412 lines
56 KiB
XML
412 lines
56 KiB
XML
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<mods:title id="448B115674B44E6656129D205E9C4B35">Integrative description of new giant pill-millipedes from southern Thailand (Diplopoda, Sphaerotheriida, Zephroniidae)</mods:title>
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<mods:namePart id="8233ACA4918ED84A2CD17A5447AF58C0">Rosenmejer, Trine</mods:namePart>
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<mods:affiliation id="8D10E9A117258304D9EECC2733A1FA16">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 Copenhagen OE, Denmark.</mods:affiliation>
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<mods:nameIdentifier id="93A40CFE7DE6D2EB01DDC4211AA7B49F" type="email">trinehansen@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="2B150F68BD6CB558C9A5C787B1DC8986">Enghoff, Henrik</mods:namePart>
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<mods:affiliation id="50259A51E4C0DBE772D402D3DE36B97E">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 Copenhagen OE, Denmark.</mods:affiliation>
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<mods:nameIdentifier id="21E6263ABBE10314D055AED8443A5467" type="email">henghoff@snm.ku.dk</mods:nameIdentifier>
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<mods:namePart id="DCCF984A2F98D1392AE86D5EE089CF3F">Moritz, Leif</mods:namePart>
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<mods:affiliation id="E1DD2026125E5BC047C7C83AC01EA1CE">Zoological Research Museum Alexander Koenig, Leibniz Institute for Animal Biodiversity, Adenauerallee 160, D- 53113 Bonn, Germany. & Institute of Evolutionary Biology and Ecology, University of Bonn, An der Immenburg 1, 53121 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="AE71332F3F7B9BC69A8497A88A684EFF" type="email">moritz.leif@gmail.com</mods:nameIdentifier>
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<mods:namePart id="EB9518AB7A889AADC2530F1B9E74347E">Wesener, Thomas</mods:namePart>
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<mods:affiliation id="4F2FAF3EB0CC6535789A2F00474FF791">Zoological Research Museum Alexander Koenig, Leibniz Institute for Animal Biodiversity, Adenauerallee 160, D- 53113 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="D6EB6D3E453EB8434177A3E0D712C2AD" type="email">twesener@leibniz-zfmk.de</mods:nameIdentifier>
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<mods:title id="FF86F34F2063157E4DEC350F94030FE6">European Journal of Taxonomy</mods:title>
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<mods:date id="CEA59FB1C1B555F2BCF52038E727AAEA">2021</mods:date>
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<treatment id="03D487ADFFB0FFD695C3FC75A1CEFDC7" ID-DOI="http://doi.org/10.5281/zenodo.5572835" ID-GBIF-Taxon="184570871" ID-Zenodo-Dep="5572835" LSID="urn:lsid:zoobank.org:act:84D3714E-5CF8-4063-8A4D-40BF8CBEBFB0" httpUri="http://treatment.plazi.org/id/03D487ADFFB0FFD695C3FC75A1CEFDC7" lastPageId="12" lastPageNumber="120" pageId="6" pageNumber="114">
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<heading id="D08A81D7FFB0FFDC95C3FC75A75AFBC1" box="[459,1129,1001,1028]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="114" reason="2">
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<taxonomicName id="4C7D4D38FFB0FFDC95C3FC75A734FBC1" ID-CoL="9BDG4" authority="Rosenmejer & Wesener" authorityName="Rosenmejer & Enghoff & Moritz & Wesener" authorityYear="2021" box="[459,1031,1001,1028]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="6" pageNumber="114" phylum="Arthropoda" rank="species" species="aesculus" status="sp. nov.">
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<emphasis id="B909EAA9FFB0FFDC95C3FC75A1D9FBC1" bold="true" box="[459,746,1001,1028]" italics="true" pageId="6" pageNumber="114">Sphaerobelum aesculus</emphasis>
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Rosenmejer & Wesener
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<taxonomicNameLabel id="A23A57D2FFB0FFDC9006FC76A75AFBC1" box="[1038,1129,1002,1028]" pageId="6" pageNumber="114" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C3676530FFB0FFDC9563FB91A0B3FB8F" pageId="6" pageNumber="114" type="description">
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<paragraph id="8BC236BBFFB0FFDC9563FB91A7FBFBED" blockId="6.[363,1224,1001,1099]" box="[363,1224,1037,1064]" pageId="6" pageNumber="114">
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urn:lsid:zoobank.org:act:
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</uri>
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<paragraph id="8BC236BBFFB0FFDC96BDFBACA0B3FB8F" blockId="6.[363,1224,1001,1099]" box="[693,896,1072,1099]" pageId="6" pageNumber="114">
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<figureCitation id="13462A3EFFB0FFDC96BDFBACA026FB8F" box="[693,789,1072,1098]" captionStart="Fig" captionStartId="7.[189,232,1882,1908]" captionTargetBox="[202,1381,333,1766]" captionTargetId="figure-16@7.[189,1398,321,1784]" captionTargetPageId="7" captionText="Fig. 2. Map of Thailand, collection localities of Sphaerobelum aesculus Rosenmejer & Wesener sp. nov. and Zephronia viridisoma Rosenmejer & Wesener sp. nov., and habitus photographs: S. aesculus Rosenmejer & Wesener sp. nov., ♀ (NHMD 621694); Z. viridisoma Rosenmejer & Wesener sp. nov., paratype, ♀ (ZFMK MYR8787). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5211731" httpUri="https://zenodo.org/record/5211731/files/figure.png" pageId="6" pageNumber="114">Figs 2A</figureCitation>
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,
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<figureCitation id="13462A3EFFB0FFDC9729FBACA07CFB8E" box="[801,847,1072,1099]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="9.[189,232,1882,1908]" captionStartId-1="10.[189,232,1814,1840]" captionStartId-2="11.[189,232,1846,1872]" captionTargetBox-0="[190,1397,310,1797]" captionTargetBox-1="[285,1308,283,1724]" captionTargetBox-2="[308,1338,326,1743]" captionTargetId-0="figure-15@9.[189,1398,309,1798]" captionTargetId-1="figure-11@10.[258,1329,265,1777]" captionTargetId-2="figure-15@11.[245,1342,265,1805]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="Fig. 3. Sphaerobelum aesculus Rosenmejer & Wesener sp. nov., holotype, ♂ (NHMD 621693), scanning electron micrographs. A. Right antenna, lateral view. B. Right antenna, disc. C. Detail of apical cone. D. Tergite surface of midbody ring. E. Tergite surface at very high magnification. Abbreviations: ac = apical cone; sb = sensilla basiconica." captionText-1="Fig. 4. Sphaerobelum aesculus Rosenmejer & Wesenersp. nov., holotype, ♂ (NHMD 621693), drawings. A. First left coxa with stigmatic plate, posterior view. B. Second left coxa with gonopore, posterior view. C. Left leg 9, posterior view. D. Left anterior telopod, posterior view. E. Left anterior telopod, anterior view. F. Left anterior telopod, lateral view on podomeres 2–4. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; St = stigmatic plate; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." captionText-2="Fig. 5. Sphaerobelum aesculus Rosenmejer & Wesener sp. nov. drawings. A–B. Holotype, ♂ (NHMD 621693), C. ♀ (NHMD 621694). A. Left posterior telopod, anterior view. B. Left posterior telopod, posterior view, arrow points to swelling of immovable finger. C. Left second coxa and prefemur with female vulva. Abbreviations: Cx = coxa; EP = external, lateral plate; IP = inner, mesal plate; Op = operculum; Pre = prefemur. Scale bars = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5211733" figureDoi-1="http://doi.org/10.5281/zenodo.5211739" figureDoi-2="http://doi.org/10.5281/zenodo.5211741" httpUri-0="https://zenodo.org/record/5211733/files/figure.png" httpUri-1="https://zenodo.org/record/5211739/files/figure.png" httpUri-2="https://zenodo.org/record/5211741/files/figure.png" pageId="6" pageNumber="114">3–5</figureCitation>
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,
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<figureCitation id="13462A3EFFB0FFDC9754FBACA0B3FB8F" box="[860,896,1072,1098]" captionStart="Fig" captionStartId="12.[189,232,1916,1942]" captionTargetBox="[194,1394,1255,1873]" captionTargetId="figure-181@12.[189,1398,1247,1878]" captionTargetPageId="12" captionText="Fig. 6. Endoterga of midbody tergites, scanning electron micrographs. A. Sphaerobelum aesculus Rosenmejer & Wesener sp. nov., holotype (NHMD 621693). B. Zephronia viridisoma Rosenmejer & Wesener sp. nov., holotype, ♂ (NHMD 621695). Abbreviations: cp = cuticular patterns; iA = inner area." figureDoi="http://doi.org/10.5281/zenodo.5211743" httpUri="https://zenodo.org/record/5211743/files/figure.png" pageId="6" pageNumber="114">6A</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3676530FFB0FFDC94B5FBEDA673FAC5" pageId="6" pageNumber="114" type="diagnosis">
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<paragraph id="8BC236BBFFB0FFDC94B5FBEDA205FB4E" blockId="6.[189,310,1137,1163]" box="[189,310,1137,1163]" pageId="6" pageNumber="114">
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<heading id="D08A81D7FFB0FFDC94B5FBEDA205FB4E" bold="true" box="[189,310,1137,1163]" fontSize="11" level="3" pageId="6" pageNumber="114" reason="3">
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<emphasis id="B909EAA9FFB0FFDC94B5FBEDA205FB4E" bold="true" box="[189,310,1137,1163]" pageId="6" pageNumber="114">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC236BBFFB0FFDC94B5FB3CA673FAC5" blockId="6.[189,1400,1184,1281]" pageId="6" pageNumber="114">
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Differs from all other species of the genus
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<taxonomicName id="4C7D4D38FFB0FFDC96C8FB3CA05BFB7F" authorityName="Verhoeff" authorityYear="1924" box="[704,872,1184,1210]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="6" pageNumber="114" phylum="Arthropoda" rank="genus">
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<emphasis id="B909EAA9FFB0FFDC96C8FB3CA05BFB7F" box="[704,872,1184,1210]" italics="true" pageId="6" pageNumber="114">Sphaerobelum</emphasis>
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</taxonomicName>
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in the shape of the posterior telopod, where there is a swelling at the tip of the immovable finger, but the swelling does not extend above the margin (
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<figureCitation id="13462A3EFFB0FFDC94CEFB7AA22CFAC4" box="[198,287,1254,1281]" captionStart="Fig" captionStartId="11.[189,232,1846,1872]" captionTargetBox="[308,1338,326,1743]" captionTargetId="figure-15@11.[245,1342,265,1805]" captionTargetPageId="11" captionText="Fig. 5. Sphaerobelum aesculus Rosenmejer & Wesener sp. nov. drawings. A–B. Holotype, ♂ (NHMD 621693), C. ♀ (NHMD 621694). A. Left posterior telopod, anterior view. B. Left posterior telopod, posterior view, arrow points to swelling of immovable finger. C. Left second coxa and prefemur with female vulva. Abbreviations: Cx = coxa; EP = external, lateral plate; IP = inner, mesal plate; Op = operculum; Pre = prefemur. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5211741" httpUri="https://zenodo.org/record/5211741/files/figure.png" pageId="6" pageNumber="114">Fig. 5B</figureCitation>
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arrow). Such a swelling is currently unknown from any other giant pill-millipede species.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3676530FFB0FFDC94B5FABBA1DAFA56" pageId="6" pageNumber="114" type="etymology">
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<paragraph id="8BC236BBFFB0FFDC94B5FABBA2AEFA84" blockId="6.[189,413,1319,1345]" box="[189,413,1319,1345]" pageId="6" pageNumber="114">
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<heading id="D08A81D7FFB0FFDC94B5FABBA2AEFA84" bold="true" box="[189,413,1319,1345]" fontSize="11" level="3" pageId="6" pageNumber="114" reason="3">
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<emphasis id="B909EAA9FFB0FFDC94B5FABBA2AEFA84" bold="true" box="[189,413,1319,1345]" pageId="6" pageNumber="114">Derivatio nominis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC236BBFFB0FFDC94B5FACAA1DAFA56" blockId="6.[189,1401,1366,1428]" pageId="6" pageNumber="114">
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Named after the horse chestnut tree
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<emphasis id="B909EAA9FFB0FFDC967AFACAA0A5FAB5" box="[626,918,1366,1392]" italics="true" pageId="6" pageNumber="114">Aesculus hippocastanum</emphasis>
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L., for the resemblance of the rolled-up female to a horse chestnut. Noun in apposition.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3676530FFB0FFDC94B5FA27A0D8F917" pageId="6" pageNumber="114" type="materials_examined">
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<paragraph id="8BC236BBFFB0FFDC94B5FA27A299FA10" blockId="6.[189,426,1467,1493]" box="[189,426,1467,1493]" pageId="6" pageNumber="114">
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<heading id="D08A81D7FFB0FFDC94B5FA27A299FA10" bold="true" box="[189,426,1467,1493]" fontSize="11" level="3" pageId="6" pageNumber="114" reason="3">
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<emphasis id="B909EAA9FFB0FFDC94B5FA27A299FA10" bold="true" box="[189,426,1467,1493]" pageId="6" pageNumber="114">Material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC236BBFFB0FFDC94E2FA76A18DF98F" blockId="6.[187,1398,1514,1610]" pageId="6" pageNumber="114">
|
||
<materialsCitation id="3B153CE6FFB0FFDC94E2FA76A18DF98F" ID-GBIF-Occurrence="3335033302" collectedFrom="in and under old logs" collectingDate="1989-02-14" collectionCode="NHMD" collectorName="M. Andersen & A. R. Rasmussen" country="Thailand" county="Island" location="Forest in Kathu district" municipality="Forest in Kathu" pageId="6" pageNumber="114" specimenCode="NHMD 621693" specimenCount="1" specimenCount-male="1" stateProvince="Phuket Island" typeStatus="holotype">
|
||
<emphasis id="B909EAA9FFB0FFDC94E2FA76A26FF9C1" bold="true" box="[234,348,1514,1540]" pageId="6" pageNumber="114">
|
||
<typeStatus id="54C68819FFB0FFDC94E2FA76A26FF9C1" box="[234,348,1514,1540]" pageId="6" pageNumber="114" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F36A762BFFB0FFDC94B5F991A266F9E2" box="[189,341,1549,1575]" name="Thailand" pageId="6" pageNumber="114">THAILAND</collectingCountry>
|
||
•
|
||
<specimenCount id="9D7BFD32FFB0FFDC9562F991A2B0F9E2" box="[362,387,1549,1575]" pageId="6" pageNumber="114" type="male">♂</specimenCount>
|
||
;
|
||
<collectingRegion id="49B9F859FFB0FFDC9585F991A11CF9E2" box="[397,559,1549,1575]" pageId="6" pageNumber="114">Phuket Island</collectingRegion>
|
||
,
|
||
<location id="8EA26060FFB0FFDC9630F991A076F9E2" LSID="urn:lsid:plazi:treatment:03D487ADFFB0FFD695C3FC75A1CEFDC7:8EA26060FFB0FFDC9630F991A076F9E2" box="[568,837,1549,1575]" country="Thailand" county="Island" municipality="Forest in Kathu" name="Forest in Kathu district" pageId="6" pageNumber="114" stateProvince="Phuket Island">Forest in Kathu district</location>
|
||
;
|
||
<date id="FFC3107BFFB0FFDC9747F991A0D0F9E2" box="[847,995,1549,1575]" pageId="6" pageNumber="114" value="1989-02-14">
|
||
<collectingDate id="EF87E993FFB0FFDC9747F991A0D0F9E2" box="[847,995,1549,1575]" pageId="6" pageNumber="114" value="1989-02-14">14 Feb.1989</collectingDate>
|
||
</date>
|
||
.
|
||
<collectorName id="2688536DFFB0FFDC97E5F991A7B1F9E2" box="[1005,1154,1549,1575]" pageId="6" pageNumber="114">M. Andersen</collectorName>
|
||
and
|
||
<collectorName id="2688536DFFB0FFDC90BEF991A645F9E2" box="[1206,1398,1549,1575]" pageId="6" pageNumber="114">A.R. Rasmussen</collectorName>
|
||
leg.;
|
||
<collectedFrom id="38617546FFB0FFDC94FEF9ACA2C3F98F" box="[246,496,1584,1610]" pageId="6" pageNumber="114">in and under old logs</collectedFrom>
|
||
;
|
||
<specimenCode id="DBDB9EC0FFB0FFDC95F5F9ACA189F98F" box="[509,698,1584,1610]" collectionCode="NHMD" pageId="6" pageNumber="114">NHMD 621693</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC236BBFFB0FFDC94E2F9EDA295F94E" blockId="6.[189,1399,1649,1746]" box="[234,422,1649,1675]" pageId="6" pageNumber="114">
|
||
<emphasis id="B909EAA9FFB0FFDC94E2F9EDA295F94E" bold="true" box="[234,422,1649,1675]" pageId="6" pageNumber="114">Other material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC236BBFFB0FFDC94B5F909A0D8F917" blockId="6.[189,1399,1649,1746]" pageId="6" pageNumber="114">
|
||
<materialsCitation id="3B153CE6FFB0FFDC94B5F909A0DBF917" ID-GBIF-Occurrence="3335033306" collectingDate="2003-10-10" collectingDateMax="2003-10-12" collectingDateMin="2003-10-10" collectionCode="ATOL" collectorName="ATOL Expedition 2003" country="Thailand" elevation="355" latitude="8.723666" location="Khao Luang NP" longLatPrecision="1" longitude="99.6688" pageId="6" pageNumber="114" specimenCode="NHMD 621694" specimenCount="1" specimenCount-female="1" stateProvince="Nakhon Si Thammarat Prov.">
|
||
<collectingCountry id="F36A762BFFB0FFDC94B5F909A266F96A" box="[189,341,1685,1711]" name="Thailand" pageId="6" pageNumber="114">THAILAND</collectingCountry>
|
||
•
|
||
<specimenCount id="9D7BFD32FFB0FFDC9567F909A2AFF96A" box="[367,412,1685,1711]" pageId="6" pageNumber="114" type="female">1 ♀</specimenCount>
|
||
;
|
||
<collectingRegion id="49B9F859FFB0FFDC95A3F908A1CCF96A" box="[427,767,1684,1711]" country="Thailand" name="Nakhon Si Thammarat" pageId="6" pageNumber="114">Nakhon Si Thammarat Prov.</collectingRegion>
|
||
,
|
||
<location id="8EA26060FFB0FFDC9703F908A0FDF96A" LSID="urn:lsid:plazi:treatment:03D487ADFFB0FFD695C3FC75A1CEFDC7:8EA26060FFB0FFDC9703F908A0FDF96A" box="[779,974,1684,1711]" country="Thailand" latitude="8.723666" longLatPrecision="1" longitude="99.6688" name="Khao Luang NP" pageId="6" pageNumber="114" stateProvince="Nakhon Si Thammarat Prov.">Khao Luang NP</location>
|
||
;
|
||
<geoCoordinate id="EE49507CFFB0FFDC97D3F90FA74AF96A" box="[987,1145,1683,1711]" degrees="8" direction="north" minutes="43" orientation="latitude" pageId="6" pageNumber="114" precision="1" seconds="25.2" value="8.723666">8°43′25.2″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE49507CFFB0FFDC908DF90FA62CF96A" box="[1157,1311,1683,1711]" degrees="99" direction="east" minutes="40" orientation="longitude" pageId="6" pageNumber="114" precision="1" seconds="7.7" value="99.6688">99°40′7.7″ E</geoCoordinate>
|
||
,
|
||
<elevation id="0050D188FFB0FFDC9124F908A3C5F917" metricMagnitude="2" metricUnit="m" metricValue="3.55" pageId="6" pageNumber="114" unit="m" value="355.0">
|
||
<quantity id="4C859B5EFFB0FFDC9124F908A644F96A" box="[1324,1399,1684,1711]" metricMagnitude="2" metricUnit="m" metricValue="3.55" pageId="6" pageNumber="114" unit="m" value="355.0">355 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="FFC3107BFFB0FFDC950BF924A2F8F917" box="[259,459,1720,1746]" pageId="6" pageNumber="114" value="2003-10-10" valueMax="2003-10-12" valueMin="2003-10-10">
|
||
<collectingDate id="EF87E993FFB0FFDC950BF924A2F8F917" box="[259,459,1720,1746]" pageId="6" pageNumber="114" value="2003-10-10" valueMax="2003-10-12" valueMin="2003-10-10">10–12 Oct. 2003</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="2688536DFFB0FFDC95DDF924A1DAF917" box="[469,745,1720,1746]" pageId="6" pageNumber="114">ATOL Expedition 2003</collectorName>
|
||
leg.;
|
||
<specimenCode id="DBDB9EC0FFB0FFDC9723F924A0DBF917" box="[811,1000,1720,1746]" collectionCode="NHMD" pageId="6" pageNumber="114">NHMD 621694</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3676530FFB0FFD294B5F965A646F807" lastPageId="8" lastPageNumber="116" pageId="6" pageNumber="114" type="description">
|
||
<paragraph id="8BC236BBFFB0FFDC94B5F965A10DF8D1" blockId="6.[189,574,1785,1812]" box="[189,574,1785,1812]" pageId="6" pageNumber="114">
|
||
<emphasis id="B909EAA9FFB0FFDC94B5F965A27DF8D6" bold="true" box="[189,334,1785,1811]" pageId="6" pageNumber="114">Description</emphasis>
|
||
(based on
|
||
<typeStatus id="54C68819FFB0FFDC95C6F965A104F8D1" box="[462,567,1785,1812]" pageId="6" pageNumber="114" type="holotype">holotype</typeStatus>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BC236BBFFB0FFDC94B5F8B4A0BFF8A0" blockId="6.[189,1398,1831,1893]" pageId="6" pageNumber="114">
|
||
SIZE. Length
|
||
<quantity id="4C859B5EFFB0FFDC9553F8B4A2FAF887" box="[347,457,1832,1858]" metricMagnitude="-2" metricUnit="m" metricValue="2.38" pageId="6" pageNumber="114" unit="mm" value="23.8">23.8 mm</quantity>
|
||
. Width of thoracic shield
|
||
<quantity id="4C859B5EFFB0FFDC9709F8BBA05CF887" box="[769,879,1831,1858]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="6" pageNumber="114" unit="mm" value="10.5">10.5 mm</quantity>
|
||
, of widest segment (9)
|
||
<quantity id="4C859B5EFFB0FFDC9083F8B4A7C4F887" box="[1163,1271,1832,1858]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="6" pageNumber="114" unit="mm" value="11.1">11.1 mm</quantity>
|
||
. Height of thoracic shield
|
||
<quantity id="4C859B5EFFB0FFDC957AF8D7A2FCF8A0" box="[370,463,1867,1893]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="6" pageNumber="114" unit="mm" value="6.2">6.2 mm</quantity>
|
||
, Height of highest segment (7)
|
||
<quantity id="4C859B5EFFB0FFDC9748F8D7A0B4F8A0" box="[832,903,1867,1893]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="114" unit="mm" value="7.0">7 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC236BBFFB0FFD294B5F810A12FFE49" blockId="6.[188,1399,1932,1994]" lastBlockId="8.[187,1399,265,397]" lastPageId="8" lastPageNumber="116" pageId="6" pageNumber="114">
|
||
COLOUR. Head, antenna, legs and tergites golden light brown. Posterior margin of tergites and paratergite tips medium brown. Anal shield medially brown, with lighter colour at edges (
|
||
<figureCitation id="13462A3EFFB0FFDC905BF833A781F80F" box="[1107,1202,1967,1994]" captionStart="Fig" captionStartId="7.[189,232,1882,1908]" captionTargetBox="[202,1381,333,1766]" captionTargetId="figure-16@7.[189,1398,321,1784]" captionTargetPageId="7" captionText="Fig. 2. Map of Thailand, collection localities of Sphaerobelum aesculus Rosenmejer & Wesener sp. nov. and Zephronia viridisoma Rosenmejer & Wesener sp. nov., and habitus photographs: S. aesculus Rosenmejer & Wesener sp. nov., ♀ (NHMD 621694); Z. viridisoma Rosenmejer & Wesener sp. nov., paratype, ♀ (ZFMK MYR8787). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5211731" httpUri="https://zenodo.org/record/5211731/files/figure.png" pageId="6" pageNumber="114">Fig. 2A</figureCitation>
|
||
). HEAD. Number of ommatidia 55. Antennae (
|
||
<figureCitation id="13462A3EFFBEFFD296B0FE95A00BFEE6" box="[696,824,265,291]" captionStart="Fig" captionStartId="9.[189,232,1882,1908]" captionTargetBox="[190,1397,310,1797]" captionTargetId="figure-15@9.[189,1398,309,1798]" captionTargetPageId="9" captionText="Fig. 3. Sphaerobelum aesculus Rosenmejer & Wesener sp. nov., holotype, ♂ (NHMD 621693), scanning electron micrographs. A. Right antenna, lateral view. B. Right antenna, disc. C. Detail of apical cone. D. Tergite surface of midbody ring. E. Tergite surface at very high magnification. Abbreviations: ac = apical cone; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.5211733" httpUri="https://zenodo.org/record/5211733/files/figure.png" pageId="8" pageNumber="116">Fig. 3A–C</figureCitation>
|
||
): Antennomere lengths: 6>1>2=3=4=5. Antennae short, barely reaching first leg pair. Sixth antennomere apically slightly swollen (
|
||
<figureCitation id="13462A3EFFBEFFD2908BFEB0A7D7FE83" box="[1155,1252,300,326]" captionStart="Fig" captionStartId="9.[189,232,1882,1908]" captionTargetBox="[190,1397,310,1797]" captionTargetId="figure-15@9.[189,1398,309,1798]" captionTargetPageId="9" captionText="Fig. 3. Sphaerobelum aesculus Rosenmejer & Wesener sp. nov., holotype, ♂ (NHMD 621693), scanning electron micrographs. A. Right antenna, lateral view. B. Right antenna, disc. C. Detail of apical cone. D. Tergite surface of midbody ring. E. Tergite surface at very high magnification. Abbreviations: ac = apical cone; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.5211733" httpUri="https://zenodo.org/record/5211733/files/figure.png" pageId="8" pageNumber="116">Fig. 3A</figureCitation>
|
||
), number of apical cones 52/56 (
|
||
<figureCitation id="13462A3EFFBEFFD295AEFED3A132FEAC" box="[422,513,335,361]" captionStart="Fig" captionStartId="9.[189,232,1882,1908]" captionTargetBox="[190,1397,310,1797]" captionTargetId="figure-15@9.[189,1398,309,1798]" captionTargetPageId="9" captionText="Fig. 3. Sphaerobelum aesculus Rosenmejer & Wesener sp. nov., holotype, ♂ (NHMD 621693), scanning electron micrographs. A. Right antenna, lateral view. B. Right antenna, disc. C. Detail of apical cone. D. Tergite surface of midbody ring. E. Tergite surface at very high magnification. Abbreviations: ac = apical cone; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.5211733" httpUri="https://zenodo.org/record/5211733/files/figure.png" pageId="8" pageNumber="116">Fig. 3B</figureCitation>
|
||
). Mandible: not dissected. Gnathochilarium: lingual lamella with numerous long setae, medially glabrous.
|
||
</paragraph>
|
||
<caption id="DF026633FFB1FFDD94B5F8C6A012F81A" ID-DOI="http://doi.org/10.5281/zenodo.5211731" ID-Zenodo-Dep="5211731" httpUri="https://zenodo.org/record/5211731/files/figure.png" pageId="7" pageNumber="115" startId="7.[189,232,1882,1908]" targetBox="[202,1381,333,1766]" targetPageId="7">
|
||
<paragraph id="8BC236BBFFB1FFDD94B5F8C6A012F81A" blockId="7.[188,1400,1882,2015]" pageId="7" pageNumber="115">
|
||
<emphasis id="B909EAA9FFB1FFDD94B5F8C6A239F8B1" bold="true" box="[189,266,1882,1908]" pageId="7" pageNumber="115">Fig. 2.</emphasis>
|
||
Map of Thailand, collection localities of
|
||
<taxonomicName id="4C7D4D38FFB1FFDD96E1F8C7A624F8B0" authority="Rosenmejer & Wesener" authorityName="Rosenmejer & Enghoff & Moritz & Wesener" authorityYear="2021" box="[745,1303,1883,1909]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="7" pageNumber="115" phylum="Arthropoda" rank="species" species="aesculus" status="sp. nov.">
|
||
<emphasis id="B909EAA9FFB1FFDD96E1F8C7A0CFF8B0" box="[745,1020,1883,1909]" italics="true" pageId="7" pageNumber="115">Sphaerobelum aesculus</emphasis>
|
||
Rosenmejer & Wesener
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23A57D2FFB1FFDD9114F8C7A646F8B0" box="[1308,1397,1883,1909]" pageId="7" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C7D4D38FFB1FFDD94FDF8E2A01AF85D" authority="Rosenmejer & Wesener" authorityName="Rosenmejer & Enghoff & Moritz & Wesener" authorityYear="2021" box="[245,809,1918,1944]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="7" pageNumber="115" phylum="Arthropoda" rank="species" species="viridisoma" status="sp. nov.">
|
||
<emphasis id="B909EAA9FFB1FFDD94FDF8E2A2CAF85D" box="[245,505,1918,1944]" italics="true" pageId="7" pageNumber="115">Zephronia viridisoma</emphasis>
|
||
Rosenmejer & Wesener
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23A57D2FFB1FFDD973FF8E2A0A9F85D" box="[823,922,1918,1944]" pageId="7" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and habitus photographs:
|
||
<taxonomicName id="4C7D4D38FFB1FFDD90E6F8E2A2E5F879" authority="Rosenmejer & Wesener" authorityName="Rosenmejer & Enghoff & Moritz & Wesener" authorityYear="2021" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="7" pageNumber="115" phylum="Arthropoda" rank="species" species="aesculus" status="sp. nov.">
|
||
<emphasis id="B909EAA9FFB1FFDD90E6F8E2A645F85D" box="[1262,1398,1918,1944]" italics="true" pageId="7" pageNumber="115">S. aesculus</emphasis>
|
||
Rosenmejer & Wesener
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23A57D2FFB1FFDD95E8F83EA10DF879" box="[480,574,1954,1980]" pageId="7" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, ♀ (NHMD 621694);
|
||
<taxonomicName id="4C7D4D38FFB1FFDD9742F83EA639F879" authority="Rosenmejer & Wesener" authorityName="Rosenmejer & Enghoff & Moritz & Wesener" authorityYear="2021" box="[842,1290,1953,1980]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="7" pageNumber="115" phylum="Arthropoda" rank="species" species="viridisoma" status="sp. nov.">
|
||
<emphasis id="B909EAA9FFB1FFDD9742F83EA0DBF87E" box="[842,1000,1953,1979]" italics="true" pageId="7" pageNumber="115">Z. viridisoma</emphasis>
|
||
Rosenmejer & Wesener
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23A57D2FFB1FFDD911CF83EA640F879" box="[1300,1395,1954,1980]" pageId="7" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, ♀ (ZFMK MYR8787). Scale bars = 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC236BBFFBEFFD294B5FE28A014FE0B" blockId="8.[189,807,436,463]" box="[189,807,436,463]" pageId="8" pageNumber="116">COLLUM. With few setae spread thinly along borders.</paragraph>
|
||
<paragraph id="8BC236BBFFBEFFD294B5FE6AA653FDCA" blockId="8.[189,1376,501,528]" box="[189,1376,501,528]" pageId="8" pageNumber="116">THORACIC SHIELD. Thoracic shield grooves wide and deep, with 3 sclerotized ledges along inner ridge.</paragraph>
|
||
<paragraph id="8BC236BBFFBEFFD294B5FDABA0D8FDB1" blockId="8.[188,1399,567,629]" pageId="8" pageNumber="116">
|
||
TERGITES. Paratergite tips on posterior half projecting backwards. Tergites glabrous with dull leather-like surface (
|
||
<figureCitation id="13462A3EFFBEFFD29529FDC6A292FDB1" box="[289,417,602,628]" captionStart="Fig" captionStartId="9.[189,232,1882,1908]" captionTargetBox="[190,1397,310,1797]" captionTargetId="figure-15@9.[189,1398,309,1798]" captionTargetPageId="9" captionText="Fig. 3. Sphaerobelum aesculus Rosenmejer & Wesener sp. nov., holotype, ♂ (NHMD 621693), scanning electron micrographs. A. Right antenna, lateral view. B. Right antenna, disc. C. Detail of apical cone. D. Tergite surface of midbody ring. E. Tergite surface at very high magnification. Abbreviations: ac = apical cone; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.5211733" httpUri="https://zenodo.org/record/5211733/files/figure.png" pageId="8" pageNumber="116">Fig. 3D–E</figureCitation>
|
||
). At high magnification tiny setae become visible.
|
||
</paragraph>
|
||
<paragraph id="8BC236BBFFBEFFD294B5FD00A7E2FD70" blockId="8.[189,1233,667,694]" box="[189,1233,667,694]" pageId="8" pageNumber="116">
|
||
ENDOTERGUM (
|
||
<figureCitation id="13462A3EFFBEFFD2956DFD07A2F7FD73" box="[357,452,667,694]" captionStart="Fig" captionStartId="12.[189,232,1916,1942]" captionTargetBox="[194,1394,1255,1873]" captionTargetId="figure-181@12.[189,1398,1247,1878]" captionTargetPageId="12" captionText="Fig. 6. Endoterga of midbody tergites, scanning electron micrographs. A. Sphaerobelum aesculus Rosenmejer & Wesener sp. nov., holotype (NHMD 621693). B. Zephronia viridisoma Rosenmejer & Wesener sp. nov., holotype, ♂ (NHMD 621695). Abbreviations: cp = cuticular patterns; iA = inner area." figureDoi="http://doi.org/10.5281/zenodo.5211743" httpUri="https://zenodo.org/record/5211743/files/figure.png" pageId="8" pageNumber="116">Fig. 6A</figureCitation>
|
||
). With a regular flat margin. Outer zone with two rows of irregular
|
||
</paragraph>
|
||
<paragraph id="8BC236BBFFBEFFD294B5FD7EA29BFCFB" blockId="8.[189,1399,731,830]" pageId="8" pageNumber="116">MARGINAL SETAE. Not extending beyond posterior margin, but reaching ⅔ of the outer area. Anterior part of marginal ridge flat. Intersegmental membrane smooth, without cones and with a row of setae with large glabrous gaps.</paragraph>
|
||
<paragraph id="8BC236BBFFBEFFD294B5FCF9A041FCBA" blockId="8.[189,882,869,895]" box="[189,882,869,895]" pageId="8" pageNumber="116">
|
||
FIRST STIGMATIC PLATE (
|
||
<figureCitation id="13462A3EFFBEFFD295C0FCF9A115FCBA" box="[456,550,869,895]" captionStart="Fig" captionStartId="10.[189,232,1814,1840]" captionTargetBox="[285,1308,283,1724]" captionTargetId="figure-11@10.[258,1329,265,1777]" captionTargetPageId="10" captionText="Fig. 4. Sphaerobelum aesculus Rosenmejer & Wesenersp. nov., holotype, ♂ (NHMD 621693), drawings. A. First left coxa with stigmatic plate, posterior view. B. Second left coxa with gonopore, posterior view. C. Left leg 9, posterior view. D. Left anterior telopod, posterior view. E. Left anterior telopod, anterior view. F. Left anterior telopod, lateral view on podomeres 2–4. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; St = stigmatic plate; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5211739" httpUri="https://zenodo.org/record/5211739/files/figure.png" pageId="8" pageNumber="116">Fig. 4A</figureCitation>
|
||
). With a well-rounded apex.
|
||
</paragraph>
|
||
<paragraph id="8BC236BBFFBEFFD294B5FC3BA228FC26" blockId="8.[189,1398,934,995]" pageId="8" pageNumber="116">PLEURITES. Pleurite 1 projecting posteriorly with sharp apex. Pleurite 2 with rounded apex projecting slightly.</paragraph>
|
||
<paragraph id="8BC236BBFFBEFFD294B5FB97A077FBAE" blockId="8.[189,1399,1035,1132]" pageId="8" pageNumber="116">
|
||
LEGS. Ventral spines on leg 1 4/4, on leg 2 3/5, on leg 3 3/4. Apical spine absent on leg 3. Three apical and 4 or 5 ventral spines on midbody legs (
|
||
<figureCitation id="13462A3EFFBEFFD296C8FBB2A02DFB8D" box="[704,798,1070,1096]" captionStart="Fig" captionStartId="10.[189,232,1814,1840]" captionTargetBox="[285,1308,283,1724]" captionTargetId="figure-11@10.[258,1329,265,1777]" captionTargetPageId="10" captionText="Fig. 4. Sphaerobelum aesculus Rosenmejer & Wesenersp. nov., holotype, ♂ (NHMD 621693), drawings. A. First left coxa with stigmatic plate, posterior view. B. Second left coxa with gonopore, posterior view. C. Left leg 9, posterior view. D. Left anterior telopod, posterior view. E. Left anterior telopod, anterior view. F. Left anterior telopod, lateral view on podomeres 2–4. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; St = stigmatic plate; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5211739" httpUri="https://zenodo.org/record/5211739/files/figure.png" pageId="8" pageNumber="116">Fig. 4C</figureCitation>
|
||
). Inner margin of femur with 8–12 small teeth, but not excavated. Femur 2, tarsus 4 times as long as wide.
|
||
</paragraph>
|
||
<paragraph id="8BC236BBFFBEFFD294B5FB0FA22DFB15" blockId="8.[189,1398,1169,1232]" pageId="8" pageNumber="116">ANAL SHIELD. Well-rounded and glabrous. Ventral side with a single small locking carina, placed ⅓ from pleurite.</paragraph>
|
||
<paragraph id="8BC236BBFFBEFFD294B5FB64A67EFAD4" blockId="8.[189,1357,1271,1298]" box="[189,1357,1271,1298]" pageId="8" pageNumber="116">
|
||
MALE GONOPORE. Located at mesal margin of coxa (
|
||
<figureCitation id="13462A3EFFBEFFD2971EFB6BA041FAD4" box="[790,882,1271,1297]" captionStart="Fig" captionStartId="10.[189,232,1814,1840]" captionTargetBox="[285,1308,283,1724]" captionTargetId="figure-11@10.[258,1329,265,1777]" captionTargetPageId="10" captionText="Fig. 4. Sphaerobelum aesculus Rosenmejer & Wesenersp. nov., holotype, ♂ (NHMD 621693), drawings. A. First left coxa with stigmatic plate, posterior view. B. Second left coxa with gonopore, posterior view. C. Left leg 9, posterior view. D. Left anterior telopod, posterior view. E. Left anterior telopod, anterior view. F. Left anterior telopod, lateral view on podomeres 2–4. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; St = stigmatic plate; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5211739" httpUri="https://zenodo.org/record/5211739/files/figure.png" pageId="8" pageNumber="116">Fig. 4B</figureCitation>
|
||
), large, covered by a membranous plate.
|
||
</paragraph>
|
||
<paragraph id="8BC236BBFFBEFFD294B5FAA5A7A5FA25" blockId="8.[188,1399,1337,1504]" pageId="8" pageNumber="116">
|
||
ANTERIOR TELOPODS (
|
||
<figureCitation id="13462A3EFFBEFFD295A6FAA5A118FA96" box="[430,555,1337,1363]" captionStart="Fig" captionStartId="10.[189,232,1814,1840]" captionTargetBox="[285,1308,283,1724]" captionTargetId="figure-11@10.[258,1329,265,1777]" captionTargetPageId="10" captionText="Fig. 4. Sphaerobelum aesculus Rosenmejer & Wesenersp. nov., holotype, ♂ (NHMD 621693), drawings. A. First left coxa with stigmatic plate, posterior view. B. Second left coxa with gonopore, posterior view. C. Left leg 9, posterior view. D. Left anterior telopod, posterior view. E. Left anterior telopod, anterior view. F. Left anterior telopod, lateral view on podomeres 2–4. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; St = stigmatic plate; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5211739" httpUri="https://zenodo.org/record/5211739/files/figure.png" pageId="8" pageNumber="116">Fig. 4D–F</figureCitation>
|
||
). Podomere 1 rectangular, as long as wide, covered with setae anteriorly, with extra long setae medially, and posteriorly glabrous. Podomere 2 with long immovable finger visible laterally in anterior view. Finger curving inwards against podomere 3. Podomere 3 cylindrical twice as long as wide. Suture between podomere 3 and 4 barely visible in posterior and lateral views. Podomere 4 very short, just ¼ of length of podomere 3, with one short and dark spine on apex.
|
||
</paragraph>
|
||
<paragraph id="8BC236BBFFBEFFD294B5F994A0FFF931" blockId="8.[189,1400,1543,1781]" pageId="8" pageNumber="116">
|
||
POSTERIOR TELOPODS (
|
||
<figureCitation id="13462A3EFFBEFFD295B1F99BA10FF9E7" box="[441,572,1543,1570]" captionStart="Fig" captionStartId="11.[189,232,1846,1872]" captionTargetBox="[308,1338,326,1743]" captionTargetId="figure-15@11.[245,1342,265,1805]" captionTargetPageId="11" captionText="Fig. 5. Sphaerobelum aesculus Rosenmejer & Wesener sp. nov. drawings. A–B. Holotype, ♂ (NHMD 621693), C. ♀ (NHMD 621694). A. Left posterior telopod, anterior view. B. Left posterior telopod, posterior view, arrow points to swelling of immovable finger. C. Left second coxa and prefemur with female vulva. Abbreviations: Cx = coxa; EP = external, lateral plate; IP = inner, mesal plate; Op = operculum; Pre = prefemur. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5211741" httpUri="https://zenodo.org/record/5211741/files/figure.png" pageId="8" pageNumber="116">Fig. 5A–B</figureCitation>
|
||
). Podomere 1 as long as wide, covered in setae anteriorly, and nearly glabrous posteriorly. Podomere 2 with wide immovable finger. Immovable finger straight, only slightly tapering towards apex, bearing an oval membranous spot on posterior side. The membrane between podomere 2 and 3 with single membranous lobe, angled. Podomere 3 with sparse setation. Podomere 3 ca 3 times longer than wide, towards immovable finger with membranous ledge, a single spine, posterior side with row of 9 or 10 crenulated teeth. Podomere 4 short, straight, ca 2.5 times as long as wide, with a short membranous ledge and 2 spines towards immovable finger.
|
||
</paragraph>
|
||
<paragraph id="8BC236BBFFBEFFD294E2F880A645F89C" blockId="8.[187,1401,1819,1987]" pageId="8" pageNumber="116">
|
||
<emphasis id="B909EAA9FFBEFFD294E2F880A16EF8F3" bold="true" box="[234,605,1819,1846]" pageId="8" pageNumber="116">Female from Khao Luang NP</emphasis>
|
||
Length
|
||
<quantity id="4C859B5EFFBEFFD29512F8A3A241F89C" box="[282,370,1855,1882]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="116" unit="mm" value="45.0">45 mm</quantity>
|
||
. Width of thoracic shield
|
||
<quantity id="4C859B5EFFBEFFD296A3F8A3A030F89C" box="[683,771,1855,1881]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="116" unit="mm" value="23.0">23 mm</quantity>
|
||
, of widest segment (9)
|
||
<quantity id="4C859B5EFFBEFFD29029F8A3A7BCF89C" box="[1057,1167,1855,1881]" metricMagnitude="-2" metricUnit="m" metricValue="2.46" pageId="8" pageNumber="116" unit="mm" value="24.6">24.6 mm</quantity>
|
||
. Height of thoracic
|
||
</paragraph>
|
||
<paragraph id="8BC236BBFFBEFFD294B7F8FEA646F807" blockId="8.[187,1401,1819,1987]" pageId="8" pageNumber="116">
|
||
shield
|
||
<quantity id="4C859B5EFFBEFFD29503F8FEA246F8B8" box="[267,373,1890,1917]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="8" pageNumber="116" unit="mm" value="12.6">12.6 mm</quantity>
|
||
, of highest segment (8)
|
||
<quantity id="4C859B5EFFBEFFD2968FF8FEA1C2F8B8" box="[647,753,1890,1917]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="8" pageNumber="116" unit="mm" value="15.2">15.2 mm</quantity>
|
||
.Antennae reaching leg pair 2. Apical cones on antennae 43/34. Subanal plate well rounded. Vulvae (
|
||
<figureCitation id="13462A3EFFBEFFD296BAF819A03EF865" box="[690,781,1925,1952]" captionStart="Fig" captionStartId="11.[189,232,1846,1872]" captionTargetBox="[308,1338,326,1743]" captionTargetId="figure-15@11.[245,1342,265,1805]" captionTargetPageId="11" captionText="Fig. 5. Sphaerobelum aesculus Rosenmejer & Wesener sp. nov. drawings. A–B. Holotype, ♂ (NHMD 621693), C. ♀ (NHMD 621694). A. Left posterior telopod, anterior view. B. Left posterior telopod, posterior view, arrow points to swelling of immovable finger. C. Left second coxa and prefemur with female vulva. Abbreviations: Cx = coxa; EP = external, lateral plate; IP = inner, mesal plate; Op = operculum; Pre = prefemur. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5211741" httpUri="https://zenodo.org/record/5211741/files/figure.png" pageId="8" pageNumber="116">Fig. 5C</figureCitation>
|
||
): small, with a narrow operculum (Op), only reaching bottom part of prefemur. External lateral plate (EP) and inner mesal plate (IP) almost completely fused.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF026633FFBFFFD394B5F8C6A15DF81B" ID-DOI="http://doi.org/10.5281/zenodo.5211733" ID-Zenodo-Dep="5211733" httpUri="https://zenodo.org/record/5211733/files/figure.png" pageId="9" pageNumber="117" startId="9.[189,232,1882,1908]" targetBox="[190,1397,310,1797]" targetPageId="9">
|
||
<paragraph id="8BC236BBFFBFFFD394B5F8C6A15DF81B" blockId="9.[189,1399,1882,2015]" pageId="9" pageNumber="117">
|
||
<emphasis id="B909EAA9FFBFFFD394B5F8C6A239F8B1" bold="true" box="[189,266,1882,1908]" pageId="9" pageNumber="117">Fig. 3.</emphasis>
|
||
<taxonomicName id="4C7D4D38FFBFFFD39518F8C7A008F8B0" authority="Rosenmejer & Wesener" authorityName="Rosenmejer & Enghoff & Moritz & Wesener" authorityYear="2021" box="[272,827,1883,1909]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="9" pageNumber="117" phylum="Arthropoda" rank="species" species="aesculus" status="sp. nov.">
|
||
<emphasis id="B909EAA9FFBFFFD39518F8C7A110F8B0" box="[272,547,1883,1909]" italics="true" pageId="9" pageNumber="117">Sphaerobelum aesculus</emphasis>
|
||
Rosenmejer & Wesener
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23A57D2FFBFFFD39737F8C7A0AFF8B0" box="[831,924,1883,1909]" pageId="9" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂ (NHMD 621693), scanning electron micrographs.
|
||
<emphasis id="B909EAA9FFBFFFD395C0F8E2A2EFF85D" bold="true" box="[456,476,1918,1944]" pageId="9" pageNumber="117">A</emphasis>
|
||
. Right antenna, lateral view.
|
||
<emphasis id="B909EAA9FFBFFFD39734F8E2A07DF85D" bold="true" box="[828,846,1918,1944]" pageId="9" pageNumber="117">B</emphasis>
|
||
. Right antenna, disc.
|
||
<emphasis id="B909EAA9FFBFFFD3905CF8E2A75BF85D" bold="true" box="[1108,1128,1918,1944]" pageId="9" pageNumber="117">C</emphasis>
|
||
. Detail of apical cone.
|
||
<emphasis id="B909EAA9FFBFFFD394B6F83DA3E1F87E" bold="true" box="[190,210,1953,1979]" pageId="9" pageNumber="117">D</emphasis>
|
||
. Tergite surface of midbody ring.
|
||
<emphasis id="B909EAA9FFBFFFD39664F83DA14DF87E" bold="true" box="[620,638,1953,1979]" pageId="9" pageNumber="117">E</emphasis>
|
||
. Tergite surface at very high magnification. Abbreviations: ac = apical cone; sb = sensilla basiconica.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF026633FFBCFFD094B5F88AA06BF824" ID-DOI="http://doi.org/10.5281/zenodo.5211739" ID-Zenodo-Dep="5211739" httpUri="https://zenodo.org/record/5211739/files/figure.png" pageId="10" pageNumber="118" startId="10.[189,232,1814,1840]" targetBox="[285,1308,283,1724]" targetPageId="10">
|
||
<paragraph id="8BC236BBFFBCFFD094B5F88AA06BF824" blockId="10.[188,1401,1814,2017]" pageId="10" pageNumber="118">
|
||
<emphasis id="B909EAA9FFBCFFD094B5F88AA23AF8F5" bold="true" box="[189,265,1814,1840]" pageId="10" pageNumber="118">Fig. 4.</emphasis>
|
||
<taxonomicName id="4C7D4D38FFBCFFD09506F88AA06AF8F4" authority="Rosenmejer & Wesenersp." authorityName="Rosenmejer & Wesenersp." box="[270,857,1814,1841]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="118" phylum="Arthropoda" rank="species" species="aesculus">
|
||
<emphasis id="B909EAA9FFBCFFD09506F88AA113F8F5" box="[270,544,1814,1840]" italics="true" pageId="10" pageNumber="118">Sphaerobelum aesculus</emphasis>
|
||
Rosenmejer & Wesenersp.
|
||
</taxonomicName>
|
||
nov., holotype, ♂ (NHMD 621693), drawings.
|
||
<emphasis id="B909EAA9FFBCFFD094B5F8A5A3E1F896" bold="true" box="[189,210,1849,1875]" pageId="10" pageNumber="118">A</emphasis>
|
||
. First left coxa with stigmatic plate, posterior view.
|
||
<emphasis id="B909EAA9FFBCFFD09722F8A6A00DF891" bold="true" box="[810,830,1850,1876]" pageId="10" pageNumber="118">B</emphasis>
|
||
. Second left coxa with gonopore, posterior view.
|
||
<emphasis id="B909EAA9FFBCFFD094B4F8C1A3E2F8B2" bold="true" box="[188,209,1885,1911]" pageId="10" pageNumber="118">C</emphasis>
|
||
. Left leg 9, posterior view.
|
||
<emphasis id="B909EAA9FFBCFFD09612F8C1A11CF8B2" bold="true" box="[538,559,1885,1911]" pageId="10" pageNumber="118">D</emphasis>
|
||
. Left anterior telopod, posterior view.
|
||
<emphasis id="B909EAA9FFBCFFD097FEF8C1A73AF8B2" bold="true" box="[1014,1033,1885,1911]" pageId="10" pageNumber="118">E</emphasis>
|
||
. Left anterior telopod, anterior view.
|
||
<emphasis id="B909EAA9FFBCFFD0950DF81CA224F85F" bold="true" box="[261,279,1920,1946]" pageId="10" pageNumber="118">F</emphasis>
|
||
. Left anterior telopod, lateral view on podomeres 2–4. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; St = stigmatic plate; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF026633FFBDFFD194B5F8AAA1D8F81B" ID-DOI="http://doi.org/10.5281/zenodo.5211741" ID-Zenodo-Dep="5211741" httpUri="https://zenodo.org/record/5211741/files/figure.png" pageId="11" pageNumber="119" startId="11.[189,232,1846,1872]" targetBox="[308,1338,326,1743]" targetPageId="11">
|
||
<paragraph id="8BC236BBFFBDFFD194B5F8AAA1D8F81B" blockId="11.[189,1401,1846,2014]" pageId="11" pageNumber="119">
|
||
<emphasis id="B909EAA9FFBDFFD194B5F8AAA23FF894" bold="true" box="[189,268,1846,1873]" pageId="11" pageNumber="119">Fig. 5.</emphasis>
|
||
<taxonomicName id="4C7D4D38FFBDFFD1951DF8ABA079F894" authority="Rosenmejer & Wesener" authorityName="Rosenmejer & Enghoff & Moritz & Wesener" authorityYear="2021" box="[277,842,1847,1873]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="119" phylum="Arthropoda" rank="species" species="aesculus" status="sp. nov.">
|
||
<emphasis id="B909EAA9FFBDFFD1951DF8ABA119F894" box="[277,554,1847,1873]" italics="true" pageId="11" pageNumber="119">Sphaerobelum aesculus</emphasis>
|
||
Rosenmejer & Wesener
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23A57D2FFBDFFD1975BF8ABA09DF894" box="[851,942,1847,1873]" pageId="11" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
|
||
drawings.
|
||
<emphasis id="B909EAA9FFBDFFD1903AF8AAA75BF895" bold="true" box="[1074,1128,1846,1872]" pageId="11" pageNumber="119">A–B</emphasis>
|
||
. Holotype, ♂ (NHMD 621693),
|
||
<emphasis id="B909EAA9FFBDFFD19538F8C6A276F8B1" bold="true" box="[304,325,1882,1908]" pageId="11" pageNumber="119">C</emphasis>
|
||
. ♀ (NHMD 621694).
|
||
<emphasis id="B909EAA9FFBDFFD19650F8C6A15EF8B1" bold="true" box="[600,621,1882,1908]" pageId="11" pageNumber="119">A</emphasis>
|
||
. Left posterior telopod, anterior view.
|
||
<emphasis id="B909EAA9FFBDFFD1904CF8C6A76BF8B1" bold="true" box="[1092,1112,1882,1908]" pageId="11" pageNumber="119">B</emphasis>
|
||
. Left posterior telopod, posterior view, arrow points to swelling of immovable finger.
|
||
<emphasis id="B909EAA9FFBDFFD197A9F8E1A085F852" bold="true" box="[929,950,1917,1943]" pageId="11" pageNumber="119">C</emphasis>
|
||
. Left second coxa and prefemur with female vulva. Abbreviations: Cx = coxa; EP = external, lateral plate; IP = inner, mesal plate; Op = operculum; Pre = prefemur. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3676530FFBAFFD694B5FE94A1CEFDC7" pageId="12" pageNumber="120" type="distribution">
|
||
<paragraph id="8BC236BBFFBAFFD694B5FE94A265FEE7" blockId="12.[189,342,264,290]" box="[189,342,264,290]" pageId="12" pageNumber="120">
|
||
<heading id="D08A81D7FFBAFFD694B5FE94A265FEE7" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="12" pageNumber="120" reason="3">
|
||
<emphasis id="B909EAA9FFBAFFD694B5FE94A265FEE7" bold="true" box="[189,342,264,290]" pageId="12" pageNumber="120">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC236BBFFBAFFD694B5FEABA1CEFDC7" blockId="12.[189,1399,311,514]" pageId="12" pageNumber="120">
|
||
If the female from Khao Luang (
|
||
<figureCitation id="13462A3EFFBAFFD69659FEABA182FE94" box="[593,689,311,337]" captionStart="Fig" captionStartId="7.[189,232,1882,1908]" captionTargetBox="[202,1381,333,1766]" captionTargetId="figure-16@7.[189,1398,321,1784]" captionTargetPageId="7" captionText="Fig. 2. Map of Thailand, collection localities of Sphaerobelum aesculus Rosenmejer & Wesener sp. nov. and Zephronia viridisoma Rosenmejer & Wesener sp. nov., and habitus photographs: S. aesculus Rosenmejer & Wesener sp. nov., ♀ (NHMD 621694); Z. viridisoma Rosenmejer & Wesener sp. nov., paratype, ♀ (ZFMK MYR8787). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5211731" httpUri="https://zenodo.org/record/5211731/files/figure.png" pageId="12" pageNumber="120">Fig. 2A</figureCitation>
|
||
) is indeed conspecific with the male
|
||
<typeStatus id="54C68819FFBAFFD69072FEABA7D7FE94" box="[1146,1252,311,337]" pageId="12" pageNumber="120" type="holotype">holotype</typeStatus>
|
||
, the species appears to have a wide area of distribution stretching from
|
||
<collectingRegion id="49B9F859FFBAFFD69782FEC6A0E8FEB1" box="[906,987,346,372]" country="Thailand" name="Phuket" pageId="12" pageNumber="120">Phuket</collectingRegion>
|
||
Island at least
|
||
<quantity id="4C859B5EFFBAFFD6909DFEC6A7C3FEB1" box="[1173,1264,346,372]" metricMagnitude="5" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="120" unit="km" value="160.0">160 km</quantity>
|
||
to the east (
|
||
<figureCitation id="13462A3EFFBAFFD694CEFEE1A221FE5D" box="[198,274,381,408]" captionStart="Fig" captionStartId="7.[189,232,1882,1908]" captionTargetBox="[202,1381,333,1766]" captionTargetId="figure-16@7.[189,1398,321,1784]" captionTargetPageId="7" captionText="Fig. 2. Map of Thailand, collection localities of Sphaerobelum aesculus Rosenmejer & Wesener sp. nov. and Zephronia viridisoma Rosenmejer & Wesener sp. nov., and habitus photographs: S. aesculus Rosenmejer & Wesener sp. nov., ♀ (NHMD 621694); Z. viridisoma Rosenmejer & Wesener sp. nov., paratype, ♀ (ZFMK MYR8787). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5211731" httpUri="https://zenodo.org/record/5211731/files/figure.png" pageId="12" pageNumber="120">Fig. 2</figureCitation>
|
||
). Unpublished data from a larger inventory project (see, e.g., Wesener
|
||
<emphasis id="B909EAA9FFBAFFD69076FEE2A78EFE52" box="[1150,1213,381,407]" italics="true" pageId="12" pageNumber="120">et al.</emphasis>
|
||
2021) of giant pill-millipedes in the surroundings of
|
||
<collectingRegion id="49B9F859FFBAFFD69684FE3DA1E1FE7E" box="[652,722,417,443]" country="Thailand" name="Krabi" pageId="12" pageNumber="120">Krabi</collectingRegion>
|
||
, half way between
|
||
<collectingRegion id="49B9F859FFBAFFD697CCFE3DA726FE7E" box="[964,1045,417,443]" country="Thailand" name="Phuket" pageId="12" pageNumber="120">Phuket</collectingRegion>
|
||
Island and Khao Luang, did not recover this species among the numerous specimens, hinting at a patchy distribution and specific microhabitat requirements of
|
||
<taxonomicName id="4C7D4D38FFBAFFD69611FE7BA1A8FDC4" authority="Rosenmejer & Enghoff & Moritz & Wesener, 2021" authorityName="Rosenmejer & Enghoff & Moritz & Wesener" authorityYear="2021" box="[537,667,487,513]" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="120" phylum="Arthropoda" rank="species" species="aesculus" status="sp. nov.">
|
||
<emphasis id="B909EAA9FFBAFFD69611FE7BA1A8FDC4" box="[537,667,487,513]" italics="true" pageId="12" pageNumber="120">S. aesculus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23A57D2FFBAFFD696AAFE74A1CEFDC7" box="[674,765,488,514]" pageId="12" pageNumber="120" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |