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<document id="109BCE98C0DEB5DD174281E9F8D58BE3" ID-CLB-Dataset="46493" ID-DOI="10.11646/zootaxa.3764.2.3" ID-GBIF-Dataset="7f3952c3-0f54-40f8-bb3e-a17fb9cb6a49" ID-ISSN="1175-5326" ID-Zenodo-Dep="228666" ID-ZooBank="82E13B7A-C007-4345-B4CF-D4921194DBA2" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460381030042" checkinUser="plazi" docAuthor="Podeniene, Virginija, Naseviciene, Nijole &amp; Podenas, Sigitas" docDate="2014" docId="03A61B45A247FF88DECB60EC9148FEC7" docLanguage="en" docName="zt03764p168.pdf" docOrigin="Zootaxa 3764 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nephrotoma submaculosa Edwards 1928" docType="treatment" docVersion="8" lastPageNumber="165" masterDocId="FF9F633DA24BFF85DE5C631E9231FFD8" masterDocTitle="Notes on the first instar larvae of Ctenophora and Nephrotoma (Diptera, Tipulidae)" masterLastPageNumber="168" masterPageNumber="152" pageNumber="164" updateTime="1698296849864" updateUser="ExternalLinkService">
<mods:mods id="586F99B0DF363A491DE91E671EF62F3C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AE275D3461B16A7B5C7BABFBB6AAE047">
<mods:title id="2F261A4C280C2FC06882E1B37FF3C391">Notes on the first instar larvae of Ctenophora and Nephrotoma (Diptera, Tipulidae)</mods:title>
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<mods:namePart id="B0A2B7F6CDC6E53F24856DB439B333B3">Podeniene, Virginija</mods:namePart>
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<mods:namePart id="B37535A8DE0D638F9EC30C9609AFEA7B">Naseviciene, Nijole</mods:namePart>
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<mods:role id="66EA4DD0538A5622FE2845426F442A4B">
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<mods:namePart id="E17E3716C3B2A3BB4E338AB9DFB21790">Podenas, Sigitas</mods:namePart>
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<mods:title id="D50CAE7E7D1ABEC566CB02B2A9864F6E">Zootaxa</mods:title>
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<mods:part id="6ACA9554C93DEEA3712E3692F7444B01">
<mods:date id="85AE8005F589D03E00F6388192D61EBD">2014</mods:date>
<mods:detail id="841DC8EC765A43CE1D7B1F5772D3C4D2" type="volume">
<mods:number id="BF7A283BF566A08A0FB80FF2D4C14A6C">3764</mods:number>
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<mods:number id="B09D3BD13A139F28071150310EBDCDCF">2</mods:number>
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<mods:classification id="CCCB48D6587D7444BC81E856EF30FEF1">journal article</mods:classification>
<mods:identifier id="BD91D0682F683B9BC7E40373113A786C" type="CLB-Dataset">46493</mods:identifier>
<mods:identifier id="EC41F31E10D6E642B7CD53A2CA78B75F" type="DOI">10.11646/zootaxa.3764.2.3</mods:identifier>
<mods:identifier id="01FBDDF7627B4B25CA98F4903CBC146B" type="GBIF-Dataset">7f3952c3-0f54-40f8-bb3e-a17fb9cb6a49</mods:identifier>
<mods:identifier id="6AE4B3E63326483CDF92A9EF7F6837CC" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="05F7B5AA6263E824B37A9B9FF534424B" type="ZooBank">82E13B7A-C007-4345-B4CF-D4921194DBA2</mods:identifier>
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<treatment id="03A61B45A247FF88DECB60EC9148FEC7" ID-DOI="http://doi.org/10.5281/zenodo.6136184" ID-GBIF-Taxon="119414189" ID-Zenodo-Dep="6136184" LSID="urn:lsid:plazi:treatment:03A61B45A247FF88DECB60EC9148FEC7" httpUri="http://treatment.plazi.org/id/03A61B45A247FF88DECB60EC9148FEC7" lastPageId="13" lastPageNumber="165" pageId="12" pageNumber="164">
<subSubSection id="C315F9D8A247FF88DECB60EC9148FEC7" lastPageId="13" lastPageNumber="165" pageId="12" pageNumber="164" type="nomenclature">
<paragraph id="8BB0AA53A247FF89DECB60EC904CFBD4" blockId="12.[151,637,1010,1071]" box="[151,637,1010,1036]" pageId="12" pageNumber="164">
<heading id="D0F81D3FA247FF89DECB60EC904CFBD4" bold="true" box="[151,637,1010,1036]" fontSize="10" level="2" pageId="12" pageNumber="164" reason="2">
<taxonomicName id="4C0FD1D0A247FF89DECB60EC904CFBD4" ID-CoL="472PM" authority="Edwards, 1928" authorityName="Edwards" authorityYear="1928" box="[151,637,1010,1036]" class="Insecta" family="Tipulidae" genus="Nephrotoma" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="164" phylum="Arthropoda" rank="species" species="submaculosa">
<emphasis id="B97B7641A247FF89DECB60EC904CFBD4" bold="true" box="[151,637,1010,1036]" pageId="12" pageNumber="164">
<emphasis id="B97B7641A247FF89DECB60EC93F4FBD3" bold="true" box="[151,453,1010,1035]" italics="true" pageId="12" pageNumber="164">Nephrotoma submaculosa</emphasis>
<bibRefCitation id="EF9ED7A2A247FF89DF9060ED904CFBD4" author="Edwards" box="[460,637,1011,1036]" pageId="12" pageNumber="164" refString="Edwards, F. W. (1928) The nematocerous Diptera of Corsica. Encyclopedie Entomologique, (B II), Diptera, 4,157 - 189." type="book" year="1928">Edwards, 1928</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BB0AA53A247FF89DECB6708931FFBF7" blockId="12.[151,637,1010,1071]" box="[151,302,1046,1071]" pageId="12" pageNumber="164">
(
<figureCitation id="1334B6D6A247FF89DEC367089314FBF7" box="[159,293,1046,1071]" captionStart="FIGURES 57 – 64" captionStartId="13.[151,264,1337,1360]" captionTargetBox="[160,1421,350,1309]" captionTargetId="figure@13.[151,1436,342,1315]" captionTargetPageId="13" captionText="FIGURES 57 – 64. Nephrotoma submaculosa, first instar larva: 57. Hypostomium. 58. Mandible. 59. Right antenna. 60. General view of labrum. 61. Dorsal abdominal setae (segments II – VII). 62. Ventral abdominal setae (segments II – VII). 63. Lateral abdominal setae (segments II – VII). 64. Spiracular field." httpUri="https://zenodo.org/record/228674/files/figure.png" pageId="12" pageNumber="164">Figs. 57–64</figureCitation>
)
</paragraph>
<paragraph id="8BB0AA53A247FF89DECB6747932AFB4C" blockId="12.[151,1437,1112,2037]" pageId="12" pageNumber="164">
<emphasis id="B97B7641A247FF89DECB6747935EFBA9" box="[151,367,1112,1137]" italics="true" pageId="12" pageNumber="164">First instar larvae.</emphasis>
Length
<quantity id="4CF707B6A247FF89DF9367469046FBA8" box="[463,631,1112,1137]" metricMagnitude="-3" metricUnit="m" metricValue="1.595" metricValueMax="1.6" metricValueMin="1.59" pageId="12" pageNumber="164" unit="mm" value="1.595" valueMax="1.6" valueMin="1.59">1.59–1.60 mm</quantity>
; width
<quantity id="4CF707B6A247FF89DC9067479145FBA8" box="[716,884,1113,1137]" metricMagnitude="-4" metricUnit="m" metricValue="3.15" metricValueMax="3.2" metricValueMin="3.1" pageId="12" pageNumber="164" unit="mm" value="0.315" valueMax="0.32" valueMin="0.31">0.31–0.32 mm</quantity>
. Body is covered by pale pubescence. Cuticle is transparent.
</paragraph>
<paragraph id="8BB0AA53A247FF89DE9B67BE91A4FA49" blockId="12.[151,1437,1112,2037]" pageId="12" pageNumber="164">
<emphasis id="B97B7641A247FF89DE9B67BE935DFB61" box="[199,364,1184,1209]" italics="true" pageId="12" pageNumber="164">Head capsule.</emphasis>
Length
<quantity id="4CF707B6A247FF89DF8F67BF904CFB60" box="[467,637,1185,1209]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="3.8" metricValueMin="3.7" pageId="12" pageNumber="164" unit="mm" value="0.375" valueMax="0.38" valueMin="0.37">0.37–0.38 mm</quantity>
, width
<quantity id="4CF707B6A247FF89DC8667BF91B4FB60" box="[730,901,1185,1209]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.7999999999999998" pageId="12" pageNumber="164" unit="mm" value="0.19" valueMax="0.2" valueMin="0.18">0.18–0.20 mm</quantity>
. In general it is very similar to that of other
<taxonomicName id="4C0FD1D0A247FF89DECB67DA9313FB05" box="[151,290,1220,1245]" class="Insecta" family="Tipulidae" genus="Nephrotoma" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="B97B7641A247FF89DECB67DA9313FB05" box="[151,290,1220,1245]" italics="true" pageId="12" pageNumber="164">Nephrotoma</emphasis>
</taxonomicName>
species. Minor differences were noticed in shape of hypostomium, mandible and antenna. Hypostomium bears five sharp teeth. Three middle teeth are very similar is size and shape; the outermost teeth are much smaller. There is a prominent incision on posterior part of hypostomium (
<figureCitation id="1334B6D6A247FF89DA72661296B5FAFD" box="[1070,1156,1292,1317]" captionStart="FIGURES 57 – 64" captionStartId="13.[151,264,1337,1360]" captionTargetBox="[160,1421,350,1309]" captionTargetId="figure@13.[151,1436,342,1315]" captionTargetPageId="13" captionText="FIGURES 57 – 64. Nephrotoma submaculosa, first instar larva: 57. Hypostomium. 58. Mandible. 59. Right antenna. 60. General view of labrum. 61. Dorsal abdominal setae (segments II – VII). 62. Ventral abdominal setae (segments II – VII). 63. Lateral abdominal setae (segments II – VII). 64. Spiracular field." httpUri="https://zenodo.org/record/228674/files/figure.png" pageId="12" pageNumber="164">Fig. 57</figureCitation>
). Mandible in general is very similar to that of
<taxonomicName id="4C0FD1D0A247FF89DFC9662F901BFA91" box="[405,554,1328,1353]" class="Insecta" family="Tipulidae" genus="Nephrotoma" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="164" phylum="Arthropoda" rank="species" species="flavescens">
<emphasis id="B97B7641A247FF89DFC9662F901BFA91" box="[405,554,1328,1353]" italics="true" pageId="12" pageNumber="164">N. flavescens</emphasis>
</taxonomicName>
, but shapes of teeth are slightly different (
<figureCitation id="1334B6D6A247FF89DA50662E9650FA91" box="[1036,1121,1328,1353]" captionStart="FIGURES 57 – 64" captionStartId="13.[151,264,1337,1360]" captionTargetBox="[160,1421,350,1309]" captionTargetId="figure@13.[151,1436,342,1315]" captionTargetPageId="13" captionText="FIGURES 57 – 64. Nephrotoma submaculosa, first instar larva: 57. Hypostomium. 58. Mandible. 59. Right antenna. 60. General view of labrum. 61. Dorsal abdominal setae (segments II – VII). 62. Ventral abdominal setae (segments II – VII). 63. Lateral abdominal setae (segments II – VII). 64. Spiracular field." httpUri="https://zenodo.org/record/228674/files/figure.png" pageId="12" pageNumber="164">Fig. 58</figureCitation>
). Antenna is very similar to that of
<emphasis id="B97B7641A247FF89DEBA664B9359FAB5" box="[230,360,1364,1389]" italics="true" pageId="12" pageNumber="164">
<taxonomicName id="4C0FD1D0A247FF89DEBA664B9355FAB5" box="[230,356,1364,1389]" class="Insecta" family="Tipulidae" genus="Nephrotoma" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="164" phylum="Arthropoda" rank="species" species="dorsalis">N. dorsalis</taxonomicName>
,
</emphasis>
but shape of basal segment is different (
<figureCitation id="1334B6D6A247FF89DD73664A91B2FAB5" box="[815,899,1364,1389]" captionStart="FIGURES 57 – 64" captionStartId="13.[151,264,1337,1360]" captionTargetBox="[160,1421,350,1309]" captionTargetId="figure@13.[151,1436,342,1315]" captionTargetPageId="13" captionText="FIGURES 57 – 64. Nephrotoma submaculosa, first instar larva: 57. Hypostomium. 58. Mandible. 59. Right antenna. 60. General view of labrum. 61. Dorsal abdominal setae (segments II – VII). 62. Ventral abdominal setae (segments II – VII). 63. Lateral abdominal setae (segments II – VII). 64. Spiracular field." httpUri="https://zenodo.org/record/228674/files/figure.png" pageId="12" pageNumber="164">Fig. 59</figureCitation>
). Labrum in general is very similar to that of
<taxonomicName id="4C0FD1D0A247FF89DBDE664B92C0FA49" class="Insecta" family="Tipulidae" genus="Nephrotoma" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="164" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="B97B7641A247FF89DBDE664B92C0FA49" italics="true" pageId="12" pageNumber="164">N. dorsalis</emphasis>
</taxonomicName>
, but arrangement of sensory structures is different (
<figureCitation id="1334B6D6A247FF89DD68666691B8FA49" box="[820,905,1400,1425]" captionStart="FIGURES 57 – 64" captionStartId="13.[151,264,1337,1360]" captionTargetBox="[160,1421,350,1309]" captionTargetId="figure@13.[151,1436,342,1315]" captionTargetPageId="13" captionText="FIGURES 57 – 64. Nephrotoma submaculosa, first instar larva: 57. Hypostomium. 58. Mandible. 59. Right antenna. 60. General view of labrum. 61. Dorsal abdominal setae (segments II – VII). 62. Ventral abdominal setae (segments II – VII). 63. Lateral abdominal setae (segments II – VII). 64. Spiracular field." httpUri="https://zenodo.org/record/228674/files/figure.png" pageId="12" pageNumber="164">Fig. 60</figureCitation>
).
</paragraph>
<paragraph id="8BB0AA53A247FF89DE9B668291EDFA6D" blockId="12.[151,1437,1112,2037]" box="[199,988,1436,1461]" pageId="12" pageNumber="164">
<emphasis id="B97B7641A247FF89DE9B66829329FA6D" box="[199,280,1436,1461]" italics="true" pageId="12" pageNumber="164">Thorax</emphasis>
. In general is very similar to that of other
<taxonomicName id="4C0FD1D0A247FF89DCAD6682914DFA6D" box="[753,892,1436,1461]" class="Insecta" family="Tipulidae" genus="Nephrotoma" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="B97B7641A247FF89DCAD6682914DFA6D" box="[753,892,1436,1461]" italics="true" pageId="12" pageNumber="164">Nephrotoma</emphasis>
</taxonomicName>
species.
</paragraph>
<paragraph id="8BB0AA53A247FF89DE9B66DE961AF968" blockId="12.[151,1437,1112,2037]" pageId="12" pageNumber="164">
<emphasis id="B97B7641A247FF89DE9B66DE9300FA01" box="[199,305,1472,1497]" italics="true" pageId="12" pageNumber="164">Abdomen</emphasis>
. In general is very similar to that of other
<taxonomicName id="4C0FD1D0A247FF89DD5066DE91A6FA01" box="[780,919,1472,1497]" class="Insecta" family="Tipulidae" genus="Nephrotoma" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="B97B7641A247FF89DD5066DE91A6FA01" box="[780,919,1472,1497]" italics="true" pageId="12" pageNumber="164">Nephrotoma</emphasis>
</taxonomicName>
species. Seta D1 is very short; D2 is long and four times as long as D1 (
<figureCitation id="1334B6D6A247FF89DFEA66FA9038FA25" box="[438,521,1508,1533]" captionStart="FIGURES 57 – 64" captionStartId="13.[151,264,1337,1360]" captionTargetBox="[160,1421,350,1309]" captionTargetId="figure@13.[151,1436,342,1315]" captionTargetPageId="13" captionText="FIGURES 57 – 64. Nephrotoma submaculosa, first instar larva: 57. Hypostomium. 58. Mandible. 59. Right antenna. 60. General view of labrum. 61. Dorsal abdominal setae (segments II – VII). 62. Ventral abdominal setae (segments II – VII). 63. Lateral abdominal setae (segments II – VII). 64. Spiracular field." httpUri="https://zenodo.org/record/228674/files/figure.png" pageId="12" pageNumber="164">Fig. 61</figureCitation>
); D3 is slightly shorter than D2; D4 and D5 are very similar in length and slightly longer than D1; D6 is the shortest seta and twice as short as D1. Setae D2, D3 and D4, D5 make pairs and are very close to each other. Seta
<date id="FFB18C93A247FF89DFF6653393FDF99D" box="[426,460,1581,1605]" pageId="12" pageNumber="164">V1</date>
is long;
<date id="FFB18C93A247FF89DC716533907EF99D" box="[557,591,1581,1605]" pageId="12" pageNumber="164">V2</date>
is slightly longer than
<date id="FFB18C93A247FF89DD1365339140F99D" box="[847,881,1581,1605]" pageId="12" pageNumber="164">V1</date>
and is the longest ventral seta (
<figureCitation id="1334B6D6A247FF89DA8C65329712F99D" box="[1232,1315,1580,1605]" captionStart="FIGURES 57 – 64" captionStartId="13.[151,264,1337,1360]" captionTargetBox="[160,1421,350,1309]" captionTargetId="figure@13.[151,1436,342,1315]" captionTargetPageId="13" captionText="FIGURES 57 – 64. Nephrotoma submaculosa, first instar larva: 57. Hypostomium. 58. Mandible. 59. Right antenna. 60. General view of labrum. 61. Dorsal abdominal setae (segments II – VII). 62. Ventral abdominal setae (segments II – VII). 63. Lateral abdominal setae (segments II – VII). 64. Spiracular field." httpUri="https://zenodo.org/record/228674/files/figure.png" pageId="12" pageNumber="164">Fig. 62</figureCitation>
). Setae
<date id="FFB18C93A247FF89DB26653392DBF9B1" pageId="12" pageNumber="164">V3 and V4</date>
are very similar in length and half the length of
<date id="FFB18C93A247FF89DD4F654F9109F9B1" box="[787,824,1617,1641]" pageId="12" pageNumber="164">V1</date>
;
<date id="FFB18C93A247FF89DD19654F915BF9B1" box="[837,874,1617,1641]" pageId="12" pageNumber="164">V1</date>
,
<date id="FFB18C93A247FF89DD2A654F91C7F9B1" box="[886,1014,1616,1641]" pageId="12" pageNumber="164">V2 and V3</date>
,
<date id="FFB18C93A247FF89DA5F654F9614F9B1" box="[1027,1061,1617,1641]" pageId="12" pageNumber="164">V4</date>
make pairs and are very close to each other. Lateral setae are arranged in a line. Setae L1, L4 are very similar in length and three times as long as L2 (
<figureCitation id="1334B6D6A247FF89DEC3658692C2F969" box="[159,243,1688,1713]" captionStart="FIGURES 57 – 64" captionStartId="13.[151,264,1337,1360]" captionTargetBox="[160,1421,350,1309]" captionTargetId="figure@13.[151,1436,342,1315]" captionTargetPageId="13" captionText="FIGURES 57 – 64. Nephrotoma submaculosa, first instar larva: 57. Hypostomium. 58. Mandible. 59. Right antenna. 60. General view of labrum. 61. Dorsal abdominal setae (segments II – VII). 62. Ventral abdominal setae (segments II – VII). 63. Lateral abdominal setae (segments II – VII). 64. Spiracular field." httpUri="https://zenodo.org/record/228674/files/figure.png" pageId="12" pageNumber="164">Fig. 63</figureCitation>
). Seta L3 is twice as long as L2; L4 is situated away from the other setae.
</paragraph>
<paragraph id="8BB0AA53A247FF89DE9B65A291F2F82D" blockId="12.[151,1437,1112,2037]" pageId="12" pageNumber="164">
<emphasis id="B97B7641A247FF89DE9B65A2934BF90D" box="[199,378,1724,1749]" italics="true" pageId="12" pageNumber="164">Spiracular disc.</emphasis>
Spiracular field is surrounded by four almost equal, flat, elongate and round-tipped lobes (
<figureCitation id="1334B6D6A247FF89DB3265A29286F921" captionStart="FIGURES 57 – 64" captionStartId="13.[151,264,1337,1360]" captionTargetBox="[160,1421,350,1309]" captionTargetId="figure@13.[151,1436,342,1315]" captionTargetPageId="13" captionText="FIGURES 57 – 64. Nephrotoma submaculosa, first instar larva: 57. Hypostomium. 58. Mandible. 59. Right antenna. 60. General view of labrum. 61. Dorsal abdominal setae (segments II – VII). 62. Ventral abdominal setae (segments II – VII). 63. Lateral abdominal setae (segments II – VII). 64. Spiracular field." httpUri="https://zenodo.org/record/228674/files/figure.png" pageId="12" pageNumber="164">Fig. 64</figureCitation>
). Lobes are almost as long as their width at the base. Lateral lobe has two long stout apical bristles. Each bristle is almost as long as one-third of the whole body. Base of the inner bristle extends into an irregular stripe-shaped sclerite, which ends above the spiracle. A tuft of two long bristles and a tuft of three long bristles are situated above that sclerite on dorsal part of the lobe. Above the spiracles there are three tufts, each consisting of three bristles of different length. Ventral lobe has an apical tuft of four long bristles; three bristles, similar in length, are situated on inner margin; a tuft of three short bristles is on outer margin; a tuft of two short bristles close to it; a tuft of four long bristles is at the base. The lobe bears very pale oval-shaped sclerite. Spiracles are large, oval and close to each other. Distance between them is the same as diameter of a spiracle itself.
</paragraph>
<paragraph id="8BB0AA53A246FF88DE9B638992CDFF0D" blockId="13.[151,1436,151,288]" pageId="13" pageNumber="165">
<emphasis id="B97B7641A246FF88DE9B63899370FF68" box="[199,321,151,176]" italics="true" pageId="13" pageNumber="165">Anal field.</emphasis>
Anus is surrounded by four short, white and fleshy anal papillae. Few long setae are on anal segment.
</paragraph>
<paragraph id="8BB0AA53A246FF88DE9B63FE9148FEC7" blockId="13.[151,1436,151,288]" pageId="13" pageNumber="165">
<emphasis id="B97B7641A246FF88DE9B63FE9389FF20" box="[199,440,223,248]" italics="true" pageId="13" pageNumber="165">Specimens examined.</emphasis>
15 egg-larvae from female captured near Lazdijai,
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,
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,
<date id="FFB18C93A246FF88DB4863FE97A9FF20" box="[1300,1432,224,248]" pageId="13" pageNumber="165" value="2011-06-23">23 VI 2011</date>
, eggs were laid on
<date id="FFB18C93A246FF88DF3F621893D5FEC7" box="[355,484,262,287]" pageId="13" pageNumber="165" value="2011-06-25">25 VI 2011</date>
, egg-larvae hatched on 0
<date id="FFB18C93A246FF88DCA762199147FEC7" box="[763,886,263,287]" pageId="13" pageNumber="165" value="2011-07-02">2 VII 2011</date>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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