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<document id="B2764112E1A8D58725927D14C2DEE193" ID-DOI="10.4081/nhs.2017.310" ID-ISSN="2385-0922" ID-Zenodo-Dep="12753805" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1721184724886" checkinUser="felipe" docAuthor="Pasini, Giovanni &amp; Garassino, Alessandro" docDate="2017" docId="03CD830F7B046B6BFCAEFCE08673A2A9" docLanguage="en" docName="NatHistSci.4.1.43-72.pdf" docOrigin="Natural History Sciences 4 (1)" docSource="http://dx.doi.org/10.4081/nhs.2017.310" docStyle="DocumentStyle:926963A4B76E17DFFD817EF231EB8DA9.3:NatHistSci.2014-.journal_article" docStyleId="926963A4B76E17DFFD817EF231EB8DA9" docStyleName="NatHistSci.2014-.journal_article" docStyleVersion="3" docTitle="Ranina elegans Rathbun 1945" docType="treatment" docVersion="1" lastPageNumber="60" masterDocId="FFF4FB777B146B7AFF91FF998441A166" masterDocTitle="A preliminary review of the fossil species of Ranina Lamarck, 1801 (Decapoda, Brachyura, Raninidae), with systematic remarks" masterLastPageNumber="72" masterPageNumber="43" pageNumber="59" updateTime="1726747683517" updateUser="guilherme" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="5D40B0A53B65D38DB60750CFC0FD35CC">A preliminary review of the fossil species of Ranina Lamarck, 1801 (Decapoda, Brachyura, Raninidae), with systematic remarks</mods:title>
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<mods:namePart id="12CEF43373FEA529B4C16CB768334866">Pasini, Giovanni</mods:namePart>
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<mods:date id="23F91AEB63B3EAB6507AE2D305574CBA">2017</mods:date>
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<paragraph id="8BDB32197B046B6AFCAEFCE080D8A2F5" blockId="16.[831,1177,888,945]" box="[831,1177,888,915]" pageId="16" pageNumber="59">
<heading id="D09385757B046B6AFCAEFCE080D8A2F5" box="[831,1177,888,915]" fontSize="10" level="2" pageId="16" pageNumber="59" reason="3">
<taxonomicName id="4C64499A7B046B6AFCAEFCE080D8A2F5" ID-CoL="9ZPLJ" authority="Rathbun, 1945" authorityName="Rathbun" authorityYear="1945" box="[831,1177,888,915]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="59" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B910EE0B7B046B6AFCAEFCE087ACA2F4" box="[831,1005,888,914]" italics="true" pageId="16" pageNumber="59">Ranina elegans</emphasis>
<bibRefCitation id="EFF54FE87B046B6AFC65FCE180D8A2F5" author="Rathbun M. J." box="[1012,1177,888,915]" pageId="16" pageNumber="59" pagination="373 - 383" refId="ref24791" refString="Rathbun M. J., 1945 - Decapod Crustacea. In: Geology of Lau, Fiji. Ladd H. S. &amp; Hoffmeister J. E. (eds.). Bernice Pauahi Bishop Museum Bulletin, 181: 373 - 383." type="journal article" year="1945">Rathbun, 1945</bibRefCitation>
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<paragraph id="8BDB32197B046B6AFCAEFC0F87CEA2D7" blockId="16.[831,1177,888,945]" box="[831,911,918,945]" pageId="16" pageNumber="59">
<figureCitation id="135F2E9C7B046B6AFCAEFC0F87CEA2D7" box="[831,911,918,945]" captionStart="Fig" captionStartId="16.[831,864,1920,1942]" captionTargetBox="[841,1465,1179,1892]" captionTargetId="figure-856@16.[832,1467,1170,1898]" captionTargetPageId="16" captionText="Fig. 13 - Ranina elegans Rathbun, 1945, holotype, USNM 498404 (x 5)." figureDoi="http://doi.org/10.5281/zenodo.12753829" httpUri="https://zenodo.org/record/12753829/files/figure.png" pageId="16" pageNumber="59">Fig. 13</figureCitation>
</paragraph>
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<subSubSection id="C37E61927B046B6AFCAEFC4D80D3A503" pageId="16" pageNumber="59" type="reference_group">
<paragraph id="8BDB32197B046B6AFCAEFC4D81DCA28B" blockId="16.[831,1437,979,1006]" box="[831,1437,979,1006]" pageId="16" pageNumber="59">
<heading id="D09385757B046B6AFCAEFC4D81DCA28B" box="[831,1437,979,1006]" fontSize="10" level="2" pageId="16" pageNumber="59" reason="3">
<treatmentCitationGroup id="AB7415377B046B6AFCAEFC4D81DCA28B" box="[831,1437,979,1006]" pageId="16" pageNumber="59">
<treatmentCitation id="0AC514087B046B6AFCAEFC4D8094A288" author="Rathbun M. J." box="[831,1237,979,1006]" page="375" pageId="16" pageNumber="59" year="1945">
<taxonomicName id="4C64499A7B046B6AFCAEFC4D8094A288" ID-CoL="9ZPLJ" authority="Rathbun, 1945: 375" authorityName="Rathbun" authorityPageNumber="375" authorityYear="1945" box="[831,1237,979,1006]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="59" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B910EE0B7B046B6AFCAEFC4D87ACA28B" box="[831,1005,979,1005]" italics="true" pageId="16" pageNumber="59">Ranina elegans</emphasis>
<bibRefCitation id="EFF54FE87B046B6AFC65FC4A8094A288" author="Rathbun M. J." box="[1012,1237,979,1006]" pageId="16" pageNumber="59" pagination="373 - 383" refId="ref24791" refString="Rathbun M. J., 1945 - Decapod Crustacea. In: Geology of Lau, Fiji. Ladd H. S. &amp; Hoffmeister J. E. (eds.). Bernice Pauahi Bishop Museum Bulletin, 181: 373 - 383." type="journal article" year="1945">Rathbun, 1945: 375</bibRefCitation>
</taxonomicName>
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, Pl. 55, Figs F, G.
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</heading>
</paragraph>
<paragraph id="8BDB32197B046B6AFCAEFB8980D3A503" blockId="16.[831,1474,1039,1126]" pageId="16" pageNumber="59">
<heading id="D09385757B046B6AFCAEFB8980D3A503" centered="true" fontSize="10" level="2" pageId="16" pageNumber="59" reason="3">
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<taxonomicName id="4C64499A7B046B6AFCAEFB8987AFA54F" ID-CoL="9ZPLJ" authorityName="Rathbun" authorityYear="1945" box="[831,1006,1039,1065]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="59" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B910EE0B7B046B6AFCAEFB8987AFA54F" box="[831,1006,1039,1065]" italics="true" pageId="16" pageNumber="59">Ranina elegans</emphasis>
</taxonomicName>
-
<treatmentCitation id="0AC514087B046B6AFB94FB96816EA54F" author="Schweitzer C. &amp; Feldmann R. M. &amp; Garassino A. &amp; Karasawa H. &amp; Schweigert G." box="[1029,1327,1039,1065]" page="74" pageId="16" pageNumber="59" year="2010">
<bibRefCitation id="EFF54FE87B046B6AFB94FB96816EA54F" author="Schweitzer C. &amp; Feldmann R. M. &amp; Garassino A. &amp; Karasawa H. &amp; Schweigert G." box="[1029,1327,1039,1065]" pageId="16" pageNumber="59" pagination="1 - 222" refId="ref24966" refString="Schweitzer C. E, Feldmann R. M., Garassino A., Karasawa H. &amp; Schweigert G., 2010 - Systematic list of fossil decapod crustacean species. Crustaceana Monographs, 10: 1 - 222." type="journal article" year="2010">
Schweitzer
<emphasis id="B910EE0B7B046B6AFB18FB898083A54F" box="[1161,1218,1039,1065]" italics="true" pageId="16" pageNumber="59">et al.</emphasis>
2010: 74
</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AC514087B046B6AFADEFB89800CA521" author="De Angeli A. &amp; Beschin C." page="13" pageId="16" pageNumber="59" year="2011">
<bibRefCitation id="EFF54FE87B046B6AFADEFB89800CA521" author="De Angeli A. &amp; Beschin C." pageId="16" pageNumber="59" pagination="11 - 20" refId="ref23611" refString="De Angeli A. &amp; Beschin C., 2011 - Il genere Ranina Lamarck, 1801 (Crustacea, Decapoda, Raninidae) nel Terziario del Vicentino, con descrizione di due nuove specie. Studi e Ricerche - Associazione Amici del Museo - Museo Civico &quot; G. Zannato &quot;, 18: 11 - 20." type="book chapter" year="2011">De Angeli &amp; Beschin 2011: 13</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AC514087B046B6AFBE0FBB781E6A521" author="Van Bakel W. M. &amp; Guinot D. &amp; Artal P. &amp; Fraaije R. H. B. &amp; Jagt J. W. M." box="[1137,1447,1069,1096]" page="209" pageId="16" pageNumber="59" year="2012">
<bibRefCitation id="EFF54FE87B046B6AFBE0FBB781E6A521" author="Van Bakel W. M. &amp; Guinot D. &amp; Artal P. &amp; Fraaije R. H. B. &amp; Jagt J. W. M." box="[1137,1447,1069,1096]" pageId="16" pageNumber="59" pagination="1 - 216" refId="ref25151" refString="Van Bakel W. M., Guinot D., Artal P., Fraaije R. H. B. &amp; Jagt J. W. M., 2012 - A revision of the Palaeocorystoidea and the phylogeny of raninoidian crabs (Crustacea, Decapoda, Brachyura, Podotremata). Zootaxa, 3215: 1 - 216." type="journal article" year="2012">
Van Bakel
<emphasis id="B910EE0B7B046B6AFB7EFBB78168A521" box="[1263,1321,1069,1095]" italics="true" pageId="16" pageNumber="59">et al.</emphasis>
2012: 209
</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AC514087B046B6AFCF4FBD580CFA503" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M. &amp; Luque J." box="[869,1166,1099,1126]" page="260" pageId="16" pageNumber="59" year="2014">
<bibRefCitation id="EFF54FE87B046B6AFCF4FBD580CFA503" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M. &amp; Luque J." box="[869,1166,1099,1126]" pageId="16" pageNumber="59" pagination="216 - 272" refId="ref24222" refString="Karasawa H., Schweitzer C. E., Feldmann R. M. &amp; Luque J., 2014 - Phylogeny and classification of Raninoida (Decapoda: Brachyura). Journal of Crustacean Biology, 34 (2): 216 - 272." type="journal article" year="2014">
Karasawa
<emphasis id="B910EE0B7B046B6AFC4AFBD58052A503" box="[987,1043,1099,1125]" italics="true" pageId="16" pageNumber="59">et al.</emphasis>
2014: 260
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</heading>
</paragraph>
</subSubSection>
<caption id="DF1B62917B046B6AFCAEF8198730A6D7" ID-DOI="http://doi.org/10.5281/zenodo.12753829" ID-Zenodo-Dep="12753829" httpUri="https://zenodo.org/record/12753829/files/figure.png" pageId="16" pageNumber="59" startId="16.[831,864,1920,1942]" targetBox="[841,1465,1179,1892]" targetPageId="16" targetType="figure">
<paragraph id="8BDB32197B046B6AFCAEF8198730A6D7" blockId="16.[831,1474,1920,1969]" pageId="16" pageNumber="59">
Fig. 13 -
<taxonomicName id="4C64499A7B046B6AFC31F8188087A6F0" authority="Rathbun, 1945" authorityName="Rathbun" authorityYear="1945" box="[928,1222,1920,1942]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="59" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B910EE0B7B046B6AFC31F8188071A6F3" box="[928,1072,1920,1942]" italics="true" pageId="16" pageNumber="59">Ranina elegans</emphasis>
<bibRefCitation id="EFF54FE87B046B6AFBABF8198087A6F0" author="Rathbun M. J." box="[1082,1222,1920,1942]" pageId="16" pageNumber="59" pagination="373 - 383" refId="ref24791" refString="Rathbun M. J., 1945 - Decapod Crustacea. In: Geology of Lau, Fiji. Ladd H. S. &amp; Hoffmeister J. E. (eds.). Bernice Pauahi Bishop Museum Bulletin, 181: 373 - 383." type="journal article" year="1945">Rathbun, 1945</bibRefCitation>
</taxonomicName>
, holotype, USNM 498404 (x 5).
</paragraph>
</caption>
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<paragraph id="8BDB32197B056B6BFF06FF1185B9A1A6" blockId="17.[113,756,135,975]" pageId="17" pageNumber="60">
<emphasis id="B910EE0B7B056B6BFF06FF118502A1C7" bold="true" box="[151,323,135,162]" pageId="17" pageNumber="60">Type material</emphasis>
:
<materialsCitation id="3B0C38447B056B6BFEC9FF1E86C2A1C4" box="[344,643,135,162]" collectionCode="USNM" pageId="17" pageNumber="60" specimenCode="USNM 498404" specimenCount="1" typeStatus="holotype">
<typeStatus id="54DF8CBB7B056B6BFEC9FF1E85FCA1C7" box="[344,445,135,161]" pageId="17" pageNumber="60" type="holotype">holotype</typeStatus>
,
<specimenCode id="DBC29A627B056B6BFE5FFF1186C2A1C4" box="[462,643,136,162]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="60" type="Museum">USNM 498404</specimenCode>
</materialsCitation>
;
<typeStatus id="54DF8CBB7B056B6BFD02FF1186B2A1C4" box="[659,755,136,162]" pageId="17" pageNumber="60" type="paratype">paratype</typeStatus>
lost (K. Hollis pers. comm., 2015).
</paragraph>
<paragraph id="8BDB32197B056B6BFF06FF5D8533A19A" blockId="17.[113,756,135,975]" pageId="17" pageNumber="60">
<emphasis id="B910EE0B7B056B6BFF06FF5D8574A1BB" bold="true" box="[151,309,195,222]" pageId="17" pageNumber="60">
<typeStatus id="54DF8CBB7B056B6BFF06FF5D8493A1B8" box="[151,210,196,222]" pageId="17" pageNumber="60">Type</typeStatus>
locality
</emphasis>
: Vanua Mbalavu (station 110B), Lau,
<collectingCountry id="F37372897B056B6BFFE0FF7B84DDA19A" box="[113,156,226,252]" name="Fiji" pageId="17" pageNumber="60">Fiji</collectingCountry>
; Futuna formation.
</paragraph>
<paragraph id="8BDB32197B056B6BFF06FE9985FCA07C" blockId="17.[113,756,135,975]" box="[151,445,256,283]" pageId="17" pageNumber="60">
<emphasis id="B910EE0B7B056B6BFF06FE998505A07C" bold="true" box="[151,324,256,282]" pageId="17" pageNumber="60">Geological age</emphasis>
: Miocene.
</paragraph>
<paragraph id="8BDB32197B056B6BFF06FE8785B4A05E" blockId="17.[113,756,135,975]" box="[151,501,286,313]" pageId="17" pageNumber="60">
<emphasis id="B910EE0B7B056B6BFF06FE87853CA05E" bold="true" box="[151,381,286,312]" pageId="17" pageNumber="60">Examined material</emphasis>
:
<typeStatus id="54DF8CBB7B056B6BFE1AFE8785B0A05E" box="[395,497,286,312]" pageId="17" pageNumber="60" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E61927B056B6BFF06FEA586B5A3E2" pageId="17" pageNumber="60" type="description">
<paragraph id="8BDB32197B056B6BFF06FEA586B5A3E2" blockId="17.[113,756,135,975]" pageId="17" pageNumber="60">
<emphasis id="B910EE0B7B056B6BFF06FEA58642A030" bold="true" box="[151,515,316,342]" pageId="17" pageNumber="60">
Description by
<bibRefCitation id="EFF54FE87B056B6BFED8FEA58642A030" author="Rathbun M. J." box="[329,515,316,342]" pageId="17" pageNumber="60" pagination="373 - 383" refId="ref24791" refString="Rathbun M. J., 1945 - Decapod Crustacea. In: Geology of Lau, Fiji. Ladd H. S. &amp; Hoffmeister J. E. (eds.). Bernice Pauahi Bishop Museum Bulletin, 181: 373 - 383." type="journal article" year="1945">Rathbun (1945)</bibRefCitation>
</emphasis>
: “
<emphasis id="B910EE0B7B056B6BFD8DFEA586A9A3E5" italics="true" pageId="17" pageNumber="60">
<typeStatus id="54DF8CBB7B056B6BFD8DFEA586C4A030" box="[540,645,316,342]" pageId="17" pageNumber="60" type="holotype">Holotype</typeStatus>
, a portion of carapace, showing the surface of the left posterior side; surface almost level, very slightly convex; the major part is covered with short, irregular, transverse raised lines, composed of rounded granules directed forward; the anterior and anterolateral regions are more closely roughened; posterior end bluntly thickened; in front of it a long curved row of granules which trends inward a little toward a median gap; the ornamentation of the exposed surface is longitudinally oblique to the marginal line and consists of slightly arched lines edged with minute granules.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37E61927B056B6BFF06FD1E8673A2A9" pageId="17" pageNumber="60" type="discussion">
<paragraph id="8BDB32197B056B6BFF06FD1E8611A231" blockId="17.[113,756,135,975]" pageId="17" pageNumber="60">
<emphasis id="B910EE0B7B056B6BFF06FD1E8554A3C7" bold="true" box="[151,277,647,673]" pageId="17" pageNumber="60">Discussion</emphasis>
. Based upon Rathbuns description and figures, it is difficult to determine a generic placement for this specimen, lacking all
<taxonomicName id="4C64499A7B056B6BFE37FD5C85B6A3B8" box="[422,503,709,734]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="B910EE0B7B056B6BFE37FD5C85B6A3B8" box="[422,503,709,734]" italics="true" pageId="17" pageNumber="60">Ranina</emphasis>
</taxonomicName>
characters. Moreover the
<typeStatus id="54DF8CBB7B056B6BFF0AFD7A84BFA39B" box="[155,254,739,765]" pageId="17" pageNumber="60" type="paratype">paratype</typeStatus>
, consisting in a left palm and figured by Rathbun (1945: Pl. 55, Fig. G) is today lost. The inability to recognize the diagnostic characters of
<taxonomicName id="4C64499A7B056B6BFDBEFC8686C1A25E" box="[559,640,799,824]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="B910EE0B7B056B6BFDBEFC8686C1A25E" box="[559,640,799,824]" italics="true" pageId="17" pageNumber="60">Ranina</emphasis>
</taxonomicName>
questions the placement of
<taxonomicName id="4C64499A7B056B6BFEA2FCA585C8A230" authorityName="Rathbun" authorityYear="1945" box="[307,393,828,854]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="60" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B910EE0B7B056B6BFEA2FCA585C8A230" box="[307,393,828,854]" italics="true" pageId="17" pageNumber="60">elegans</emphasis>
</taxonomicName>
within the genus.
</paragraph>
<paragraph id="8BDB32197B056B6BFF06FCC28673A2A9" blockId="17.[113,756,135,975]" pageId="17" pageNumber="60">
However the presence of short transverse raised lines with rounded granules does not fit into the typical ornamentation of
<taxonomicName id="4C64499A7B056B6BFE96FC018519A2D7" box="[263,344,920,945]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="B910EE0B7B056B6BFE96FC018519A2D7" box="[263,344,920,945]" italics="true" pageId="17" pageNumber="60">Ranina</emphasis>
</taxonomicName>
, remembering instead the transverse tegulate ornamentation of
<emphasis id="B910EE0B7B056B6BFE09FC2C8673A2A9" box="[408,562,949,975]" italics="true" pageId="17" pageNumber="60">
<taxonomicName id="4C64499A7B056B6BFE09FC2C866FA2A9" authorityName="Fabiani" authorityYear="1910" box="[408,558,949,975]" class="Malacostraca" family="Raninidae" genus="Lophoranina" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="60" phylum="Arthropoda" rank="genus">Lophoranina</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>