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<document id="C97FED3A5ED0996325EE79467C174440" ID-DOI="10.5281/zenodo.188831" ID-GBIF-Dataset="370305cb-7612-43e1-853a-235c328dc888" ID-ISSN="1175-5326" ID-Zenodo-Dep="188831" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461097017994" checkinUser="plazi" docAuthor="Schubart, Christoph D., Liu, Hung-Chang &amp; Ng, Peter K. L." docDate="2009" docId="03B9AB00FFD4B642FF7478C656F4140D" docLanguage="en" docName="zt02154p029.pdf" docOrigin="Zootaxa 2154" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Lithoselatium kusu Schubart, Liu &amp; Ng, 2009, n. sp." docType="treatment" docVersion="10" lastPageNumber="26" masterDocId="FF80D378FFC0B65BFFE3785154051603" masterDocTitle="Revision of Selatium Serène &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), with description of a new genus and two new species" masterLastPageNumber="29" masterPageNumber="1" pageNumber="21" updateTime="1698589813226" updateUser="plazi">
<mods:mods id="99B839E49A82369272E9238189FF0CDC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BE50721D915E5C154A38AF05FB5D5B7E">
<mods:title id="71DE86EB969FF0818E5B6CCC74521305">Revision of Selatium Serène &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), with description of a new genus and two new species</mods:title>
</mods:titleInfo>
<mods:name id="A6F36CF878A0DCF4973B56CBC5A9C180" type="personal">
<mods:role id="60FD65B683ACAB68B8F498F3EF73EE02">
<mods:roleTerm id="3BF7DF49296D9ECEBD216F9DCC3CAD9E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="18210A127F39C3B92C76465F40B0F757">Schubart, Christoph D.</mods:namePart>
</mods:name>
<mods:name id="9A237D0612B9DC8175B3A099ACCF9723" type="personal">
<mods:role id="1342842E49A3B811D63D7B33250AD045">
<mods:roleTerm id="709BFEED9E7A76D867E1854AAAE0B797">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="038505FF91A07D80DE8CC3D835D20E99">Liu, Hung-Chang</mods:namePart>
</mods:name>
<mods:name id="5084EB3A131C5971500BF298DED8F04C" type="personal">
<mods:role id="957D878B7EE3A746D70A3A9CE995F13A">
<mods:roleTerm id="631335C1028F1FA301F7A568B8056C2D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="78514B0F68CD4B27C5750985EE609577">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0B175867D3E8026919800BD92E6496D2">text</mods:typeOfResource>
<mods:relatedItem id="CAE2020EE63ECB2F7F86373EDA556626" type="host">
<mods:titleInfo id="36F3C509349058536EF31A22414741DC">
<mods:title id="53DF944AA36BEE0A0D10D0296F9BBD76">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6810D1BD7E143079A4265C1C9FC11190">
<mods:date id="29CDC995AAD6EED6054ABC9DF096F293">2009</mods:date>
<mods:detail id="B4BC807C5D7AD1735F3EB46D97FA89B4" type="volume">
<mods:number id="B005578D0E516D7F1C53D53C417F4C06">2154</mods:number>
</mods:detail>
<mods:extent id="4984C32F49302553082E7A51AA4153BB" unit="page">
<mods:start id="0DA9143E1D1A689A55E80678D4478FDE">1</mods:start>
<mods:end id="2F011D2192985BA71DE5E7115D049724">29</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="83824EA49403A9C9924F94319BD52A96">journal article</mods:classification>
<mods:identifier id="5912C2D220184A79783204182E947D30" type="DOI">10.5281/zenodo.188831</mods:identifier>
<mods:identifier id="45E08E0F74BB4DA06A76CE85839525C9" type="GBIF-Dataset">370305cb-7612-43e1-853a-235c328dc888</mods:identifier>
<mods:identifier id="449599341CC6575D0B5F8AAAC1B938CD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BF6439737121500C9041C8B5285667A3" type="Zenodo-Dep">188831</mods:identifier>
</mods:mods>
<treatment id="03B9AB00FFD4B642FF7478C656F4140D" ID-DOI="http://doi.org/10.5281/zenodo.5678013" ID-GBIF-Taxon="119620953" ID-Zenodo-Dep="5678013" LSID="urn:lsid:plazi:treatment:03B9AB00FFD4B642FF7478C656F4140D" httpUri="http://treatment.plazi.org/id/03B9AB00FFD4B642FF7478C656F4140D" lastPageId="25" lastPageNumber="26" pageId="20" pageNumber="21">
<subSubSection id="C30A499DFFD4B64FFF7478C6553016D7" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8BAF1A16FFD4B64FFF7478C655CF16B2" blockId="20.[151,458,151,212]" box="[151,458,151,177]" pageId="20" pageNumber="21">
<heading id="D0E7AD7AFFD4B64FFF7478C655CF16B2" bold="true" box="[151,458,151,177]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="B964C604FFD4B64FFF7478C655CF16B2" bold="true" box="[151,458,151,177]" pageId="20" pageNumber="21">
<taxonomicName id="4C106195FFD4B64FFF7478C6558416B2" ID-CoL="3VL7F" box="[151,385,151,177]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="kusu" status="sp. nov.">
<emphasis id="B964C604FFD4B64FFF7478C6558416B2" bold="true" box="[151,385,151,177]" italics="true" pageId="20" pageNumber="21">Lithoselatium kusu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2577B7FFFD4B64FFE6B78C655CF16B2" box="[392,458,151,177]" pageId="20" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF1A16FFD4B64FFF7478EB553016D7" blockId="20.[151,458,151,212]" box="[151,309,186,212]" pageId="20" pageNumber="21">
(
<figureCitation id="132B0693FFD4B64FFF7C78EB552916D7" box="[159,300,186,212]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStart-2="FIGURE 15" captionStart-3="FIGURE 16" captionStartId-0="20.[151,255,1604,1628]" captionStartId-1="21.[151,255,1958,1982]" captionStartId-2="22.[151,255,1944,1968]" captionStartId-3="23.[151,255,1443,1467]" captionTargetBox-0="[338,1248,258,1590]" captionTargetBox-1="[247,1343,292,1931]" captionTargetBox-2="[241,1344,194,1915]" captionTargetBox-3="[278,1289,228,1346]" captionTargetId-0="figure@20.[338,1248,258,1590]" captionTargetId-1="figure@21.[247,1344,292,1931]" captionTargetId-2="figure@22.[241,1344,194,1915]" captionTargetId-3="figure@23.[228,1358,194,1420]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionTargetPageId-2="22" captionTargetPageId-3="23" captionText-0="FIGURE 13. Lithoselatium kusu n. sp., colour in life. A, dorsal view, paratype male (29.0 x 25.8 mm) (ZRC 2009.0569), Singapore; B, dorsal view, paratype female (21.2 x 19.0 mm) (ZRC 2009.0570), Singapore. A, B, dorsal views; C, ventral view." captionText-1="FIGURE 14. Lithoselatium kusu n. sp. Holotype male (16.0 x 14.35 mm) (ZRC 2002.0149), Kusu Island, Singapore. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." captionText-2="FIGURE 15. Lithoselatium kusu n. sp. Holotype male (16.0 x 14.35 mm) (ZRC 2002.0149), Kusu Island, Singapore. A, anterior thoracic sternum and abdomen; B, outer view of right chela; C, dorsal view of ridge on palm; D, ridge on dactylus of palm." captionText-3="FIGURE 16. Lithoselatium kusu n. sp. Holotype male (16.0 x 14.35 mm) (ZRC 2002.0149), Kusu Island, Singapore. A, abdomen; B, left third maxilliped (denuded); C, dorsal view of left G 1 (denuded); D, ventral view of left G 1 (denuded); E, dorsal view of distal part of left G 1 (denuded); F, ventral view of distal part of left G 1 (denuded)." httpUri-0="https://zenodo.org/record/188846/files/figure.png" httpUri-1="https://zenodo.org/record/188848/files/figure.png" httpUri-2="https://zenodo.org/record/188849/files/figure.png" httpUri-3="https://zenodo.org/record/188850/files/figure.png" pageId="20" pageNumber="21">Figs. 13–16</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="DF6F4A9EFFD4B64FFF747E1554D410A2" httpUri="https://zenodo.org/record/188846/files/figure.png" pageId="20" pageNumber="21" targetBox="[338,1248,258,1590]" targetPageId="20">
<paragraph id="8BAF1A16FFD4B64FFF747E1554D410A2" blockId="20.[151,1436,1604,1697]" pageId="20" pageNumber="21">
<emphasis id="B964C604FFD4B64FFF747E155523105F" bold="true" box="[151,294,1604,1628]" pageId="20" pageNumber="21">FIGURE 13.</emphasis>
<taxonomicName id="4C106195FFD4B64FFECF7E1555F21058" box="[300,503,1604,1627]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="kusu" status="sp. nov.">
<emphasis id="B964C604FFD4B64FFECF7E1555F21058" box="[300,503,1604,1627]" italics="true" pageId="20" pageNumber="21">Lithoselatium kusu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2577B7FFFD4B64FFE1E7E15563D105F" box="[509,568,1604,1628]" pageId="20" pageNumber="21" rank="species">
<emphasis id="B964C604FFD4B64FFE1E7E15563D105F" bold="true" box="[509,568,1604,1628]" pageId="20" pageNumber="21">n. sp.</emphasis>
</taxonomicNameLabel>
, colour in life. A, dorsal view, paratype male (29.0 x 25.8 mm) (ZRC2009.0569), Singapore; B, dorsal view, paratype female (21.2 x 19.0 mm) (ZRC2009.0570), Singapore. A, B, dorsal views; C, ventral view.
</paragraph>
</caption>
<subSubSection id="C30A499DFFD4B64CFF747E825688104F" lastPageId="23" lastPageNumber="24" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="8BAF1A16FFD4B64EFF747E82507A16FF" blockId="20.[151,1436,1747,2034]" lastBlockId="21.[151,1436,152,252]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<emphasis id="B964C604FFD4B64FFF747E82558B10EE" bold="true" box="[151,398,1747,1773]" pageId="20" pageNumber="21">Material examined</emphasis>
.
<collectingCountry id="F3075A86FFD4B64FFE7C7E82562610EE" box="[415,547,1747,1773]" name="Singapore" pageId="20" pageNumber="21">
<emphasis id="B964C604FFD4B64FFE7C7E82562610EE" bold="true" box="[415,547,1747,1773]" pageId="20" pageNumber="21">Singapore</emphasis>
</collectingCountry>
:
<typeStatus id="54ABA4B4FFD4B64FFDD67E8256A210EE" box="[565,679,1747,1773]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
male (16.0 x
<quantity id="4CE8B7F3FFD4B64FFCB77E8257D610EE" box="[852,979,1747,1773]" metricMagnitude="-2" metricUnit="m" metricValue="1.435" pageId="20" pageNumber="21" unit="mm" value="14.35">14.35 mm</quantity>
) (
<collectionCode id="ED0182D3FFD4B64FFC0F7E82502C10EE" box="[1004,1065,1747,1773]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="21">ZRC</collectionCode>
2002.0149),
<collectingCountry id="F3075A86FFD4B64FFB2D7E82514A10EE" box="[1230,1359,1747,1773]" name="Singapore" pageId="20" pageNumber="21">Singapore</collectingCountry>
:
<taxonomicName id="4C106195FFD4B64FFABF7E82519910EE" box="[1372,1436,1747,1773]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="27" phylum="Arthropoda" rank="species" species="kusu">Kusu</taxonomicName>
Island: between loose rocks, coll. C. D. Schubart,
<date id="FFAE3CD6FFD4B64FFD3D7EA957B91111" box="[734,956,1784,1810]" pageId="20" pageNumber="21" value="1999-12-26">26 December 1999</date>
(
<collectionCode id="ED0182D3FFD4B64FFC287EA950081111" LSID="urn:lsid:biocol.org:col:15536" box="[971,1037,1784,1810]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="20" pageNumber="21">DNA</collectionCode>
voucher);
<typeStatus id="54ABA4B4FFD4B64FFB687EA951051111" box="[1163,1280,1784,1810]" pageId="20" pageNumber="21" type="paratype">paratypes</typeStatus>
:
<specimenCount id="9D16D19FFFD4B64FFAE97EA9515D1111" box="[1290,1368,1784,1810]" pageId="20" pageNumber="21" type="male">1 male</specimenCount>
(16.8 x
<quantity id="4CE8B7F3FFD4B64FFF4F7F4F5522113B" box="[172,295,1822,1848]" metricMagnitude="-2" metricUnit="m" metricValue="1.4949999999999999" pageId="20" pageNumber="21" unit="mm" value="14.95">14.95 mm</quantity>
),
<specimenCount id="9D16D19FFFD4B64FFED87F4F55AA113B" box="[315,431,1822,1848]" pageId="20" pageNumber="21" type="juvenile">2 juvenile</specimenCount>
females (14.15 x 12.4 &amp; 11.95 x
<quantity id="4CE8B7F3FFD4B64FFCD47F4F57A4113B" box="[823,929,1822,1848]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="20" pageNumber="21" unit="mm" value="10.3">10.3 mm</quantity>
) (
<collectionCode id="ED0182D3FFD4B64FFC547F4F57F7113B" box="[951,1010,1822,1848]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="21">ZRC</collectionCode>
2002.0150), same collection data as
<typeStatus id="54ABA4B4FFD4B64FFF747F125500115E" box="[151,261,1859,1885]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D16D19FFFD4B64FFEF17F12558E115E" box="[274,395,1859,1885]" pageId="20" pageNumber="21" type="juvenile">1 juvenile</specimenCount>
female (
<collectionCode id="ED0182D3FFD4B64FFE157F125637115E" box="[502,562,1859,1885]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="21">ZRC</collectionCode>
2009.0568), Sentosa Island: rock jetty, coll. C. D. Schubart,
<date id="FFAE3CD6FFD4B64FFAFD7F1254DD1181" pageId="20" pageNumber="21" value="1999-08-15">15 August 1999</date>
;
<specimenCount id="9D16D19FFFD4B64FFF057F3955431181" box="[230,326,1896,1922]" pageId="20" pageNumber="21" type="male">7 males</specimenCount>
(largest 29.0 x
<quantity id="4CE8B7F3FFD4B64FFDEF7F39567A1181" box="[524,639,1896,1922]" metricMagnitude="-2" metricUnit="m" metricValue="2.58" pageId="20" pageNumber="21" unit="mm" value="25.8">25.8 mm</quantity>
),
<specimenCount id="9D16D19FFFD4B64FFD757F39570B1181" box="[662,782,1896,1922]" pageId="20" pageNumber="21" type="female">9 females</specimenCount>
(largest 29.1 x 26.0 mm) (
<collectionCode id="ED0182D3FFD4B64FFBBD7F3950991181" box="[1118,1180,1896,1922]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="21">ZRC</collectionCode>
2009.0569), Raffles Lighthouse, coll. H. H. Tan
<emphasis id="B964C604FFD4B64FFE057FDF561E11A4" box="[486,539,1934,1959]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
.,
<date id="FFAE3CD6FFD4B64FFDD17FDF56FE11AB" box="[562,763,1934,1960]" pageId="20" pageNumber="21" value="2002-01-30">30 January 2002</date>
;
<specimenCount id="9D16D19FFFD4B64FFCEB7FDF577E11AB" box="[776,891,1934,1960]" pageId="20" pageNumber="21" type="female">5 females</specimenCount>
(20.9 x 17.8, 19.2 x 16.2, 17.0 x 14.4, 17.0 x 14.5, 16.2 x
<quantity id="4CE8B7F3FFD4B64FFECD7FE2559E11CE" box="[302,411,1971,1997]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="20" pageNumber="21" unit="mm" value="13.5">13.5 mm</quantity>
, smallest
<collectionCode id="ED0182D3FFD4B64FFDF27FE2565411CE" LSID="urn:lsid:biocol.org:col:15536" box="[529,593,1971,1997]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="20" pageNumber="21">DNA</collectionCode>
voucher),
<specimenCount id="9D16D19FFFD4B64FFD307FE2574C11CE" box="[723,841,1971,1997]" pageId="20" pageNumber="21" type="juvenile">2 juvenile</specimenCount>
males (12.1 x 10.2, 11.31 x
<quantity id="4CE8B7F3FFD4B64FFB427FE2510A11CE" box="[1185,1295,1971,1997]" metricMagnitude="-3" metricUnit="m" metricValue="9.64" pageId="20" pageNumber="21" unit="mm" value="9.64">9.64 mm</quantity>
),
<specimenCount id="9D16D19FFFD4B64FFAC67FE2519E11CE" box="[1317,1435,1971,1997]" pageId="20" pageNumber="21" type="juvenile">1 juvenile</specimenCount>
female (11.34 x
<quantity id="4CE8B7F3FFD4B64FFE837F8955CA11F1" box="[352,463,2008,2034]" metricMagnitude="-3" metricUnit="m" metricValue="9.72" pageId="20" pageNumber="21" unit="mm" value="9.72">9.72 mm</quantity>
) (SMF-33840), Labrador Beach, coll. C. D. Schubart, S. Klaus &amp; P. Koller,
<date id="FFAE3CD6FFD4B64EFA9D7F89552D16B1" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" value="2006-03-19">19 March 2006</date>
;
<specimenCount id="9D16D19FFFD5B64EFED778C9559C16B1" box="[308,409,152,178]" pageId="21" pageNumber="22" type="female">1 female</specimenCount>
(21.2 x 19.0 mm) (
<collectionCode id="ED0182D3FFD5B64EFD9978C956B016B1" box="[634,693,152,178]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="22">ZRC</collectionCode>
2009.0570), Raffles Lighthouse,
<collectingCountry id="F3075A86FFD5B64EFBDE78C950B216B1" box="[1085,1207,152,178]" name="Singapore" pageId="21" pageNumber="22">Singapore</collectingCountry>
, coll. P. K. L. Ng,
<date id="FFAE3CD6FFD5B64EFA6E78C9553416D4" pageId="21" pageNumber="22" value="2008-08-02">2 August 2008</date>
. Other material:
<emphasis id="B964C604FFD5B64EFE1678EC56C116D4" bold="true" box="[501,708,189,215]" pageId="21" pageNumber="22">
<collectingCountry id="F3075A86FFD5B64EFE1678EC566C16D4" box="[501,617,189,215]" name="Malaysia" pageId="21" pageNumber="22">Malaysia</collectingCountry>
, Sabah
</emphasis>
:
<specimenCount id="9D16D19FFFD5B64EFD3778EC572616D4" box="[724,803,189,215]" pageId="21" pageNumber="22" type="male">1 male</specimenCount>
molt (animal lost after molting in lab, photo voucher) (ca.
<quantity id="4CE8B7F3FFD5B64EFF2A78B3553716FF" box="[201,306,226,252]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="21" pageNumber="22" unit="mm" value="12.9">12.9 mm</quantity>
carapace width) (
<collectionCode id="ED0182D3FFD5B64EFDE178B3563816FF" box="[514,573,226,252]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="22">ZRC</collectionCode>
2000.1654),
<collectingCountry id="F3075A86FFD5B64EFD3A78B3571916FF" box="[729,796,226,252]" name="Iceland" pageId="21" pageNumber="22">Pulau</collectingCountry>
Manukan, coll.
<date id="FFAE3CD6FFD5B64EFC3878B3507E16FF" box="[987,1147,226,252]" pageId="21" pageNumber="22" value="2000-06-22">22 June 2000</date>
.
</paragraph>
<caption id="DF6F4A9EFFD5B64EFF747FF7577611E3" httpUri="https://zenodo.org/record/188848/files/figure.png" pageId="21" pageNumber="22" targetBox="[247,1343,292,1931]" targetPageId="21">
<paragraph id="8BAF1A16FFD5B64EFF747FF7577611E3" blockId="21.[151,1436,1958,2016]" pageId="21" pageNumber="22">
<emphasis id="B964C604FFD5B64EFF747FF7552311BD" bold="true" box="[151,294,1958,1982]" pageId="21" pageNumber="22">FIGURE 14.</emphasis>
<taxonomicName id="4C106195FFD5B64EFECE7FF655F211BD" box="[301,503,1959,1982]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="kusu" status="sp. nov.">
<emphasis id="B964C604FFD5B64EFECE7FF655F211BD" box="[301,503,1959,1982]" italics="true" pageId="21" pageNumber="22">Lithoselatium kusu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2577B7FFFD5B64EFE1E7FF7563C11BD" box="[509,569,1958,1982]" pageId="21" pageNumber="22" rank="species">
<emphasis id="B964C604FFD5B64EFE1E7FF7563C11BD" bold="true" box="[509,569,1958,1982]" pageId="21" pageNumber="22">n. sp.</emphasis>
</taxonomicNameLabel>
Holotype male (16.0 x 14.35 mm) (ZRC 2002.0149),
<taxonomicName id="4C106195FFD5B64EFB9F7FF650B111BD" box="[1148,1204,1959,1982]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="27" phylum="Arthropoda" rank="species" species="kusu">Kusu</taxonomicName>
Island, Singapore. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace.
</paragraph>
</caption>
<caption id="DF6F4A9EFFD6B64DFF747FC9555511F7" httpUri="https://zenodo.org/record/188849/files/figure.png" pageId="22" pageNumber="23" targetBox="[241,1344,194,1915]" targetPageId="22">
<paragraph id="8BAF1A16FFD6B64DFF747FC9555511F7" blockId="22.[151,1437,1944,2036]" pageId="22" pageNumber="23">
<emphasis id="B964C604FFD6B64DFF747FC9552311B3" bold="true" box="[151,294,1944,1968]" pageId="22" pageNumber="23">FIGURE 15.</emphasis>
<taxonomicName id="4C106195FFD6B64DFECE7FC955F211AC" box="[301,503,1944,1967]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="kusu" status="sp. nov.">
<emphasis id="B964C604FFD6B64DFECE7FC955F211AC" box="[301,503,1944,1967]" italics="true" pageId="22" pageNumber="23">Lithoselatium kusu</emphasis>
</taxonomicName>
<emphasis id="B964C604FFD6B64DFE1E7FC9563C11B3" bold="true" box="[509,569,1944,1968]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="A2577B7FFFD6B64DFE1E7FC9563C11B3" box="[509,569,1944,1968]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Holotype male (16.0 x 14.35 mm) (ZRC 2002.0149),
<taxonomicName id="4C106195FFD6B64DFB9F7FC950B111AC" box="[1148,1204,1944,1967]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="27" phylum="Arthropoda" rank="species" species="kusu">Kusu</taxonomicName>
Island, Singapore. A, anterior thoracic sternum and abdomen; B, outer view of right chela; C, dorsal view of ridge on palm; D, ridge on dactylus of palm.
</paragraph>
</caption>
<caption id="DF6F4A9EFFD7B64CFF747DF250C41003" httpUri="https://zenodo.org/record/188850/files/figure.png" pageId="23" pageNumber="24" targetBox="[278,1289,228,1346]" targetPageId="23">
<paragraph id="8BAF1A16FFD7B64CFF747DF250C41003" blockId="23.[151,1436,1443,1536]" pageId="23" pageNumber="24">
<emphasis id="B964C604FFD7B64CFF747DF2552313B8" bold="true" box="[151,294,1443,1467]" pageId="23" pageNumber="24">FIGURE 16.</emphasis>
<taxonomicName id="4C106195FFD7B64CFECE7DF555F213B8" box="[301,503,1444,1467]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="kusu" status="sp. nov.">
<emphasis id="B964C604FFD7B64CFECE7DF555F213B8" box="[301,503,1444,1467]" italics="true" pageId="23" pageNumber="24">Lithoselatium kusu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2577B7FFFD7B64CFE1E7DF2563C13B8" box="[509,569,1443,1467]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B964C604FFD7B64CFE1E7DF2563C13B8" bold="true" box="[509,569,1443,1467]" pageId="23" pageNumber="24">n. sp.</emphasis>
</taxonomicNameLabel>
Holotype male (16.0 x 14.35 mm) (ZRC 2002.0149),
<taxonomicName id="4C106195FFD7B64CFB9F7DF550B113B8" box="[1148,1204,1444,1467]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="27" phylum="Arthropoda" rank="species" species="kusu">Kusu</taxonomicName>
Island, Singapore. A, abdomen; B, left third maxilliped (denuded); C, dorsal view of left G1 (denuded); D, ventral view of left G1 (denuded); E, dorsal view of distal part of left G1 (denuded); F, ventral view of distal part of left G1 (denuded).
</paragraph>
</caption>
<paragraph id="8BAF1A16FFD7B64CFF257E635688104F" blockId="23.[151,1436,1586,2038]" box="[198,653,1586,1612]" pageId="23" pageNumber="24">
<emphasis id="B964C604FFD7B64CFF257E635561104F" bold="true" box="[198,356,1586,1612]" pageId="23" pageNumber="24">
<typeStatus id="54ABA4B4FFD7B64CFF257E635501104F" box="[198,260,1586,1612]" pageId="23" pageNumber="24">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="F3075A86FFD7B64CFE907E6355E8104F" box="[371,493,1586,1612]" name="Singapore" pageId="23" pageNumber="24">Singapore</collectingCountry>
,
<taxonomicName id="4C106195FFD7B64CFE1B7E635633104F" box="[504,566,1586,1612]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="27" phylum="Arthropoda" rank="species" species="kusu">Kusu</taxonomicName>
Island.
</paragraph>
</subSubSection>
<subSubSection id="C30A499DFFD7B64CFF257E08576B11F5" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="8BAF1A16FFD7B64CFF257E08576B11F5" blockId="23.[151,1436,1586,2038]" pageId="23" pageNumber="24">
<emphasis id="B964C604FFD7B64CFF257E08553A1070" bold="true" box="[198,319,1625,1651]" pageId="23" pageNumber="24">Diagnosis</emphasis>
. Carapace slightly trapezoidal, diverging toward posterior margin; lateral margins entire, without trace of epibranchial tooth (
<figureCitation id="132B0693FFD7B64CFDB37ED156AD1099" box="[592,680,1664,1690]" captionStart="FIGURE 14" captionStartId="21.[151,255,1958,1982]" captionTargetBox="[247,1343,292,1931]" captionTargetId="figure@21.[247,1344,292,1931]" captionTargetPageId="21" captionText="FIGURE 14. Lithoselatium kusu n. sp. Holotype male (16.0 x 14.35 mm) (ZRC 2002.0149), Kusu Island, Singapore. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/188848/files/figure.png" pageId="23" pageNumber="24">Fig. 14</figureCitation>
A, B); frontal margin with anterior lobes prominent, posterior lobes weakly demarcated (
<figureCitation id="132B0693FFD7B64CFE317EF6562210C2" box="[466,551,1703,1729]" captionStart="FIGURE 14" captionStartId="21.[151,255,1958,1982]" captionTargetBox="[247,1343,292,1931]" captionTargetId="figure@21.[247,1344,292,1931]" captionTargetPageId="21" captionText="FIGURE 14. Lithoselatium kusu n. sp. Holotype male (16.0 x 14.35 mm) (ZRC 2002.0149), Kusu Island, Singapore. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/188848/files/figure.png" pageId="23" pageNumber="24">Fig. 14</figureCitation>
A, B); outer and inner surfaces of chelae smooth (
<figureCitation id="132B0693FFD7B64CFB947EF650CB10C2" box="[1143,1230,1703,1729]" captionStart="FIGURE 15" captionStartId="22.[151,255,1944,1968]" captionTargetBox="[241,1344,194,1915]" captionTargetId="figure@22.[241,1344,194,1915]" captionTargetPageId="22" captionText="FIGURE 15. Lithoselatium kusu n. sp. Holotype male (16.0 x 14.35 mm) (ZRC 2002.0149), Kusu Island, Singapore. A, anterior thoracic sternum and abdomen; B, outer view of right chela; C, dorsal view of ridge on palm; D, ridge on dactylus of palm." httpUri="https://zenodo.org/record/188849/files/figure.png" pageId="23" pageNumber="24">Fig. 15</figureCitation>
B); dorsal margin of palm with one continuous longitudinal crest of proximal tubercles, followed by a pectinated crest consisting of 33 or 34 teeth, and approximately 5 or 6 partly fused distal tubercles (fewer, broader and shorter teeth in females); 3–5 short and oblique subsidiary rows of tubercles on inner side (
<figureCitation id="132B0693FFD7B64CFB957F4B50C91137" box="[1142,1228,1818,1844]" captionStart="FIGURE 15" captionStartId="22.[151,255,1944,1968]" captionTargetBox="[241,1344,194,1915]" captionTargetId="figure@22.[241,1344,194,1915]" captionTargetPageId="22" captionText="FIGURE 15. Lithoselatium kusu n. sp. Holotype male (16.0 x 14.35 mm) (ZRC 2002.0149), Kusu Island, Singapore. A, anterior thoracic sternum and abdomen; B, outer view of right chela; C, dorsal view of ridge on palm; D, ridge on dactylus of palm." httpUri="https://zenodo.org/record/188849/files/figure.png" pageId="23" pageNumber="24">Fig. 15</figureCitation>
C); dorsal margin of dactylar finger with a row of 30–32 tubercles, with circular rings and a central conical apex, more widely spaced towards distal part (
<figureCitation id="132B0693FFD7B64CFE057F39563A1181" box="[486,575,1896,1922]" captionStart="FIGURE 15" captionStartId="22.[151,255,1944,1968]" captionTargetBox="[241,1344,194,1915]" captionTargetId="figure@22.[241,1344,194,1915]" captionTargetPageId="22" captionText="FIGURE 15. Lithoselatium kusu n. sp. Holotype male (16.0 x 14.35 mm) (ZRC 2002.0149), Kusu Island, Singapore. A, anterior thoracic sternum and abdomen; B, outer view of right chela; C, dorsal view of ridge on palm; D, ridge on dactylus of palm." httpUri="https://zenodo.org/record/188849/files/figure.png" pageId="23" pageNumber="24">Fig. 15</figureCitation>
D); cutting edge of with 2 larger teeth proximally and one larger tooth subdistally; cutting edge of pollex with 2 larger teeth proximally to medially and one larger tooth subdistally (
<figureCitation id="132B0693FFD7B64CFF7C7FE454F011CC" box="[159,245,1973,1999]" captionStart="FIGURE 15" captionStartId="22.[151,255,1944,1968]" captionTargetBox="[241,1344,194,1915]" captionTargetId="figure@22.[241,1344,194,1915]" captionTargetPageId="22" captionText="FIGURE 15. Lithoselatium kusu n. sp. Holotype male (16.0 x 14.35 mm) (ZRC 2002.0149), Kusu Island, Singapore. A, anterior thoracic sternum and abdomen; B, outer view of right chela; C, dorsal view of ridge on palm; D, ridge on dactylus of palm." httpUri="https://zenodo.org/record/188849/files/figure.png" pageId="23" pageNumber="24">Fig. 15</figureCitation>
B); G1 with pectinated distal part subtruncate, tip gently clefted, lower part larger than upper part (
<figureCitation id="132B0693FFD7B64CFA8E7FE454B111F5" captionStart="FIGURE 16" captionStartId="23.[151,255,1443,1467]" captionTargetBox="[278,1289,228,1346]" captionTargetId="figure@23.[228,1358,194,1420]" captionTargetPageId="23" captionText="FIGURE 16. Lithoselatium kusu n. sp. Holotype male (16.0 x 14.35 mm) (ZRC 2002.0149), Kusu Island, Singapore. A, abdomen; B, left third maxilliped (denuded); C, dorsal view of left G 1 (denuded); D, ventral view of left G 1 (denuded); E, dorsal view of distal part of left G 1 (denuded); F, ventral view of distal part of left G 1 (denuded)." httpUri="https://zenodo.org/record/188850/files/figure.png" pageId="23" pageNumber="24">Fig. 16</figureCitation>
C–F). Proportionally shorter legs compared to
<taxonomicName id="4C106195FFD7B64CFD3B7F8D576211F6" box="[728,871,2012,2037]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="pulchrum">
<emphasis id="B964C604FFD7B64CFD3B7F8D576211F6" box="[728,871,2012,2037]" italics="true" pageId="23" pageNumber="24">L. pulchrum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF6F4A9EFFD8B643FF747F3A519611F3" httpUri="https://zenodo.org/record/188851/files/figure.png" pageId="24" pageNumber="25" targetBox="[229,1367,182,1873]" targetPageId="24">
<paragraph id="8BAF1A16FFD8B643FF747F3A519611F3" blockId="24.[151,1436,1899,2032]" pageId="24" pageNumber="25">
<emphasis id="B964C604FFD8B643FF747F3A552D1180" bold="true" box="[151,296,1899,1923]" pageId="24" pageNumber="25">FIGURE 17.</emphasis>
Male thoracic sternum and abdomen. A,
<taxonomicName id="4C106195FFD8B643FD097F3D50561180" authority="De Man, 1887" authorityName="De Man" authorityYear="1887" box="[746,1107,1900,1923]" class="Malacostraca" family="Sesarmidae" genus="Selatium" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="brockii">
<emphasis id="B964C604FFD8B643FD097F3D579E1180" box="[746,923,1900,1923]" italics="true" pageId="24" pageNumber="25">Selatium brockii</emphasis>
(De Man, 1887)
</taxonomicName>
, male (29.2 x 26.9 mm) (ZRC 1989.2875), Mandai, Singapore; B,
<taxonomicName id="4C106195FFD8B643FDFF7FC1571811A4" box="[540,797,1936,1959]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pulchrum" status="sp. nov.">
<emphasis id="B964C604FFD8B643FDFF7FC1571811A4" box="[540,797,1936,1959]" italics="true" pageId="24" pageNumber="25">Lithoselatium pulchrum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2577B7FFFD8B643FCC67FDE576711A4" box="[805,866,1935,1959]" pageId="24" pageNumber="25" rank="species">
<emphasis id="B964C604FFD8B643FCC67FDE576711A4" bold="true" box="[805,866,1935,1959]" pageId="24" pageNumber="25">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype male (23.5 x 20.8 mm) (ZRC 2002.0152), Kenting, Taiwan; C,
<taxonomicName id="4C106195FFD8B643FE677FE5565F11C8" box="[388,602,1972,1995]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="kusu" status="sp. nov.">
<emphasis id="B964C604FFD8B643FE677FE5565F11C8" box="[388,602,1972,1995]" italics="true" pageId="24" pageNumber="25">Lithoselatium kusu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2577B7FFFD8B643FD817FE256A711C8" box="[610,674,1971,1995]" pageId="24" pageNumber="25" rank="species">
<emphasis id="B964C604FFD8B643FD817FE256A711C8" bold="true" box="[610,674,1971,1995]" pageId="24" pageNumber="25">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype male (16.0 x 14.35 mm) (ZRC 2002.0149),
<taxonomicName id="4C106195FFD8B643FAEE7FE5514D11C8" box="[1293,1352,1972,1995]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="27" phylum="Arthropoda" rank="species" species="kusu">Kusu</taxonomicName>
Island, Singapore. a2–6 = abdominal somites 2–6; st5–8 = sternites 5–8; es5–7 = episternites 5–7; c2–5 = ambulatory coxa 2–5
</paragraph>
</caption>
<subSubSection id="C30A499DFFD9B642FF2578C655001725" pageId="25" pageNumber="26" type="etymology">
<paragraph id="8BAF1A16FFD9B642FF2578C655001725" blockId="25.[151,1436,151,526]" pageId="25" pageNumber="26">
<emphasis id="B964C604FFD9B642FF2578C6554E16B2" bold="true" box="[198,331,151,177]" pageId="25" pageNumber="26">Etymology</emphasis>
. The name
<taxonomicName id="4C106195FFD9B642FE3478C9560B16B1" box="[471,526,152,178]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kusu">kusu</taxonomicName>
is used as a noun and refers to the
<typeStatus id="54ABA4B4FFD9B642FC4D78C957E416B1" box="[942,993,152,178]" pageId="25" pageNumber="26">type</typeStatus>
locality of the new species, the small island south of
<collectingCountry id="F3075A86FFD9B642FEA878EF55C316DB" box="[331,454,190,216]" name="Singapore" pageId="25" pageNumber="26">Singapore</collectingCountry>
. In Chinese
<taxonomicName id="4C106195FFD9B642FDB678EF568916DB" box="[597,652,190,216]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kusu">kusu</taxonomicName>
means “turtle”, describing the shape of the island.
<emphasis id="B964C604FFD9B642FF2578B4551916FC" bold="true" box="[198,284,229,255]" pageId="25" pageNumber="26">Colour</emphasis>
. In life, carapace and legs purplish; carapace with yellowish-green mottling; chelae orangish-red (
<figureCitation id="132B0693FFD9B642FF7C795D54F31725" box="[159,246,268,294]" captionStart="FIGURE 13" captionStartId="20.[151,255,1604,1628]" captionTargetBox="[338,1248,258,1590]" captionTargetId="figure@20.[338,1248,258,1590]" captionTargetPageId="20" captionText="FIGURE 13. Lithoselatium kusu n. sp., colour in life. A, dorsal view, paratype male (29.0 x 25.8 mm) (ZRC 2009.0569), Singapore; B, dorsal view, paratype female (21.2 x 19.0 mm) (ZRC 2009.0570), Singapore. A, B, dorsal views; C, ventral view." httpUri="https://zenodo.org/record/188846/files/figure.png" pageId="25" pageNumber="26">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A499DFFD9B642FF25796356F4140D" pageId="25" pageNumber="26" type="discussion">
<paragraph id="8BAF1A16FFD9B642FF25796356F4140D" blockId="25.[151,1436,151,526]" pageId="25" pageNumber="26">
<emphasis id="B964C604FFD9B642FF257963553D174F" bold="true" box="[198,312,306,332]" pageId="25" pageNumber="26">Remarks</emphasis>
.
<taxonomicName id="4C106195FFD9B642FEA9796356291748" box="[330,556,306,331]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kusu" status="sp. nov.">
<emphasis id="B964C604FFD9B642FEA9796356291748" box="[330,556,306,331]" italics="true" pageId="25" pageNumber="26">Lithoselatium kusu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2577B7FFFD9B642FDD479635678174F" box="[567,637,306,332]" pageId="25" pageNumber="26" rank="species">
<emphasis id="B964C604FFD9B642FDD479635678174F" bold="true" box="[567,637,306,332]" pageId="25" pageNumber="26">n. sp.</emphasis>
</taxonomicNameLabel>
can be morphologically separated from
<taxonomicName id="4C106195FFD9B642FB89796350F91748" box="[1130,1276,306,331]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pulchrum" status="sp. nov.">
<emphasis id="B964C604FFD9B642FB89796350F91748" box="[1130,1276,306,331]" italics="true" pageId="25" pageNumber="26">L. pulchrum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2577B7FFFD9B642FAEB7963514B174F" box="[1288,1358,306,332]" pageId="25" pageNumber="26" rank="species">
<emphasis id="B964C604FFD9B642FAEB7963514B174F" bold="true" box="[1288,1358,306,332]" pageId="25" pageNumber="26">n. sp.</emphasis>
</taxonomicNameLabel>
by its proportionately shorter ambulatory legs and, if alive, by the strikingly different colour pattern.
<emphasis id="B964C604FFD9B642FF25792E552D179A" bold="true" box="[198,296,383,409]" pageId="25" pageNumber="26">Ecology</emphasis>
.
<taxonomicName id="4C106195FFD9B642FEDF79D15625179A" box="[316,544,384,409]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kusu">
<emphasis id="B964C604FFD9B642FEDF79D15625179A" box="[316,544,384,409]" italics="true" pageId="25" pageNumber="26">Lithoselatium kusu</emphasis>
</taxonomicName>
is so far known from Labrador Beach,
<taxonomicName id="4C106195FFD9B642FBF979D1505D1799" box="[1050,1112,384,410]" class="Malacostraca" family="Sesarmidae" genus="Lithoselatium" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="27" phylum="Arthropoda" rank="species" species="kusu">Kusu</taxonomicName>
I., Sentosa I. and Raffles Lighthouse (
<collectingCountry id="F3075A86FFD9B642FECE79F755AC17C3" box="[301,425,422,448]" name="Singapore" pageId="25" pageNumber="26">Singapore</collectingCountry>
) as well as from an island off Sabah. The species was found among coarse rock and coral rubble well above the water line in all localities. When chased, it moves rapidly between the rocks without necessarily seeking protection in the water.
</paragraph>
</subSubSection>
</treatment>
</document>
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