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<document id="1C4A5C785922734E0924B65D3D22F6E8" ID-DOI="10.5281/zenodo.185451" ID-GBIF-Dataset="b9b090a4-6bfe-4939-b2eb-174c92dea859" ID-ISSN="1175-5326" ID-Zenodo-Dep="185451" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459987272300" checkinUser="plazi" docAuthor="De, Claudio Augusto R., Candido, Denise M., Lucas, Sylvia Marlene &amp; Brescovit, Antonio Domingos" docDate="2009" docId="03C4F714FFC9FFA057A9FB61D08CD45A" docLanguage="en" docName="zt01987p038.pdf" docOrigin="Zootaxa 1987" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tityus cylindricus Karsch 1879" docType="treatment" docVersion="9" lastPageNumber="35" masterDocId="FFFD8F6CFFEBFF82573EFFE0D151D374" masterDocTitle="On the Tityus stigmurus complex (Scorpiones, Buthidae)" masterLastPageNumber="38" masterPageNumber="1" pageNumber="35" updateTime="1698232407615" updateUser="plazi">
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<mods:title id="46D7F63F0AD85061C7D1A5D436AC4E5E">On the Tityus stigmurus complex (Scorpiones, Buthidae)</mods:title>
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<mods:namePart id="8E41C70C6EFECA68ACA6AF464C9E2A9C">De, Claudio Augusto R.</mods:namePart>
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<mods:namePart id="8F029521F5C62586610EC7AA9E39AE42">Candido, Denise M.</mods:namePart>
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<mods:namePart id="4C470A148C26FBC1F509077A07F57DEF">Lucas, Sylvia Marlene</mods:namePart>
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<mods:namePart id="3449BA832EA7132C2B986D228DF48FB1">Brescovit, Antonio Domingos</mods:namePart>
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<mods:date id="06D31C7AF4EAE4123C6ED0AFE8824CD0">2009</mods:date>
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<mods:number id="58E49F599F9751C1B57A1C1C08DB715B">1987</mods:number>
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<treatment id="03C4F714FFC9FFA057A9FB61D08CD45A" ID-DOI="http://doi.org/10.5281/zenodo.6225623" ID-GBIF-Taxon="119379506" ID-Zenodo-Dep="6225623" LSID="urn:lsid:plazi:treatment:03C4F714FFC9FFA057A9FB61D08CD45A" httpUri="http://treatment.plazi.org/id/03C4F714FFC9FFA057A9FB61D08CD45A" lastPageNumber="35" pageId="34" pageNumber="35">
<subSubSection id="C3771589FFC9FFA057A9FB61D277D7EE" box="[151,806,1152,1179]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="8BD24602FFC9FFA057A9FB61D277D7EE" blockId="34.[151,806,1152,1179]" box="[151,806,1152,1179]" pageId="34" pageNumber="35">
<heading id="D09AF16EFFC9FFA057A9FB61D277D7EE" bold="true" box="[151,806,1152,1179]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
<emphasis id="B9199A10FFC9FFA057A9FB61D277D7EE" bold="true" box="[151,806,1152,1179]" pageId="34" pageNumber="35">
<taxonomicName id="4C6D3D81FFC9FFA057A9FB61D37FD7EE" ID-CoL="5799C" authority="Karsch, 1879" authorityName="Karsch" authorityYear="1879" box="[151,558,1152,1179]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="cylindricus">
<emphasis id="B9199A10FFC9FFA057A9FB61D03CD7EF" bold="true" box="[151,365,1153,1179]" italics="true" pageId="34" pageNumber="35">Tityus cylindricus</emphasis>
(
<bibRefCitation id="EFFC3BF3FFC9FFA05643FB60D374D7EE" author="Karsch" box="[381,549,1152,1178]" pageId="34" pageNumber="37" refString="Karsch, F. (1879) Scorpionologische Beitrage. Part I. Mitteilungem des Munchener Entomologischen Vereins, 3, 6 - 22." type="journal article" year="1879">Karsch, 1879</bibRefCitation>
)
</taxonomicName>
, species revalidated
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3771589FFC9FFA057A9FB2CD3A0D677" pageId="34" pageNumber="35" type="reference_group">
<paragraph id="8BD24602FFC9FFA057A9FB2CD3A0D677" blockId="34.[151,1231,1227,1283]" pageId="34" pageNumber="35">
<treatmentCitationGroup id="AB7D612CFFC9FFA057A9FB2CD3A0D677" pageId="34" pageNumber="35">
<taxonomicName id="4C6D3D81FFC9FFA057A9FB2CD319D797" authority="Karsch, 1879: 114" authorityName="Karsch" authorityPageNumber="114" authorityYear="1879" box="[151,584,1228,1251]" class="Arachnida" family="Buthidae" genus="Isometrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="cylindricus">
<emphasis id="B9199A10FFC9FFA057A9FB2CD02CD797" box="[151,381,1228,1251]" italics="true" pageId="34" pageNumber="35">Isometrus cylindricus</emphasis>
<treatmentCitation id="0ACC6013FFC9FFA056BAFB2CD319D797" author="Karsch" box="[388,584,1228,1251]" page="114" pageId="34" pageNumber="35" year="1879">
<bibRefCitation id="EFFC3BF3FFC9FFA056BAFB2CD347D797" author="Karsch" box="[388,534,1228,1251]" pageId="34" pageNumber="37" refString="Karsch, F. (1879) Scorpionologische Beitrage. Part I. Mitteilungem des Munchener Entomologischen Vereins, 3, 6 - 22." type="journal article" year="1879">Karsch, 1879</bibRefCitation>
: 114
</treatmentCitation>
</taxonomicName>
115.
<typeStatus id="54D6F8A0FFC9FFA055B7FB2CD3BDD797" box="[649,748,1228,1251]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
♂ from Bahia,
<collectingCountry id="F37A0692FFC9FFA054AFFB2CD283D797" box="[913,978,1228,1251]" name="Brazil" pageId="34" pageNumber="35">Brazil</collectingCountry>
(ZMB 105, examined).
<taxonomicName id="4C6D3D81FFC9FFA057A9FB0CD341D677" ID-CoL="5799C" authority="Kraepelin, 1899" authorityName="Kraepelin" authorityYear="1899" box="[151,528,1260,1283]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="cylindricus">
<emphasis id="B9199A10FFC9FFA057A9FB0CD002D677" box="[151,339,1260,1283]" italics="true" pageId="34" pageNumber="35">Tityus cylindricus</emphasis>
:
<bibRefCitation id="EFFC3BF3FFC9FFA0565CFB0CD341D677" author="Kraepelin" box="[354,528,1260,1283]" pageId="34" pageNumber="37" refString="Kraepelin, K. (1899) Scorpiones und Pedipalpi. In F. Dahl (ed.), Das Tierreich. Herausgegeben von der Deutschen Zoologishen Gesellschaft. Berlin: R. Friedlander und Sohn Verlag, 8 (Arachnoidea), 1 - 265." type="journal article" year="1899">Kraepelin, 1899</bibRefCitation>
</taxonomicName>
; Fet et al, 2000: 262.
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</paragraph>
</subSubSection>
<subSubSection id="C3771589FFC9FFA057A9FAD3D08CD45A" pageId="34" pageNumber="35" type="discussion">
<paragraph id="8BD24602FFC9FFA057A9FAD3D08CD45A" blockId="34.[151,1436,1331,1838]" pageId="34" pageNumber="35">
<emphasis id="B9199A10FFC9FFA057A9FAD3D074D639" bold="true" box="[151,293,1331,1357]" pageId="34" pageNumber="35">Comments.</emphasis>
According to Fet et al. (2000) “In his original description”,
<bibRefCitation id="EFFC3BF3FFC9FFA05339FAD4D5A1D63A" author="Karsch" box="[1031,1264,1332,1358]" pageId="34" pageNumber="37" refString="Karsch, F. (1879) Scorpionologische Beitrage. Part I. Mitteilungem des Munchener Entomologischen Vereins, 3, 6 - 22." type="journal article" year="1879">Karsch (1879: 115)</bibRefCitation>
indicated that
<taxonomicName id="4C6D3D81FFC9FFA057A9FABCD0C2D601" box="[151,403,1372,1397]" class="Arachnida" family="Buthidae" genus="Isometrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="cylindricus">
<emphasis id="B9199A10FFC9FFA057A9FABCD0C2D601" box="[151,403,1372,1397]" italics="true" pageId="34" pageNumber="35">Isometrus cylindricus</emphasis>
</taxonomicName>
was perhaps the male of
<taxonomicName id="4C6D3D81FFC9FFA05585FABCD503D602" authority="Thorell, 1876" authorityName="Thorell" authorityYear="1876" box="[699,1106,1372,1398]" class="Arachnida" family="Buthidae" genus="Isometrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="stigmurus">
<emphasis id="B9199A10FFC9FFA05585FABCD2F8D601" box="[699,937,1372,1397]" italics="true" pageId="34" pageNumber="35">Isometrus stigmurus</emphasis>
Thorell, 1876
</taxonomicName>
. Kraepelin, in 1891, listed it as a questionable synonym of
<taxonomicName id="4C6D3D81FFC9FFA056C4FA64D3E4D6E9" box="[506,693,1412,1437]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="bahiensis">
<emphasis id="B9199A10FFC9FFA056C4FA64D3E4D6E9" box="[506,693,1412,1437]" italics="true" pageId="34" pageNumber="35">Tityus bahiensis</emphasis>
</taxonomicName>
, but in 1899 reverted to listing it as a questionable synonym of
<taxonomicName id="4C6D3D81FFC9FFA057A9FA4CD009D6B1" box="[151,344,1452,1477]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="stigmurus">
<emphasis id="B9199A10FFC9FFA057A9FA4CD009D6B1" box="[151,344,1452,1477]" italics="true" pageId="34" pageNumber="35">Tityus stigmurus</emphasis>
</taxonomicName>
” (Fet et al, 2000). The
<typeStatus id="54D6F8A0FFC9FFA05553FA4CD3CED6B2" box="[621,671,1452,1478]" pageId="34" pageNumber="35">type</typeStatus>
specimen (
<figureCitation id="13565A87FFC9FFA05418FA4CD22BD6B2" box="[806,890,1452,1478]" captionStart="FIGURE 23. T" captionStartId="35.[151,255,1959,1983]" captionTargetBox="[228,1366,186,1887]" captionTargetId="figure@35.[214,1387,178,1935]" captionTargetPageId="35" captionText="FIGURE 23. T. cylindricus. (Holotype ♂ ZMB 105; Bahia, Brazil). Dorsal (A) and ventral (B) view; Right palp in dorsal view (C); Metasomal segments and telson in lateral view (D E)." httpUri="https://zenodo.org/record/185474/files/figure.png" pageId="34" pageNumber="35">Fig. 23</figureCitation>
AB) is in poor condition, probably due to the initial preservation in a dry collection. This species is not a synonym of
<taxonomicName id="4C6D3D81FFC9FFA054CAFA34D5D2D699" box="[1012,1155,1492,1517]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="stigmurus">
<emphasis id="B9199A10FFC9FFA054CAFA34D5D2D699" box="[1012,1155,1492,1517]" italics="true" pageId="34" pageNumber="35">T. stigmurus</emphasis>
</taxonomicName>
and does not belong to the
<taxonomicName id="4C6D3D81FFC9FFA057FDFA1CD005D561" box="[195,340,1532,1557]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="stigmurus">
<emphasis id="B9199A10FFC9FFA057FDFA1CD005D561" box="[195,340,1532,1557]" italics="true" pageId="34" pageNumber="35">T. stigmurus</emphasis>
</taxonomicName>
complex. It has blackish coloration, absence of posterior spinoid granules on the dorsal carinae of metasomal segments IIIV (
<figureCitation id="13565A87FFC9FFA05507F9C4D3DED54A" box="[569,655,1572,1598]" captionStart="FIGURE 23. T" captionStartId="35.[151,255,1959,1983]" captionTargetBox="[228,1366,186,1887]" captionTargetId="figure@35.[214,1387,178,1935]" captionTargetPageId="35" captionText="FIGURE 23. T. cylindricus. (Holotype ♂ ZMB 105; Bahia, Brazil). Dorsal (A) and ventral (B) view; Right palp in dorsal view (C); Metasomal segments and telson in lateral view (D E)." httpUri="https://zenodo.org/record/185474/files/figure.png" pageId="34" pageNumber="35">Fig. 23</figureCitation>
DE), metasomal segments and telson very slender and rhomboid shape of subaculear granule (
<figureCitation id="13565A87FFC9FFA056D5F9ACD311D512" box="[491,576,1612,1638]" captionStart="FIGURE 23. T" captionStartId="35.[151,255,1959,1983]" captionTargetBox="[228,1366,186,1887]" captionTargetId="figure@35.[214,1387,178,1935]" captionTargetPageId="35" captionText="FIGURE 23. T. cylindricus. (Holotype ♂ ZMB 105; Bahia, Brazil). Dorsal (A) and ventral (B) view; Right palp in dorsal view (C); Metasomal segments and telson in lateral view (D E)." httpUri="https://zenodo.org/record/185474/files/figure.png" pageId="34" pageNumber="35">Fig. 23</figureCitation>
DE), whereas in species of the
<taxonomicName id="4C6D3D81FFC9FFA05489F9ACD517D511" box="[951,1094,1612,1637]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="stigmurus">
<emphasis id="B9199A10FFC9FFA05489F9ACD517D511" box="[951,1094,1612,1637]" italics="true" pageId="34" pageNumber="35">T. stigmurus</emphasis>
</taxonomicName>
complex the metasomal segments are stouter and the subaculear granule is spiniform. The
<typeStatus id="54D6F8A0FFC9FFA0544DF994D2F7D5FA" box="[883,934,1652,1678]" pageId="34" pageNumber="35">type</typeStatus>
locality is uncertain, it is stated as “Bahia” in
<collectingCountry id="F37A0692FFC9FFA05786F97CD053D5C2" box="[184,258,1692,1718]" name="Brazil" pageId="34" pageNumber="35">Brazil</collectingCountry>
, without concise location data.
<taxonomicName id="4C6D3D81FFC9FFA055BCF97CD274D5C1" box="[642,805,1692,1717]" class="Arachnida" family="Buthidae" genus="Isometrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="cylindricus">
<emphasis id="B9199A10FFC9FFA055BCF97CD274D5C1" box="[642,805,1692,1717]" italics="true" pageId="34" pageNumber="35">T. cylindricus</emphasis>
</taxonomicName>
it is associated with he subgenus
<taxonomicName id="4C6D3D81FFC9FFA053FBF97CD18FD5AA" authority="Lourenco, 2006" authorityName="Lourenco" authorityYear="2006" class="Arachnida" family="Buthidae" genus="Atreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B9199A10FFC9FFA053FBF97CD441D5C1" box="[1221,1296,1692,1717]" italics="true" pageId="34" pageNumber="35">Atreus</emphasis>
(Lourenço, 2006)
</taxonomicName>
, by having blackish coloration, metasomal segments and telson very slender (
<figureCitation id="13565A87FFC9FFA053BCF924D589D5AA" box="[1154,1240,1732,1758]" captionStart="FIGURE 23. T" captionStartId="35.[151,255,1959,1983]" captionTargetBox="[228,1366,186,1887]" captionTargetId="figure@35.[214,1387,178,1935]" captionTargetPageId="35" captionText="FIGURE 23. T. cylindricus. (Holotype ♂ ZMB 105; Bahia, Brazil). Dorsal (A) and ventral (B) view; Right palp in dorsal view (C); Metasomal segments and telson in lateral view (D E)." httpUri="https://zenodo.org/record/185474/files/figure.png" pageId="34" pageNumber="35">Fig. 23</figureCitation>
DE) and rhomboid shape of subaculear granule. More details will be given in a future work which will encompass the species of the subgenus
<taxonomicName id="4C6D3D81FFC9FFA056B5F8F4D087D459" box="[395,470,1812,1837]" class="Arachnida" family="Buthidae" genus="Atreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B9199A10FFC9FFA056B5F8F4D087D459" box="[395,470,1812,1837]" italics="true" pageId="34" pageNumber="35">Atreus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>