" pageId="6" pageNumber="387">India</collectingCountry>
,
<date id="FFC9A9FCE906FFFDDB22FECDC833FE28" box="[377,562,367,393]" pageId="6" pageNumber="387" value="2013-08-22">
22 August
<quantity id="4C8F22D9E906FFFDDBACFECDC833FE28" box="[503,562,367,393]" metricMagnitude="6" metricUnit="m" metricValue="2.0134739999999995" pageId="6" pageNumber="387" unit="m" value="2013474.0">2013</quantity>
</date>
, 474 m a.s.l., S.P. Narayanan,
<collectionCode id="ED6617F9E906FFFDD9C9FECDC9A5FE28" box="[914,932,367,393]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="387">T</collectionCode>
. Augustine, A. Sasi, S. Sathrumithra and D. Kuriakose; 1 clitellate (
<collectionCode id="ED6617F9E906FFFDDBFAFE31C80EFE0C" box="[417,527,403,429]" pageId="6" pageNumber="387">ACESSD</collectionCode>
/EW/733), evergreen forest, near to Ambanad estate in Achankovil Reserve Forest (
<geoCoordinate id="EE43E9FBE906FFFDDA92FE15CB56FE70" box="[201,343,439,465]" degrees="9" direction="north" minutes="02" orientation="latitude" pageId="6" pageNumber="387" precision="1" seconds="26.4" value="9.040668">9°02'26.4&quot;N</geoCoordinate>
,
<geoCoordinate id="EE43E9FBE906FFFDDB3CFE15C801FE73" box="[359,512,439,466]" degrees="77" direction="east" minutes="07" orientation="longitude" pageId="6" pageNumber="387" precision="1" seconds="06.3" value="77.118416">77°07'06.3&quot;E</geoCoordinate>
), Kollam District,
<collectingRegion id="49B341DEE906FFFDD880FE15C929FE70" box="[731,808,439,465]" country="India" name="Kerala" pageId="6" pageNumber="387">Kerala</collectingRegion>
,
<collectingCountry id="F360CFACE906FFFDD963FE15C975FE70" box="[824,884,439,465]" name="India" pageId="6" pageNumber="387">India</collectingCountry>
,
<date id="FFC9A9FCE906FFFDD9D8FE15CE3FFE70" box="[899,1086,439,465]" pageId="6" pageNumber="387" value="2013-08-24">
24 August
<quantity id="4C8F22D9E906FFFDDE58FE15CE3FFE70" box="[1027,1086,439,465]" metricMagnitude="6" metricUnit="m" metricValue="2.013795" pageId="6" pageNumber="387" unit="m" value="2013795.0">2013</quantity>
</date>
, 795 m a.s.l., S.P. Narayanan, D. Kuriakose, S. Sathrumithra,
<collectionCode id="ED6617F9E906FFFDD858FE79C814FE54" box="[515,533,475,501]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="387">T</collectionCode>
. Augustine and A. Sasi; 1 clitellate (
<collectionCode id="ED6617F9E906FFFDD9E4FE79CE2CFE54" box="[959,1069,475,501]" pageId="6" pageNumber="387">ACESSD</collectionCode>
/EW/734), mixed crop area, Peruvanthanam (
<geoCoordinate id="EE43E9FBE906FFFDDB60FE5CCBC8FDB8" box="[315,457,510,537]" degrees="9" direction="north" minutes="33" orientation="latitude" pageId="6" pageNumber="387" precision="1" seconds="55.3" value="9.565361">9°33'55.3&quot;N</geoCoordinate>
,
<geoCoordinate id="EE43E9FBE906FFFDDB8CFE5CC862FDB8" box="[471,611,510,537]" degrees="76" direction="east" minutes="56" orientation="longitude" pageId="6" pageNumber="387" precision="1" seconds="7.7" value="76.93547">76°56'7.7&quot;E</geoCoordinate>
), Idukki District,
<collectingRegion id="49B341DEE906FFFDD96BFE5DC97CFDB8" box="[816,893,511,537]" country="India" name="Kerala" pageId="6" pageNumber="387">Kerala</collectingRegion>
,
<collectingCountry id="F360CFACE906FFFDD9D0FE5DC9C6FDB8" box="[907,967,511,537]" name="India" pageId="6" pageNumber="387">India</collectingCountry>
,
<date id="FFC9A9FCE906FFFDD98EFE5DCE6CFDB8" box="[981,1133,511,537]" pageId="6" pageNumber="387" value="2014-07-22">
22 July
<quantity id="4C8F22D9E906FFFDDE69FE5DCE6CFDB8" box="[1074,1133,511,537]" metricMagnitude="6" metricUnit="m" metricValue="2.014213" pageId="6" pageNumber="387" unit="m" value="2014213.0">2014</quantity>
</date>
, 213 m a.s.l., J.K. Joseph; 1 clitellate and 4 aclitellates (
<collectionCode id="ED6617F9E906FFFDDBB6FD80C85AFD9C" box="[493,603,546,573]" pageId="6" pageNumber="387">ACESSD</collectionCode>
/EW/735), semi evergreen forest, Thannithodumoozhy in Konni Reserve Forest (
<geoCoordinate id="EE43E9FBE906FFFDDAB4FDE4CB7DFDC0" box="[239,380,582,609]" degrees="9" direction="north" minutes="15" orientation="latitude" pageId="6" pageNumber="387" precision="1" seconds="19.0" value="9.255278">9°15'19.0&quot;N</geoCoordinate>
,
<geoCoordinate id="EE43E9FBE906FFFDDBD1FDE4C822FDC0" box="[394,547,582,609]" degrees="76" direction="east" minutes="55" orientation="longitude" pageId="6" pageNumber="387" precision="1" seconds="36.4" value="76.92677">76°55'36.4&quot;E</geoCoordinate>
), Kollam District,
<collectingRegion id="49B341DEE906FFFDD8A2FDE5C947FDC0" box="[761,838,583,609]" country="India" name="Kerala" pageId="6" pageNumber="387">Kerala</collectingRegion>
,
<collectingCountry id="F360CFACE906FFFDD908FDE5C98EFDC0" box="[851,911,583,609]" name="India" pageId="6" pageNumber="387">India</collectingCountry>
,
<date id="FFC9A9FCE906FFFDD9C6FDE5CE39FDC0" box="[925,1080,583,609]" pageId="6" pageNumber="387" value="2016-06-10">10 June 2016</date>
, 94 m a.s.l., S. Sathrumithra; 1 clitellate and 4 aclitellates (
<collectionCode id="ED6617F9E906FFFDDB8BFDC9C83FFD27" box="[464,574,619,646]" pageId="6" pageNumber="387">ACESSD</collectionCode>
/EW/736),
<taxonomicName id="4C77F4BFE906FFFDD8E0FDC9C970FD24" box="[699,881,619,645]" class="Magnoliopsida" family="Lamiaceae" genus="Tectona" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="387" phylum="Tracheophyta" rank="species" species="grandis">
<emphasis id="B903532EE906FFFDD8E0FDC9C970FD24" box="[699,881,619,645]" italics="true" pageId="6" pageNumber="387">Tectona grandis</emphasis>
</taxonomicName>
plantation, Pananthodu in Konni Reserve Forest (
<geoCoordinate id="EE43E9FBE906FFFDDAFBFD2DCB32FD08" box="[160,307,655,681]" degrees="9" direction="north" minutes="13" orientation="latitude" pageId="6" pageNumber="387" precision="1" seconds="37.4" value="9.227056">9°1337.4”N</geoCoordinate>
,
<geoCoordinate id="EE43E9FBE906FFFDDB64FD2CCBDCFD08" box="[319,477,654,681]" degrees="76" direction="east" minutes="55" orientation="longitude" pageId="6" pageNumber="387" precision="1" seconds="26.8" value="76.92411">76°5526.8”E</geoCoordinate>
), Pathanamthitta District,
<collectingRegion id="49B341DEE906FFFDD959FD2DC94EFD08" box="[770,847,655,681]" country="India" name="Kerala" pageId="6" pageNumber="387">Kerala</collectingRegion>
,
<collectingCountry id="F360CFACE906FFFDD901FD2DC997FD08" box="[858,918,655,681]" name="India" pageId="6" pageNumber="387">India</collectingCountry>
,
<date id="FFC9A9FCE906FFFDD9F9FD2DCE34FD08" box="[930,1077,655,681]" pageId="6" pageNumber="387" value="2016-07-10">
10 July
<quantity id="4C8F22D9E906FFFDD9A1FD2DCE34FD08" box="[1018,1077,655,681]" metricMagnitude="6" metricUnit="m" metricValue="2.016105" pageId="6" pageNumber="387" unit="m" value="2016105.0">2016</quantity>
</date>
, 105 m a.s.l., S. Sathrumithra; 1 clitellate and 6 aclitellates (
<collectionCode id="ED6617F9E906FFFDDB9FFD11C833FD6C" box="[452,562,691,717]" pageId="6" pageNumber="387">ACESSD</collectionCode>
/EW/737),
<taxonomicName id="4C77F4BFE906FFFDD8F7FD11C97BFD6C" box="[684,890,691,717]" class="Magnoliopsida" family="Euphorbiaceae" genus="Hevea" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="387" phylum="Tracheophyta" rank="species" species="brasiliensis">
<emphasis id="B903532EE906FFFDD8F7FD11C97BFD6C" box="[684,890,691,717]" italics="true" pageId="6" pageNumber="387">Hevea brasiliensis</emphasis>
</taxonomicName>
plantation, Chengara (
<geoCoordinate id="EE43E9FBE906FFFDDE2CFD10CEFAFD6C" box="[1143,1275,690,717]" degrees="9" direction="north" minutes="15" orientation="latitude" pageId="6" pageNumber="387" precision="1" seconds="2.1" value="9.250584">9°152.1”N</geoCoordinate>
,
<geoCoordinate id="EE43E9FBE906FFFDDF5DFD10CF8EFD6C" box="[1286,1423,690,717]" degrees="76" direction="east" minutes="51" orientation="longitude" pageId="6" pageNumber="387" precision="15" seconds="54" value="76.865">76°5154”E</geoCoordinate>
), Pathanamthitta District,
<collectingRegion id="49B341DEE906FFFDDBF3FD75CBF4FD50" box="[424,501,727,753]" country="India" name="Kerala" pageId="6" pageNumber="387">Kerala</collectingRegion>
,
<collectingCountry id="F360CFACE906FFFDD85AFD75C83CFD50" box="[513,573,727,753]" name="India" pageId="6" pageNumber="387">India</collectingCountry>
,
<date id="FFC9A9FCE906FFFDD813FD75C8FCFD50" box="[584,765,727,753]" pageId="6" pageNumber="387" value="2016-08-23">23 August 2016</date>
, 88 m a.s.l., S. Sathrumithra.
</paragraph>
</subSubSection>
<caption id="DF08DFB4E906FFFDDACCF953C8A2F83B" ID-DOI="http://doi.org/10.5281/zenodo.4636246" ID-Zenodo-Dep="4636246" httpUri="https://zenodo.org/record/4636246/files/figure.png" pageId="6" pageNumber="387" startId="6.[151,250,1777,1801]" targetBox="[169,1422,783,1746]" targetPageId="6">
<paragraph id="8BC88F3CE906FFFDDACCF953C8A2F83B" blockId="6.[151,1437,1777,1946]" pageId="6" pageNumber="387">
<emphasis id="B903532EE906FFFDDACCF953CB15F8A8" bold="true" box="[151,276,1777,1801]" pageId="6" pageNumber="387">FIGURE 3.</emphasis>
<taxonomicName id="4C77F4BFE906FFFDDB40F950C8D8F8AB" authority="Narayanan &amp; Julka" authorityName="Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka" authorityYear="2021" box="[283,729,1778,1802]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Moniligastrida" pageId="6" pageNumber="387" phylum="Annelida" rank="species" species="keralensis" status="sp. nov.">
<emphasis id="B903532EE906FFFDDB40F950C80CF8AB" box="[283,525,1778,1802]" italics="true" pageId="6" pageNumber="387">Moniligaster keralensis</emphasis>
Narayanan &amp; Julka
</taxonomicName>
<emphasis id="B903532EE906FFFDD8BAF950C953F8A8" bold="true" box="[737,850,1777,1802]" pageId="6" pageNumber="387">
<taxonomicNameLabel id="A230EE55E906FFFDD8BAF950C932F8AB" box="[737,819,1778,1802]" pageId="6" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Ventral view.
<emphasis id="B903532EE906FFFDD9BDF950C9FFF8AB" bold="true" box="[998,1022,1778,1802]" pageId="6" pageNumber="387">B.</emphasis>
Prostate of right hand side, dorsal view.
<emphasis id="B903532EE906FFFDDACCF8B7CAB1F88C" bold="true" box="[151,176,1813,1837]" pageId="6" pageNumber="387">C.</emphasis>
Prostatic capsule of right hand side, dorsal view.
<emphasis id="B903532EE906FFFDD8FDF8B4C8BEF88F" bold="true" box="[678,703,1814,1838]" pageId="6" pageNumber="387">D.</emphasis>
Spermatheca of left hand side. Abbreviations. M.H.L.—Mass of hairpin loops; Oe.—Oesophagus; Pr.C.—Prostatic capsule; Pr.—Prostate; Pr.D.—Prostatic duct; Sep.—Septum; Sp.A.—Spermathecal ampulla; Sp.Atr.G.—Spermathecal atrial gland; Sp.D.—Spermathecal duct; S.M.A.—Secondary male aperture; Sp.P.—Spermathecal pore; T.S.—Testis sac; Vd—Vas deferens.
</paragraph>
</caption>
<subSubSection id="C36DDCB7E907FFFCDA9CFF34C9E6FE29" pageId="7" pageNumber="388" type="diagnosis">
<paragraph id="8BC88F3CE907FFFCDA9CFF34C9E6FE29" blockId="7.[151,1437,150,1869]" pageId="7" pageNumber="388">
<emphasis id="B903532EE907FFFCDA9CFF34CB40FF10" bold="true" box="[199,321,150,177]" pageId="7" pageNumber="388">Diagnosis.</emphasis>
Colour bluish. Length
<quantity id="4C8F22D9E907FFFCD815FF34C8DBFF10" box="[590,730,150,177]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="1.29" metricValueMin="0.71" pageId="7" pageNumber="388" unit="mm" value="100.0" valueMax="129.0" valueMin="71.0">71129 mm</quantity>
, diameter
<quantity id="4C8F22D9E907FFFCD909FF34C9C9FF10" box="[850,968,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.5" metricValueMin="4.0" pageId="7" pageNumber="388" unit="mm" value="4.75" valueMax="5.5" valueMin="4.0">45.5 mm</quantity>
, segments 109195. Secondary male apertures paired, large transverse slits, slightly lateral to
<emphasis id="B903532EE907FFFCD95AFF18C90EFF75" box="[769,783,186,212]" italics="true" pageId="7" pageNumber="388">b</emphasis>
setal lines, at intersegmental furrow 10/11. Spermathecal pores paired, small transverse slits, close to
<emphasis id="B903532EE907FFFCD8D6FF7CC8A6FF59" box="[653,679,222,248]" italics="true" pageId="7" pageNumber="388">cd</emphasis>
setal lines, at intersegmental furrow 7/8. Genital markings absent. Gizzards four in segments 1419. Vas deferens a mass of hairpin loops, mass as large as testis sac, entering prostate directly, near the ectal end. Prostates glandular, tubular with irregular margins, slightly flattened, prostatic capsule tubular, prostatic duct bent downwards and narrowed. Spermathecal atrial glands bilobed in segment 7, glands bound together, duct of each gland about as long as the common atrial duct.
</paragraph>
</subSubSection>
<subSubSection id="C36DDCB7E907FFFCDA9CFE30C909FB65" pageId="7" pageNumber="388" type="description">
<paragraph id="8BC88F3CE907FFFCDA9CFE30CBAEFD51" blockId="7.[151,1437,150,1869]" pageId="7" pageNumber="388">
<emphasis id="B903532EE907FFFCDA9CFE30CB58FE0C" bold="true" box="[199,345,402,429]" pageId="7" pageNumber="388">Description.</emphasis>
External: Colour bluish; body circular in cross section. Dimensions:
<typeStatus id="54CC319EE907FFFCDE01FE30CEC6FE0D" box="[1114,1223,402,428]" pageId="7" pageNumber="388" type="holotype">Holotype</typeStatus>
:
<quantity id="4C8F22D9E907FFFCDE8BFE30CF1EFE0C" box="[1232,1311,402,429]" metricMagnitude="-2" metricUnit="m" metricValue="7.1" pageId="7" pageNumber="388" unit="mm" value="71.0">71 mm</quantity>
by
<quantity id="4C8F22D9E907FFFCDF1CFE30CF9CFE0C" box="[1351,1437,402,429]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="388" unit="mm" value="4.5">4.5 mm</quantity>
at segment 9, segments 109;
<typeStatus id="54CC319EE907FFFCDBBAFE15C850FE70" box="[481,593,439,465]" pageId="7" pageNumber="388" type="paratype">paratypes</typeStatus>
:
<quantity id="4C8F22D9E907FFFCD806FE14C8D6FE70" box="[605,727,438,465]" metricMagnitude="-2" metricUnit="m" metricValue="8.55" metricValueMax="8.9" metricValueMin="8.2" pageId="7" pageNumber="388" unit="mm" value="85.5" valueMax="89.0" valueMin="82.0">8289 mm</quantity>
by
<quantity id="4C8F22D9E907FFFCD958FE14C963FE70" box="[771,866,438,465]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="7" pageNumber="388" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
at segment 9, segments 135149; other materials:
<quantity id="4C8F22D9E907FFFCDACCFE78CB1EFE54" box="[151,287,474,501]" metricMagnitude="-1" metricUnit="m" metricValue="1.105" metricValueMax="1.29" metricValueMin="0.9199999999999999" pageId="7" pageNumber="388" unit="mm" value="110.5" valueMax="129.0" valueMin="92.0">92129 mm</quantity>
by
<quantity id="4C8F22D9E907FFFCDB10FE78CBD2FE54" box="[331,467,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="7" pageNumber="388" unit="mm" value="5.0" valueMax="5.5" valueMin="4.5">4.55.5 mm</quantity>
at segment 9, segments 166195. Setae lumbricine, closely paired, present from segment 2; setal formula
<emphasis id="B903532EE907FFFCDBC8FE5DCBAEFDB9" box="[403,431,511,536]" italics="true" pageId="7" pageNumber="388">aa</emphasis>
= 78.75
<emphasis id="B903532EE907FFFCD87EFE5CC840FDB9" box="[549,577,510,536]" italics="true" pageId="7" pageNumber="388">ab</emphasis>
= 1.081.46
<emphasis id="B903532EE907FFFCD880FE5CC8F4FDB9" box="[731,757,510,536]" italics="true" pageId="7" pageNumber="388">bc</emphasis>
= 9.3311.67
<emphasis id="B903532EE907FFFCD9C0FE5CC9B4FDB9" box="[923,949,510,536]" italics="true" pageId="7" pageNumber="388">cd</emphasis>
= 0.20.27
<emphasis id="B903532EE907FFFCDE62FE5CCE54FDB9" box="[1081,1109,510,536]" italics="true" pageId="7" pageNumber="388">dd</emphasis>
at segment 8,
<emphasis id="B903532EE907FFFCDEA6FE5DCF18FDB9" box="[1277,1305,511,536]" italics="true" pageId="7" pageNumber="388">aa</emphasis>
= 6.28.75
<emphasis id="B903532EE907FFFCDACCFD80CAB2FD9D" box="[151,179,546,572]" italics="true" pageId="7" pageNumber="388">ab</emphasis>
= 0.911
<emphasis id="B903532EE907FFFCDB7DFD80CB41FD9D" box="[294,320,546,572]" italics="true" pageId="7" pageNumber="388">bc</emphasis>
= 7.7511.67
<emphasis id="B903532EE907FFFCDBB8FD80CBFCFD9D" box="[483,509,546,572]" italics="true" pageId="7" pageNumber="388">cd</emphasis>
= 0.310.44
<emphasis id="B903532EE907FFFCD8C8FD80C8AEFD9D" box="[659,687,546,572]" italics="true" pageId="7" pageNumber="388">dd</emphasis>
at segment 20 (n=3). Clitellum annular, on segments 1013, intersegmental furrows distinct, colour reddish. Spermathecal pores small transverse slits, aligned between setae
<emphasis id="B903532EE907FFFCDF04FDE4CF78FDC1" box="[1375,1401,582,608]" italics="true" pageId="7" pageNumber="388">cd</emphasis>
, at intersegmental furrow 7/8. Male pores paired, minute, at the centre of short, eversible penes, secondary male apertures oval-shaped at intersegmental furrow 10/11 (
<figureCitation id="134C93B9E907FFFCD896FD2CC926FD09" box="[717,807,654,680]" captionStart="FIGURE 3" captionStartId="6.[151,250,1777,1801]" captionTargetBox="[169,1422,783,1746]" captionTargetId="figure-269@6.[160,1428,777,1754]" captionTargetPageId="6" captionText="FIGURE 3. Moniligaster keralensis Narayanan &amp; Julka sp. nov. A. Ventral view. B. Prostate of right hand side, dorsal view. C. Prostatic capsule of right hand side, dorsal view. D. Spermatheca of left hand side. Abbreviations. M.H.L.—Mass of hairpin loops; Oe.—Oesophagus; Pr.C.—Prostatic capsule; Pr.—Prostate; Pr.D.—Prostatic duct; Sep.—Septum; Sp.A.—Spermathecal ampulla; Sp.Atr.G.—Spermathecal atrial gland; Sp.D.—Spermathecal duct; S.M.A.—Secondary male aperture; Sp.P.—Spermathecal pore; T.S.—Testis sac; Vd—Vas deferens." figureDoi="http://doi.org/10.5281/zenodo.4636246" httpUri="https://zenodo.org/record/4636246/files/figure.png" pageId="7" pageNumber="388">Fig. 3A</figureCitation>
). Female pores paired, minute, at intersegmental furrow 11/12, slightly median to
<emphasis id="B903532EE907FFFCDBE2FD10CBC6FD6D" box="[441,455,690,716]" italics="true" pageId="7" pageNumber="388">b</emphasis>
setal lines. Nephridiopores minute, present from segment 3, aligned with
<emphasis id="B903532EE907FFFCDF54FD10CF1CFD6D" box="[1295,1309,690,716]" italics="true" pageId="7" pageNumber="388">d</emphasis>
setal lines. Genital markings absent.
</paragraph>
<paragraph id="8BC88F3CE907FFFCDA9CFD58C909FB65" blockId="7.[151,1437,150,1869]" pageId="7" pageNumber="388">
Internal: Septum 5/6 slightly muscular, septa 6/7/8/9 muscular, septum 9/10 delicate. Gizzards four in segments 1419; intestinal origin in segments 2729. Commissures of extra-oesophageal vessels present on posterior faces of septa 8/9 and 9/10. Testis sacs paired, in segments 911 on one side and dislocated to segments 1314 on the other side but retaining connection with septum 9/10 through a narrow tube like membranous structure; vas deferens in segments 910, coiled into a mass of hairpin loops, mass as large as testis sac, entering near the ectal end of the prostate gland directly without entering musculature. Prostates paired, extending from segment 11 to segments 1314, glandular, tubular and somewhat flattened with irregular margins (
<figureCitation id="134C93B9E907FFFCD928FC70C9CBFC4D" box="[883,970,978,1004]" captionStart="FIGURE 3" captionStartId="6.[151,250,1777,1801]" captionTargetBox="[169,1422,783,1746]" captionTargetId="figure-269@6.[160,1428,777,1754]" captionTargetPageId="6" captionText="FIGURE 3. Moniligaster keralensis Narayanan &amp; Julka sp. nov. A. Ventral view. B. Prostate of right hand side, dorsal view. C. Prostatic capsule of right hand side, dorsal view. D. Spermatheca of left hand side. Abbreviations. M.H.L.—Mass of hairpin loops; Oe.—Oesophagus; Pr.C.—Prostatic capsule; Pr.—Prostate; Pr.D.—Prostatic duct; Sep.—Septum; Sp.A.—Spermathecal ampulla; Sp.Atr.G.—Spermathecal atrial gland; Sp.D.—Spermathecal duct; S.M.A.—Secondary male aperture; Sp.P.—Spermathecal pore; T.S.—Testis sac; Vd—Vas deferens." figureDoi="http://doi.org/10.5281/zenodo.4636246" httpUri="https://zenodo.org/record/4636246/files/figure.png" pageId="7" pageNumber="388">Fig. 3B</figureCitation>
); capsule tubular, slightly enlarged ectally before its junction with prostatic duct (
<figureCitation id="134C93B9E907FFFCD80FFC54C8AFFBB1" box="[596,686,1014,1040]" captionStart="FIGURE 3" captionStartId="6.[151,250,1777,1801]" captionTargetBox="[169,1422,783,1746]" captionTargetId="figure-269@6.[160,1428,777,1754]" captionTargetPageId="6" captionText="FIGURE 3. Moniligaster keralensis Narayanan &amp; Julka sp. nov. A. Ventral view. B. Prostate of right hand side, dorsal view. C. Prostatic capsule of right hand side, dorsal view. D. Spermatheca of left hand side. Abbreviations. M.H.L.—Mass of hairpin loops; Oe.—Oesophagus; Pr.C.—Prostatic capsule; Pr.—Prostate; Pr.D.—Prostatic duct; Sep.—Septum; Sp.A.—Spermathecal ampulla; Sp.Atr.G.—Spermathecal atrial gland; Sp.D.—Spermathecal duct; S.M.A.—Secondary male aperture; Sp.P.—Spermathecal pore; T.S.—Testis sac; Vd—Vas deferens." figureDoi="http://doi.org/10.5281/zenodo.4636246" httpUri="https://zenodo.org/record/4636246/files/figure.png" pageId="7" pageNumber="388">Fig. 3C</figureCitation>
); prostatic duct bent downwards, narrowed and slightly muscular, much shorter than gland, about one ninth of the combined length of the gland and duct. Spermathecae paired in segment 8, ampulla elongate-ovoidal, each with a long coiled duct penetrating septum 7/8 to discharge at junction of ducts of dichotomous atrial glands in segment 7; atrial glands bound together with one on top of the other (
<figureCitation id="134C93B9E907FFFCDF34FBC0CABDFB01" captionStart="FIGURE 3" captionStartId="6.[151,250,1777,1801]" captionTargetBox="[169,1422,783,1746]" captionTargetId="figure-269@6.[160,1428,777,1754]" captionTargetPageId="6" captionText="FIGURE 3. Moniligaster keralensis Narayanan &amp; Julka sp. nov. A. Ventral view. B. Prostate of right hand side, dorsal view. C. Prostatic capsule of right hand side, dorsal view. D. Spermatheca of left hand side. Abbreviations. M.H.L.—Mass of hairpin loops; Oe.—Oesophagus; Pr.C.—Prostatic capsule; Pr.—Prostate; Pr.D.—Prostatic duct; Sep.—Septum; Sp.A.—Spermathecal ampulla; Sp.Atr.G.—Spermathecal atrial gland; Sp.D.—Spermathecal duct; S.M.A.—Secondary male aperture; Sp.P.—Spermathecal pore; T.S.—Testis sac; Vd—Vas deferens." figureDoi="http://doi.org/10.5281/zenodo.4636246" httpUri="https://zenodo.org/record/4636246/files/figure.png" pageId="7" pageNumber="388">Fig. 3D</figureCitation>
), duct of each gland, almost as long as the common atrial duct. Ovarian chamber complete, ovisacs in segments 1216. Nephridia avesiculate; functional at segment 10.
</paragraph>
</subSubSection>
<subSubSection id="C36DDCB7E907FFFCDA9CFB6CCF84FAF5" pageId="7" pageNumber="388" type="etymology">
<paragraph id="8BC88F3CE907FFFCDA9CFB6CCB8FFAAD" blockId="7.[151,1437,150,1869]" pageId="7" pageNumber="388">
<emphasis id="B903532EE907FFFCDA9CFB6CCB4EFB48" bold="true" box="[199,335,1230,1257]" pageId="7" pageNumber="388">Etymology.</emphasis>
The specific epithet
<emphasis id="B903532EE907FFFCD818FB6CC8BFFB49" box="[579,702,1230,1256]" italics="true" pageId="7" pageNumber="388">keralensis</emphasis>
is derived from the name of the Indian state of
<collectingRegion id="49B341DEE907FFFCDE8BFB6CCF1CFB49" box="[1232,1309,1230,1256]" country="India" name="Kerala" pageId="7" pageNumber="388">Kerala</collectingRegion>
, where this species was collected.
</paragraph>
<paragraph id="8BC88F3CE907FFFCDA9CFAB4CEA7FA91" blockId="7.[151,1437,150,1869]" box="[199,1190,1302,1329]" pageId="7" pageNumber="388">
<emphasis id="B903532EE907FFFCDA9CFAB4CB24FA90" bold="true" box="[199,293,1302,1329]" pageId="7" pageNumber="388">Ingesta.</emphasis>
Colloid of fine soil (silt) with tiny pieces of quartz and brown organic particles.
</paragraph>
<paragraph id="8BC88F3CE907FFFCDA9CFA98CF84FAF5" blockId="7.[151,1437,150,1869]" box="[199,1413,1338,1365]" pageId="7" pageNumber="388">
<emphasis id="B903532EE907FFFCDA9CFA98CB2BFAF4" bold="true" box="[199,298,1338,1365]" pageId="7" pageNumber="388">Habitat.</emphasis>
Evergreen and semi-evergreen forests and plantations of cardamom, black pepper, teak and rubber.
</paragraph>
</subSubSection>
<subSubSection id="C36DDCB7E907FFFCDA9CFAFCCB77FA3C" pageId="7" pageNumber="388" type="biology_ecology">
<paragraph id="8BC88F3CE907FFFCDA9CFAFCCB77FA3C" blockId="7.[151,1437,150,1869]" pageId="7" pageNumber="388">
<emphasis id="B903532EE907FFFCDA9CFAFCCB29FAD8" bold="true" box="[199,296,1374,1401]" pageId="7" pageNumber="388">Biology.</emphasis>
The species shows serpentine movement when disturbed. Autotomy and regeneration of lost parts appear to be common.
</paragraph>
</subSubSection>
<subSubSection id="C36DDCB7E907FFFCDA9CFA04CEFDF9A9" pageId="7" pageNumber="388" type="distribution">
<paragraph id="8BC88F3CE907FFFCDA9CFA04CEFDF9A9" blockId="7.[151,1437,150,1869]" pageId="7" pageNumber="388">
<emphasis id="B903532EE907FFFCDA9CFA04CB60FA61" bold="true" box="[199,353,1446,1472]" pageId="7" pageNumber="388">Distribution.</emphasis>
<collectingCountry id="F360CFACE907FFFCDB3CFA04CBA4FA61" box="[359,421,1446,1472]" name="India" pageId="7" pageNumber="388">India</collectingCountry>
:
<collectingRegion id="49B341DEE907FFFCDBF4FA04C83DFA61" box="[431,572,1446,1472]" country="India" name="Kerala" pageId="7" pageNumber="388">Kerala State</collectingRegion>
: widespread in the middle and higher altitude regions of Idukki District: Chathurangappara, Peruvanthanam; Pathanamthitta District: Aluvamkudy, Chengara, Kurichikanam, Mundomoozhy, Pananthodu, Thannithodumoozhy; and Kollam District: Kottavasal, near to Ambanad estate (
<figureCitation id="134C93B9E907FFFCDEF2FA4CCEEEF9A9" box="[1193,1263,1518,1544]" captionStart="FIGURE 4" captionStartId="8.[151,250,1672,1696]" captionTargetBox="[218,1369,192,1640]" captionTargetId="figure-21@8.[206,1381,181,1648]" captionTargetPageId="8" captionText="FIGURE 4. Map of sampling localities of species of Moniligaster recorded in this study." figureDoi="http://doi.org/10.5281/zenodo.4636250" httpUri="https://zenodo.org/record/4636250/files/figure.png" pageId="7" pageNumber="388">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36DDCB7E907FFFCDA9CF9B0C977F8ED" pageId="7" pageNumber="388" type="discussion">
<paragraph id="8BC88F3CE907FFFCDA9CF9B0C977F8ED" blockId="7.[151,1437,150,1869]" pageId="7" pageNumber="388">
<emphasis id="B903532EE907FFFCDA9CF9B0CB3AF98D" bold="true" box="[199,315,1554,1580]" pageId="7" pageNumber="388">Remarks.</emphasis>
<taxonomicName id="4C77F4BFE907FFFCDB1EF9B0C853F98D" authority="Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka, 2021" authorityName="Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka" authorityYear="2021" box="[325,594,1554,1580]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Moniligastrida" pageId="7" pageNumber="388" phylum="Annelida" rank="species" species="keralensis" status="sp. nov.">
<emphasis id="B903532EE907FFFCDB1EF9B0C853F98D" box="[325,594,1554,1580]" italics="true" pageId="7" pageNumber="388">Moniligaster keralensis</emphasis>
</taxonomicName>
<emphasis id="B903532EE907FFFCD807F9B0C8B9F98D" bold="true" box="[604,696,1554,1580]" pageId="7" pageNumber="388">
<taxonomicNameLabel id="A230EE55E907FFFCD807F9B0C8B9F98D" box="[604,696,1554,1580]" pageId="7" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to the same group of species as
<taxonomicName id="4C77F4BFE907FFFCDEC7F9B1CEFEF98D" authority="Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka, 2021" authorityName="Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka" authorityYear="2021" box="[1180,1279,1554,1580]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Moniligastrida" pageId="7" pageNumber="388" phylum="Annelida" rank="species" species="bahli" status="sp. nov.">
<emphasis id="B903532EE907FFFCDEC7F9B1CEFEF98D" box="[1180,1279,1554,1580]" italics="true" pageId="7" pageNumber="388">M. bahli</emphasis>
</taxonomicName>
<emphasis id="B903532EE907FFFCDF53F9B0CF64F98D" bold="true" box="[1288,1381,1554,1580]" pageId="7" pageNumber="388">
<taxonomicNameLabel id="A230EE55E907FFFCDF53F9B0CF64F98D" box="[1288,1381,1554,1580]" pageId="7" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(see remarks there) and
<taxonomicName id="4C77F4BFE907FFFCDB28F995C817F9F1" authority="Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka, 2021" authorityName="Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka" authorityYear="2021" box="[371,534,1590,1616]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Moniligastrida" pageId="7" pageNumber="388" phylum="Annelida" rank="species" species="blakemorei" status="sp. nov.">
<emphasis id="B903532EE907FFFCDB28F995C817F9F1" box="[371,534,1590,1616]" italics="true" pageId="7" pageNumber="388">M. blakemorei</emphasis>
</taxonomicName>
<emphasis id="B903532EE907FFFCD845F994C879F9F1" bold="true" box="[542,632,1590,1616]" pageId="7" pageNumber="388">
<taxonomicNameLabel id="A230EE55E907FFFCD845F994C879F9F1" box="[542,632,1590,1616]" pageId="7" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. The species are compared in
<tableCitation id="C6F5BA87E907FFFCD98BF994CE25F9F1" box="[976,1060,1590,1616]" captionStart="TABLE 1" captionStartId="9.[130,218,623,649]" captionTargetBox="[144,2009,736,1524]" captionTargetPageId="9" captionText="TABLE 1.Comparison of characters among the three new Moniligaster species and two closely related species. These five species share the following characters: (1) vas deferens" httpUri="http://table.plazi.org/id/DF08DFB4E909FFF2DAD9FDCDCAACFD70" pageId="7" pageNumber="388" tableUuid="DF08DFB4E909FFF2DAD9FDCDCAACFD70">Table 1</tableCitation>
. Differences of
<taxonomicName id="4C77F4BFE907FFFCDE80F995CF72F9F1" authority="Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka, 2021" authorityName="Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka" authorityYear="2021" box="[1243,1395,1590,1616]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Moniligastrida" pageId="7" pageNumber="388" phylum="Annelida" rank="species" species="keralensis" status="sp. nov.">
<emphasis id="B903532EE907FFFCDE80F995CF72F9F1" box="[1243,1395,1590,1616]" italics="true" pageId="7" pageNumber="388">M. keralensis</emphasis>
</taxonomicName>
<emphasis id="B903532EE907FFFCDF20F994CAC9F9D5" bold="true" pageId="7" pageNumber="388">
<taxonomicNameLabel id="A230EE55E907FFFCDF20F994CAC9F9D5" pageId="7" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
to other species of the group are as follows:
<taxonomicName id="4C77F4BFE907FFFCD895F9F9C966F9D5" authority="Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka, 2021" authorityName="Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka" authorityYear="2021" box="[718,871,1626,1652]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Moniligastrida" pageId="7" pageNumber="388" phylum="Annelida" rank="species" species="keralensis" status="sp. nov.">
<emphasis id="B903532EE907FFFCD895F9F9C966F9D5" box="[718,871,1626,1652]" italics="true" pageId="7" pageNumber="388">M. keralensis</emphasis>
</taxonomicName>
<emphasis id="B903532EE907FFFCD934F9F8C9CBF9D5" bold="true" box="[879,970,1626,1652]" pageId="7" pageNumber="388">
<taxonomicNameLabel id="A230EE55E907FFFCD934F9F8C9CBF9D5" box="[879,970,1626,1652]" pageId="7" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from
<taxonomicName id="4C77F4BFE907FFFCDE91F9F9CF4CF9D5" authorityName="Stephenson" authorityYear="1915" box="[1226,1357,1626,1652]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Moniligastrida" pageId="7" pageNumber="388" phylum="Annelida" rank="species" species="gravelyi">
<emphasis id="B903532EE907FFFCDE91F9F9CF4CF9D5" box="[1226,1357,1626,1652]" italics="true" pageId="7" pageNumber="388">M. gravelyi</emphasis>
</taxonomicName>
by the junction of vas deferens with the prostate glands at ectal rather than at ental end and intestinal origin (in segments 2729 vs segment 25).
<taxonomicName id="4C77F4BFE907FFFCDBFAF901C838F91D" authority="Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka, 2021" authorityName="Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka" authorityYear="2021" box="[417,569,1698,1724]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Moniligastrida" pageId="7" pageNumber="388" phylum="Annelida" rank="species" species="keralensis" status="sp. nov.">
<emphasis id="B903532EE907FFFCDBFAF901C838F91D" box="[417,569,1698,1724]" italics="true" pageId="7" pageNumber="388">M. keralensis</emphasis>
</taxonomicName>
<emphasis id="B903532EE907FFFCD81AF900C89DF91D" bold="true" box="[577,668,1698,1724]" pageId="7" pageNumber="388">
<taxonomicNameLabel id="A230EE55E907FFFCD81AF900C89DF91D" box="[577,668,1698,1724]" pageId="7" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C77F4BFE907FFFCD968F901C98FF91D" authorityName="Jamieson" authorityYear="1977" box="[819,910,1699,1724]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Moniligastrida" pageId="7" pageNumber="388" phylum="Annelida" rank="species" species="troyi">
<emphasis id="B903532EE907FFFCD968F901C98FF91D" box="[819,910,1699,1724]" italics="true" pageId="7" pageNumber="388">M. troyi</emphasis>
</taxonomicName>
in the shape of the prostate glands (tubular vs club-shaped), body dimensions (length
<quantity id="4C8F22D9E907FFFCD80AF964C8D9F940" box="[593,728,1734,1761]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="1.29" metricValueMin="0.71" pageId="7" pageNumber="388" unit="mm" value="100.0" valueMax="129.0" valueMin="71.0">71129 mm</quantity>
vs
<quantity id="4C8F22D9E907FFFCD8A5F964C978F940" box="[766,889,1734,1761]" metricMagnitude="-2" metricUnit="m" metricValue="6.15" metricValueMax="6.3" metricValueMin="6.0" pageId="7" pageNumber="388" unit="mm" value="61.5" valueMax="63.0" valueMin="60.0">6063 mm</quantity>
; diameter
<quantity id="4C8F22D9E907FFFCD9B6F964CE5EF940" box="[1005,1119,1734,1761]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.5" metricValueMin="4.0" pageId="7" pageNumber="388" unit="mm" value="4.75" valueMax="5.5" valueMin="4.0">45.5 mm</quantity>
vs
<quantity id="4C8F22D9E907FFFCDEDEF964CF11F940" box="[1157,1296,1734,1761]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" metricValueMax="3.7" metricValueMin="3.5" pageId="7" pageNumber="388" unit="mm" value="3.6" valueMax="3.7" valueMin="3.5">3.53.7 mm</quantity>
), junction of vas deferens with the prostate glands (at ectal end vs about middle) and length of atrial gland duct (almost as long as common atrial duct vs about one fourth the length of common atrial duct). For differences of
<taxonomicName id="4C77F4BFE907FFFCDEF6F8ADCF0CF889" authority="Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka, 2021" authorityName="Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka" authorityYear="2021" box="[1197,1293,1806,1832]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Moniligastrida" pageId="7" pageNumber="388" phylum="Annelida" rank="species" species="bahli" status="sp. nov.">
<emphasis id="B903532EE907FFFCDEF6F8ADCF0CF889" box="[1197,1293,1806,1832]" italics="true" pageId="7" pageNumber="388">M. bahli</emphasis>
</taxonomicName>
<emphasis id="B903532EE907FFFCDF48F8ACCF6CF889" bold="true" box="[1299,1389,1806,1832]" pageId="7" pageNumber="388">
<taxonomicNameLabel id="A230EE55E907FFFCDF48F8ACCF6CF889" box="[1299,1389,1806,1832]" pageId="7" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C77F4BFE907FFFCDACCF891CB3BF8ED" authority="Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka, 2021" authorityName="Narayanan &amp; Sathrumithra &amp; Anuja &amp; Christopher &amp; Thomas &amp; Julka" authorityYear="2021" box="[151,314,1842,1868]" class="Clitellata" family="Moniligastridae" genus="Moniligaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Moniligastrida" pageId="7" pageNumber="388" phylum="Annelida" rank="species" species="blakemorei" status="sp. nov.">
<emphasis id="B903532EE907FFFCDACCF891CB3BF8ED" box="[151,314,1842,1868]" italics="true" pageId="7" pageNumber="388">M. blakemorei</emphasis>
</taxonomicName>
<emphasis id="B903532EE907FFFCDB1BF890CB9BF8ED" bold="true" box="[320,410,1842,1868]" pageId="7" pageNumber="388">
<taxonomicNameLabel id="A230EE55E907FFFCDB1BF890CB9BF8ED" box="[320,410,1842,1868]" pageId="7" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, see the remarks sections of these species.
</paragraph>
</subSubSection>
</treatment>
</document>