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<document id="2B7EFCFF32AF3A7C03D2F79FDC5ED840" ID-DOI="10.11646/zootaxa.5447.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="11150401" ID-ZooBank="DFD56548-A978-4D66-BCC9-BAC911C10234" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1715181675979" checkinUser="plazi" docAuthor="Sideleva, Valentina G. &amp; Zhidkov, Zakhar V." docDate="2024" docId="03A85444FFEC1856FF6AFA05907636D5" docLanguage="en" docName="zootaxa.5447.4.2.pdf" docOrigin="Zootaxa 5447 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cottus dorofeevi Sideleva &amp; Zhidkov 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="491" masterDocId="FF912C3CFFE21844FFFDFF9692323555" masterDocTitle="Description of a new species of Cottus Linnaeus (Cottidae) from the Western Dvina / Daugava River system, Baltic Sea basin, based on integrative taxonomy" masterLastPageNumber="495" masterPageNumber="473" pageNumber="487" updateTime="1715275075009" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E0052E48F20C335604C93E54DDDD165F">Description of a new species of Cottus Linnaeus (Cottidae) from the Western Dvina / Daugava River system, Baltic Sea basin, based on integrative taxonomy</mods:title>
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<mods:name id="41505074BE3FE8A6FAA67E7DD4959D4F" type="personal">
<mods:role id="42BDC8D281714B017CF301E6D0DB2206">
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<mods:namePart id="83CC49937134557BC2848E30B716D78B">Sideleva, Valentina G.</mods:namePart>
<mods:nameIdentifier id="6770652E9CB6543F2EDC5E2CAD77C6DF" type="ORCID">0000-0002-4711-1379</mods:nameIdentifier>
<mods:affiliation id="E5217198800A994ECA6DB9C9E183B99C">Zoological Institute, Russian Academy of Sciences, Universitetskaya emb., 1, St. Petersburg, 199034 Russia.</mods:affiliation>
<mods:nameIdentifier id="DFF95297B409662D4296E17995770225" type="email">vsideleva@gmail.com</mods:nameIdentifier>
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<mods:name id="914CA638A1D9D46A85F54A0B7BBA6B7B" type="personal">
<mods:role id="4A7B28ABA40E12107CFA01107C22C1EB">
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<mods:namePart id="31003ADFBAE04CAD687D3CFCC148D3D1">Zhidkov, Zakhar V.</mods:namePart>
<mods:nameIdentifier id="F348C56BFDE907B798DBF1D5DC30A0B4" type="ORCID">0000-0002-8866-7779</mods:nameIdentifier>
<mods:affiliation id="25F760200713AD717A2942EDE25EB427">Zoological Institute, Russian Academy of Sciences, Universitetskaya emb., 1, St. Petersburg, 199034 Russia.</mods:affiliation>
<mods:nameIdentifier id="C318E83FEA708F4C9E33CA107764976B" type="email">zhidkovz@gmail.com</mods:nameIdentifier>
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<mods:title id="A148461E4AF6CFCB9ED666B9880A84D4">Zootaxa</mods:title>
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<mods:part id="0F36E77AD0685AFDABABD74DA29F7E5A">
<mods:date id="380EA13B91BBD891F48E9F92DD8C67B2">2024</mods:date>
<mods:detail id="3720022416DF170A0F5B282A132CBD3F" type="pubDate">
<mods:number id="F3FA0424841EA4E24D031CF3118EB538">2024-05-08</mods:number>
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<mods:detail id="DA396AE07B88F1ECEF821687A4D3806B" type="volume">
<mods:number id="D27FF1E3408CBABF0A1B161C75FC802B">5447</mods:number>
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<mods:number id="40962D9A3A2E1620DBBAA0BE33BC2BBB">4</mods:number>
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<mods:identifier id="9BA79B262C4FFABD7CC10BC7722F14B1" type="DOI">10.11646/zootaxa.5447.4.2</mods:identifier>
<mods:identifier id="4E6AE53FDC81D17267A82AB0D6536FD3" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03A85444FFEC1856FF6AFA05907636D5" ID-DOI="http://doi.org/10.5281/zenodo.11165789" ID-Zenodo-Dep="11165789" LSID="urn:lsid:plazi:treatment:03A85444FFEC1856FF6AFA05907636D5" httpUri="http://treatment.plazi.org/id/03A85444FFEC1856FF6AFA05907636D5" lastPageId="18" lastPageNumber="491" pageId="14" pageNumber="487">
<subSubSection id="C31BB6D9FFEC184AFF6AFA0593F030F8" box="[151,450,1426,1453]" pageId="14" pageNumber="487" type="nomenclature">
<paragraph id="8BBEE552FFEC184AFF6AFA0593F030F8" blockId="14.[151,467,1426,1489]" box="[151,450,1426,1453]" pageId="14" pageNumber="487">
<heading id="D0F6523EFFEC184AFF6AFA0593F030F8" bold="true" box="[151,450,1426,1453]" fontSize="11" level="1" pageId="14" pageNumber="487" reason="1">
<emphasis id="B9753940FFEC184AFF6AFA0593F030F8" bold="true" box="[151,450,1426,1453]" pageId="14" pageNumber="487">
<taxonomicName id="4C019ED1FFEC184AFF6AFA05936F30F8" authority="Sideleva &amp; Zhidkov, 2024" authorityName="Sideleva &amp; Zhidkov" authorityYear="2024" box="[151,349,1426,1453]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="14" pageNumber="487" phylum="Chordata" rank="species" species="dorofeevi" status="sp. nov.">
<emphasis id="B9753940FFEC184AFF6AFA05936F30F8" bold="true" box="[151,349,1426,1453]" italics="true" pageId="14" pageNumber="487">Cottus dorofeevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246843BFFEC184AFE98FA0593F030F8" box="[357,450,1427,1453]" pageId="14" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31BB6D9FFEC184AFF6AFA2193E13084" box="[151,467,1463,1489]" pageId="14" pageNumber="487" type="description">
<paragraph id="8BBEE552FFEC184AFF6AFA2193E13084" blockId="14.[151,467,1426,1489]" box="[151,467,1463,1489]" pageId="14" pageNumber="487">
<figureCitation id="133AF9D7FFEC184AFF6AFA2192EC3084" box="[151,222,1463,1489]" captionStart="FIGURE 2" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[158,1429,181,1919]" captionTargetId="figure-29@4.[158,1429,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Cottus sp. from the Western Dvina / Daugava River system, ♁, ZIN 56916, SL 68 mm, TL 79 mm; the Ovsyanka River, tributary of the Western Dvina / Daugava, Belarus, 55.5907° N, 30.4080° E; August 24, 2022; collectors Z.V. Zhidkov and S.A. Dorofeev. (a) lateral view; (b) dorsal view; (c) ventral view." figureDoi="http://doi.org/10.5281/zenodo.11150405" httpUri="https://zenodo.org/record/11150405/files/figure.png" pageId="14" pageNumber="487">Figs 2</figureCitation>
,
<figureCitation id="133AF9D7FFEC184AFF17FA2192C83084" box="[234,250,1463,1489]" captionStart="FIGURE 9" captionStartId="15.[152,255,1913,1937]" captionTargetBox="[156,1432,996,1878]" captionTargetId="figure-357@15.[151,1435,993,1890]" captionTargetPageId="15" captionText="FIGURE 9. Illustration of the holotype of C. dorofeevi (ZIN 56916): (a) lateral view; (b) dorsal view of the head." figureDoi="http://doi.org/10.5281/zenodo.11150423" httpUri="https://zenodo.org/record/11150423/files/figure.png" pageId="14" pageNumber="487">9</figureCitation>
,
<figureCitation id="133AF9D7FFEC184AFEFBFA2193163084" box="[262,292,1463,1489]" captionStart="FIGURE 10" captionStartId="16.[152,255,1511,1535]" captionTargetBox="[151,1436,921,1487]" captionTargetId="figure-350@16.[151,1436,921,1487]" captionTargetPageId="16" captionText="FIGURE 10. The skeleton of Cottus dorofeevi, ZIN 56917, Ovsyanka River, tributary of the Western Dvina / Daugava." figureDoi="http://doi.org/10.5281/zenodo.11150425" httpUri="https://zenodo.org/record/11150425/files/figure.png" pageId="14" pageNumber="487">10</figureCitation>
,
<figureCitation id="133AF9D7FFEC184AFECDFA21937C3084" box="[304,334,1463,1489]" captionStart="FIGURE 11" captionStartId="17.[152,255,1940,1964]" captionTargetBox="[163,1423,357,1904]" captionTargetId="figure-101@17.[151,1435,345,1917]" captionTargetPageId="17" captionText="FIGURE 11. Lateral line system on the head of Cottus dorofeevi (ZIN 56916). CSO, supraorabital sensory canal; CIO, infraorbital canal, CT, temporal canal; CPM, praeoperculo-mandibular canal; CLL, trunk (lateral) canal; CMT, temporal (postorbital) commissure." figureDoi="http://doi.org/10.5281/zenodo.11150427" httpUri="https://zenodo.org/record/11150427/files/figure.png" pageId="14" pageNumber="487">11</figureCitation>
;
<tableCitation id="C683D0E9FFEC184AFEA7FA2193E13084" box="[346,467,1463,1489]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="5.[152,245,291,315]" captionStartId-1="6.[152,245,151,175]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="TABLE 1. Proportional measurements as percentages of standard length for holotype and paratypes of Cottus sp. “dorofeevi” from the Western Dvina / Daugava River system." captionText-1="TABLE 2. Frequency of occurrence of fin-rays counts for Cottus sp. “dorofeevi” from the Western Dvina / Daugava system, C. koshewnikowi (ZIN 55582), C. microstomus, and C. gobio (UK 1889)." httpUri-0="http://table.plazi.org/id/DF7EB5DAFFE71841FF65FEB591153435" httpUri-1="http://table.plazi.org/id/DF7EB5DAFFE41842FF65FF0191CC3586" pageId="14" pageNumber="487" tableUuid-0="DF7EB5DAFFE71841FF65FEB591153435" tableUuid-1="DF7EB5DAFFE41842FF65FF0191CC3586">Tables 1–2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31BB6D9FFEC184AFF6AFA6A911C3304" pageId="14" pageNumber="487" type="reference_group">
<paragraph id="8BBEE552FFEC184AFF6AFA6A906E3346" blockId="14.[151,925,1531,1617]" box="[151,604,1531,1556]" pageId="14" pageNumber="487">
<taxonomicName id="4C019ED1FFEC184AFF6AFA6A93283346" baseAuthorityName="UK" baseAuthorityYear="1889" box="[151,282,1531,1555]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="14" pageNumber="487" phylum="Chordata" rank="species" species="gobio">
<emphasis id="B9753940FFEC184AFF6AFA6A93283346" box="[151,282,1531,1555]" italics="true" pageId="14" pageNumber="487">Cottus gobio</emphasis>
</taxonomicName>
—
<bibRefCitation id="EF9098A3FFEC184AFEC9FA6D93FB3346" author="Sapunov, A. P." box="[308,457,1531,1555]" pageId="14" pageNumber="487" refId="ref12177" refString="Sapunov, A. P. (1893) Western Dvina River. s. n., Vitebsk, 512 pp. [in Russian]" type="book" year="1893">Sapunov, 1893</bibRefCitation>
(in part): 238;
</paragraph>
<paragraph id="8BBEE552FFEC184AFF6AF98C91AF3367" blockId="14.[151,925,1531,1617]" box="[151,925,1562,1587]" pageId="14" pageNumber="487">
<treatmentCitationGroup id="AB11C27CFFEC184AFF6AF98C91AF3367" box="[151,925,1562,1587]" pageId="14" pageNumber="487">
<taxonomicName id="4C019ED1FFEC184AFF6AF98C939C3367" ID-CoL="YYJ9" box="[151,430,1562,1586]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="14" pageNumber="487" phylum="Chordata" rank="subSpecies" species="gobio" subSpecies="koshewnikowi">
<emphasis id="B9753940FFEC184AFF6AF98C939C3367" box="[151,430,1562,1586]" italics="true" pageId="14" pageNumber="487">Cottus gobio koshewnikowi</emphasis>
</taxonomicName>
—
<bibRefCitation id="EF9098A3FFEC184AFE35F98D90033367" author="Berg, L. S." box="[456,561,1562,1587]" pageId="14" pageNumber="487" refId="ref10874" refString="Berg, L. S. (1949) Freshwater Fishes of Soviet Union and Adjacent Countries. Part 3. Academy of Sciences of USSR, Moscow- Leningrad, 452 pp. [in Russian]" type="book" year="1949">Berg 1949</bibRefCitation>
(in part): 1147;
<treatmentCitation id="0AA0C343FFEC184AFD2BF98C91AB3367" author="Zhukov, P. I." box="[726,921,1562,1586]" page="397" pageId="14" pageNumber="487" year="1965">
<bibRefCitation id="EF9098A3FFEC184AFD2BF98C91AB3367" author="Zhukov, P. I." box="[726,921,1562,1586]" pageId="14" pageNumber="487" refId="ref13085" refString="Zhukov, P. I. (1965) Fishes of Belorussia. Nauka i tekhnika, Minsk, 415 pp. [in Russian]" type="book" year="1965">Zhukov, 1965: 397</bibRefCitation>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BBEE552FFEC184AFF6AF9AF911C3304" blockId="14.[151,925,1531,1617]" box="[151,814,1593,1617]" pageId="14" pageNumber="487">
<taxonomicName id="4C019ED1FFEC184AFF6AF9AF93283304" ID-CoL="YYGV" baseAuthorityName="UK" baseAuthorityYear="1889" box="[151,282,1593,1617]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="14" pageNumber="487" phylum="Chordata" rank="species" species="gobio">
<emphasis id="B9753940FFEC184AFF6AF9AF93283304" box="[151,282,1593,1617]" italics="true" pageId="14" pageNumber="487">Cottus gobio</emphasis>
</taxonomicName>
—
<bibRefCitation id="EF9098A3FFEC184AFECEF9AF909A3304" author="Leshchenko, A. V. &amp; Palchevskaya, K. I." box="[307,680,1593,1617]" pageId="14" pageNumber="487" pagination="79 - 81" refId="ref11776" refString="Leshchenko, A. V. &amp; Palchevskaya, K. I. (2012) The current state of the fish fauna of the water bodies and water courses of the national park &quot; Braslavskie ozera &quot;. In: Yanchurevich, O. V. (Ed.), &quot; Zoological Readings 2012 &quot;. Proceedings of the Republican scientific and practical conference dedicated to the 250 th anniversary of Professor S. B. Yundzill (1761 - 1847). Grodno State University, Grodno, pp. 79 - 81. [in Russian]" type="book chapter" year="2012">Leshchenko and Palchevskaya, 2012</bibRefCitation>
(in part): 79.
</paragraph>
</subSubSection>
<subSubSection id="C31BB6D9FFEC184AFF6AF9E991FC3224" pageId="14" pageNumber="487" type="materials_examined">
<paragraph id="8BBEE552FFEC184AFF6AF9E9967633E8" blockId="14.[151,1436,1663,2013]" pageId="14" pageNumber="487">
<materialsCitation id="3B69EF0FFFEC184AFF6AF9E9967633E8" collectingDate="2022-08-24" collectionCode="V" collectorName="Zhidkov &amp; S. A. Dorofeev." country="Belarus" latitude="55.5907" location="Ovsyanka River" longLatPrecision="6" longitude="30.408" municipality="Western" pageId="14" pageNumber="487" specimenCode="ZIN 56916" specimenCount="1" stateProvince="Vitsyebsk" typeStatus="holotype">
<emphasis id="B9753940FFEC184AFF6AF9E9933633CC" bold="true" box="[151,260,1663,1689]" pageId="14" pageNumber="487">
<typeStatus id="54BA5BF0FFEC184AFF6AF9E9933633CC" box="[151,260,1663,1689]" pageId="14" pageNumber="487" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCode id="DBA74D29FFEC184AFEE8F9E993A433CF" box="[277,406,1663,1690]" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="14" pageNumber="487" type="Museum">ZIN 56916</specimenCode>
, SL 68.0 mm,
<location id="8EDEB389FFEC184AFDBCF9E990C833CC" LSID="urn:lsid:plazi:treatment:03A85444FFEC1856FF6AFA05907636D5:8EDEB389FFEC184AFDBCF9E990C833CC" box="[577,762,1663,1689]" country="Belarus" latitude="55.5907" longLatPrecision="6" longitude="30.408" municipality="Western" name="Ovsyanka River" pageId="14" pageNumber="487" stateProvince="Vitsyebsk">Ovsyanka River</location>
, a tributary of the
<collectingMunicipality id="6BDA7F28FFEC184AFC2FF9E9961F33CC" box="[978,1069,1663,1689]" pageId="14" pageNumber="487">Western</collectingMunicipality>
Dvina,
<collectingRegion id="49C52BB0FFEC184AFB77F9E9970033CC" box="[1162,1330,1663,1689]" country="Belarus" name="Vitsyebsk" pageId="14" pageNumber="487">Vitebsk region</collectingRegion>
,
<collectingCountry id="F316A5C2FFEC184AFABDF9E997AA33CC" box="[1344,1432,1663,1689]" name="Belarus" pageId="14" pageNumber="487">Belarus</collectingCountry>
,
<geoCoordinate id="EE358395FFEC184AFF6AF935933F33E8" box="[151,269,1699,1726]" degrees="55.5907" direction="north" orientation="latitude" pageId="14" pageNumber="487" precision="5" value="55.5907">55.5907 N</geoCoordinate>
<geoCoordinate id="EE358395FFEC184AFEE9F93593BB33E8" box="[276,393,1699,1725]" degrees="30.4080" direction="east" orientation="longitude" pageId="14" pageNumber="487" precision="5" value="30.408">30.4080 E</geoCoordinate>
,
<date id="FFBFC392FFEC184AFE69F935906333E8" box="[404,593,1699,1725]" pageId="14" pageNumber="487" value="2022-08-24">
<collectingDate id="EFFB3A7AFFEC184AFE69F935906333E8" box="[404,593,1699,1725]" pageId="14" pageNumber="487" value="2022-08-24">August 24, 2022</collectingDate>
</date>
, collectors Z.
<collectionCode id="ED107D97FFEC184AFD1BF93590CE33E8" box="[742,764,1699,1725]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="487" type="Museum">V</collectionCode>
.
<collectorName id="26F48084FFEC184AFCFBF935915733E8" box="[774,869,1699,1725]" pageId="14" pageNumber="487">Zhidkov</collectorName>
and
<collectorName id="26F48084FFEC184AFC61F935967633E8" box="[924,1092,1699,1725]" pageId="14" pageNumber="487">S.A. Dorofeev.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BBEE552FFEC184AFF3AF95191FC3224" blockId="14.[151,1436,1663,2013]" pageId="14" pageNumber="487">
<materialsCitation id="3B69EF0FFFEC184AFF3AF95190F33250" collectingDate="2022-08-24" collectionCode="ZIN" country="Belarus" latitude="55.5907" location="Western" longLatPrecision="6" longitude="30.408" municipality="Ovsyanka River" pageId="14" pageNumber="487" specimenCode="ZIN 56917" specimenCount="1" stateProvince="Vitsyebsk" typeStatus="paratype">
<emphasis id="B9753940FFEC184AFF3AF951930D33B4" bold="true" box="[199,319,1735,1761]" pageId="14" pageNumber="487">
<typeStatus id="54BA5BF0FFEC184AFF3AF951930D33B4" box="[199,319,1735,1761]" pageId="14" pageNumber="487" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCode id="DBA74D29FFEC184AFEB1F95193F833B7" box="[332,458,1735,1762]" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="14" pageNumber="487" type="Museum">ZIN 56917</specimenCode>
, n = 4,
<quantity id="4CF948B7FFEC184AFDEAF951908933B4" box="[535,699,1735,1762]" metricMagnitude="-2" metricUnit="m" metricValue="4.695" metricValueMax="6.64" metricValueMin="2.75" pageId="14" pageNumber="487" unit="mm" value="46.95" valueMax="66.4" valueMin="27.5">27.5–66.4 mm</quantity>
,
<collectingMunicipality id="6BDA7F28FFEC184AFD38F951914933B4" box="[709,891,1735,1761]" pageId="14" pageNumber="487">Ovsyanka River</collectingMunicipality>
, a tributary of the
<location id="8EDEB389FFEC184AFBBCF95196AE33B4" LSID="urn:lsid:plazi:treatment:03A85444FFEC1856FF6AFA05907636D5:8EDEB389FFEC184AFBBCF95196AE33B4" box="[1089,1180,1735,1761]" country="Belarus" latitude="55.5907" longLatPrecision="6" longitude="30.408" municipality="Ovsyanka River" name="Western" pageId="14" pageNumber="487" stateProvince="Vitsyebsk">Western</location>
Dvina,
<collectingRegion id="49C52BB0FFEC184AFB0EF95197AA33B4" box="[1267,1432,1735,1761]" country="Belarus" name="Vitsyebsk" pageId="14" pageNumber="487">Vitebsk region</collectingRegion>
,
<collectingCountry id="F316A5C2FFEC184AFF6AF97D92DD3250" box="[151,239,1771,1797]" name="Belarus" pageId="14" pageNumber="487">Belarus</collectingCountry>
,
<geoCoordinate id="EE358395FFEC184AFF00F97D93463250" box="[253,372,1771,1798]" degrees="55.5907" direction="north" orientation="latitude" pageId="14" pageNumber="487" precision="5" value="55.5907">55.5907 N</geoCoordinate>
<geoCoordinate id="EE358395FFEC184AFE80F97D93C63250" box="[381,500,1771,1797]" degrees="30.4080" direction="east" orientation="longitude" pageId="14" pageNumber="487" precision="5" value="30.408">30.4080 E</geoCoordinate>
,
<date id="FFBFC392FFEC184AFDFCF97D90F33250" box="[513,705,1771,1797]" pageId="14" pageNumber="487" value="2022-08-24">
<collectingDate id="EFFB3A7AFFEC184AFDFCF97D90F33250" box="[513,705,1771,1797]" pageId="14" pageNumber="487" value="2022-08-24">August 24, 2022</collectingDate>
</date>
</materialsCitation>
, collectors Z.
<materialsCitation id="3B69EF0FFFEC184AFCA7F97D968F3250" box="[858,1213,1771,1797]" collectionCode="V" collectorName="S. A. Dorofeev" country="Belarus" location="Zhidkov" pageId="14" pageNumber="487" specimenCount="1" stateProvince="Vitsyebsk" typeStatus="paratype">
<collectionCode id="ED107D97FFEC184AFCA7F97D91423250" box="[858,880,1771,1797]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="487" type="Museum">V</collectionCode>
.
<location id="8EDEB389FFEC184AFC81F97D91E93250" LSID="urn:lsid:plazi:treatment:03A85444FFEC1856FF6AFA05907636D5:8EDEB389FFEC184AFC81F97D91E93250" box="[892,987,1771,1797]" country="Belarus" name="Zhidkov" pageId="14" pageNumber="487" stateProvince="Vitsyebsk">Zhidkov</location>
and
<collectorName id="26F48084FFEC184AFBE8F97D968F3250" box="[1045,1213,1771,1797]" pageId="14" pageNumber="487">S.A. Dorofeev</collectorName>
</materialsCitation>
;
<materialsCitation id="3B69EF0FFFEC184AFB37F97D90533218" collectingDate="2022-08-22" collectionCode="ZIN" country="Belarus" elevation="5" latitude="55.2527" location="Western" longLatPrecision="6" longitude="30.1431" municipality="Luzhesnyanka River" pageId="14" pageNumber="487" specimenCode="ZIN 56918" specimenCount="1" stateProvince="Vitsyebsk" typeStatus="paratype">
<specimenCode id="DBA74D29FFEC184AFB37F97D97793253" box="[1226,1355,1771,1798]" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="14" pageNumber="487" type="Museum">ZIN 56918</specimenCode>
, n = 2, SL 26.0 and
<quantity id="4CF948B7FFEC184AFED5F89993BD327C" box="[296,399,1807,1834]" metricMagnitude="-2" metricUnit="m" metricValue="5.529999999999999" pageId="14" pageNumber="487" unit="mm" value="55.3">55.3 mm</quantity>
,
<collectingMunicipality id="6BDA7F28FFEC184AFE61F89990B7327C" box="[412,645,1807,1833]" pageId="14" pageNumber="487">Luzhesnyanka River</collectingMunicipality>
, a tributary of the
<location id="8EDEB389FFEC184AFCAAF8999180327C" LSID="urn:lsid:plazi:treatment:03A85444FFEC1856FF6AFA05907636D5:8EDEB389FFEC184AFCAAF8999180327C" box="[855,946,1807,1833]" country="Belarus" latitude="55.2527" longLatPrecision="6" longitude="30.1431" municipality="Luzhesnyanka River" name="Western" pageId="14" pageNumber="487" stateProvince="Vitsyebsk">Western</location>
Dvina,
<collectingCountry id="F316A5C2FFEC184AFBF0F8999657327C" box="[1037,1125,1807,1833]" name="Belarus" pageId="14" pageNumber="487">Belarus</collectingCountry>
,
<collectingRegion id="49C52BB0FFEC184AFB8CF899972B327C" box="[1137,1305,1807,1833]" country="Belarus" name="Vitsyebsk" pageId="14" pageNumber="487">Vitebsk region</collectingRegion>
,
<geoCoordinate id="EE358395FFEC184AFAD8F89997AE327C" box="[1317,1436,1807,1834]" degrees="55.2527" direction="north" orientation="latitude" pageId="14" pageNumber="487" precision="5" value="55.2527">55.2527 N</geoCoordinate>
<geoCoordinate id="EE358395FFEC184AFF6AF8A593393218" box="[151,267,1843,1869]" degrees="30.1431" direction="east" orientation="longitude" pageId="14" pageNumber="487" precision="5" value="30.1431">30.1431 E</geoCoordinate>
, depth
<quantity id="4CF948B7FFEC184AFEA6F8A593AE3218" box="[347,412,1843,1870]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="487" unit="m" value="0.5">
<elevation id="002C0261FFEC184AFEA6F8A593AE3218" box="[347,412,1843,1870]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="487" unit="m" value="0.5">0.5 m</elevation>
</quantity>
,
<date id="FFBFC392FFEC184AFE5BF8A590533218" box="[422,609,1843,1869]" pageId="14" pageNumber="487" value="2022-08-22">
<collectingDate id="EFFB3A7AFFEC184AFE5BF8A590533218" box="[422,609,1843,1869]" pageId="14" pageNumber="487" value="2022-08-22">August 22, 2022</collectingDate>
</date>
</materialsCitation>
, collectors Z.
<materialsCitation id="3B69EF0FFFEC184AFD08F8A5967F3218" box="[757,1101,1843,1869]" collectionCode="V" collectorName="S. A. Dorofeev" country="Belarus" location="Zhidkov" pageId="14" pageNumber="487" specimenCount="1" stateProvince="Vitsyebsk" typeStatus="paratype">
<collectionCode id="ED107D97FFEC184AFD08F8A591393218" box="[757,779,1843,1869]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="487" type="Museum">V</collectionCode>
.
<location id="8EDEB389FFEC184AFCE9F8A591413218" LSID="urn:lsid:plazi:treatment:03A85444FFEC1856FF6AFA05907636D5:8EDEB389FFEC184AFCE9F8A591413218" box="[788,883,1843,1869]" country="Belarus" name="Zhidkov" pageId="14" pageNumber="487" stateProvince="Vitsyebsk">Zhidkov</location>
and
<collectorName id="26F48084FFEC184AFC55F8A5967F3218" box="[936,1101,1843,1869]" pageId="14" pageNumber="487">S.A. Dorofeev</collectorName>
</materialsCitation>
;
<materialsCitation id="3B69EF0FFFEC184AFBAAF8A591FC3224" collectingDate="1886-08" collectionCode="ZIN" collectorName="Radivanovsky." country="Belarus" location="Lake Ostrovito" pageId="14" pageNumber="487" specimenCode="ZIN 7156" specimenCount="1" stateProvince="Vitsyebsk" typeStatus="paratype">
<specimenCode id="DBA74D29FFEC184AFBAAF8A596F5321B" box="[1111,1223,1843,1870]" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="14" pageNumber="487" type="Museum">ZIN 7156</specimenCode>
, n = 2, SL 71.7 and 74.0 mm,
<location id="8EDEB389FFEC184AFEF4F8C193873224" LSID="urn:lsid:plazi:treatment:03A85444FFEC1856FF6AFA05907636D5:8EDEB389FFEC184AFEF4F8C193873224" box="[265,437,1879,1905]" country="Belarus" name="Lake Ostrovito" pageId="14" pageNumber="487" stateProvince="Vitsyebsk">Lake Ostrovito</location>
,
<collectingRegion id="49C52BB0FFEC184AFE42F8C190253224" box="[447,535,1879,1905]" country="Belarus" name="Vitsyebsk" pageId="14" pageNumber="487">Vitebsk</collectingRegion>
,
<date id="FFBFC392FFEC184AFDDCF8C190863224" box="[545,692,1879,1905]" pageId="14" pageNumber="487" value="1886-08">
<collectingDate id="EFFB3A7AFFEC184AFDDCF8C190863224" box="[545,692,1879,1905]" pageId="14" pageNumber="487" value="1886-08">August 1886</collectingDate>
</date>
, collector
<collectorName id="26F48084FFEC184AFCD5F8C191FC3224" box="[808,974,1879,1905]" pageId="14" pageNumber="487">Radivanovsky.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31BB6D9FFEC184BFF3AF8ED9731369C" lastPageId="15" lastPageNumber="488" pageId="14" pageNumber="487" type="diagnosis">
<paragraph id="8BBEE552FFEC184BFF3AF8ED904C3580" blockId="14.[151,1436,1663,2013]" lastBlockId="15.[151,1437,150,969]" lastPageId="15" lastPageNumber="488" pageId="14" pageNumber="487">
<emphasis id="B9753940FFEC184AFF3AF8ED93C832C0" bold="true" box="[199,506,1915,1941]" pageId="14" pageNumber="487">Morphological diagnosis.</emphasis>
Нead pointed anteriorly and considerably flattened in dorso-ventral direction; presence of dermal papillae on top and sides of head; large bony prickles under pectoral fins; long second dorsal fin (length of its base 40% SL); long pelvic fins reaching anus; incomplete trunk canal of seismosensory system with 25–27 pores, its end on vertical of last rays of anal fin; semicircular shape of genital papilla in males; all fins (including pelvic and anal) with dark spots.
</paragraph>
<paragraph id="8BBEE552FFED184BFF3AFF49968F34A0" blockId="15.[151,1437,150,969]" pageId="15" pageNumber="488">
The new species differs from the closely related
<taxonomicName id="4C019ED1FFED184BFCE2FF4991D635AC" box="[799,996,223,249]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="species" species="koshewnikowi">
<emphasis id="B9753940FFED184BFCE2FF4991D635AC" box="[799,996,223,249]" italics="true" pageId="15" pageNumber="488">C. koshewnikowi</emphasis>
</taxonomicName>
in following features: abundance of melanophores on ventral side of body (vs. absence of melanophores); presence of dermal papillae on upper and lateral parts of head (vs. absence of papillae and presence of dermal wrinkles); longer caudal peduncle (14.6±2.1 vs. 8.3±0.5% SL); large eye diameter (23.1±1.1 vs. 19.1±0.6% of head length); wider interorbital space (16.3±3.4 vs. 9.8±0.5% of head length).
<taxonomicName id="4C019ED1FFED184BFE08FEF9908234DC" authorityName="Sideleva &amp; Zhidkov" authorityYear="2024" box="[501,688,367,393]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="species" species="dorofeevi">
<emphasis id="B9753940FFED184BFE08FEF9908234DC" box="[501,688,367,393]" italics="true" pageId="15" pageNumber="488">Cottus dorofeevi</emphasis>
</taxonomicName>
differs from another Eastern European species
<taxonomicName id="4C019ED1FFED184BFB29FEF9974F34DD" authorityName="Sideleva, Naseka &amp; Zhidkov" authorityYear="2015" box="[1236,1405,367,393]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="species" species="gratzianowi">
<emphasis id="B9753940FFED184BFB29FEF9974F34DD" box="[1236,1405,367,393]" italics="true" pageId="15" pageNumber="488">C. gratzianowi</emphasis>
</taxonomicName>
in following characters: absence of branched rays in pectoral fin (vs. frequent presence in
<taxonomicName id="4C019ED1FFED184BFB9AFE05973C34F9" authorityName="Sideleva, Naseka &amp; Zhidkov" authorityYear="2015" box="[1127,1294,403,429]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="species" species="gratzianowi">
<emphasis id="B9753940FFED184BFB9AFE05973C34F9" box="[1127,1294,403,429]" italics="true" pageId="15" pageNumber="488">C. gratzianowi</emphasis>
</taxonomicName>
); absence of gap and presence of three pores in occipital commissure of lateral line system (vs. presence of gap and four pores in
<taxonomicName id="4C019ED1FFED184BFF6AFE4D930C34A1" authorityName="Sideleva, Naseka &amp; Zhidkov" authorityYear="2015" box="[151,318,475,501]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="species" species="gratzianowi">
<emphasis id="B9753940FFED184BFF6AFE4D930C34A1" box="[151,318,475,501]" italics="true" pageId="15" pageNumber="488">C. gratzianowi</emphasis>
</taxonomicName>
); poorly developed lower lobes of mouth (vs. massive lobes in
<taxonomicName id="4C019ED1FFED184BFBFBFE4D969F34A1" authorityName="Sideleva, Naseka &amp; Zhidkov" authorityYear="2015" box="[1030,1197,475,501]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="species" species="gratzianowi">
<emphasis id="B9753940FFED184BFBFBFE4D969F34A1" box="[1030,1197,475,501]" italics="true" pageId="15" pageNumber="488">C. gratzianowi</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BBEE552FFED184BFF3AFE6990133768" blockId="15.[151,1437,150,969]" pageId="15" pageNumber="488">
A more detailed morphological comparison of
<taxonomicName id="4C019ED1FFED184BFD29FE69916C374C" authorityName="Sideleva &amp; Zhidkov" authorityYear="2024" box="[724,862,511,537]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="species" species="dorofeevi">
<emphasis id="B9753940FFED184BFD29FE69916C374C" box="[724,862,511,537]" italics="true" pageId="15" pageNumber="488">C. dorofeevi</emphasis>
</taxonomicName>
with other European species of the genus
<taxonomicName id="4C019ED1FFED184BFAC4FE699745374C" box="[1337,1399,511,537]" genus="Cottu" pageId="15" pageNumber="488" rank="genus">
<emphasis id="B9753940FFED184BFAC4FE699745374C" box="[1337,1399,511,537]" italics="true" pageId="15" pageNumber="488">Cottu</emphasis>
</taxonomicName>
s is provided in the Discussion section.
</paragraph>
<paragraph id="8BBEE552FFED184BFF3AFDD190D337D0" blockId="15.[151,1437,150,969]" pageId="15" pageNumber="488">
<emphasis id="B9753940FFED184BFF3AFDD193F23735" bold="true" box="[199,448,582,609]" pageId="15" pageNumber="488">Molecular diagnosis.</emphasis>
<taxonomicName id="4C019ED1FFED184BFE34FDD190B63734" authorityName="Sideleva &amp; Zhidkov" authorityYear="2024" box="[457,644,583,609]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="species" species="dorofeevi">
<emphasis id="B9753940FFED184BFE34FDD190B63734" box="[457,644,583,609]" italics="true" pageId="15" pageNumber="488">Cottus dorofeevi</emphasis>
</taxonomicName>
differs from closely related
<taxonomicName id="4C019ED1FFED184BFC37FDD196BB3734" box="[970,1161,583,609]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="species" species="koshewnikowi">
<emphasis id="B9753940FFED184BFC37FDD196BB3734" box="[970,1161,583,609]" italics="true" pageId="15" pageNumber="488">C. koshewnikowi</emphasis>
</taxonomicName>
by a combination of 11 nucleotide substitutions (three of them are unique*).
</paragraph>
<paragraph id="8BBEE552FFED184BFF3AFD19976D37FC" blockId="15.[151,1437,150,969]" box="[199,1375,655,681]" pageId="15" pageNumber="488">
Gene of cytochrome
<emphasis id="B9753940FFED184BFE4EFD19938D37FD" box="[435,447,655,680]" italics="true" pageId="15" pageNumber="488">c</emphasis>
oxidase subunit 1 (mtDNA, partial sequence):
<emphasis id="B9753940FFED184BFC2DFD19963237FC" box="[976,1024,655,681]" italics="true" pageId="15" pageNumber="488">COI</emphasis>
223 (
<collectionCode id="ED107D97FFED184BFBBCFD19966637FC" box="[1089,1108,655,681]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="488">T</collectionCode>
→C),
<emphasis id="B9753940FFED184BFB64FD1996FB37FC" box="[1177,1225,655,681]" italics="true" pageId="15" pageNumber="488">COI</emphasis>
430 (G→A).
</paragraph>
<paragraph id="8BBEE552FFED184BFF3AFD2596BC37A4" blockId="15.[151,1437,150,969]" pageId="15" pageNumber="488">
<emphasis id="B9753940FFED184BFF3AFD25935E3799" box="[199,364,691,717]" italics="true" pageId="15" pageNumber="488">Control region</emphasis>
(mtDNA, complete sequence):
<emphasis id="B9753940FFED184BFD2FFD2590C43798" box="[722,758,691,717]" italics="true" pageId="15" pageNumber="488">CR</emphasis>
18 (C→G*),
<emphasis id="B9753940FFED184BFC6CFD2591873798" box="[913,949,691,717]" italics="true" pageId="15" pageNumber="488">CR</emphasis>
31 (A→C*),
<emphasis id="B9753940FFED184BFBACFD2596473798" box="[1105,1141,691,717]" italics="true" pageId="15" pageNumber="488">CR</emphasis>
38 (gap→G),
<emphasis id="B9753940FFED184BFAE5FD25970E3798" box="[1304,1340,691,717]" italics="true" pageId="15" pageNumber="488">CR</emphasis>
37 (
<collectionCode id="ED107D97FFED184BFA93FD2597B33798" box="[1390,1409,691,717]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="488">T</collectionCode>
→ gap),
<emphasis id="B9753940FFED184BFF2AFD4192C937A4" box="[215,251,727,753]" italics="true" pageId="15" pageNumber="488">CR</emphasis>
52 (A→C*),
<emphasis id="B9753940FFED184BFE6BFD41938837A4" box="[406,442,727,753]" italics="true" pageId="15" pageNumber="488">CR</emphasis>
232 (G→A),
<emphasis id="B9753940FFED184BFDAAFD41904937A4" box="[599,635,727,753]" italics="true" pageId="15" pageNumber="488">CR</emphasis>
233 (A→G),
<emphasis id="B9753940FFED184BFCE5FD41910E37A4" box="[792,828,727,753]" italics="true" pageId="15" pageNumber="488">CR</emphasis>
412 (
<collectionCode id="ED107D97FFED184BFC81FD4191BD37A4" box="[892,911,727,753]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="488">T</collectionCode>
→C),
<emphasis id="B9753940FFED184BFC29FD4191CA37A4" box="[980,1016,727,753]" italics="true" pageId="15" pageNumber="488">CR</emphasis>
771 (G→A).
</paragraph>
<paragraph id="8BBEE552FFED184BFF3AFD6D917A3608" blockId="15.[151,1437,150,969]" pageId="15" pageNumber="488">
<taxonomicName id="4C019ED1FFED184BFF3AFD6D93B33640" authorityName="Sideleva &amp; Zhidkov" authorityYear="2024" box="[199,385,763,789]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="species" species="dorofeevi">
<emphasis id="B9753940FFED184BFF3AFD6D93B33640" box="[199,385,763,789]" italics="true" pageId="15" pageNumber="488">Cottus dorofeevi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C019ED1FFED184BFE45FD6D90443640" box="[440,630,763,789]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="species" species="koshewnikowi">
<emphasis id="B9753940FFED184BFE45FD6D90443640" box="[440,630,763,789]" italics="true" pageId="15" pageNumber="488">C. koshewnikowi</emphasis>
</taxonomicName>
have the following seven nucleotide substitutions that distinguish them from other European and Siberian
<taxonomicName id="4C019ED1FFED184BFDE6FC899056366C" box="[539,612,799,825]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="genus">
<emphasis id="B9753940FFED184BFDE6FC899056366C" box="[539,612,799,825]" italics="true" pageId="15" pageNumber="488">Cottus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C019ED1FFED184BFD88FC8990E4366C" baseAuthorityName="UK" baseAuthorityYear="1889" box="[629,726,799,825]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="species" species="gobio">
<emphasis id="B9753940FFED184BFD88FC8990E4366C" box="[629,726,799,825]" italics="true" pageId="15" pageNumber="488">C. gobio</emphasis>
</taxonomicName>
,
<taxonomicName id="4C019ED1FFED184BFD1EFC899186366C" authorityName="Sideleva, Kesminas &amp; Zhidkov" authorityYear="2022" box="[739,948,799,825]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="species" species="cyclophthalmus">
<emphasis id="B9753940FFED184BFD1EFC899186366C" box="[739,948,799,825]" italics="true" pageId="15" pageNumber="488">C. cyclophthalmus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C019ED1FFED184BFC3FFC899745366C" authority="Freyhof, Kottelat &amp; Nolte" authorityName="Freyhof, Kottelat &amp; Nolte" authorityYear="2005" box="[962,1399,799,825]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="species" species="rhenanus">
<emphasis id="B9753940FFED184BFC3FFC899679366C" box="[962,1099,799,825]" italics="true" pageId="15" pageNumber="488">C. rhenanus</emphasis>
Freyhof, Kottelat &amp; Nolte
</taxonomicName>
,
<taxonomicName id="4C019ED1FFED184BFA7EFC8992C53608" authorityName="Kessler" authorityYear="1889" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="species" species="sibiricus">
<emphasis id="B9753940FFED184BFA7EFC8992C53608" italics="true" pageId="15" pageNumber="488">C. sibiricus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C019ED1FFED184BFEFBFCD5935B3609" authorityName="Freyhof, Kottelat &amp; Nolte" authorityYear="2005" box="[262,361,835,861]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="species" species="metae">
<emphasis id="B9753940FFED184BFEFBFCD5935B3609" box="[262,361,835,861]" italics="true" pageId="15" pageNumber="488">C. metae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C019ED1FFED184BFE8AFCD5910E3608" authority="Freyhof, Kottelat &amp; Nolte" authorityName="Freyhof, Kottelat &amp; Nolte" authorityYear="2005" box="[375,828,835,861]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="species" species="perifretum">
<emphasis id="B9753940FFED184BFE8AFCD5903F3608" box="[375,525,835,861]" italics="true" pageId="15" pageNumber="488">C. perifretum</emphasis>
Freyhof, Kottelat &amp; Nolte
</taxonomicName>
).
</paragraph>
<paragraph id="8BBEE552FFED184BFF3AFCF1938C36F0" blockId="15.[151,1437,150,969]" pageId="15" pageNumber="488">
Gene of cytochrome
<emphasis id="B9753940FFED184BFE4DFCF1938E36D5" box="[432,444,871,896]" italics="true" pageId="15" pageNumber="488">c</emphasis>
oxidase subunit 1 (mtDNA, partial sequence):
<emphasis id="B9753940FFED184BFC3AFCF191C536D4" box="[967,1015,871,897]" italics="true" pageId="15" pageNumber="488">COI</emphasis>
91 (
<collectionCode id="ED107D97FFED184BFBD5FCF1960936D4" box="[1064,1083,871,897]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="488">T</collectionCode>
→C),
<emphasis id="B9753940FFED184BFB82FCF1969D36D4" box="[1151,1199,871,897]" italics="true" pageId="15" pageNumber="488">COI</emphasis>
97 (G→A),
<emphasis id="B9753940FFED184BFAC1FCF1975E36D4" box="[1340,1388,871,897]" italics="true" pageId="15" pageNumber="488">COI</emphasis>
454 (C→
<collectionCode id="ED107D97FFED184BFF2DFC1D92D136F0" box="[208,227,907,933]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="488">T</collectionCode>
),
<emphasis id="B9753940FFED184BFF05FC1D931A36F0" box="[248,296,907,933]" italics="true" pageId="15" pageNumber="488">COI</emphasis>
496 (G→A).
</paragraph>
<paragraph id="8BBEE552FFED184BFF3AFC399731369C" blockId="15.[151,1437,150,969]" box="[199,1283,943,969]" pageId="15" pageNumber="488">
<emphasis id="B9753940FFED184BFF3AFC39935E369D" box="[199,364,943,969]" italics="true" pageId="15" pageNumber="488">Control region</emphasis>
(mtDNA, complete sequence):
<emphasis id="B9753940FFED184BFD2CFC3990C7369C" box="[721,757,943,969]" italics="true" pageId="15" pageNumber="488">CR</emphasis>
119 (C→
<collectionCode id="ED107D97FFED184BFC98FC39914A369C" box="[869,888,943,969]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="488">T</collectionCode>
),
<emphasis id="B9753940FFED184BFC71FC399182369C" box="[908,944,943,969]" italics="true" pageId="15" pageNumber="488">CR</emphasis>
268 (
<collectionCode id="ED107D97FFED184BFC0CFC399636369C" box="[1009,1028,943,969]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="488">T</collectionCode>
→C),
<emphasis id="B9753940FFED184BFBB4FC39965F369C" box="[1097,1133,943,969]" italics="true" pageId="15" pageNumber="488">CR</emphasis>
778 (G→A).
</paragraph>
</subSubSection>
<caption id="DF7EB5DAFFED184BFF65F8EF972C32C4" ID-DOI="http://doi.org/10.5281/zenodo.11150423" ID-Zenodo-Dep="11150423" box="[152,1310,1913,1938]" httpUri="https://zenodo.org/record/11150423/files/figure.png" pageId="15" pageNumber="488" startId="15.[152,255,1913,1937]" targetBox="[156,1432,996,1878]" targetPageId="15" targetType="figure">
<paragraph id="8BBEE552FFED184BFF65F8EF972C32C4" blockId="15.[152,1310,1913,1938]" box="[152,1310,1913,1938]" pageId="15" pageNumber="488">
<emphasis id="B9753940FFED184BFF65F8EF932A32C4" bold="true" box="[152,280,1913,1937]" pageId="15" pageNumber="488">FIGURE 9.</emphasis>
Illustration of the holotype of
<taxonomicName id="4C019ED1FFED184BFDB2F8EC90FE32C4" authorityName="Sideleva &amp; Zhidkov" authorityYear="2024" box="[591,716,1913,1937]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="488" phylum="Chordata" rank="species" species="dorofeevi">
<emphasis id="B9753940FFED184BFDB2F8EC90FE32C4" box="[591,716,1913,1937]" italics="true" pageId="15" pageNumber="488">C. dorofeevi</emphasis>
</taxonomicName>
(ZIN 56916): (a) lateral view; (b) dorsal view of the head.
</paragraph>
</caption>
<subSubSection id="C31BB6D9FFF21856FF6AFF01977F37FD" lastPageId="18" lastPageNumber="491" pageId="16" pageNumber="489" type="description">
<paragraph id="8BBEE552FFF21854FF6AFF01931A35E4" blockId="16.[151,296,151,177]" box="[151,296,151,177]" pageId="16" pageNumber="489">
<heading id="D0F6523EFFF21854FF6AFF01931A35E4" bold="true" box="[151,296,151,177]" fontSize="11" level="1" pageId="16" pageNumber="489" reason="1">
<emphasis id="B9753940FFF21854FF6AFF01931A35E4" bold="true" box="[151,296,151,177]" pageId="16" pageNumber="489">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BBEE552FFF21854FF6AFF49977A374C" blockId="16.[151,1437,223,898]" pageId="16" pageNumber="489">
<emphasis id="B9753940FFF21854FF6AFF4993AB35AC" bold="true" box="[151,409,223,249]" pageId="16" pageNumber="489">General morphology.</emphasis>
<taxonomicName id="4C019ED1FFF21854FE59FF76905335AC" authorityName="Sideleva &amp; Zhidkov" authorityYear="2024" box="[420,609,223,250]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="16" pageNumber="489" phylum="Chordata" rank="species" species="dorofeevi">
<emphasis id="B9753940FFF21854FE59FF76905335AC" box="[420,609,223,250]" italics="true" pageId="16" pageNumber="489">Cottus dorofeevi</emphasis>
</taxonomicName>
from the Western Dvina have a large head, 1/3 SL (
<tableCitation id="C683D0E9FFF21854FB2EFF49971935AC" box="[1235,1323,223,249]" captionStart="TABLE 1" captionStartId="5.[152,245,291,315]" captionTargetPageId="5" captionText="TABLE 1. Proportional measurements as percentages of standard length for holotype and paratypes of Cottus sp. “dorofeevi” from the Western Dvina / Daugava River system." httpUri="http://table.plazi.org/id/DF7EB5DAFFE71841FF65FEB591153435" pageId="16" pageNumber="489" tableUuid="DF7EB5DAFFE71841FF65FEB591153435">Table 1</tableCitation>
); pointed anteriorly and considerably flattened dorso-ventrally. On average, height of head at occiput is 1.8 times less than length and width of head. Presence of dermal papillae on top and sides of head. Anterior nostrils as pigmented cylindrical tubules directed laterally; posterior nostrils as pores with raised margins. Eyes small; 21.3–25.4% of head length in
<typeStatus id="54BA5BF0FFF21854FEBEFEE6939C34DF" box="[323,430,368,394]" pageId="16" pageNumber="489" type="paratype">paratypes</typeStatus>
(in
<typeStatus id="54BA5BF0FFF21854FE1DFEF9907734DC" box="[480,581,367,393]" pageId="16" pageNumber="489" type="holotype">holotype</typeStatus>
, 22.5%). Interorbital space narrow; on average, 1.4 times smaller than eye diameter. Upper jaw 10.3–11.5% SL in
<typeStatus id="54BA5BF0FFF21854FDA3FE0290FB34FB" box="[606,713,404,430]" pageId="16" pageNumber="489" type="paratype">paratypes</typeStatus>
(in
<typeStatus id="54BA5BF0FFF21854FD07FE05916D34F8" box="[762,863,403,429]" pageId="16" pageNumber="489" type="holotype">holotype</typeStatus>
, 11.6%); corner of mouth on vertical to middle of eye. Teeth on jaws and vomer conical, numerous, small, uniform in shape and size. Preoperculum with three spines; upper one directed backwards and bent inwards; second spine smaller than upper one, but well visible; all spines hidden under skin. Interbranchial space wide, 26.5–33.9% of head length in
<typeStatus id="54BA5BF0FFF21854FC13FD96966B374F" box="[1006,1113,512,538]" pageId="16" pageNumber="489" type="paratype">paratypes</typeStatus>
(38.4% in
<typeStatus id="54BA5BF0FFF21854FB2BFE69970F374C" box="[1238,1341,511,537]" pageId="16" pageNumber="489" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BBEE552FFF21854FF3AFDB590AC37FC" blockId="16.[151,1437,223,898]" pageId="16" pageNumber="489">Dorsal profile straight; head passes into body smoothly (without hump). Trunk shortened; preanal distance longer than postanal; maximum body height at beginning of first dorsal fin 6–6.5 times in SL. Caudal peduncle short, 11.9–16.7% SL (on average, 14%), 1.7 times as deep as long. Skin with bony prickles under pectoral fins. Semicircular shape of genital papilla in males.</paragraph>
<paragraph id="8BBEE552FFF21854FF3AFD2290C536D4" blockId="16.[151,1437,223,898]" pageId="16" pageNumber="489">
No gap between dorsal fins. First dorsal fin short and low; its base length 14.2–16.6% SL in
<typeStatus id="54BA5BF0FFF21854FAF5FD229741379B" box="[1288,1395,692,718]" pageId="16" pageNumber="489" type="paratype">paratypes</typeStatus>
(in
<typeStatus id="54BA5BF0FFF21854FF6AFD4192CE37A4" box="[151,252,727,753]" pageId="16" pageNumber="489" type="holotype">holotype</typeStatus>
, 17%); length of its longest ray 8.3–11.0% SL in
<typeStatus id="54BA5BF0FFF21854FCC5FD4E919137A7" box="[824,931,728,754]" pageId="16" pageNumber="489" type="paratype">paratypes</typeStatus>
(in
<typeStatus id="54BA5BF0FFF21854FC29FD41960B37A4" box="[980,1081,727,753]" pageId="16" pageNumber="489" type="holotype">holotype</typeStatus>
, 8.6%). Second dorsal fin very long; length of its base 37.1–41% SL in
<typeStatus id="54BA5BF0FFF21854FD97FD6A90E73643" box="[618,725,764,790]" pageId="16" pageNumber="489" type="paratype">paratypes</typeStatus>
(in
<typeStatus id="54BA5BF0FFF21854FCFBFD6D91593640" box="[774,875,763,789]" pageId="16" pageNumber="489" type="holotype">holotype</typeStatus>
, 42.6%). Beginning of anal fin 4% SL from anal opening. Rays in anal fin long, equal to or greater than those in second dorsal fin. Pectoral fins short, 26.1–31.3% SL in
<typeStatus id="54BA5BF0FFF21854FF49FCD2932D360B" box="[180,287,836,862]" pageId="16" pageNumber="489" type="paratype">paratypes</typeStatus>
(in
<typeStatus id="54BA5BF0FFF21854FEB3FCD593813608" box="[334,435,835,861]" pageId="16" pageNumber="489" type="holotype">holotype</typeStatus>
, 28.5%); its end on vertical of second ray of second dorsal fin. Pelvic fins long, reaching anus, 19.7–22% SL in
<typeStatus id="54BA5BF0FFF21854FE68FCFE903236D7" box="[405,512,872,898]" pageId="16" pageNumber="489" type="paratype">paratypes</typeStatus>
(in
<typeStatus id="54BA5BF0FFF21854FDD3FCF190A136D4" box="[558,659,871,897]" pageId="16" pageNumber="489" type="holotype">holotype</typeStatus>
, 20.3%).
</paragraph>
<caption id="DF7EB5DAFFF21854FF65FA71976A30AA" ID-DOI="http://doi.org/10.5281/zenodo.11150425" ID-Zenodo-Dep="11150425" box="[152,1368,1511,1535]" httpUri="https://zenodo.org/record/11150425/files/figure.png" pageId="16" pageNumber="489" startId="16.[152,255,1511,1535]" targetBox="[151,1436,921,1487]" targetPageId="16" targetType="figure">
<paragraph id="8BBEE552FFF21854FF65FA71976A30AA" blockId="16.[152,1368,1511,1535]" box="[152,1368,1511,1535]" pageId="16" pageNumber="489">
<emphasis id="B9753940FFF21854FF65FA71931730AA" bold="true" box="[152,293,1511,1535]" pageId="16" pageNumber="489">FIGURE 10.</emphasis>
The skeleton of
<taxonomicName id="4C019ED1FFF21854FE32FA71909F30AB" authority=", ZIN" authorityName="ZIN" box="[463,685,1511,1535]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="16" pageNumber="489" phylum="Chordata" rank="species" species="dorofeevi">
<emphasis id="B9753940FFF21854FE32FA71904430AA" box="[463,630,1511,1535]" italics="true" pageId="16" pageNumber="489">Cottus dorofeevi</emphasis>
, ZIN
</taxonomicName>
56917, Ovsyanka River, tributary of the Western Dvina / Daugava.
</paragraph>
</caption>
<paragraph id="8BBEE552FFF21854FF3AF9BB965232A2" blockId="16.[151,1437,1581,2040]" pageId="16" pageNumber="489">
<emphasis id="B9753940FFF21854FF3AF9BB935D3312" bold="true" box="[199,367,1581,1607]" pageId="16" pageNumber="489">Axial skeleton</emphasis>
(
<figureCitation id="133AF9D7FFF21854FE7CF9BB93E33312" box="[385,465,1581,1607]" captionStart="FIGURE 10" captionStartId="16.[152,255,1511,1535]" captionTargetBox="[151,1436,921,1487]" captionTargetId="figure-350@16.[151,1436,921,1487]" captionTargetPageId="16" captionText="FIGURE 10. The skeleton of Cottus dorofeevi, ZIN 56917, Ovsyanka River, tributary of the Western Dvina / Daugava." figureDoi="http://doi.org/10.5281/zenodo.11150425" httpUri="https://zenodo.org/record/11150425/files/figure.png" pageId="16" pageNumber="489">Fig. 10</figureCitation>
). In total, 31–34 vertebrae; 10–11 abdominal, 20–23 caudal (in
<typeStatus id="54BA5BF0FFF21854FB69F9BB96CB3312" box="[1172,1273,1581,1607]" pageId="16" pageNumber="489" type="holotype">holotype</typeStatus>
, 11 + 21 = 32). First four abdominal vertebrae shorter than all subsequent ones. First vertebra with well-developed neural process. Last abdominal vertebrae (from eighth–ninth to 10th–11th) with two–four pairs of pleural ribs (in
<typeStatus id="54BA5BF0FFF21854FB0EF9E3976A33DA" box="[1267,1368,1653,1679]" pageId="16" pageNumber="489" type="holotype">holotype</typeStatus>
, three pairs). First proximal dorsal pterygiophore between first and second abdominal vertebrae, supporting first spine of first dorsal fin. One or two interdorsal pterygiophores (not supporting rays of fins) between first and second dorsal fins. Last pterygiophore of second dorsal fin before neural process of seventh preural vertebra, supporting one or two rays (in
<typeStatus id="54BA5BF0FFF21854FEDBF89393B9324A" box="[294,395,1797,1823]" pageId="16" pageNumber="489" type="holotype">holotype</typeStatus>
, two rays) of second dorsal fin. First ray of anal fin supernumerary; its pterygiophore before first haemal process. Last pterygiophore of anal fin before haemal process of ninth preural vertebra, supporting one or two rays (in
<typeStatus id="54BA5BF0FFF21854FEC0F8DB93903232" box="[317,418,1869,1895]" pageId="16" pageNumber="489" type="holotype">holotype</typeStatus>
, two rays) of anal fin. Caudal fin skeleton with single hypural-parhypural complex bone and three epural bones; complex bone with deep posteromedial notch. Hypural-parhypural complex bone supporting main rays of caudal fin. Seven or eight main medial rays of caudal fin branched (in
<typeStatus id="54BA5BF0FFF21854FBD0F80396A032FA" box="[1069,1170,1941,1967]" pageId="16" pageNumber="489" type="holotype">holotype</typeStatus>
, seven). Additional rays of caudal fin in dorsal row supported by epural bones and neural processes of second and third preural vertebrae; additional rays in ventral row supported by haemal process of second preural vertebra.
</paragraph>
<paragraph id="8BBEE552FFF31855FF3AFF0092873414" blockId="17.[151,1436,150,321]" pageId="17" pageNumber="490">
<emphasis id="B9753940FFF31855FF3AFF0093B035E5" bold="true" box="[199,386,150,176]" pageId="17" pageNumber="490">Skeleton of fins</emphasis>
(
<tableCitation id="C683D0E9FFF31855FE6CFF0193DB35E4" box="[401,489,151,177]" captionStart="TABLE 2" captionStartId="6.[152,245,151,175]" captionTargetPageId="6" captionText="TABLE 2. Frequency of occurrence of fin-rays counts for Cottus sp. “dorofeevi” from the Western Dvina / Daugava system, C. koshewnikowi (ZIN 55582), C. microstomus, and C. gobio (UK 1889)." httpUri="http://table.plazi.org/id/DF7EB5DAFFE41842FF65FF0191CC3586" pageId="17" pageNumber="490" tableUuid="DF7EB5DAFFE41842FF65FF0191CC3586">Table 2</tableCitation>
,
<figureCitation id="133AF9D7FFF31855FE0BFF01907B35E4" box="[502,585,151,177]" captionStart="FIGURE 10" captionStartId="16.[152,255,1511,1535]" captionTargetBox="[151,1436,921,1487]" captionTargetId="figure-350@16.[151,1436,921,1487]" captionTargetPageId="16" captionText="FIGURE 10. The skeleton of Cottus dorofeevi, ZIN 56917, Ovsyanka River, tributary of the Western Dvina / Daugava." figureDoi="http://doi.org/10.5281/zenodo.11150425" httpUri="https://zenodo.org/record/11150425/files/figure.png" pageId="17" pageNumber="490">Fig. 10</figureCitation>
).
<taxonomicName id="4C019ED1FFF31855FDA2FF01912B35E4" box="[607,793,151,177]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="17" pageNumber="490" phylum="Chordata" rank="species" species="dorofeevi">
<emphasis id="B9753940FFF31855FDA2FF01912B35E4" box="[607,793,151,177]" italics="true" pageId="17" pageNumber="490">Cottus dorofeevi</emphasis>
</taxonomicName>
has almost similar modal values of fin-ray counts as the closely related European congeners (
<emphasis id="B9753940FFF31855FDC1FF2D91463580" box="[572,884,187,213]" italics="true" pageId="17" pageNumber="490">
<taxonomicName id="4C019ED1FFF31855FDC1FF2D90CC3580" box="[572,766,187,213]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="17" pageNumber="490" phylum="Chordata" rank="species" species="koshewnikowi">C. koshewnikowi</taxonomicName>
,
<taxonomicName id="4C019ED1FFF31855FCF6FF2D91423580" baseAuthorityName="UK" baseAuthorityYear="1889" box="[779,880,187,213]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="17" pageNumber="490" phylum="Chordata" rank="species" species="gobio">C. gobio</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C019ED1FFF31855FC52FF2D966D3581" authorityName="Heckel" authorityYear="1837" box="[943,1119,187,213]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="17" pageNumber="490" phylum="Chordata" rank="species" species="microstomus">
<emphasis id="B9753940FFF31855FC52FF2D966D3581" box="[943,1119,187,213]" italics="true" pageId="17" pageNumber="490">C. microstomus</emphasis>
</taxonomicName>
). The
<typeStatus id="54BA5BF0FFF31855FB51FF2D96EE3580" box="[1196,1244,187,213]" pageId="17" pageNumber="490">type</typeStatus>
specimens of
<taxonomicName id="4C019ED1FFF31855FA7EFF2D933235AC" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="17" pageNumber="490" phylum="Chordata" rank="species" species="dorofeevi">
<emphasis id="B9753940FFF31855FA7EFF2D933235AC" italics="true" pageId="17" pageNumber="490">C. dorofeevi</emphasis>
</taxonomicName>
has 7–8 rays in the first dorsal fin, 16–18 rays in the second dorsal fin, 13–14 rays in the anal fin, 13–15 rays in the pectoral fin. The ranges of the number of fin rays are close to that of the other European
<taxonomicName id="4C019ED1FFF31855FAF9FE95977F3448" box="[1284,1357,259,285]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="17" pageNumber="490" phylum="Chordata" rank="genus">
<emphasis id="B9753940FFF31855FAF9FE95977F3448" box="[1284,1357,259,285]" italics="true" pageId="17" pageNumber="490">Cottus</emphasis>
</taxonomicName>
(
<tableCitation id="C683D0E9FFF31855FAA1FE95929A3414" captionStart="TABLE 2" captionStartId="6.[152,245,151,175]" captionTargetPageId="6" captionText="TABLE 2. Frequency of occurrence of fin-rays counts for Cottus sp. “dorofeevi” from the Western Dvina / Daugava system, C. koshewnikowi (ZIN 55582), C. microstomus, and C. gobio (UK 1889)." httpUri="http://table.plazi.org/id/DF7EB5DAFFE41842FF65FF0191CC3586" pageId="17" pageNumber="490" tableUuid="DF7EB5DAFFE41842FF65FF0191CC3586">Table 2</tableCitation>
).
</paragraph>
<caption id="DF7EB5DAFFF31855FF65F80293AF32A0" ID-DOI="http://doi.org/10.5281/zenodo.11150427" ID-Zenodo-Dep="11150427" httpUri="https://zenodo.org/record/11150427/files/figure.png" pageId="17" pageNumber="490" startId="17.[152,255,1940,1964]" targetBox="[163,1423,357,1904]" targetPageId="17" targetType="figure">
<paragraph id="8BBEE552FFF31855FF65F80293AF32A0" blockId="17.[151,1436,1940,2037]" pageId="17" pageNumber="490">
<emphasis id="B9753940FFF31855FF65F802931032F8" bold="true" box="[152,290,1940,1965]" pageId="17" pageNumber="490">FIGURE 11</emphasis>
. Lateral line system on the head of
<taxonomicName id="4C019ED1FFF31855FD53F803916832F8" box="[686,858,1941,1965]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="17" pageNumber="490" phylum="Chordata" rank="species" species="dorofeevi">
<emphasis id="B9753940FFF31855FD53F803916832F8" box="[686,858,1941,1965]" italics="true" pageId="17" pageNumber="490">Cottus dorofeevi</emphasis>
</taxonomicName>
(ZIN 56916). CSO, supraorabital sensory canal; CIO, infraorbital canal, CT, temporal canal; CPM, praeoperculo-mandibular canal; CLL, trunk (lateral) canal; CMT, temporal (postorbital) commissure.
</paragraph>
</caption>
<paragraph id="8BBEE552FFF01856FF3AFF0090AB374D" blockId="18.[151,1437,150,897]" pageId="18" pageNumber="491">
<emphasis id="B9753940FFF01856FF3AFF00939D35E5" bold="true" box="[199,431,150,176]" pageId="18" pageNumber="491">Lateral line system</emphasis>
(
<figureCitation id="133AF9D7FFF01856FE3CFF00902735E5" box="[449,533,150,176]" captionStart="FIGURE 11" captionStartId="17.[152,255,1940,1964]" captionTargetBox="[163,1423,357,1904]" captionTargetId="figure-101@17.[151,1435,345,1917]" captionTargetPageId="17" captionText="FIGURE 11. Lateral line system on the head of Cottus dorofeevi (ZIN 56916). CSO, supraorabital sensory canal; CIO, infraorbital canal, CT, temporal canal; CPM, praeoperculo-mandibular canal; CLL, trunk (lateral) canal; CMT, temporal (postorbital) commissure." figureDoi="http://doi.org/10.5281/zenodo.11150427" httpUri="https://zenodo.org/record/11150427/files/figure.png" pageId="18" pageNumber="491">Fig. 11</figureCitation>
). Sensory canals arrangement typical for
<taxonomicName id="4C019ED1FFF01856FC08FF01960C35E4" box="[1013,1086,151,177]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="18" pageNumber="491" phylum="Chordata" rank="genus">
<emphasis id="B9753940FFF01856FC08FF01960C35E4" box="[1013,1086,151,177]" italics="true" pageId="18" pageNumber="491">Cottus</emphasis>
</taxonomicName>
. All canals (with exception of preoperculo-mandibular) interconnected in single system. In supraorbital canal, two small pores in front and behind nasale. Supraorbital canals on left and right sides connected by coronal commissure with one small pore in center. Infraorbital canal with nine pores of similar size; seventh pore on long dermal canaliculus. Temporal canal and occipital commissure with three pores. Preoperculo-mandibular canal with 10–11 pores; mandibular parts of canals interconnected in majority (seven) of specimens with one common oval pore on chin. In
<specimenCount id="9D072EDBFFF01856FB4CFEDD97273430" box="[1201,1301,331,357]" count="2" pageId="18" pageNumber="491" type="generic">two type</specimenCount>
specimens, mandibular parts of preoperculo-mandibular canals not connected with two pores on chin. Presence of additional fifth pore in reoperculo-mandibular canal variable; absent in 55% of specimens (including
<typeStatus id="54BA5BF0FFF01856FB61FE04973134F9" box="[1180,1283,402,428]" pageId="18" pageNumber="491" type="holotype">holotype</typeStatus>
). Trunk canal not full, close to medial line of body; ending at vertical of penultimate ray of anal fin. In seven individuals, end of trunk canal with one–three autonomous segments (in
<typeStatus id="54BA5BF0FFF01856FD12FE4C916634A1" box="[751,852,474,500]" pageId="18" pageNumber="491" type="holotype">holotype</typeStatus>
, two segments). Trunk canal with 19-29 small pores (modally, 25-27 pores; in
<typeStatus id="54BA5BF0FFF01856FE44FE68902C374D" box="[441,542,510,536]" pageId="18" pageNumber="491" type="holotype">holotype</typeStatus>
, 27 pores).
</paragraph>
<paragraph id="8BBEE552FFF01856FF3AFDB4977F37FD" blockId="18.[151,1437,150,897]" pageId="18" pageNumber="491">
<emphasis id="B9753940FFF01856FF3AFDB4937A3769" bold="true" box="[199,328,546,572]" pageId="18" pageNumber="491">Coloration</emphasis>
(
<figureCitation id="133AF9D7FFF01856FEA7FDB493933769" box="[346,417,546,572]" captionStart="FIGURE 2" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[158,1429,181,1919]" captionTargetId="figure-29@4.[158,1429,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Cottus sp. from the Western Dvina / Daugava River system, ♁, ZIN 56916, SL 68 mm, TL 79 mm; the Ovsyanka River, tributary of the Western Dvina / Daugava, Belarus, 55.5907° N, 30.4080° E; August 24, 2022; collectors Z.V. Zhidkov and S.A. Dorofeev. (a) lateral view; (b) dorsal view; (c) ventral view." figureDoi="http://doi.org/10.5281/zenodo.11150405" httpUri="https://zenodo.org/record/11150405/files/figure.png" pageId="18" pageNumber="491">Fig. 2</figureCitation>
). Body and head dark gray with black spots. Two dark spots on upper part of body from base of second dorsal fin to medial line. Uniform dark gray coloration of upper and lateral parts of head, including opercular region; accumulations of melanophores on underside of head. On anterior part of body, dark pigment distributed more evenly. Unpigmented stripe on ventral side of body between pelvic fins and anus (
<figureCitation id="133AF9D7FFF01856FB13FD18977237FD" box="[1262,1344,654,680]" captionStart="FIGURE 2" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[158,1429,181,1919]" captionTargetId="figure-29@4.[158,1429,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Cottus sp. from the Western Dvina / Daugava River system, ♁, ZIN 56916, SL 68 mm, TL 79 mm; the Ovsyanka River, tributary of the Western Dvina / Daugava, Belarus, 55.5907° N, 30.4080° E; August 24, 2022; collectors Z.V. Zhidkov and S.A. Dorofeev. (a) lateral view; (b) dorsal view; (c) ventral view." figureDoi="http://doi.org/10.5281/zenodo.11150405" httpUri="https://zenodo.org/record/11150405/files/figure.png" pageId="18" pageNumber="491">Fig. 2c</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31BB6D9FFF01856FF3AFD24914E3641" pageId="18" pageNumber="491" type="etymology">
<paragraph id="8BBEE552FFF01856FF3AFD24914E3641" blockId="18.[151,1437,150,897]" pageId="18" pageNumber="491">
<emphasis id="B9753940FFF01856FF3AFD24937A3799" bold="true" box="[199,328,690,716]" pageId="18" pageNumber="491">Etymology</emphasis>
. The name of new species was given in honor of zoologist Sergei A. Dorofeev (
<collectingRegion id="49C52BB0FFF01856FAFFFD2497AE3799" box="[1282,1436,690,716]" country="Belarus" name="Vitsyebsk" pageId="18" pageNumber="491">Vitebsk State</collectingRegion>
University) for his remarkable contribution to advances in the field. He organized and took part in the collection of
<typeStatus id="54BA5BF0FFF01856FF6AFD6D92F53640" box="[151,199,763,789]" pageId="18" pageNumber="491">type</typeStatus>
specimens in tributaries of the Western Dvina River,
<collectingCountry id="F316A5C2FFF01856FCE2FD6C914A3641" box="[799,888,762,788]" name="Belarus" pageId="18" pageNumber="491">Belarus</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31BB6D9FFF01856FF3AFC88907636D5" pageId="18" pageNumber="491" type="distribution">
<paragraph id="8BBEE552FFF01856FF3AFC88907636D5" blockId="18.[151,1437,150,897]" pageId="18" pageNumber="491">
<emphasis id="B9753940FFF01856FF3AFC88936B366D" bold="true" box="[199,345,798,824]" pageId="18" pageNumber="491">Distribution</emphasis>
. This sculpin is distributed in the Western Dvina / Daugava River system within the territory of
<collectingCountry id="F316A5C2FFF01856FF6AFCD492C23609" box="[151,240,834,860]" name="Belarus" pageId="18" pageNumber="491">Belarus</collectingCountry>
. The
<typeStatus id="54BA5BF0FFF01856FED1FCD5936E3608" box="[300,348,835,861]" pageId="18" pageNumber="491">type</typeStatus>
locality of
<taxonomicName id="4C019ED1FFF01856FE21FCD590573609" box="[476,613,834,861]" family="Cottidae" genus="Cottus" kingdom="Animalia" order="Scorpaeniformes" pageId="18" pageNumber="491" phylum="Chordata" rank="species" species="dorofeevi">
<emphasis id="B9753940FFF01856FE21FCD590573609" box="[476,613,834,861]" italics="true" pageId="18" pageNumber="491">C. dorofeevi</emphasis>
</taxonomicName>
is middle course of the Western Dvina / Daugava River and its tributaries near city of
<collectingRegion id="49C52BB0FFF01856FEE0FCF0934036D5" box="[285,370,870,896]" country="Belarus" name="Vitsyebsk" pageId="18" pageNumber="491">Vitebsk</collectingRegion>
(
<collectingCountry id="F316A5C2FFF01856FE7CFCF093E836D5" box="[385,474,870,896]" name="Belarus" pageId="18" pageNumber="491">Belarus</collectingCountry>
) (
<figureCitation id="133AF9D7FFF01856FE0CFCF0900036D5" box="[497,562,870,896]" captionStart="FIGURE 1" captionStartId="1.[152,255,1947,1971]" captionTargetBox="[153,1436,887,1920]" captionTargetId="figure-252@1.[151,1436,882,1920]" captionTargetPageId="1" captionText="FIGURE 1. The map showing the distribution of Cottus sp. from the Western Dvina / Daugava River system. The numbers indicate sampling sites: 1, Ovsyanka River; 2, Luzhesnyanka River; 3, Lake Ostrovito near Vitebsk (Western Dvina / Daugava River system, Belarus)." figureDoi="http://doi.org/10.5281/zenodo.11150403" httpUri="https://zenodo.org/record/11150403/files/figure.png" pageId="18" pageNumber="491">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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