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<document id="672ADFB29FE6BA27FCDC241AE35F5769" ID-DOI="10.5281/zenodo.180818" ID-GBIF-Dataset="850fc9b3-67a6-4ced-b77b-3cc2bebe8857" ID-ISSN="1175-5326" ID-Zenodo-Dep="180818" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459816636239" checkinUser="plazi" docAuthor="Ivie, Michael A. &amp; Philips, Keith" docDate="2008" docId="03BC87DCAF39FFB7FF68FB4FFD674BAE" docLanguage="en" docName="zt01701p014.pdf" docOrigin="Zootaxa 1701" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Canthonella sikesi Ivie and Philips, new species" docType="treatment" docVersion="8" lastPageNumber="7" masterDocId="FF85FFA4AF3DFFB1FFFFFFE8FFEC4D74" masterDocTitle="Three new species of Canthonella Chapin from Hispaniola, with new records and Nomenclatural changes for West Indian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="14" masterPageNumber="1" pageNumber="5" updateTime="1698221501712" updateUser="plazi">
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<mods:title id="ECD4567DFD556742E809E6C9A2BE1B7B">Three new species of Canthonella Chapin from Hispaniola, with new records and Nomenclatural changes for West Indian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="F4A12C180C49C9BBF4F1B111F9D1A75E" type="personal">
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<mods:namePart id="2520842C8562E9C3F47A8AB11EFAB85F">Ivie, Michael A.</mods:namePart>
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<mods:namePart id="CD3A5D17ABCC15006059A5B521D4BA6B">Philips, Keith</mods:namePart>
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<mods:date id="9BBACB2259D825724004693D1EB6025E">2008</mods:date>
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<mods:number id="57823E35206B09B7C7E2FE343716EC44">1701</mods:number>
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<mods:identifier id="2EFCEDB98F955D7D47B1BF8FFF6CE42B" type="DOI">10.5281/zenodo.180818</mods:identifier>
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<treatment id="03BC87DCAF39FFB7FF68FB4FFD674BAE" ID-DOI="http://doi.org/10.5281/zenodo.6233223" ID-GBIF-Taxon="119354879" ID-Zenodo-Dep="6233223" LSID="urn:lsid:plazi:treatment:03BC87DCAF39FFB7FF68FB4FFD674BAE" httpUri="http://treatment.plazi.org/id/03BC87DCAF39FFB7FF68FB4FFD674BAE" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
<subSubSection id="C30F6541AF39FFB5FF68FB4FFE3C4990" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BAA36CAAF39FFB5FF68FB4FFD0C49B5" blockId="4.[151,736,1191,1252]" box="[151,736,1191,1217]" pageId="4" pageNumber="5">
<heading id="D0E281A6AF39FFB5FF68FB4FFD0C49B5" bold="true" box="[151,736,1191,1217]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="B961EAD8AF39FFB5FF68FB4FFD0C49B5" bold="true" box="[151,736,1191,1217]" pageId="4" pageNumber="5">
<taxonomicName id="4C154D49AF39FFB5FF68FB4FFDAF49B5" ID-CoL="QPFV" authority="Ivie and Philips" authorityName="Ivie and Philips" box="[151,579,1191,1217]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sikesi" status="sp. nov.">
<emphasis id="B961EAD8AF39FFB5FF68FB4FFE9B49B5" bold="true" box="[151,375,1191,1217]" italics="true" pageId="4" pageNumber="5">Canthonella sikesi</emphasis>
Ivie and Philips
</taxonomicName>
,
<taxonomicNameLabel id="A25257A3AF39FFB5FDAFFB4FFD0C49B5" box="[592,736,1191,1217]" pageId="4" pageNumber="5" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAA36CAAF39FFB5FF68FB22FE3C4990" blockId="4.[151,736,1191,1252]" box="[151,464,1226,1252]" pageId="4" pageNumber="5">
<figureCitation id="132E2A4FAF39FFB5FF68FB22FECF4990" box="[151,291,1226,1252]" captionStart="FIGURES 1 9. 1 3" captionStartId="3.[151,269,1762,1786]" captionTargetBox="[185,1381,185,1729]" captionTargetId="figure@3.[151,1436,176,1756]" captionTargetPageId="3" captionText="FIGURES 1 9. 1 3. Labia. 1. Canthonella jarmilea; 2. Canthonella quesquaya; 3. Canthonella sikesi. 4 6. Heads 4. Canthonella jarmilea; 5. Canthonella quesquaya; 6: Canthonella sikesi. 7 9. Left elytra 7, Canthonella jarmilea; 8. Canthonella quesquaya; 9. Canthonella sikesi." httpUri="https://zenodo.org/record/180819/files/figure.png" pageId="4" pageNumber="5">Figs. 3, 6, 9</figureCitation>
, 12, 15, 18, 21
</paragraph>
</subSubSection>
<subSubSection id="C30F6541AF39FFB5FF68FAF2FE3548A3" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BAA36CAAF39FFB5FF68FAF2FE3548A3" blockId="4.[151,1436,1306,1975]" pageId="4" pageNumber="5">
<emphasis id="B961EAD8AF39FFB5FF68FAF2FEFB4840" bold="true" box="[151,279,1306,1332]" pageId="4" pageNumber="5">Diagnosis.</emphasis>
This small species is very similar to
<taxonomicName id="4C154D49AF39FFB5FD3BFAF3FC4B4840" authority="Harold" authorityName="Harold" box="[708,935,1306,1332]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pygmaea">
<emphasis id="B961EAD8AF39FFB5FD3BFAF3FCBC4840" box="[708,848,1307,1332]" italics="true" pageId="4" pageNumber="5">C. pygmaea</emphasis>
Harold
</taxonomicName>
and
<taxonomicName id="4C154D49AF39FFB5FC20FAF3FB044840" authority="Matthews" authorityName="Matthews" box="[991,1256,1306,1332]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="isabellae">
<emphasis id="B961EAD8AF39FFB5FC20FAF3FB864840" box="[991,1130,1307,1332]" italics="true" pageId="4" pageNumber="5">C. isabellae</emphasis>
Matthews
</taxonomicName>
, but can be distinguished from those and all other described
<taxonomicName id="4C154D49AF39FFB5FD52FAABFCD24828" box="[685,830,1347,1372]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAD8AF39FFB5FD52FAABFCD24828" box="[685,830,1347,1372]" italics="true" pageId="4" pageNumber="5">Canthonella</emphasis>
</taxonomicName>
species by the combination of a yellow-orange elytral macula that reaches the basal margin (separate in
<taxonomicName id="4C154D49AF39FFB5FCC5FA86FC3B48F3" box="[826,983,1390,1415]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B961EAD8AF39FFB5FCC5FA86FC3B48F3" box="[826,983,1390,1415]" italics="true" pageId="4" pageNumber="5">C. pygmaeus</emphasis>
</taxonomicName>
), and the impunctate 8th elytral stria (strongly punctate in
<taxonomicName id="4C154D49AF39FFB5FE6FFA7EFDF748DB" box="[400,539,1430,1455]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="isabellae">
<emphasis id="B961EAD8AF39FFB5FE6FFA7EFDF748DB" box="[400,539,1430,1455]" italics="true" pageId="4" pageNumber="5">C. isabellae</emphasis>
</taxonomicName>
). The unique shape of the apical portions of the aedeagus will further distinguish this species (Fig. 12).
</paragraph>
</subSubSection>
<subSubSection id="C30F6541AF39FFB4FF39FA0DFD834AA9" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
<paragraph id="8BAA36CAAF39FFB5FF39FA0DFC684AC3" blockId="4.[151,1436,1306,1975]" pageId="4" pageNumber="5">
<emphasis id="B961EAD8AF39FFB5FF39FA0DFE55488B" bold="true" box="[198,441,1509,1535]" pageId="4" pageNumber="5">Description (male).</emphasis>
Oval (
<figureCitation id="132E2A4FAF39FFB5FDEDFA0DFD81488B" box="[530,621,1509,1535]" captionStart="FIGURES 19 22" captionStartId="6.[151,269,1258,1282]" captionTargetBox="[291,1311,196,1233]" captionTargetId="figure@6.[275,1312,168,1234]" captionTargetPageId="6" captionText="FIGURES 19 22. Dorsal habiti. 19. Canthonella jarmilea; 20. Canthonella quesquaya; 21. Canthonella sikesi; 22. Canthonella constans." httpUri="https://zenodo.org/record/180820/files/figure.png" pageId="4" pageNumber="5">Fig. 21</figureCitation>
), strongly convex, finely pilose, shining black, yellow-orange spot along anterior margin of elytron from third elytral stria to humeral angle and across epipleuron (occasionally and faintly extending onto second interval), extending to posterior edge of metasternum (
<figureCitation id="132E2A4FAF39FFB5FB39F9DDFAFC4B3B" box="[1222,1296,1589,1615]" captionStart="FIGURES 1 9. 1 3" captionStartId="3.[151,269,1762,1786]" captionTargetBox="[185,1381,185,1729]" captionTargetId="figure@3.[151,1436,176,1756]" captionTargetPageId="3" captionText="FIGURES 1 9. 1 3. Labia. 1. Canthonella jarmilea; 2. Canthonella quesquaya; 3. Canthonella sikesi. 4 6. Heads 4. Canthonella jarmilea; 5. Canthonella quesquaya; 6: Canthonella sikesi. 7 9. Left elytra 7, Canthonella jarmilea; 8. Canthonella quesquaya; 9. Canthonella sikesi." httpUri="https://zenodo.org/record/180819/files/figure.png" pageId="4" pageNumber="5">Fig. 9</figureCitation>
). Eyes dorsally separated by
<date id="FFAB100AAF39FFB5FE8EF9B5FE794B03" box="[369,405,1629,1655]" pageId="4" pageNumber="5">8X</date>
maximum width of dorsal lobe (
<figureCitation id="132E2A4FAF39FFB5FCE6F9B5FC8D4B03" box="[793,865,1629,1655]" captionStart="FIGURES 1 9. 1 3" captionStartId="3.[151,269,1762,1786]" captionTargetBox="[185,1381,185,1729]" captionTargetId="figure@3.[151,1436,176,1756]" captionTargetPageId="3" captionText="FIGURES 1 9. 1 3. Labia. 1. Canthonella jarmilea; 2. Canthonella quesquaya; 3. Canthonella sikesi. 4 6. Heads 4. Canthonella jarmilea; 5. Canthonella quesquaya; 6: Canthonella sikesi. 7 9. Left elytra 7, Canthonella jarmilea; 8. Canthonella quesquaya; 9. Canthonella sikesi." httpUri="https://zenodo.org/record/180819/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
). Labium with apex slightly raised; slightly produced and angulate in hind view; anterior margin truncate to slightly emarginate in ventral view; palpi bearded with long setae medially; base of mentum bearing 2 very long setae (
<figureCitation id="132E2A4FAF39FFB5FBDDF945FB854BB3" box="[1058,1129,1709,1735]" captionStart="FIGURES 1 9. 1 3" captionStartId="3.[151,269,1762,1786]" captionTargetBox="[185,1381,185,1729]" captionTargetId="figure@3.[151,1436,176,1756]" captionTargetPageId="3" captionText="FIGURES 1 9. 1 3. Labia. 1. Canthonella jarmilea; 2. Canthonella quesquaya; 3. Canthonella sikesi. 4 6. Heads 4. Canthonella jarmilea; 5. Canthonella quesquaya; 6: Canthonella sikesi. 7 9. Left elytra 7, Canthonella jarmilea; 8. Canthonella quesquaya; 9. Canthonella sikesi." httpUri="https://zenodo.org/record/180819/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
). Pronotum at base nearly as wide as base of elytra, parallel sided for basal 3/4, evenly rounded at anterior 1/4; basal margin very fine to obsolete medially. Elytral striae 18 impunctate, consisting of 2 fine parallel lines, 7 obsolete on humerus; interstriae flat, conforming to shape of elytron. Foretibia transversely truncate apically, with 2 acute teeth externally; serrulate between and proximal to teeth. Metatibia weakly serrulate on internal margin (Fig. 18). Adeagus with shape of phallobase apex and parameres unique, as in Fig. 12. Length: 2.502.51mm, measured along midline from anterior margin of pronotum to tip of elytra.
</paragraph>
<paragraph id="8BAA36CAAF38FFB4FE48FAE0FE8F48F3" blockId="5.[151,1436,1288,1415]" pageId="5" pageNumber="6">
<collectingCountry id="F302765AAF38FFB4FE48FAE0FE02486B" box="[439,494,1288,1311]" name="Malta" pageId="5" pageNumber="6">Male</collectingCountry>
genitalia, lateral view. 10.
<taxonomicName id="4C154D49AF38FFB4FCEBFAE0FC15486B" box="[788,1017,1288,1311]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="jarmilea">
<emphasis id="B961EAD8AF38FFB4FCEBFAE0FC15486B" box="[788,1017,1288,1311]" italics="true" pageId="5" pageNumber="6">Canthonella jarmilea</emphasis>
</taxonomicName>
; 11.
<taxonomicName id="4C154D49AF38FFB4FBCEFAE0FAC2486B" box="[1073,1326,1288,1311]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="quesquaya">
<emphasis id="B961EAD8AF38FFB4FBCEFAE0FAC2486B" box="[1073,1326,1288,1311]" italics="true" pageId="5" pageNumber="6">Canthonella quesquaya</emphasis>
</taxonomicName>
; 12.
<emphasis id="B961EAD8AF38FFB4FA99FAE0FED54836" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="4C154D49AF38FFB4FA99FAE0FED94836" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sikesi">Canthonella sikesi</taxonomicName>
.
</emphasis>
<emphasis id="B961EAD8AF38FFB4FEBFFAC2FE654836" bold="true" box="[320,393,1322,1346]" pageId="5" pageNumber="6">1315.</emphasis>
Right profemur and tarsus Fig. 13:
<taxonomicName id="4C154D49AF38FFB4FCEEFAC3FC1B4836" box="[785,1015,1323,1346]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="jarmilea">
<emphasis id="B961EAD8AF38FFB4FCEEFAC3FC1B4836" box="[785,1015,1323,1346]" italics="true" pageId="5" pageNumber="6">Canthonella jarmilea</emphasis>
</taxonomicName>
; 14.
<taxonomicName id="4C154D49AF38FFB4FBCFFAC3FAC24836" box="[1072,1326,1323,1346]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="quesquaya">
<emphasis id="B961EAD8AF38FFB4FBCFFAC3FAC24836" box="[1072,1326,1323,1346]" italics="true" pageId="5" pageNumber="6">Canthonella quesquaya</emphasis>
</taxonomicName>
; 15.
<taxonomicName id="4C154D49AF38FFB4FA99FAC3FED44811" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sikesi">
<emphasis id="B961EAD8AF38FFB4FA99FAC3FED44811" italics="true" pageId="5" pageNumber="6">Canthonella sikesi</emphasis>
</taxonomicName>
;
<emphasis id="B961EAD8AF38FFB4FEB0FAA5FE754811" bold="true" box="[335,409,1357,1381]" pageId="5" pageNumber="6">1618.</emphasis>
Right metafemur and tarsus 16.
<taxonomicName id="4C154D49AF38FFB4FCD5FAA6FBFB4811" box="[810,1047,1358,1381]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="jarmilea">
<emphasis id="B961EAD8AF38FFB4FCD5FAA6FBFB4811" box="[810,1047,1358,1381]" italics="true" pageId="5" pageNumber="6">Canthonella jarmilea</emphasis>
</taxonomicName>
; 17.
<taxonomicName id="4C154D49AF38FFB4FBA0FAA6FA884811" box="[1119,1380,1358,1381]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="quesquaya">
<emphasis id="B961EAD8AF38FFB4FBA0FAA6FA884811" box="[1119,1380,1358,1381]" italics="true" pageId="5" pageNumber="6">Canthonella quesquaya</emphasis>
</taxonomicName>
; 18.
<taxonomicName id="4C154D49AF38FFB4FF68FA98FEB048F3" box="[151,348,1392,1415]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sikesi">
<emphasis id="B961EAD8AF38FFB4FF68FA98FEB048F3" box="[151,348,1392,1415]" italics="true" pageId="5" pageNumber="6">Canthonella sikesi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAA36CAAF38FFB4FF39FA53FBD74889" blockId="5.[151,1436,1467,2013]" pageId="5" pageNumber="6">
Female. Differs as in other Hispaniolan
<taxonomicName id="4C154D49AF38FFB4FD60FA53FCDC48A0" box="[671,816,1467,1492]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAD8AF38FFB4FD60FA53FCDC48A0" box="[671,816,1467,1492]" italics="true" pageId="5" pageNumber="6">Canthonella</emphasis>
</taxonomicName>
, i.e. the apical spine of the protibia is less stout than that of the male, and the interior margin of the female metatibia is not serrulate.
</paragraph>
<paragraph id="8BAA36CAAF38FFB4FF39F9E2FD9A4B01" blockId="5.[151,1436,1467,2013]" pageId="5" pageNumber="6">
<emphasis id="B961EAD8AF38FFB4FF39F9E2FEC04B50" bold="true" box="[198,300,1546,1572]" pageId="5" pageNumber="6">
<typeStatus id="54AE8868AF38FFB4FF39F9E2FECB4B50" box="[198,295,1546,1572]" pageId="5" pageNumber="6">TYPES</typeStatus>
.
</emphasis>
<typeStatus id="54AE8868AF38FFB4FEC1F9E3FE354B51" box="[318,473,1547,1573]" pageId="5" pageNumber="6" type="holotype">HOLOTYPE</typeStatus>
MALE: DOM REP: Prov Hato Mayor; Par. Nac. Los Haitises;
<date id="FFAB100AAF38FFB4FAC8F9E3FEDB4B39" pageId="5" pageNumber="6" value="1992-04-16" valueMax="1992-07-01" valueMin="1992-04-16">16APR 01JULY1992</date>
; F.I.T.#1, bosque humido; M.Ivie, D.Sikes, Lanier [from WIBF, deposited in NMNH]. ALLO-
<typeStatus id="54AE8868AF38FFB4FF68F9B3FF0C4B01" box="[151,224,1627,1653]" pageId="5" pageNumber="6">TYPE</typeStatus>
FEMALE: same data as
<typeStatus id="54AE8868AF38FFB4FDF8F9B3FD9D4B01" box="[519,625,1627,1653]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BAA36CAAF38FFB4FF3AF96BFB5D4AF9" blockId="5.[151,1436,1467,2013]" pageId="5" pageNumber="6">
<typeStatus id="54AE8868AF38FFB4FF3AF96BFE874BE9" box="[197,363,1667,1693]" pageId="5" pageNumber="6" type="paratype">PARATYPES</typeStatus>
: 12 MALES, 10 FEMALES — same data as
<typeStatus id="54AE8868AF38FFB4FC77F96BFC024BE9" box="[904,1006,1667,1693]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
[CMNC, MHND, TKPC, WIBF]. 1 MALE — DOM.REP: Prov. Hato Mayor; Par. Nac. Los Haitises;
<date id="FFAB100AAF38FFB4FC3BF943FB4A4BB1" box="[964,1190,1707,1733]" pageId="5" pageNumber="6" value="1993-07-18" valueMax="1993-07-21" valueMin="1993-07-18">1821 JULY 1993</date>
; D. S. Sikes and R. Rosenfeld; flight intercept trap [TKPC]. 3 MALES, 1 FEMALE — DOM.REP: Pr. Hato Mayor; Par. Nac. Los Haitises; 0
<date id="FFAB100AAF38FFB4FEE6F913FE2E4A61" box="[281,450,1787,1813]" pageId="5" pageNumber="6" value="1992-07-02">2 JULY 1992</date>
<date id="FFAB100AAF38FFB4FE15F913FD484A61" box="[490,676,1787,1813]" pageId="5" pageNumber="6" value="1993-07-16">16 JULY 1993</date>
, D. Sikes and R. Rosenfield; flight intercept trap [WIBF]. 1 FEMALE — DOM REP: Prov Hato Mayor; Par. Nac. Los Haitises;
<date id="FFAB100AAF38FFB4FC1EF8CBFB4F4A49" box="[993,1187,1827,1853]" pageId="5" pageNumber="6" value="1992-04-01" valueMax="1992-04-16" valueMin="1992-04-01">0116APR1992</date>
; dung bait; pit. trap, bosque humido; M.Ivie, D.Sikes, Lanier [WIBF]. 1 FEMALE — Samana; Samana RD;
<date id="FFAB100AAF38FFB4FB43F8A3FABF4A11" box="[1212,1363,1867,1893]" pageId="5" pageNumber="6" value="1978-07-29">VII 29 1978</date>
; R. 0. Schuster [UCDC]. 1 MALE — Cabrera; Rep. Dom;
<date id="FFAB100AAF38FFB4FD00F89BFC614AF9" box="[767,909,1907,1933]" pageId="5" pageNumber="6" value="1978-08-01">VIII 1 1978</date>
; R. 0. Schuster [UCDC].
</paragraph>
<paragraph id="8BAA36CAAF38FFB4FF3AF873FD834AA9" blockId="5.[151,1436,1467,2013]" pageId="5" pageNumber="6">
ADDITIONAL MATERIAL STUDIED BUT NOT INCLUDED IN
<typeStatus id="54AE8868AF38FFB4FC07F873FBAD4AC1" box="[1016,1089,1947,1973]" pageId="5" pageNumber="6">TYPE</typeStatus>
SERIES:
<specimenCount id="9D13FD43AF38FFB4FB42F873FABE4AC1" box="[1213,1362,1947,1973]" pageId="5" pageNumber="6" type="generic">2 specimens</specimenCount>
, same data as
<typeStatus id="54AE8868AF38FFB4FF10F82BFEB44AA9" box="[239,344,1987,2013]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
, disarticulated [TKPC].
</paragraph>
</subSubSection>
<caption id="DF6A6642AF3BFFB7FF68FB02FEB64850" httpUri="https://zenodo.org/record/180820/files/figure.png" pageId="6" pageNumber="7" targetBox="[291,1311,196,1233]" targetPageId="6">
<paragraph id="8BAA36CAAF3BFFB7FF68FB02FEB64850" blockId="6.[151,1435,1258,1316]" pageId="6" pageNumber="7">
<emphasis id="B961EAD8AF3BFFB7FF68FB02FEB14876" bold="true" box="[151,349,1258,1282]" pageId="6" pageNumber="7">FIGURES 1922.</emphasis>
Dorsal habiti. 19.
<taxonomicName id="4C154D49AF3BFFB7FDDEFB03FCE94876" box="[545,773,1259,1282]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="jarmilea">
<emphasis id="B961EAD8AF3BFFB7FDDEFB03FCE94876" box="[545,773,1259,1282]" italics="true" pageId="6" pageNumber="7">Canthonella jarmilea</emphasis>
</taxonomicName>
; 20.
<taxonomicName id="4C154D49AF3BFFB7FCC4FB03FBDB4876" box="[827,1079,1259,1282]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="quesquaya">
<emphasis id="B961EAD8AF3BFFB7FCC4FB03FBDB4876" box="[827,1079,1259,1282]" italics="true" pageId="6" pageNumber="7">Canthonella quesquaya</emphasis>
</taxonomicName>
; 21.
<taxonomicName id="4C154D49AF3BFFB7FB93FB03FADD4876" box="[1132,1329,1259,1282]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="sikesi">
<emphasis id="B961EAD8AF3BFFB7FB93FB03FADD4876" box="[1132,1329,1259,1282]" italics="true" pageId="6" pageNumber="7">Canthonella sikesi</emphasis>
</taxonomicName>
; 22.
<taxonomicName id="4C154D49AF3BFFB7FA99FB03FEBA4850" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="constans">Canthonella constans</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C30F6541AF3BFFB7FF3AFAB0FD674BAE" pageId="6" pageNumber="7" type="etymology">
<paragraph id="8BAA36CAAF3BFFB7FF3AFAB0FE7648EE" blockId="6.[151,1436,1368,1754]" pageId="6" pageNumber="7">
<emphasis id="B961EAD8AF3BFFB7FF3AFAB0FEBE4806" bold="true" box="[197,338,1368,1394]" pageId="6" pageNumber="7">Etymology.</emphasis>
Named after Derek S. Sikes, a valued student, colleague, and friend, as well as one of the collectors of this species.
</paragraph>
<paragraph id="8BAA36CAAF3BFFB7FF3AFA40FA82489E" blockId="6.[151,1436,1368,1754]" pageId="6" pageNumber="7">
<emphasis id="B961EAD8AF3BFFB7FF3AFA40FEFE48B6" bold="true" box="[197,274,1448,1474]" pageId="6" pageNumber="7">Notes.</emphasis>
This species is very similar to, and possibly a vicariant sister-species of,
<taxonomicName id="4C154D49AF3BFFB7FB8CFA40FA7048B6" authority="Matthews. It" authorityName="Matthews. It" box="[1139,1436,1448,1474]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="isabellae">
<emphasis id="B961EAD8AF3BFFB7FB8CFA40FAEC48B5" box="[1139,1280,1448,1473]" italics="true" pageId="6" pageNumber="7">C. isabellae</emphasis>
Matthews. It
</taxonomicName>
is known only from the mesic north coast, while
<taxonomicName id="4C154D49AF3BFFB7FD2BFA38FCB3489D" box="[724,863,1488,1513]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="isabellae">
<emphasis id="B961EAD8AF3BFFB7FD2BFA38FCB3489D" box="[724,863,1488,1513]" italics="true" pageId="6" pageNumber="7">C. isabellae</emphasis>
</taxonomicName>
is known only from the drier southern areas.
</paragraph>
<paragraph id="8BAA36CAAF3BFFB7FF3AFA10FD674BAE" blockId="6.[151,1436,1368,1754]" pageId="6" pageNumber="7">
The
<typeStatus id="54AE8868AF3BFFB7FF02FA10FEDC4B66" box="[253,304,1528,1554]" pageId="6" pageNumber="7">type</typeStatus>
locality is in Parque Nacional de Los Haïtises, a karst area west of Sabana de la Mar, on the Bahia de San Lorenzo (
<bibRefCitation id="EF844B3BAF3BFFB7FE52F9C8FDA84B4E" author="Hoppe" box="[429,580,1568,1594]" pageId="6" pageNumber="13" refString="Hoppe, J. (1989) Los Parques Nacionales de la Republica Dominicana. Editora Corripio C. por A., Santo Domingo, 137 pp." type="book" year="1989">Hoppe 1989</bibRefCitation>
). The flight intercept trap (FIT) was located in an area of soil accumulation between limestone &quot;magote&quot; hills, covered in mesic forest. Interestingly, another FIT a very short distance away, but on the rock of the magote, did not capture any of this species. The 2
<typeStatus id="54AE8868AF3BFFB7FB6FF998FAEC4BFE" box="[1168,1280,1648,1674]" pageId="6" pageNumber="7" type="paratype">paratypes</typeStatus>
not from the Los Haïtises FITs are from the area across Samana Bay from Los Haïtises and on the north coast a short distance to the west of the Samana Peninsula.
</paragraph>
</subSubSection>
</treatment>
</document>