treatments-xml/data/03/D4/87/03D487F6FFCD187268C05B8CFD7EFCF0.xml

<document id="CA7BA7AD22745E3AD77CEB07869877CB" ID-DOI="10.11646/phytotaxa.284.3.7" ID-ISSN="1179-3163" ID-Zenodo-Dep="13647834" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725378612314" checkinUser="felipe" docAuthor="Pereira-Silva, Luciana, Hefler, Sonia Marisa &amp; Trevisan, Rafael" docDate="2016" docId="03D487F6FFCD187268C05B8CFD7EFCF0" docLanguage="en" docName="phytotaxa.284.3.7.pdf" docOrigin="Phytotaxa 284 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.284.3.7" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Cyperus tuckerianus L. P. Silva, Hefler &amp; R. Trevisan 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="222" masterDocId="FFEDFF8EFFCC187668485878FFCBFFC0" masterDocTitle="Cyperus tuckerianus (Cyperaceae), a new species from the Central-West Region of Brazil" masterLastPageNumber="224" masterPageNumber="218" pageNumber="219" updateTime="1725385488073" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C881B2B407B35868DBF611BE272BD7A4">Cyperus tuckerianus (Cyperaceae), a new species from the Central-West Region of Brazil</mods:title>
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<mods:namePart id="43997066BDD6E91814D2CB9BB4D51842">Pereira-Silva, Luciana</mods:namePart>
<mods:affiliation id="DAC26416E744B523294699E375EC720D">Programa de Pós-graduação em Biologia de Fungos, Algas e Plantas, Centro de Ciências Biológicas, Departamento de Botânica, Universidade Federal de Santa Catarina, Campus Reitor João David Ferreira Lima, 88040 - 900, Florianópolis, Santa Catarina, Brazil.</mods:affiliation>
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<mods:namePart id="FCCB97C62825E74A5B661B1236F453C5">Hefler, Sonia Marisa</mods:namePart>
<mods:affiliation id="F64B6B5959E962EF609CF1A945696B8D">Universidade Federal do Rio Grande, Instituto de Ciências Biológicas, 96203 - 900, Rio Grande, Rio Grande do Sul, Brazil.</mods:affiliation>
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<mods:namePart id="11BD954C7E925CBA5E942CD526A85B5D">Trevisan, Rafael</mods:namePart>
<mods:affiliation id="1BCD85A7498D0BEFD7F210092F40B739">Programa de Pós-graduação em Biologia de Fungos, Algas e Plantas, Centro de Ciências Biológicas, Departamento de Botânica, Universidade Federal de Santa Catarina, Campus Reitor João David Ferreira Lima, 88040 - 900, Florianópolis, Santa Catarina, Brazil.</mods:affiliation>
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<subSubSection id="C367656BFFCD187768C05B8CFC8CFBCF" box="[136,839,1012,1039]" pageId="1" pageNumber="219" type="nomenclature">
<paragraph id="8BC236E0FFCD187768C05B8CFC8CFBCF" blockId="1.[136,839,1012,1039]" box="[136,839,1012,1039]" pageId="1" pageNumber="219">
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<taxonomicName id="4C7D4D63FFCD187768C05B8CFD23FBCF" ID-CoL="33DR9" authority="L.P. Silva, Hefler &amp; R. Trevisan" authorityName="L. P. Silva, Hefler &amp; R. Trevisan" authorityYear="2016" box="[136,744,1012,1039]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="1" pageNumber="219" phylum="Tracheophyta" rank="species" species="tuckerianus" status="sp. nov.">
<emphasis id="B909EAF2FFCD187768C05B8CFEBCFBCF" bold="true" box="[136,375,1012,1039]" italics="true" pageId="1" pageNumber="219">Cyperus tuckerianus</emphasis>
L.P. Silva, Hefler &amp; R. Trevisan
</taxonomicName>
, 
<emphasis id="B909EAF2FFCD18776ABA5B8EFC8CFBCF" box="[754,839,1014,1039]" italics="true" pageId="1" pageNumber="219">
<taxonomicNameLabel id="A23A5789FFCD18776ABA5B8EFC8CFBCF" box="[754,839,1014,1039]" pageId="1" pageNumber="219" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C367656BFFCD187768C05C38FD7BFBB9" pageId="1" pageNumber="219" type="materials_examined">
<paragraph id="8BC236E0FFCD187768C05C38FD7BFBB9" blockId="1.[136,1452,1088,1253]" pageId="1" pageNumber="219">
<emphasis id="B909EAF2FFCD187768C05C38FF09FB95" bold="true" box="[136,194,1088,1109]" pageId="1" pageNumber="219">Type:</emphasis>
— 
<materialsCitation id="3B153CBDFFCD187768935C38FD7BFBB9" collectingDate="2016-01-16" collectorName="L. P. Silva" country="Brazil" latitude="-15.845278" location="Poxoreu" pageId="1" pageNumber="219" specimenCount="2" stateProvince="Mato Grosso" typeStatus="holotype">
<collectingCountry id="F36A7670FFCD187768935C38FEFFFB95" box="[219,308,1088,1109]" name="Brazil" pageId="1" pageNumber="219">BRAZIL</collectingCountry>
: 
<collectingRegion id="49B9F802FFCD187769765C38FE74FB95" box="[318,447,1088,1109]" country="Brazil" name="Mato Grosso" pageId="1" pageNumber="219">Mato Grosso</collectingRegion>
: 
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: flooded site, 
<geoCoordinate id="EE495027FFCD18776AEB5C38FCDEFB95" box="[675,789,1088,1109]" degrees="15" direction="south" minutes="50" orientation="latitude" pageId="1" pageNumber="219" precision="15" seconds="43" value="-15.845278">15°50’43”S</geoCoordinate>
54°24.1’54”W, 
<date id="FFC31020FFCD18776BFB5C38FB9FFB95" box="[947,1108,1088,1109]" pageId="1" pageNumber="219" value="2016-01-16">
<collectingDate id="EF87E9C8FFCD18776BFB5C38FB9FFB95" box="[947,1108,1088,1109]" pageId="1" pageNumber="219" value="2016-01-16">16 January 2016</collectingDate>
</date>
, fl. and fr., 
<emphasis id="B909EAF2FFCD18776C8D5C38FA82FB95" box="[1221,1353,1088,1109]" italics="true" pageId="1" pageNumber="219">
<collectorName id="26885336FFCD18776C8D5C38FAEBFB95" box="[1221,1312,1088,1109]" pageId="1" pageNumber="219">L.P. Silva</collectorName>
119
</emphasis>
(
<typeStatus id="54C68842FFCD18776D1E5C38FA67FB95" box="[1366,1452,1088,1109]" pageId="1" pageNumber="219" type="holotype">holotype</typeStatus>
FLOR!; 
<typeStatus id="54C68842FFCD187769475C1CFE95FBB9" box="[271,350,1124,1145]" pageId="1" pageNumber="219" type="isotype">isotypes</typeStatus>
UFMT!, RB!, NY!). (
<figureCitation id="13462A65FFCD18776A7C5C1CFD63FBB9" box="[564,680,1124,1145]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[136,229,1799,1820]" captionStartId-1="3.[136,229,1238,1259]" captionTargetBox-0="[155,1432,353,1774]" captionTargetBox-1="[153,1435,191,1213]" captionTargetId-0="figure-104@2.[155,1432,353,1774]" captionTargetId-1="figure-18@3.[152,1436,190,1213]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Illustrations of Cyperus tuckerianus (based on L.P. Silva 119). A. Habit. B. Spikelet. C. Achene. D. Surface of achene. E. Detail of surface of the achene with isodiametric cells and prominent silica bodies. Illustrated by Luciana P. Silva." captionText-1="FIGURE 2. Photographs of Cyperus tuckerianus in its natural habitat in the state of Mato Grosso, Brazil. A. Habit. B. Inflorescence. C. Detail of the culm base (paratype). D. Detail of the elongated internode (black arrow) at the culm base (holotype). All photographs by Luciana P. Silva." figureDoi-0="http://doi.org/10.5281/zenodo.13647838" figureDoi-1="http://doi.org/10.5281/zenodo.13647842" httpUri-0="https://zenodo.org/record/13647838/files/figure.png" httpUri-1="https://zenodo.org/record/13647842/files/figure.png" pageId="1" pageNumber="219">Figures 1–2</figureCitation>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C367656BFFCD187768C05CF0FDAAFB25" pageId="1" pageNumber="219" type="diagnosis">
<paragraph id="8BC236E0FFCD187768C05CF0FDAAFB25" blockId="1.[136,1452,1088,1253]" pageId="1" pageNumber="219">
<emphasis id="B909EAF2FFCD187768C05CF0FF20FB5D" bold="true" box="[136,235,1160,1181]" pageId="1" pageNumber="219">Diagnosis</emphasis>
: 
<emphasis id="B909EAF2FFCD187768BB5CF0FDAAFB25" italics="true" pageId="1" pageNumber="219">
<emphasis id="B909EAF2FFCD187768BB5CF0FEC0FB5D" bold="true" box="[243,267,1160,1181]" italics="true" pageId="1" pageNumber="219">—</emphasis>
Caespitose herb, annual to short perennial; 57.0–79.0 cm high, culm trigonous; inflorescence lax anthelodium simple or compound; vinaceous prophyll; spikelets digitiform or grouped in glomerules; spikelets 7.5–13.0× 
<quantity id="4C859B05FFCD18776C385CD4FB2EFB01" box="[1136,1253,1196,1217]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.2" metricValueMin="2.3" pageId="1" pageNumber="219" unit="mm" value="2.75" valueMax="3.2" valueMin="2.3">2.3–3.2 mm</quantity>
; glumes 
<quantity id="4C859B05FFCD18776D735CD4FA67FB01" box="[1339,1452,1196,1217]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.7" metricValueMin="2.1" pageId="1" pageNumber="219" unit="mm" value="2.4" valueMax="2.7" valueMin="2.1">2.1–2.7 mm</quantity>
long; achenes obovoid to elliptic, biconvex.
</emphasis>
</paragraph>
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<subSubSection id="C367656BFFCD187768C05D6EFC9EF88B" pageId="1" pageNumber="219" type="description">
<paragraph id="8BC236E0FFCD187768C05D6EFC9EF88B" blockId="1.[136,1452,1302,2083]" pageId="1" pageNumber="219">
<emphasis id="B909EAF2FFCD187768C05D6EFED9FAEF" bold="true" box="[136,274,1302,1327]" pageId="1" pageNumber="219">Description</emphasis>
: 
<emphasis id="B909EAF2FFCD187769525D6EFEFDFAEF" bold="true" box="[282,310,1302,1327]" pageId="1" pageNumber="219">—</emphasis>
Caespitose herb, annual to short perennial; 57.0–79.0 cm high, with thin roots 0.2–1.0 mm diam. Culm 
<quantity id="4C859B05FFCD1877689A5D42FEA4FA93" box="[210,367,1338,1363]" metricMagnitude="-1" metricUnit="m" metricValue="5.55" metricValueMax="6.75" metricValueMin="4.35" pageId="1" pageNumber="219" unit="cm" value="55.5" valueMax="67.5" valueMin="43.5">43.5–67.5 cm</quantity>
long, 2.0–3.0 mm diam., trigonous, smooth, glabrous, acute angles, transverse septa absent. Leaf blade (1.5–) 
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× 3.4–5.0 mm, linear-lanceolate, flat, papyraceous-chartaceous, glabrous, greenish, with scaber antrorse in the apical portion of the margin and in the abaxial central vein, acute apex; basal leaves reduced to a sheath; leaf sheath 6.0–14.0 cm long, membranous to papyraceous, glabrous, grooved surface. Bracts of the inflorescence 3–4, (1.7–)2.5–24.0 cm × (1.1–)1.6–5.0 mm. Terminal anthelodium lax from simple to compound, 4.8–14.5 x 4.0–16.0 cm. Primary rays (2–)4–8; the basal one 3.5–12.0 cm long, surrounded by prophyllum 
<quantity id="4C859B05FFCD18776D015D96FF7FF9EB" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="17.5" metricValueMin="0.5" pageId="1" pageNumber="219" unit="mm" value="9.0" valueMax="17.5" valueMin="0.5">0.5–17.5 mm</quantity>
long, vinaceous. Secondary bracts inconspicuous. Secondary rays present or absent, 
<quantity id="4C859B05FFCD18776C325E6AFB2FF9EB" box="[1146,1252,1554,1579]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="3.0" metricValueMin="0.7" pageId="1" pageNumber="219" unit="cm" value="1.85" valueMax="3.0" valueMin="0.7">0.7–3 cm</quantity>
long, surrounded by vinaceous prophyllum, inconspicuous. Spikelets 7–25, digitiform or grouped in glomerules 1.1–2.5 × 
<quantity id="4C859B05FFCD18776D1F5E4EFF65F9B3" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.5" metricValueMin="1.5" pageId="1" pageNumber="219" unit="cm" value="2.5" valueMax="3.5" valueMin="1.5">1.5–3.5 cm</quantity>
. Spikelets 7.5–13.0 × 
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, narrow-elliptical to lanceolate. Rachilla 
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long, slightly flexuose, not articulate, not winged, visible or inconspicuous. Glume 9–22, 2.1–2.7 × 
<quantity id="4C859B05FFCD18776BBD5E06FB4BF957" box="[1013,1152,1662,1687]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="1" pageNumber="219" unit="mm" value="1.5" valueMax="1.7" valueMin="1.3">1.3–1.7 mm</quantity>
, 
<quantity id="4C859B05FFCD18776CC65E06FADCF957" box="[1166,1303,1662,1687]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="11.0" metricValueMin="7.0" pageId="1" pageNumber="219" unit="mm" value="0.9" valueMax="1.1" valueMin="0.7">0.7–1.1 mm</quantity>
wide in side view, ovate-lanceolate, navicular, glabrous, brown-yellow to dark-gold, 5–6 veins (including the keel), keels usually greenish or sometimes with fine vinaceous lines, apex acute muticous; sterile glumes present or absent, membranous, vinaceous. Stamen 2, anthers 0.5–0.8 x 
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, yellow. Styles (0.3–) 
<quantity id="4C859B05FFCD18776B805E92FB9AF8C3" box="[968,1105,1770,1795]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="1" pageNumber="219" unit="mm" value="1.0" valueMax="1.5" valueMin="0.5">0.5–1.5 mm</quantity>
long; stigmas 2, (1.1–)2.0– 
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long. Achene 1.1–1.3 × 0.7–1.0 mm, obovoid to elliptic, biconvex, apex slightly apiculate, dark-brown to black, surface puncticulate, shinny, completely covered by the glumes.
</paragraph>
</subSubSection>
<subSubSection id="C367656BFFCD187768F55F2EFB06F877" pageId="1" pageNumber="219" type="distribution">
<paragraph id="8BC236E0FFCD187768F55F2EFB06F877" blockId="1.[136,1452,1302,2083]" pageId="1" pageNumber="219">
<emphasis id="B909EAF2FFCD187768F55F2EFDCEF8AF" bold="true" box="[189,517,1878,1903]" pageId="1" pageNumber="219">Distribution and habitat:—</emphasis>
Presently this species is known only for the states of 
<collectingRegion id="49B9F802FFCD18776C1B5F2EFB2FF8AF" box="[1107,1252,1878,1903]" country="Brazil" name="Mato Grosso" pageId="1" pageNumber="219">Mato Grosso</collectingRegion>
and 
<collectingRegion id="49B9F802FFCD18776D525F2EFF1FF853" country="Brazil" name="Mato Grosso do Sul" pageId="1" pageNumber="219">Mato Grosso do Sul</collectingRegion>
in the Central-West Region of 
<collectingCountry id="F36A7670FFCD18776A015F02FD46F853" box="[585,653,1914,1939]" name="Brazil" pageId="1" pageNumber="219">Brazil</collectingCountry>
(
<figureCitation id="13462A65FFCD18776AE85F02FCCDF853" box="[672,774,1914,1939]" captionStart="FIGURE 3" captionStartId="4.[136,229,1597,1618]" captionTargetBox="[151,1436,841,1572]" captionTargetId="figure-237@4.[151,1436,841,1572]" captionTargetPageId="4" captionText="FIGURE 3. Distribution map of Cyperus tuckerianus in the Central-West Region of Brazil." figureDoi="http://doi.org/10.5281/zenodo.13647846" httpUri="https://zenodo.org/record/13647846/files/figure.png" pageId="1" pageNumber="219">Figure 3</figureCitation>
). The specimens were collected in the phytophysiognomy 
<emphasis id="B909EAF2FFCD187768C05FE6FF2CF877" box="[136,231,1950,1975]" italics="true" pageId="1" pageNumber="219">Cerrado</emphasis>
in humid flooded areas and also in the vegetation formation regionally known as 
<emphasis id="B909EAF2FFCD18776C335FE6FB0DF877" box="[1147,1222,1950,1975]" italics="true" pageId="1" pageNumber="219">vereda</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C367656BFFCD187268F55FBAFD7EFCF0" lastPageId="4" lastPageNumber="222" pageId="1" pageNumber="219" type="etymology">
<paragraph id="8BC236E0FFCD187768F55FBAFC29F83F" blockId="1.[136,1452,1302,2083]" pageId="1" pageNumber="219">
<emphasis id="B909EAF2FFCD187768F55FBAFEAFF81B" bold="true" box="[189,356,1986,2011]" pageId="1" pageNumber="219">Etymology:—</emphasis>
The specific epithet is given in honor of Dr. Gordon Charles Tucker, researcher of Eastern Illinois University, for his great contribution to the knowledge about genus 
<taxonomicName id="4C7D4D63FFCD18776B365F9EFC10F83F" authorityName="Linnaeus" authorityYear="1753" box="[894,987,2022,2047]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="1" pageNumber="219" phylum="Tracheophyta" rank="genus">
<emphasis id="B909EAF2FFCD18776B365F9EFC10F83F" box="[894,987,2022,2047]" italics="true" pageId="1" pageNumber="219">Cyperus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC236E0FFCD187468F55072FD21FE89" blockId="1.[136,1452,1302,2083]" lastBlockId="2.[136,1452,158,329]" lastPageId="2" lastPageNumber="220" pageId="1" pageNumber="219">
<emphasis id="B909EAF2FFCD187768F55072FE7CF7E3" bold="true" box="[189,439,2058,2083]" pageId="1" pageNumber="219">Conservation status:</emphasis>
—Based on the collection sites it was possible to calculate the Extent of Occurrence of 
<quantity id="4C859B05FFCE187468C058D8FECBFF79" box="[136,256,160,185]" metricMagnitude="7" metricUnit="m" metricValue="2.1166" pageId="2" pageNumber="220" unit="km" value="21166.0">21,166 km</quantity>
<superScript id="7C089BA8FFCE1874694858E6FEC3FF6C" attach="left" box="[256,264,158,172]" fontSize="6" pageId="2" pageNumber="220">2</superScript>
and Area of Occupancy of 
<quantity id="4C859B05FFCE18746A7258D8FD4AFF79" box="[570,641,160,185]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="220" unit="km" value="12.0">12 km</quantity>
<superScript id="7C089BA8FFCE18746AC858E6FD43FF6C" attach="left" box="[640,648,158,172]" fontSize="6" pageId="2" pageNumber="220">2</superScript>
for the species. According to the criteria proposed by 
<bibRefCitation id="EFEC4B11FFCE18746CAE58D8FAB7FF79" author="IUCN" box="[1254,1404,160,185]" pageId="2" pageNumber="220" refId="ref3880" refString="IUCN (2016) Standards and Petitions Subcommittee. Guidelines for Using the IUCN Red List Categories and Criteria. Version 12. Prepared by the Standards and Petitions Subcommittee. Avaiable from: http: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed 20 June 2016)" type="book" year="2016">IUCN (2016)</bibRefCitation>
and evaluation of the habitat available in the region, we can consider that 
<taxonomicName id="4C7D4D63FFCE18746BCA58BCFBEFFF1D" authorityName="L. P. Silva, Hefler &amp; R. Trevisan" authorityYear="2016" box="[898,1060,196,221]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="220" phylum="Tracheophyta" rank="species" species="tuckerianus">
<emphasis id="B909EAF2FFCE18746BCA58BCFBEFFF1D" box="[898,1060,196,221]" italics="true" pageId="2" pageNumber="220">C. tuckerianus</emphasis>
</taxonomicName>
is a species possibly not threatened (LC). The region where the species occurs needs an exploration with better distribution of collections. We believe that the species is under-sampled in the scientific collections and an effort to explore other areas will allow to determine the degree of threat of the species with more accuracy.
</paragraph>
<caption id="DF026668FFCE187468C05F7FFB1BF880" ID-DOI="http://doi.org/10.5281/zenodo.13647838" ID-Zenodo-Dep="13647838" httpUri="https://zenodo.org/record/13647838/files/figure.png" pageId="2" pageNumber="220" startId="2.[136,229,1799,1820]" targetBox="[155,1432,353,1774]" targetPageId="2" targetType="figure">
<paragraph id="8BC236E0FFCE187468C05F7FFB1BF880" blockId="2.[136,1452,1799,1856]" pageId="2" pageNumber="220">
<emphasis id="B909EAF2FFCE187468C05F7FFF34F8DC" bold="true" box="[136,255,1799,1820]" pageId="2" pageNumber="220">FIGURE 1.</emphasis>
Illustrations of 
<taxonomicName id="4C7D4D63FFCE187469D35F7FFDA8F8DC" authorityName="L. P. Silva, Hefler &amp; R. Trevisan" authorityYear="2016" box="[411,611,1799,1820]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="220" phylum="Tracheophyta" rank="species" species="tuckerianus">
<emphasis id="B909EAF2FFCE187469D35F7FFDA8F8DC" box="[411,611,1799,1820]" italics="true" pageId="2" pageNumber="220">Cyperus tuckerianus</emphasis>
</taxonomicName>
(based on 
<emphasis id="B909EAF2FFCE18746A875F7FFC9EF8DC" box="[719,853,1799,1820]" italics="true" pageId="2" pageNumber="220">L.P. Silva 119</emphasis>
). 
<emphasis id="B909EAF2FFCE18746B235F7FFC49F8DC" bold="true" box="[875,898,1799,1820]" pageId="2" pageNumber="220">A.</emphasis>
Habit. 
<emphasis id="B909EAF2FFCE18746B845F7FFC29F8DC" bold="true" box="[972,994,1799,1820]" pageId="2" pageNumber="220">B.</emphasis>
Spikelet. 
<emphasis id="B909EAF2FFCE18746C0D5F7FFB96F8DC" bold="true" box="[1093,1117,1799,1820]" pageId="2" pageNumber="220">C.</emphasis>
Achene. 
<emphasis id="B909EAF2FFCE18746CF35F7FFB19F8DC" bold="true" box="[1211,1234,1799,1820]" pageId="2" pageNumber="220">D.</emphasis>
Surface of achene. 
<emphasis id="B909EAF2FFCE18746DDD5F7FFA67F8DC" bold="true" box="[1429,1452,1799,1820]" pageId="2" pageNumber="220">E.</emphasis>
Detail of surface of the achene with isodiametric cells and prominent silica bodies. Illustrated by Luciana P. Silva.
</paragraph>
</caption>
<paragraph id="8BC236E0FFCE187468F55F08FDD1F835" blockId="2.[136,1452,1904,2037]" pageId="2" pageNumber="220">
<emphasis id="B909EAF2FFCE187468F55F08FDBBF849" bold="true" box="[189,624,1904,1929]" pageId="2" pageNumber="220">
Additional collection (
<typeStatus id="54C68842FFCE187469825F08FD8DF849" box="[458,582,1904,1929]" pageId="2" pageNumber="220" type="paratype">paratypes</typeStatus>
):—
</emphasis>
<materialsCitation id="3B153CBDFFCE18746A385F08FA6CF86D" collectingDate="2012-12-19" collectorName="S. N. Moreira" country="Brazil" latitude="-21.2575" location="Reserva Cisalpina" longLatPrecision="21" longitude="-51.96139" municipality="Brasilandia" pageId="2" pageNumber="220" specimenCount="1" stateProvince="Mato Grosso do Sul">
<collectingCountry id="F36A7670FFCE18746A385F08FD11F849" box="[624,730,1904,1929]" name="Brazil" pageId="2" pageNumber="220">BRAZIL</collectingCountry>
: 
<collectingRegion id="49B9F802FFCE18746AAE5F08FC19F849" box="[742,978,1904,1929]" country="Brazil" name="Mato Grosso do Sul" pageId="2" pageNumber="220">Mato Grosso do Sul</collectingRegion>
, 
<collectingMunicipality id="6BA6AC9AFFCE18746BA85F08FBA9F849" box="[992,1122,1904,1929]" pageId="2" pageNumber="220">Brasilândia</collectingMunicipality>
, 
<location id="8EA2603BFFCE18746C385F08FA8AF849" LSID="urn:lsid:plazi:treatment:03D487F6FFCD187268C05B8CFD7EFCF0:8EA2603BFFCE18746C385F08FA8AF849" box="[1136,1345,1904,1929]" country="Brazil" latitude="-21.2575" longLatPrecision="21" longitude="-51.96139" municipality="Brasilandia" name="Reserva Cisalpina" pageId="2" pageNumber="220" stateProvince="Mato Grosso do Sul">Reserva Cisalpina</location>
, 
<locationDeviation id="9A702FC2FFCE18746D065F08FE60F86D" firstLocation="Brasilandia" pageId="2" pageNumber="220" secondLocation="Pauliceia">between Brasilândia and Paulicéia</locationDeviation>
, 
<geoCoordinate id="EE495027FFCE187469FF5FECFDF7F86D" box="[439,572,1940,1965]" degrees="21" direction="south" minutes="15" orientation="latitude" pageId="2" pageNumber="220" precision="15" seconds="27" value="-21.2575">21°15’27”S</geoCoordinate>
<geoCoordinate id="EE495027FFCE18746A0D5FECFD1FF86D" box="[581,724,1940,1965]" degrees="51" direction="west" minutes="57" orientation="longitude" pageId="2" pageNumber="220" precision="15" seconds="41" value="-51.96139">51°57’41”W</geoCoordinate>
, 
<date id="FFC31020FFCE18746AA95FECFC77F86D" box="[737,956,1940,1965]" pageId="2" pageNumber="220" value="2012-12-19">
<collectingDate id="EF87E9C8FFCE18746AA95FECFC77F86D" box="[737,956,1940,1965]" pageId="2" pageNumber="220" value="2012-12-19">19 December 2012</collectingDate>
</date>
, fl. e fr., 
<emphasis id="B909EAF2FFCE18746C6E5FECFAF7F86D" box="[1062,1340,1940,1965]" italics="true" pageId="2" pageNumber="220">
<collectorName id="26885336FFCE18746C6E5FECFB72F86D" box="[1062,1209,1940,1965]" pageId="2" pageNumber="220">S.N. Moreira</collectorName>
et al. 1303
</emphasis>
(BHCB)
</materialsCitation>
; 
<materialsCitation id="3B153CBDFFCE187468C05FC0FDDDF835" collectingDate="2011-02-11" collectorName="V. J. Pott" country="Brazil" county="Inocencia" latitude="-20.027082" location="Sao Pedro" longLatPrecision="1" longitude="-52.12875" municipality="Fazenda Buriti" pageId="2" pageNumber="220" specimenCount="1">
<collectingCounty id="62A34E6CFFCE187468C05FC0FF32F811" box="[136,249,1976,2001]" pageId="2" pageNumber="220">Inocência</collectingCounty>
: 
<collectingMunicipality id="6BA6AC9AFFCE1874694C5FC0FE66F811" box="[260,429,1976,2001]" pageId="2" pageNumber="220">Fazenda Buriti</collectingMunicipality>
, along the length of córrego 
<location id="8EA2603BFFCE18746ABB5FC0FCA3F811" LSID="urn:lsid:plazi:treatment:03D487F6FFCD187268C05B8CFD7EFCF0:8EA2603BFFCE18746ABB5FC0FCA3F811" box="[755,872,1976,2001]" country="Brazil" county="Inocencia" latitude="-20.027082" longLatPrecision="1" longitude="-52.12875" municipality="Fazenda Buriti" name="Sao Pedro" pageId="2" pageNumber="220">São Pedro</location>
, 
<geoCoordinate id="EE495027FFCE18746B3C5FC0FBC5F811" box="[884,1038,1976,2001]" degrees="20" direction="south" minutes="01" orientation="latitude" pageId="2" pageNumber="220" precision="1" seconds="37.5" value="-20.027082">20°01’37.5”S</geoCoordinate>
<geoCoordinate id="EE495027FFCE18746C5E5FC0FB77F811" box="[1046,1212,1976,2001]" degrees="52" direction="west" minutes="07" orientation="longitude" pageId="2" pageNumber="220" precision="1" seconds="43.5" value="-52.12875">52°07’43.5”W</geoCoordinate>
, 
<date id="FFC31020FFCE18746C885FC0FA43F811" box="[1216,1416,1976,2001]" pageId="2" pageNumber="220" value="2011-02-11">
<collectingDate id="EF87E9C8FFCE18746C885FC0FA43F811" box="[1216,1416,1976,2001]" pageId="2" pageNumber="220" value="2011-02-11">11 February 2011</collectingDate>
</date>
, fl. e fr., 
<emphasis id="B909EAF2FFCE1874688A5FA4FE62F835" box="[194,425,2012,2037]" italics="true" pageId="2" pageNumber="220">
<collectorName id="26885336FFCE1874688A5FA4FEEBF835" box="[194,288,2012,2037]" pageId="2" pageNumber="220">V.J. Pott</collectorName>
et al. 11239
</emphasis>
(CGMS)
</materialsCitation>
.
</paragraph>
<caption id="DF026668FFCF187568C05CAEFEE1FAF3" ID-DOI="http://doi.org/10.5281/zenodo.13647842" ID-Zenodo-Dep="13647842" httpUri="https://zenodo.org/record/13647842/files/figure.png" pageId="3" pageNumber="221" startId="3.[136,229,1238,1259]" targetBox="[153,1435,191,1213]" targetPageId="3" targetType="figure">
<paragraph id="8BC236E0FFCF187568C05CAEFEE1FAF3" blockId="3.[136,1452,1238,1331]" pageId="3" pageNumber="221">
<emphasis id="B909EAF2FFCF187568C05CAEFF35FB2B" bold="true" box="[136,254,1238,1259]" pageId="3" pageNumber="221">FIGURE 2.</emphasis>
Photographs of 
<taxonomicName id="4C7D4D63FFCF187569D55CAEFDA8FB2B" authorityName="L. P. Silva, Hefler &amp; R. Trevisan" authorityYear="2016" box="[413,611,1238,1259]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="221" phylum="Tracheophyta" rank="species" species="tuckerianus">
<emphasis id="B909EAF2FFCF187569D55CAEFDA8FB2B" box="[413,611,1238,1259]" italics="true" pageId="3" pageNumber="221">Cyperus tuckerianus</emphasis>
</taxonomicName>
in its natural habitat in the state of Mato Grosso, Brazil. 
<emphasis id="B909EAF2FFCF18756CC35CAEFB69FB2B" bold="true" box="[1163,1186,1238,1259]" pageId="3" pageNumber="221">A.</emphasis>
Habit. 
<emphasis id="B909EAF2FFCF18756CA25CAEFACBFB2B" bold="true" box="[1258,1280,1238,1259]" pageId="3" pageNumber="221">B.</emphasis>
Inflorescence. 
<emphasis id="B909EAF2FFCF18756DDC5CAEFA60FB2B" bold="true" box="[1428,1451,1238,1259]" pageId="3" pageNumber="221">C.</emphasis>
Detail of the culm base (paratype). 
<emphasis id="B909EAF2FFCF187569AC5C82FE37FACF" bold="true" box="[484,508,1274,1295]" pageId="3" pageNumber="221">D.</emphasis>
Detail of the elongated internode (black arrow) at the culm base (holotype). All photographs by Luciana P. Silva.
</paragraph>
</caption>
<paragraph id="8BC236E0FFCF187568F55D1BFF75F9B8" blockId="3.[136,1452,1379,2052]" pageId="3" pageNumber="221">
<emphasis id="B909EAF2FFCF187568F55D1BFEEDFABC" bold="true" box="[189,294,1379,1404]" pageId="3" pageNumber="221">Notes:—</emphasis>
The material selected as 
<typeStatus id="54C68842FFCF18756A0E5D1BFD63FABC" box="[582,680,1379,1404]" pageId="3" pageNumber="221" type="holotype">holotype</typeStatus>
presents an elongation of the basal internodes of the culm (
<figureCitation id="13462A65FFCF18756D295D1BFF66FA60" captionStart="FIGURE 2" captionStartId="3.[136,229,1238,1259]" captionTargetBox="[153,1435,191,1213]" captionTargetId="figure-18@3.[152,1436,190,1213]" captionTargetPageId="3" captionText="FIGURE 2. Photographs of Cyperus tuckerianus in its natural habitat in the state of Mato Grosso, Brazil. A. Habit. B. Inflorescence. C. Detail of the culm base (paratype). D. Detail of the elongated internode (black arrow) at the culm base (holotype). All photographs by Luciana P. Silva." figureDoi="http://doi.org/10.5281/zenodo.13647842" httpUri="https://zenodo.org/record/13647842/files/figure.png" pageId="3" pageNumber="221">Figure 2D</figureCitation>
), however this does not seem to be the typical morphology of the species. It was observed during the collection of the material that the habitat had recently suffered a burial (
<figureCitation id="13462A65FFCF18756B535DD3FC58FA04" box="[795,915,1451,1476]" captionStart="FIGURE 3" captionStartId="4.[136,229,1597,1618]" captionTargetBox="[151,1436,841,1572]" captionTargetId="figure-237@4.[151,1436,841,1572]" captionTargetPageId="4" captionText="FIGURE 3. Distribution map of Cyperus tuckerianus in the Central-West Region of Brazil." figureDoi="http://doi.org/10.5281/zenodo.13647846" httpUri="https://zenodo.org/record/13647846/files/figure.png" pageId="3" pageNumber="221">Figure 3A</figureCitation>
), which induced the elongation of the structure. Another important aspect to mention is the classification of the species life cycle as annual to short perennial. In spite of its robust aspect, we noted the plant presented little input in the development of its vegetative part and a higher focus in its reproductive structures (inflorescence). We believe this fact is related to the short life cycle of the plant, nonetheless in order to confirm this information it would be necessary to follow the plant’s development through the year.
</paragraph>
<paragraph id="8BC236E0FFCF187568F55EFBFEC1F890" blockId="3.[136,1452,1379,2052]" pageId="3" pageNumber="221">
According to the classification of 
<bibRefCitation id="EFEC4B11FFCF18756A7B5EFBFD30F95C" author="Kukenthal, G." box="[563,763,1667,1692]" pageId="3" pageNumber="221" refId="ref3959" refString="Kukenthal, G. (1936) Cyperaceae, Scirpoideae, Cypereae. In: Engler, A. (Ed.) Das Pflanzenreich: Regni Vegetabilis Conspectus. v. 4, n. 20. H. R. Hengelmann, Weinheim, 621 pp. Available from: http: // www. biodiversitylibrary. org / item / 71650 # page / 16 / mode / 1 up (accessed 1 November 2016)" type="book" year="1936">Kükenthal (1936)</bibRefCitation>
, 
<taxonomicName id="4C7D4D63FFCF18756B4D5EFBFC20F95C" authorityName="L. P. Silva, Hefler &amp; R. Trevisan" authorityYear="2016" box="[773,1003,1667,1692]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="221" phylum="Tracheophyta" rank="species" species="tuckerianus">
<emphasis id="B909EAF2FFCF18756B4D5EFBFC20F95C" box="[773,1003,1667,1692]" italics="true" pageId="3" pageNumber="221">Cyperus tuckerianus</emphasis>
</taxonomicName>
belongs to the subdivision 
<taxonomicName id="4C7D4D63FFCF18756D505EFBFE40F900" authority="Kukenthal (1936: 327)" authorityName="Kukenthal" authorityPageNumber="327" authorityYear="1936" pageId="3" pageNumber="221" rank="subPhylum" subPhylum="Isodiametrici">
Isodiametrici 
<bibRefCitation id="EFEC4B11FFCF187568C05EDFFE40F900" author="Kukenthal, G." box="[136,395,1703,1728]" pageId="3" pageNumber="221" refId="ref3959" refString="Kukenthal, G. (1936) Cyperaceae, Scirpoideae, Cypereae. In: Engler, A. (Ed.) Das Pflanzenreich: Regni Vegetabilis Conspectus. v. 4, n. 20. H. R. Hengelmann, Weinheim, 621 pp. Available from: http: // www. biodiversitylibrary. org / item / 71650 # page / 16 / mode / 1 up (accessed 1 November 2016)" type="book" year="1936">Kükenthal (1936: 327)</bibRefCitation>
</taxonomicName>
of subgenus 
<taxonomicName id="4C7D4D63FFCF18756A6D5EDFFDB5F900" box="[549,638,1703,1728]" pageId="3" pageNumber="221" rank="subGenus" subGenus="Pycreus">
<emphasis id="B909EAF2FFCF18756A6D5EDFFDB5F900" box="[549,638,1703,1728]" italics="true" pageId="3" pageNumber="221">Pycreus</emphasis>
</taxonomicName>
. This subdivision is characterized by achenes with the surface covered by hexagonal isodiametric cells (
<figureCitation id="13462A65FFCF187569B35EB3FDA4F924" box="[507,623,1739,1764]" captionStart="FIGURE 1" captionStartId="2.[136,229,1799,1820]" captionTargetBox="[155,1432,353,1774]" captionTargetId="figure-104@2.[155,1432,353,1774]" captionTargetPageId="2" captionText="FIGURE 1. Illustrations of Cyperus tuckerianus (based on L.P. Silva 119). A. Habit. B. Spikelet. C. Achene. D. Surface of achene. E. Detail of surface of the achene with isodiametric cells and prominent silica bodies. Illustrated by Luciana P. Silva." figureDoi="http://doi.org/10.5281/zenodo.13647838" httpUri="https://zenodo.org/record/13647838/files/figure.png" pageId="3" pageNumber="221">Figure 1E</figureCitation>
). However, it was not possible to place this species in any of the sections proposed by 
<bibRefCitation id="EFEC4B11FFCF187569555E97FE2DF8C8" author="Kukenthal, G." box="[285,486,1775,1800]" pageId="3" pageNumber="221" refId="ref3959" refString="Kukenthal, G. (1936) Cyperaceae, Scirpoideae, Cypereae. In: Engler, A. (Ed.) Das Pflanzenreich: Regni Vegetabilis Conspectus. v. 4, n. 20. H. R. Hengelmann, Weinheim, 621 pp. Available from: http: // www. biodiversitylibrary. org / item / 71650 # page / 16 / mode / 1 up (accessed 1 November 2016)" type="book" year="1936">Kükenthal (1936)</bibRefCitation>
for subgenus 
<taxonomicName id="4C7D4D63FFCF18756AC25E97FD28F8C8" box="[650,739,1775,1800]" pageId="3" pageNumber="221" rank="subGenus" subGenus="Pycreus">
<emphasis id="B909EAF2FFCF18756AC25E97FD28F8C8" box="[650,739,1775,1800]" italics="true" pageId="3" pageNumber="221">Pycreus</emphasis>
</taxonomicName>
, because the species presents characteristics that fit obscurely either in section 
<taxonomicName id="4C7D4D63FFCF1875690B5F6BFE7FF8EC" box="[323,436,1811,1836]" pageId="3" pageNumber="221" rank="section" section="Propinqui">
<emphasis id="B909EAF2FFCF1875690B5F6BFE7FF8EC" box="[323,436,1811,1836]" italics="true" pageId="3" pageNumber="221">Propinqui</emphasis>
</taxonomicName>
(Kükenthal 1836: 342; 
<bibRefCitation id="EFEC4B11FFCF18756A815F6BFCB4F8EC" author="Clarke, C. B." box="[713,895,1811,1836]" pageId="3" pageNumber="221" refId="ref3701" refString="Clarke, C. B. (1908) Bulletin of Miscellaneous Information Kew. 554 pp." type="book" year="1908">Clarke 1908: 95</bibRefCitation>
) or in section 
<taxonomicName id="4C7D4D63FFCF18756C6A5F6BFB55F8EC" box="[1058,1182,1811,1836]" pageId="3" pageNumber="221" rank="section" section="Chrysanthi">
<emphasis id="B909EAF2FFCF18756C6A5F6BFB55F8EC" box="[1058,1182,1811,1836]" italics="true" pageId="3" pageNumber="221">Chrysanthi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFEC4B11FFCF18756CE55F6BFAACF8EC" author="Clarke, C. B." box="[1197,1383,1811,1836]" pageId="3" pageNumber="221" refId="ref3701" refString="Clarke, C. B. (1908) Bulletin of Miscellaneous Information Kew. 554 pp." type="book" year="1908">Clarke 1908: 95</bibRefCitation>
; 
<bibRefCitation id="EFEC4B11FFCF18756D3B5F6BFF36F890" author="Kern, J. H." pageId="3" pageNumber="221" pagination="1 - 753" refId="ref3934" refString="Kern, J. H. (1974) Flora Malesiana, Series I. Spermatophyta 7 (3): 1 - 753." type="journal article" year="1974">Kern 1974: 648</bibRefCitation>
).
</paragraph>
<paragraph id="8BC236E0FFCF187568F55F23FE92F87C" blockId="3.[136,1452,1379,2052]" pageId="3" pageNumber="221">
Within the species of subgenus 
<taxonomicName id="4C7D4D63FFCF18756A6C5F23FDB6F8B4" box="[548,637,1883,1908]" pageId="3" pageNumber="221" rank="subGenus" subGenus="Pycreus">
<emphasis id="B909EAF2FFCF18756A6C5F23FDB6F8B4" box="[548,637,1883,1908]" italics="true" pageId="3" pageNumber="221">Pycreus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7D4D63FFCF18756AC45F23FCFAF8B4" authorityName="L. P. Silva, Hefler &amp; R. Trevisan" authorityYear="2016" box="[652,817,1883,1908]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="221" phylum="Tracheophyta" rank="species" species="tuckerianus">
<emphasis id="B909EAF2FFCF18756AC45F23FCFAF8B4" box="[652,817,1883,1908]" italics="true" pageId="3" pageNumber="221">C. tuckerianus</emphasis>
</taxonomicName>
seems to be closer to 
<taxonomicName id="4C7D4D63FFCF18756C665F23FA6CF8B4" authority="Brown (1810: 216)" authorityName="Brown" authorityPageNumber="216" authorityYear="1810" box="[1070,1447,1883,1908]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="221" phylum="Tracheophyta" rank="species" species="unioloides">
<emphasis id="B909EAF2FFCF18756C665F23FB0FF8B4" box="[1070,1220,1883,1908]" italics="true" pageId="3" pageNumber="221">C. unioloides</emphasis>
<bibRefCitation id="EFEC4B11FFCF18756C845F23FA6CF8B4" author="Brown, R." box="[1228,1447,1883,1908]" pageId="3" pageNumber="221" refId="ref3647" refString="Brown, R. (1810) Prodromus florae Novae Hollandiae et insulae Van-Diemen. Typis R. Taylor et socii, London. Available from: http: // www. biodiversitylibrary. org / page / 2954372 # page / 84 / mode / 1 up (accessed 1 November 2016)" type="url" year="1810">Brown (1810: 216)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C7D4D63FFCF187568C05F07FCF3F858" authority="J. Presl &amp; C. Presl (1828: 167)" authorityName="J. Presl &amp; C. Presl" authorityPageNumber="167" authorityYear="1828" box="[136,824,1919,1944]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="221" phylum="Tracheophyta" rank="variety" species="lanceolatus" variety="compositus">
<emphasis id="B909EAF2FFCF187568C05F07FEE2F858" box="[136,297,1919,1944]" italics="true" pageId="3" pageNumber="221">C. lanceolatus</emphasis>
var. 
<emphasis id="B909EAF2FFCF187569175F07FE16F858" box="[351,477,1919,1944]" italics="true" pageId="3" pageNumber="221">compositus</emphasis>
J. Presl &amp; C. 
<bibRefCitation id="EFEC4B11FFCF18756A3F5F07FCF3F858" author="Presl, J. S. &amp; Presl, C. B." box="[631,824,1919,1944]" pageId="3" pageNumber="221" refId="ref4632" refString="Presl, J. S. &amp; Presl, C. B. (1828) Encyclopedie Methodique, Botanique 1: 167. Available from: http: // www. biodiversitylibrary. org / page / 390806 # page / 183 / mode / 1 up (accessed 1 November 2016)" type="journal volume" year="1828">Presl (1828: 167)</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C7D4D63FFCF18756B265F07FF70F87C" authority="Kukenthal (1936: 351)" authorityName="Kukenthal" authorityPageNumber="351" authorityYear="1936" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="221" phylum="Tracheophyta" rank="variety" species="lanceolatus" variety="subunioloides">
<emphasis id="B909EAF2FFCF18756B265F07FBC4F858" box="[878,1039,1919,1944]" italics="true" pageId="3" pageNumber="221">C. lanceolatus</emphasis>
var. 
<emphasis id="B909EAF2FFCF18756C0D5F07FB2AF858" box="[1093,1249,1919,1944]" italics="true" pageId="3" pageNumber="221">subunioloides</emphasis>
<bibRefCitation id="EFEC4B11FFCF18756CAF5F07FF70F87C" author="Kukenthal, G." pageId="3" pageNumber="221" refId="ref3959" refString="Kukenthal, G. (1936) Cyperaceae, Scirpoideae, Cypereae. In: Engler, A. (Ed.) Das Pflanzenreich: Regni Vegetabilis Conspectus. v. 4, n. 20. H. R. Hengelmann, Weinheim, 621 pp. Available from: http: // www. biodiversitylibrary. org / item / 71650 # page / 16 / mode / 1 up (accessed 1 November 2016)" type="book" year="1936">Kükenthal (1936: 351)</bibRefCitation>
</taxonomicName>
(see 
<tableCitation id="C6FF035BFFCF187568BE5FDBFE83F87C" box="[246,328,1955,1980]" captionStart="TABLE 1" captionStartId="4.[136,216,160,181]" captionText="TABLE 1. Comparison of Cyperus tuckerianus with related species of subgenus Pycreus that occur in the Neotropical region. Source of data: *Corcoran (1941); **Kukenthal (1936) and collection Regnell 1298 (holotype: US–photo!)." pageId="3" pageNumber="221">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BC236E0FFCF187568F55FBFFB08F7C4" blockId="3.[136,1452,1379,2052]" pageId="3" pageNumber="221">
<taxonomicName id="4C7D4D63FFCF187568F55FBFFE6CF820" authorityName="L. P. Silva, Hefler &amp; R. Trevisan" authorityYear="2016" box="[189,423,1991,2016]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="221" phylum="Tracheophyta" rank="species" species="tuckerianus">
<emphasis id="B909EAF2FFCF187568F55FBFFE6CF820" box="[189,423,1991,2016]" italics="true" pageId="3" pageNumber="221">Cyperus tuckerianus</emphasis>
</taxonomicName>
may be easily distinguished from 
<taxonomicName id="4C7D4D63FFCF18756B7E5FBFFC05F820" authorityName="Brown" authorityYear="1810" box="[822,974,1991,2016]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="221" phylum="Tracheophyta" rank="species" species="unioloides">
<emphasis id="B909EAF2FFCF18756B7E5FBFFC05F820" box="[822,974,1991,2016]" italics="true" pageId="3" pageNumber="221">C. unioloides</emphasis>
</taxonomicName>
by its shorter and narrower spikelets and shorter glumes, and the anthers which are shorter 
<quantity id="4C859B05FFCF18756AFE5F93FCF6F7C4" box="[694,829,2027,2052]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="221" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.5–0.8 mm</quantity>
(vs. 2.0– 
<quantity id="4C859B05FFCF18756BED5F93FC31F7C4" box="[933,1018,2027,2052]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="221" unit="mm" value="2.4">2.4 mm</quantity>
in 
<taxonomicName id="4C7D4D63FFCF18756C565F93FB78F7C4" authorityName="Brown" authorityYear="1810" box="[1054,1203,2027,2052]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="221" phylum="Tracheophyta" rank="species" species="unioloides">
<emphasis id="B909EAF2FFCF18756C565F93FB78F7C4" box="[1054,1203,2027,2052]" italics="true" pageId="3" pageNumber="221">C. unioloides</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="DF026668FFC8187268C058D8FBFBFF19" pageId="4" pageNumber="222" startId="4.[136,216,160,181]" targetType="table">
<paragraph id="8BC236E0FFC8187268C058D8FBFBFF19" blockId="4.[136,1451,160,217]" pageId="4" pageNumber="222">
<emphasis id="B909EAF2FFC8187268C058D8FF3BFF75" bold="true" box="[136,240,160,181]" pageId="4" pageNumber="222">TABLE 1.</emphasis>
Comparison of 
<taxonomicName id="4C7D4D63FFC8187269C458D8FD98FF75" authorityName="L. P. Silva, Hefler &amp; R. Trevisan" authorityYear="2016" box="[396,595,160,181]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="222" phylum="Tracheophyta" rank="species" species="tuckerianus">
<emphasis id="B909EAF2FFC8187269C458D8FD98FF75" box="[396,595,160,181]" italics="true" pageId="4" pageNumber="222">Cyperus tuckerianus</emphasis>
</taxonomicName>
with related species of subgenus 
<taxonomicName id="4C7D4D63FFC818726BD058D8FC2FFF75" box="[920,996,160,181]" pageId="4" pageNumber="222" rank="subGenus" subGenus="Pycreus">
<emphasis id="B909EAF2FFC818726BD058D8FC2FFF75" box="[920,996,160,181]" italics="true" pageId="4" pageNumber="222">Pycreus</emphasis>
</taxonomicName>
that occur in the Neotropical region. Source of data: * 
<bibRefCitation id="EFEC4B11FFC81872688E58BCFEA1FF19" author="Corcoran, M. L." box="[198,362,196,217]" pageId="4" pageNumber="222" pagination="1 - 68" refId="ref3719" refString="Corcoran, M. L. (1941) A revision of the subgenus Pycreus in North and South America. Thesis (Ph. D.) The Catholic University of America. Biological series 37: 1 - 68." type="book chapter" year="1941">Corcoran (1941)</bibRefCitation>
; ** 
<bibRefCitation id="EFEC4B11FFC8187269C258BCFDFCFF19" author="Kukenthal, G." box="[394,567,196,217]" pageId="4" pageNumber="222" refId="ref3959" refString="Kukenthal, G. (1936) Cyperaceae, Scirpoideae, Cypereae. In: Engler, A. (Ed.) Das Pflanzenreich: Regni Vegetabilis Conspectus. v. 4, n. 20. H. R. Hengelmann, Weinheim, 621 pp. Available from: http: // www. biodiversitylibrary. org / item / 71650 # page / 16 / mode / 1 up (accessed 1 November 2016)" type="book" year="1936">Kukenthal (1936)</bibRefCitation>
and collection 
<emphasis id="B909EAF2FFC818726A8258BCFC82FF19" box="[714,841,196,217]" italics="true" pageId="4" pageNumber="222">Regnell 1298</emphasis>
(holotype: US–photo!).
</paragraph>
</caption>
<paragraph id="8BC236E0FFC8187269035897FA86FE7F" pageId="4" pageNumber="222">
<table id="F97DC440FFC8E78968DF5897FA52FE7F" box="[151,1433,239,447]" gridcols="5" gridrows="5" pageId="4" pageNumber="222">
<tr id="354D34A2FFC8E78968DF5897FA52FEC4" box="[151,1433,239,260]" gridrow="0" pageId="4" pageNumber="222" rowspan-0="1">
<th id="769C5DDEFFC8E78969035897FDDDFEC4" box="[331,534,239,260]" gridcol="1" gridrow="0" pageId="4" pageNumber="222">
<taxonomicName id="4C7D4D63FFC8187269035897FE00FEC4" authorityName="Brown" authorityYear="1810" box="[331,459,239,260]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="222" phylum="Tracheophyta" rank="species" species="unioloides">
<emphasis id="B909EAF2FFC8187269035897FE00FEC4" box="[331,459,239,260]" italics="true" pageId="4" pageNumber="222">C. unioloides</emphasis>
</taxonomicName>
</th>
<th id="769C5DDEFFC8E7896A725897FCBBFEC4" box="[570,880,239,260]" gridcol="2" gridrow="0" pageId="4" pageNumber="222">
<taxonomicName id="4C7D4D63FFC818726A765897FCADFEC4" authorityName="J. Presl &amp; C. Presl" authorityYear="1828" box="[574,870,239,260]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="222" phylum="Tracheophyta" rank="variety" species="lanceolatus" variety="compositus">
<emphasis id="B909EAF2FFC818726A765897FD02FEC4" box="[574,713,239,260]" italics="true" pageId="4" pageNumber="222">C. lanceolatus</emphasis>
var. 
<emphasis id="B909EAF2FFC818726AB05897FCADFEC4" box="[760,870,239,260]" italics="true" pageId="4" pageNumber="222">compositus</emphasis>
</taxonomicName>
<emphasis id="B909EAF2FFC818726B2E5897FCBBFEC4" box="[870,880,239,260]" italics="true" pageId="4" pageNumber="222">*</emphasis>
</th>
<th id="769C5DDEFFC8E7896BC65897FB22FEC4" box="[910,1257,239,260]" gridcol="3" gridrow="0" pageId="4" pageNumber="222">
<taxonomicName id="4C7D4D63FFC818726BDB5897FB18FEC4" authorityName="Kukenthal" authorityYear="1936" box="[915,1235,239,260]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="222" phylum="Tracheophyta" rank="variety" species="lanceolatus" variety="subunioloides">
<emphasis id="B909EAF2FFC818726BDB5897FBD5FEC4" box="[915,1054,239,260]" italics="true" pageId="4" pageNumber="222">C. lanceolatus</emphasis>
var. 
<emphasis id="B909EAF2FFC818726C045897FB18FEC4" box="[1100,1235,239,260]" italics="true" pageId="4" pageNumber="222">subunioloides</emphasis>
</taxonomicName>
<emphasis id="B909EAF2FFC818726C9B5897FB22FEC4" box="[1235,1257,239,260]" italics="true" pageId="4" pageNumber="222">**</emphasis>
</th>
<th id="769C5DDEFFC8E7896D405897FA52FEC4" box="[1288,1433,239,260]" gridcol="4" gridrow="0" pageId="4" pageNumber="222">
<taxonomicName id="4C7D4D63FFC818726D455897FA52FEC4" authority="Spikelet" authorityName="Spikelet" box="[1293,1433,239,260]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="222" phylum="Tracheophyta" rank="species" species="tuckerianus">
<emphasis id="B909EAF2FFC818726D455897FA52FEC4" box="[1293,1433,239,260]" italics="true" pageId="4" pageNumber="222">C. tuckerianus</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="354D34A2FFC8E78968DF5966FA52FEF3" box="[151,1433,286,307]" gridrow="1" pageId="4" pageNumber="222">
<th id="769C5DDEFFC8E78968DF5966FEE3FEF3" box="[151,296,286,307]" gridcol="0" gridrow="1" pageId="4" pageNumber="222">Spikelet length</th>
<td id="769C5DDEFFC8E78969035966FDDDFEF3" box="[331,534,286,307]" gridcol="1" gridrow="1" pageId="4" pageNumber="222">(7.0–) 12.0–35.0 mm</td>
<td id="769C5DDEFFC8E7896A725966FCBBFEF3" box="[570,880,286,307]" gridcol="2" gridrow="1" pageId="4" pageNumber="222">3–7 mm</td>
<td id="769C5DDEFFC8E7896BC65966FB22FEF3" box="[910,1257,286,307]" gridcol="3" gridrow="1" pageId="4" pageNumber="222">15–20 mm</td>
<td id="769C5DDEFFC8E7896D405966FA52FEF3" box="[1288,1433,286,307]" gridcol="4" gridrow="1" pageId="4" pageNumber="222">7.5–13.0 mm</td>
</tr>
<tr id="354D34A2FFC8E78968DF5934FA52FEA1" box="[151,1433,332,353]" gridrow="2" pageId="4" pageNumber="222">
<th id="769C5DDEFFC8E78968DF5934FEE3FEA1" box="[151,296,332,353]" gridcol="0" gridrow="2" pageId="4" pageNumber="222">Spikelet width</th>
<td id="769C5DDEFFC8E78969035934FDDDFEA1" box="[331,534,332,353]" gridcol="1" gridrow="2" pageId="4" pageNumber="222">(3.0–) 3.5–5.0 mm</td>
<td id="769C5DDEFFC8E7896A725934FCBBFEA1" box="[570,880,332,353]" gridcol="2" gridrow="2" pageId="4" pageNumber="222">1.5–2.0 mm</td>
<td id="769C5DDEFFC8E7896BC65934FB22FEA1" box="[910,1257,332,353]" gridcol="3" gridrow="2" pageId="4" pageNumber="222">3 mm</td>
<td id="769C5DDEFFC8E7896D405934FA52FEA1" box="[1288,1433,332,353]" gridcol="4" gridrow="2" pageId="4" pageNumber="222">2.3–3.2 mm</td>
</tr>
<tr id="354D34A2FFC8E78968DF5903FA52FE50" box="[151,1433,379,400]" gridrow="3" pageId="4" pageNumber="222">
<th id="769C5DDEFFC8E78968DF5903FEE3FE50" box="[151,296,379,400]" gridcol="0" gridrow="3" pageId="4" pageNumber="222">Glume length</th>
<td id="769C5DDEFFC8E78969035903FDDDFE50" box="[331,534,379,400]" gridcol="1" gridrow="3" pageId="4" pageNumber="222">3.5–4.5 (–5.0) mm</td>
<td id="769C5DDEFFC8E7896A725903FCBBFE50" box="[570,880,379,400]" gridcol="2" gridrow="3" pageId="4" pageNumber="222">1.5 mm</td>
<td id="769C5DDEFFC8E7896BC65903FB22FE50" box="[910,1257,379,400]" gridcol="3" gridrow="3" pageId="4" pageNumber="222">2.0–2.6 mm</td>
<td id="769C5DDEFFC8E7896D405903FA52FE50" box="[1288,1433,379,400]" gridcol="4" gridrow="3" pageId="4" pageNumber="222">2.1–2.7 mm</td>
</tr>
<tr id="354D34A2FFC8E78968DF59D2FA52FE7F" box="[151,1433,426,447]" gridrow="4" pageId="4" pageNumber="222">
<th id="769C5DDEFFC8E78968DF59D2FEE3FE7F" box="[151,296,426,447]" gridcol="0" gridrow="4" pageId="4" pageNumber="222">Rays</th>
<td id="769C5DDEFFC8E789690359D2FDDDFE7F" box="[331,534,426,447]" gridcol="1" gridrow="4" pageId="4" pageNumber="222">present</td>
<td id="769C5DDEFFC8E7896A7259D2FCBBFE7F" box="[570,880,426,447]" gridcol="2" gridrow="4" pageId="4" pageNumber="222">present</td>
<td id="769C5DDEFFC8E7896BC659D2FB22FE7F" box="[910,1257,426,447]" gridcol="3" gridrow="4" pageId="4" pageNumber="222">absent</td>
<td id="769C5DDEFFC8E7896D4059D2FA52FE7F" box="[1288,1433,426,447]" gridcol="4" gridrow="4" pageId="4" pageNumber="222">present</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC236E0FFC8187268F5598FFEF7FD98" blockId="4.[136,1452,503,816]" pageId="4" pageNumber="222">
<taxonomicName id="4C7D4D63FFC8187268F5598FFE6CFDD0" box="[189,423,503,528]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="222" phylum="Tracheophyta" rank="species" species="tuckerianus">
<emphasis id="B909EAF2FFC8187268F5598FFE6CFDD0" box="[189,423,503,528]" italics="true" pageId="4" pageNumber="222">Cyperus tuckerianus</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C7D4D63FFC818726AE1598FFBCDFDD0" authorityName="J. Presl &amp; C. Presl" authorityYear="1828" box="[681,1030,503,528]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="222" phylum="Tracheophyta" rank="variety" species="lanceolatus" variety="compositus">
<emphasis id="B909EAF2FFC818726AE1598FFC86FDD0" box="[681,845,503,528]" italics="true" pageId="4" pageNumber="222">C. lanceolatus</emphasis>
var. 
<emphasis id="B909EAF2FFC818726BC0598FFBCDFDD0" box="[904,1030,503,528]" italics="true" pageId="4" pageNumber="222">compositus</emphasis>
</taxonomicName>
by its longer and wider spikelet and longer glumes, and 
<bibRefCitation id="EFEC4B11FFC81872692C5A63FDE1FDF4" author="Kukenthal, G." box="[356,554,539,564]" pageId="4" pageNumber="222" refId="ref3959" refString="Kukenthal, G. (1936) Cyperaceae, Scirpoideae, Cypereae. In: Engler, A. (Ed.) Das Pflanzenreich: Regni Vegetabilis Conspectus. v. 4, n. 20. H. R. Hengelmann, Weinheim, 621 pp. Available from: http: // www. biodiversitylibrary. org / item / 71650 # page / 16 / mode / 1 up (accessed 1 November 2016)" type="book" year="1936">Kükenthal (1936)</bibRefCitation>
described the rays of this 
<taxonomicName id="4C7D4D63FFC818726B065A63FC1EFDF4" box="[846,981,539,564]" pageId="4" pageNumber="222" rank="variety" variety="only">variety only</taxonomicName>
<quantity id="4C859B05FFC818726B935A63FBECFDF4" box="[987,1063,539,564]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="222" unit="cm" value="2.5">2.5 cm</quantity>
long (vs. rays 3.5–12.0 cm long in 
<taxonomicName id="4C7D4D63FFC8187268C05A47FEE7FD98" box="[136,300,575,600]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="222" phylum="Tracheophyta" rank="species" species="tuckerianus">
<emphasis id="B909EAF2FFC8187268C05A47FEE7FD98" box="[136,300,575,600]" italics="true" pageId="4" pageNumber="222">C. tuckerianus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BC236E0FFC8187268F55A1BFD7EFCF0" blockId="4.[136,1452,503,816]" pageId="4" pageNumber="222">
<taxonomicName id="4C7D4D63FFC8187268F55A1BFD48FDBC" authorityName="Kukenthal" authorityYear="1936" box="[189,643,611,636]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="222" phylum="Tracheophyta" rank="variety" species="lanceolatus" variety="subunioloides">
<emphasis id="B909EAF2FFC8187268F55A1BFE6CFDBC" box="[189,423,611,636]" italics="true" pageId="4" pageNumber="222">Cyperus lanceolatus</emphasis>
var. 
<emphasis id="B909EAF2FFC8187269AF5A1BFD48FDBC" box="[487,643,611,636]" italics="true" pageId="4" pageNumber="222">subunioloides</emphasis>
</taxonomicName>
presents similarities with 
<taxonomicName id="4C7D4D63FFC818726BF35A1BFBA8FDBC" box="[955,1123,611,636]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="222" phylum="Tracheophyta" rank="species" species="tuckerianus">
<emphasis id="B909EAF2FFC818726BF35A1BFBA8FDBC" box="[955,1123,611,636]" italics="true" pageId="4" pageNumber="222">C. tuckerianus</emphasis>
</taxonomicName>
, however 
<bibRefCitation id="EFEC4B11FFC818726CA95A1BFA60FDBC" author="Kukenthal, G." box="[1249,1451,611,636]" pageId="4" pageNumber="222" refId="ref3959" refString="Kukenthal, G. (1936) Cyperaceae, Scirpoideae, Cypereae. In: Engler, A. (Ed.) Das Pflanzenreich: Regni Vegetabilis Conspectus. v. 4, n. 20. H. R. Hengelmann, Weinheim, 621 pp. Available from: http: // www. biodiversitylibrary. org / item / 71650 # page / 16 / mode / 1 up (accessed 1 November 2016)" type="book" year="1936">Kükenthal (1936)</bibRefCitation>
described this 
<taxonomicName id="4C7D4D63FFC8187269615AFFFE7BFD60" box="[297,432,647,672]" pageId="4" pageNumber="222" rank="variety" variety="with">variety with</taxonomicName>
a lax anthela and with spikelet of 
<quantity id="4C859B05FFC818726B655AFFFCB7FD60" box="[813,892,647,672]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="222" unit="mm" value="20.0">20 mm</quantity>
long (vs. 7.5–13.0 mm long in 
<taxonomicName id="4C7D4D63FFC818726C925AFFFAB6FD60" box="[1242,1405,647,672]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="222" phylum="Tracheophyta" rank="species" species="tuckerianus">
<emphasis id="B909EAF2FFC818726C925AFFFAB6FD60" box="[1242,1405,647,672]" italics="true" pageId="4" pageNumber="222">C. tuckerianus</emphasis>
</taxonomicName>
). In the work of 
<bibRefCitation id="EFEC4B11FFC81872695A5AD3FE1BFD04" author="Corcoran, M. L." box="[274,464,683,708]" pageId="4" pageNumber="222" pagination="1 - 68" refId="ref3719" refString="Corcoran, M. L. (1941) A revision of the subgenus Pycreus in North and South America. Thesis (Ph. D.) The Catholic University of America. Biological series 37: 1 - 68." type="book chapter" year="1941">Corcoran (1941)</bibRefCitation>
, the author comments that 
<taxonomicName id="4C7D4D63FFC818726B4A5AD3FB77FD04" authorityName="Kukenthal" authorityYear="1936" box="[770,1212,683,708]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="222" phylum="Tracheophyta" rank="variety" species="lanceolatus" variety="subunioloides">
<emphasis id="B909EAF2FFC818726B4A5AD3FC23FD04" box="[770,1000,683,708]" italics="true" pageId="4" pageNumber="222">Cyperus lanceolatus</emphasis>
var. 
<emphasis id="B909EAF2FFC818726C685AD3FB77FD04" box="[1056,1212,683,708]" italics="true" pageId="4" pageNumber="222">subunioloides</emphasis>
</taxonomicName>
appears to be a form of 
<taxonomicName id="4C7D4D63FFC8187268E05AB7FDDEFD28" authority="Poir. (1806: 245)" authorityName="Poir." authorityPageNumber="245" authorityYear="1806" box="[168,533,719,744]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="222" phylum="Tracheophyta" rank="species" species="lanceolatus">
<emphasis id="B909EAF2FFC8187268E05AB7FE80FD28" box="[168,331,719,744]" italics="true" pageId="4" pageNumber="222">C. lanceolatus</emphasis>
Poir. (1806: 245)
</taxonomicName>
with wider spikelets, and that this difference is not enough to recognizing at the 
<taxonomicName id="4C7D4D63FFC8187268C05A8BFEDDFCCC" box="[136,278,755,780]" pageId="4" pageNumber="222" rank="variety" variety="level">variety level</taxonomicName>
. We observed a high resolution photograph of the 
<typeStatus id="54C68842FFC818726B0A5A8BFC6FFCCC" box="[834,932,755,780]" pageId="4" pageNumber="222" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C7D4D63FFC818726B8F5A8BFAB6FCCC" authorityName="Kukenthal" authorityYear="1936" box="[967,1405,755,780]" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="222" phylum="Tracheophyta" rank="variety" species="lanceolatus" variety="subunioloides">
<emphasis id="B909EAF2FFC818726B8F5A8BFB67FCCC" box="[967,1196,755,780]" italics="true" pageId="4" pageNumber="222">Cyperus lanceolatus</emphasis>
var. 
<emphasis id="B909EAF2FFC818726CA95A8BFAB6FCCC" box="[1249,1405,755,780]" italics="true" pageId="4" pageNumber="222">subunioloides</emphasis>
</taxonomicName>
and we agree with the conclusion of 
<bibRefCitation id="EFEC4B11FFC8187269BC5B6FFD7AFCF0" author="Corcoran, M. L." box="[500,689,791,816]" pageId="4" pageNumber="222" pagination="1 - 68" refId="ref3719" refString="Corcoran, M. L. (1941) A revision of the subgenus Pycreus in North and South America. Thesis (Ph. D.) The Catholic University of America. Biological series 37: 1 - 68." type="book chapter" year="1941">Corcoran (1941)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>