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<document id="9C97FEC5B2E33ABCF7953907D5B2C052" ID-CLB-Dataset="32767" ID-DOI="10.11646/zootaxa.4286.1.1" ID-GBIF-Dataset="a614f1d9-bda9-4df0-a486-aef5af7813fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="828206" ID-ZooBank="6A22EB50-0146-465A-8E2A-523E2F8FE470" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500108241184" checkinUser="plazi" docAuthor="Galvis, William" docDate="2017" docId="0386A71957389060FF21FE30FB09F936" docLanguage="en" docName="zootaxa.4286.1.1.pdf" docOrigin="Zootaxa 4286 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphidraus sikuani Galvis, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="17" masterDocId="FFBFDF6157289070FFB6FFCFFF81FFE8" masterDocTitle="Nineteen new species of Amphidraus Simon, 1900 (Salticidae: Euophryini) from Colombia, with comments about their conservation" masterLastPageNumber="40" masterPageNumber="1" pageNumber="17" updateTime="1698465056562" updateUser="ExternalLinkService">
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<mods:title id="AB76E80EEB277053B24A87388C0DCFBC">Nineteen new species of Amphidraus Simon, 1900 (Salticidae: Euophryini) from Colombia, with comments about their conservation</mods:title>
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<mods:namePart id="50260CA695FC82BA0FC3CDE1A9542FF7">Galvis, William</mods:namePart>
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<mods:title id="BF68EEC8C7104D499AE24FF41A3D395F">Zootaxa</mods:title>
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<mods:part id="84BC1D96729FB774DDFDC0C474EF9AF5">
<mods:date id="46A03DB743A0C03AD16A739D299FE134">2017</mods:date>
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<treatment id="0386A71957389060FF21FE30FB09F936" ID-DOI="http://doi.org/10.5281/zenodo.6045654" ID-GBIF-Taxon="132548503" ID-Zenodo-Dep="6045654" LSID="urn:lsid:plazi:treatment:0386A71957389060FF21FE30FB09F936" httpUri="http://treatment.plazi.org/id/0386A71957389060FF21FE30FB09F936" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection id="C335458457389060FF21FE30FD02FDD3" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8B90160F57389060FF21FE30FE72FDF2" blockId="16.[151,643,511,571]" box="[151,499,511,538]" pageId="16" pageNumber="17">
<heading id="D0D8A16357389060FF21FE30FE72FDF2" bold="true" box="[151,499,511,538]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B95BCA1D57389060FF21FE30FE72FDF2" bold="true" box="[151,499,511,538]" pageId="16" pageNumber="17">
<taxonomicName id="4C2F6D8C57389060FF21FE30FE0FFDF1" ID-CoL="D2GT" box="[151,398,511,537]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="sikuani" status="sp. nov.">
<emphasis id="B95BCA1D57389060FF21FE30FE0FFDF1" bold="true" box="[151,398,511,537]" italics="true" pageId="16" pageNumber="17">Amphidraus sikuani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268776657389060FE23FDCFFE72FDF2" box="[405,499,512,538]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B90160F57389060FF21FDEDFD02FDD3" blockId="16.[151,643,511,571]" box="[151,643,546,571]" pageId="16" pageNumber="17">
<figureCitation id="13140A8A57389060FF21FDEDFE94FDD3" box="[151,277,546,571]" captionStart="FIGURES 10" captionStartId="17.[151,264,1888,1910]" captionTargetBox="[175,1413,209,1862]" captionTargetId="figure@17.[163,1427,193,1871]" captionTargetPageId="17" captionText="FIGURES 10 a – f. A. sikuani sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. A. somondoco sp. nov., e) epigyne, cleared, ventral view; f) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, PED = process on emblic disc, ED = embolic disc, EF = embolic filament, EL = embolic lamella, fd = fertilization duct, po = epigynal pocket, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, sp = spermatheca, TL = tegular lobe. Scale bars: 0.25 mm (a – b, e – f), 0.20 mm (c – d)." httpUri="https://zenodo.org/record/828226/files/figure.png" pageId="16" pageNumber="17">Figs 10a–d</figureCitation>
,
<figureCitation id="13140A8A57389060FE96FDEDFEE2FDD3" box="[288,355,546,571]" captionStart="FIGURES 17" captionStartId="28.[151,264,1660,1682]" captionTargetBox="[340,1244,413,1635]" captionTargetId="figure@28.[340,1247,411,1639]" captionTargetPageId="28" captionText="FIGURES 17 a – i. A. guatavita sp. nov., a) male holotype (ANDES – IN 3155). A. pae sp. nov., b) male holotype (ICN – Ar 6576). A. quimbaya sp. nov., c) male holotype (ICN – Ar 7746). A. quinini sp. nov., d) male holotype (ICN – Ar 8266); e) female paratype (ICN – Ar 8267). A. sie sp. nov., f) male holotype (ICN – Ar 8254); g) female paratype (ICN – Ar 8255). A. sikuani sp. nov., h) male holotype (ICN – Ar 6558); i) female paratype (ICN – Ar 7185). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828240/files/figure.png" pageId="16" pageNumber="17">17h–i</figureCitation>
,
<figureCitation id="13140A8A57389060FED8FDEDFE34FDD3" box="[366,437,546,571]" captionStart="FIGURES 22" captionStartId="33.[151,264,1900,1922]" captionTargetBox="[195,1386,193,1874]" captionTargetId="figure@33.[162,1426,193,1883]" captionTargetPageId="33" captionText="FIGURES 22 a – k. Amphidraus spp, left male palp. A. sikuani sp. nov., a) ventral view; b) same, retrolateral view. A. sotairensis sp. nov., c) ventral view; d) same, retrolateral view. A. sua sp. nov., e) ventral view; f) same, retrolateral view. A. tisquesusa sp. nov., g) ventral view; h) same, retrolateral view. A. tanimuca sp. nov., i) ventral view of the bulb and cymbium; j) palp without cymbium to show the RTA, retrolateral view; k) same, prolateral view; l) same, ventral view." httpUri="https://zenodo.org/record/828250/files/figure.png" pageId="16" pageNumber="17">22a–b</figureCitation>
,
<figureCitation id="13140A8A57389060FE77FDEDFDA5FDD3" box="[449,548,546,571]" captionStart="FIGURES 25" captionStartId="36.[151,264,1889,1911]" captionTargetBox="[185,1412,194,1864]" captionTargetId="figure@36.[151,1436,193,1867]" captionTargetPageId="36" captionText="FIGURES 25 a – l. Amphidraus spp, epigyne. A. sikuani sp. nov., a) ventral view; g) disposition of the internal structures. A. somondoco sp. nov., b) ventral view; h) disposition of the internal structures. A. sua sp. nov., c) ventral view; i) disposition of the internal structures. A. tundama sp. nov., d) ventral view; j) disposition of the internal structures. A. zaque sp. nov., e) ventral view; k) disposition of the internal structures. A. zipa sp. nov., f) ventral view; l) disposition of the internal structures." httpUri="https://zenodo.org/record/828256/files/figure.png" pageId="16" pageNumber="17">
25a,
<quantity id="4CD7BBEA57389060FE41FDEDFDA5FDD3" box="[503,548,546,571]" metricMagnitude="-2" metricUnit="kg" metricValue="2.5" pageId="16" pageNumber="17" unit="g" value="25.0">25g</quantity>
</figureCitation>
,
<figureCitation id="13140A8A57389060FD99FDEDFDDDFDD3" box="[559,604,546,571]" captionStart="FIGURES 26" captionStartId="37.[151,264,1181,1203]" captionTargetBox="[151,1436,193,1159]" captionTargetId="figure@37.[151,1436,193,1159]" captionTargetPageId="37" captionText="FIGURES 26 a – d. Different altitudinal ecosystems where some of the species of Amphidraus here described were found in Colombia. a) Páramo ecosystem, at Las Lagunas Encantadas Natural Reserve, Duitama, Colombia, habitat of A. tundama sp. nov. (~ 3800 m); b) Andean wet forest at Cueva de los Guácharos National Natural Park, Huila, Colombia, habitat of A. guaitipan sp. nov. (~ 2000 m); c) Andean Oak forest (Quercus humboldtii) at Quinini Natural Reserve, Cundinamarca, Colombia, habitat of A. quinini sp. nov. (~ 1700 m); d) Secondary lowland semi wet forest on the outskirts of the city of Villavicencio, Meta, Colombia, habitat of A. sikuani sp. nov. (~ 500 m)." httpUri="https://zenodo.org/record/828258/files/figure.png" pageId="16" pageNumber="17">26d</figureCitation>
,
<figureCitation id="13140A8A57389060FDD1FDECFD02FDD3" box="[615,643,547,571]" captionStart="FIGURE 27" captionStartId="38.[151,250,1317,1339]" captionTargetBox="[138,1418,182,1273]" captionTargetId="figure@38.[137,1422,179,1282]" captionTargetPageId="38" captionText="FIGURE 27. Distribution records of the species of the genus Amphidraus Simon from Colombia treated here. Symbols with a dot inside represent species that have distributions in the Cundiboyacense high-Andean plateau, in the Boyacá and Cundinamarca departments." httpUri="https://zenodo.org/record/828260/files/figure.png" pageId="16" pageNumber="17">27</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C335458457389060FF21FDA5FCF1FD07" pageId="16" pageNumber="17" type="description">
<paragraph id="8B90160F57389060FF21FDA5FCF1FD07" blockId="16.[151,1437,618,1759]" pageId="16" pageNumber="17">
<materialsCitation id="3B471C5257389060FF21FDA5FC4EFD4F" ID-GBIF-Occurrence="1572324840" collectingDate="2005-04-16" collectionCode="ICN" collectorName="Vereda La Vanguardia" country="Colombia" county="Villavicencio" elevation="375" latitude="4.16432" location="Pozo Azul" longLatPrecision="1" longitude="-73.625786" municipality="Vereda La Vanguardia" pageId="16" pageNumber="17" specimenCount="2" stateProvince="Meta" typeStatus="holotype">
<emphasis id="B95BCA1D57389060FF21FDA5FEEBFD6B" bold="true" box="[151,362,618,643]" pageId="16" pageNumber="17">
<typeStatus id="5494A8AD57389060FF21FDA5FF5EFD6B" box="[151,223,618,643]" pageId="16" pageNumber="17">Types</typeStatus>
.
<typeStatus id="5494A8AD57389060FF42FDA5FEE4FD6B" box="[244,357,618,643]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
:
</emphasis>
male from
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,
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<collectingMunicipality id="6BF48C7557389060FD17FDA5FC31FD6B" box="[673,944,618,643]" pageId="16" pageNumber="17">Vereda La Vanguardia</collectingMunicipality>
</collectorName>
,
<collectingCounty id="62F16E8357389060FC70FDA5FBDFFD6B" box="[966,1118,618,643]" pageId="16" pageNumber="17">Villavicencio</collectingCounty>
,
<collectingRegion id="49EBD8ED57389060FBC5FDA4FB31FD6A" box="[1139,1200,619,642]" country="Colombia" name="Meta" pageId="16" pageNumber="17">Meta</collectingRegion>
,
<collectingCountry id="F338569F57389060FB73FDA5FAB7FD6B" box="[1221,1334,618,643]" name="Colombia" pageId="16" pageNumber="17">Colombia</collectingCountry>
,
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[
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,
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],
<collectingDate id="EFD5C92757389060FE52FD41FDE3FD4F" box="[484,610,654,679]" pageId="16" pageNumber="17" value="2005-04-16">
<date id="FF9130CF57389060FE52FD41FDE3FD4F" box="[484,610,654,679]" pageId="16" pageNumber="17" value="2005-04-16">16.IV.2005</date>
</collectingDate>
, unknown coll. (
<collectionCode id="ED3E8ECA57389060FC95FD40FCD5FD4F" box="[803,852,655,679]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="16" pageNumber="17">ICN</collectionCode>
–Ar 6558)
</materialsCitation>
.
<materialsCitation id="3B471C5257389060FC6AFD41FCF1FD07" ID-GBIF-Occurrence="1572324841" collectionCode="ICN" collectorName="E. Florez" country="Colombia" county="Villavicencio" elevation="453" latitude="4.084531" location="Villa Lole" longLatPrecision="1" longitude="-73.56651" municipality="Vereda Apiay" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Meta" typeStatus="paratype">
<emphasis id="B95BCA1D57389060FC6AFD41FBD3FD4F" bold="true" box="[988,1106,654,679]" pageId="16" pageNumber="17">
<typeStatus id="5494A8AD57389060FC6AFD41FBCCFD4F" box="[988,1101,654,679]" pageId="16" pageNumber="17" type="paratype">Paratype</typeStatus>
:
</emphasis>
one female from
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,
<collectingMunicipality id="6BF48C7557389060FF21FD7DFEB4FD23" box="[151,309,690,715]" pageId="16" pageNumber="17">Vereda Apiay</collectingMunicipality>
,
<collectingCounty id="62F16E8357389060FEF5FD7DFE5AFD23" box="[323,475,690,715]" pageId="16" pageNumber="17">Villavicencio</collectingCounty>
,
<collectingRegion id="49EBD8ED57389060FE5FFD7CFDA7FD22" box="[489,550,691,714]" country="Colombia" name="Meta" pageId="16" pageNumber="17">Meta</collectingRegion>
,
<collectingCountry id="F338569F57389060FD82FD7DFD24FD23" box="[564,677,690,715]" name="Colombia" pageId="16" pageNumber="17">Colombia</collectingCountry>
,
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</elevation>
[
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,
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],
<collectorName id="26DA73D957389060FBD4FD7CFB4DFD23" box="[1122,1228,690,715]" pageId="16" pageNumber="17">E. Flórez</collectorName>
&amp; Estudiantes de Aracnología Universidad Nacional de
<collectingCountry id="F338569F57389060FDF0FD19FD35FD07" box="[582,692,726,751]" name="Colombia" pageId="16" pageNumber="17">Colombia</collectingCountry>
(
<collectionCode id="ED3E8ECA57389060FD74FD18FD72FD07" box="[706,755,727,751]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="16" pageNumber="17">ICN</collectionCode>
–Ar 7185).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C335458457389060FF71FD35FC4EFCDF" pageId="16" pageNumber="17" type="etymology">
<paragraph id="8B90160F57389060FF71FD35FC4EFCDF" blockId="16.[151,1437,618,1759]" pageId="16" pageNumber="17">
<emphasis id="B95BCA1D57389060FF71FD35FECFFCFB" bold="true" box="[199,334,762,787]" pageId="16" pageNumber="17">Etymology.</emphasis>
The epithet is a noun in apposition that honors the
<taxonomicName id="4C2F6D8C57389060FC22FD35FC6DFCFB" box="[916,1004,762,787]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="38" phylum="Arthropoda" rank="species" species="sikuani" status="sp. nov.">Sikuani</taxonomicName>
, or Guahibo, indigenous people of the Guajiboan language family, which inhabits the region of the
<typeStatus id="5494A8AD57389060FC88FCD0FCEFFCDF" box="[830,878,799,823]" pageId="16" pageNumber="17">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C335458457389060FF71FC8DFDBBFBDB" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8B90160F57389060FF71FC8DFDBBFBDB" blockId="16.[151,1437,618,1759]" pageId="16" pageNumber="17">
<emphasis id="B95BCA1D57389060FF71FC8DFEC0FCB3" bold="true" box="[199,321,834,859]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
Males of
<taxonomicName id="4C2F6D8C57389060FE02FC8CFDA4FCB3" box="[436,549,834,859]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="sikuani" status="sp. nov.">
<emphasis id="B95BCA1D57389060FE02FC8CFDA4FCB3" box="[436,549,834,859]" italics="true" pageId="16" pageNumber="17">A. sikuani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268776657389060FD9AFC8DFD07FCB3" box="[556,646,834,859]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B95BCA1D57389060FD9AFC8DFD07FCB3" bold="true" box="[556,646,834,859]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
are similar to those of
<taxonomicName id="4C2F6D8C57389060FC3BFC8CFB99FCB3" box="[909,1048,835,859]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="guaitipan" status="sp. nov.">
<emphasis id="B95BCA1D57389060FC3BFC8CFB99FCB3" box="[909,1048,835,859]" italics="true" pageId="16" pageNumber="17">A. guaitipan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268776657389060FB96FC8DFBFBFCB3" box="[1056,1146,834,859]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B95BCA1D57389060FB96FC8DFBFBFCB3" bold="true" box="[1056,1146,834,859]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C2F6D8C57389060FB04FC8CFABBFCB3" box="[1202,1338,835,859]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tanimuca" status="sp. nov.">
<emphasis id="B95BCA1D57389060FB04FC8CFABBFCB3" box="[1202,1338,835,859]" italics="true" pageId="16" pageNumber="17">A. tanimuca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268776657389060FAF4FC8DFA1DFCB3" box="[1346,1436,834,859]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B95BCA1D57389060FAF4FC8DFA1DFCB3" bold="true" box="[1346,1436,834,859]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
by the shape of the RTA. However, they can be recognized from those by the presence of a single bifurcated RTA, with the ventral and the dorsal lobes with the same proportion and shape (
<figureCitation id="13140A8A57389060FBB0FC45FB0BFC4B" box="[1030,1162,906,931]" captionStart="FIGURES 10" captionStartId="17.[151,264,1888,1910]" captionTargetBox="[175,1413,209,1862]" captionTargetId="figure@17.[163,1427,193,1871]" captionTargetPageId="17" captionText="FIGURES 10 a – f. A. sikuani sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. A. somondoco sp. nov., e) epigyne, cleared, ventral view; f) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, PED = process on emblic disc, ED = embolic disc, EF = embolic filament, EL = embolic lamella, fd = fertilization duct, po = epigynal pocket, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, sp = spermatheca, TL = tegular lobe. Scale bars: 0.25 mm (a – b, e – f), 0.20 mm (c – d)." httpUri="https://zenodo.org/record/828226/files/figure.png" pageId="16" pageNumber="17">Figs 10a–b</figureCitation>
,
<figureCitation id="13140A8A57389060FB2FFC45FB62FC4B" box="[1177,1251,906,931]" captionStart="FIGURES 22" captionStartId="33.[151,264,1900,1922]" captionTargetBox="[195,1386,193,1874]" captionTargetId="figure@33.[162,1426,193,1883]" captionTargetPageId="33" captionText="FIGURES 22 a – k. Amphidraus spp, left male palp. A. sikuani sp. nov., a) ventral view; b) same, retrolateral view. A. sotairensis sp. nov., c) ventral view; d) same, retrolateral view. A. sua sp. nov., e) ventral view; f) same, retrolateral view. A. tisquesusa sp. nov., g) ventral view; h) same, retrolateral view. A. tanimuca sp. nov., i) ventral view of the bulb and cymbium; j) palp without cymbium to show the RTA, retrolateral view; k) same, prolateral view; l) same, ventral view." httpUri="https://zenodo.org/record/828250/files/figure.png" pageId="16" pageNumber="17">22a–b</figureCitation>
). Females of
<taxonomicName id="4C2F6D8C57389060FA32FC44FF69FC2F" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="sikuani" status="sp. nov.">
<emphasis id="B95BCA1D57389060FA32FC44FF69FC2F" italics="true" pageId="16" pageNumber="17">A. sikuani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268776657389060FF44FC61FECFFC2F" box="[242,334,942,967]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B95BCA1D57389060FF44FC61FECFFC2F" bold="true" box="[242,334,942,967]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
are similar to those of
<taxonomicName id="4C2F6D8C57389060FDD4FC60FD5CFC2F" box="[610,733,942,967]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bochica" status="sp. nov.">
<emphasis id="B95BCA1D57389060FDD4FC60FD5CFC2F" box="[610,733,942,967]" italics="true" pageId="16" pageNumber="17">A. bochica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268776657389060FD50FC61FCC2FC2F" box="[742,835,942,967]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B95BCA1D57389060FD50FC61FCC2FC2F" bold="true" box="[742,835,942,967]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C2F6D8C57389060FCC9FC60FC51FC2F" box="[895,976,942,967]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="chie" status="sp. nov.">
<emphasis id="B95BCA1D57389060FCC9FC60FC51FC2F" box="[895,976,942,967]" italics="true" pageId="16" pageNumber="17">A. chie</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A268776657389060FC6CFC61FBB7FC2F" box="[986,1078,942,967]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B95BCA1D57389060FC6CFC61FBB7FC2F" bold="true" box="[986,1078,942,967]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
by their general disposition of internal structures of the epigyne, but can be distinguished from those by their longer copulatory ducts (cd), the presence of a single and medial pocket (po) on the posteriorly extended epigynal plate, and a posterior slender spermathecae (
<figureCitation id="13140A8A57389060FE88FBD5FE3FFBDB" box="[318,446,1050,1075]" captionStart="FIGURES 10" captionStartId="17.[151,264,1888,1910]" captionTargetBox="[175,1413,209,1862]" captionTargetId="figure@17.[163,1427,193,1871]" captionTargetPageId="17" captionText="FIGURES 10 a – f. A. sikuani sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. A. somondoco sp. nov., e) epigyne, cleared, ventral view; f) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, PED = process on emblic disc, ED = embolic disc, EF = embolic filament, EL = embolic lamella, fd = fertilization duct, po = epigynal pocket, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, sp = spermatheca, TL = tegular lobe. Scale bars: 0.25 mm (a – b, e – f), 0.20 mm (c – d)." httpUri="https://zenodo.org/record/828226/files/figure.png" pageId="16" pageNumber="17">Figs 10c–d</figureCitation>
,
<figureCitation id="13140A8A57389060FE7FFBD5FDAFFBDB" box="[457,558,1050,1075]" captionStart="FIGURES 25" captionStartId="36.[151,264,1889,1911]" captionTargetBox="[185,1412,194,1864]" captionTargetId="figure@36.[151,1436,193,1867]" captionTargetPageId="36" captionText="FIGURES 25 a – l. Amphidraus spp, epigyne. A. sikuani sp. nov., a) ventral view; g) disposition of the internal structures. A. somondoco sp. nov., b) ventral view; h) disposition of the internal structures. A. sua sp. nov., c) ventral view; i) disposition of the internal structures. A. tundama sp. nov., d) ventral view; j) disposition of the internal structures. A. zaque sp. nov., e) ventral view; k) disposition of the internal structures. A. zipa sp. nov., f) ventral view; l) disposition of the internal structures." httpUri="https://zenodo.org/record/828256/files/figure.png" pageId="16" pageNumber="17">
25a,
<quantity id="4CD7BBEA57389060FDB6FBD5FDAFFBDB" box="[512,558,1050,1075]" metricMagnitude="-2" metricUnit="kg" metricValue="2.5" pageId="16" pageNumber="17" unit="g" value="25.0">25g</quantity>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C335458457389060FF71FBF1FB81F99B" pageId="16" pageNumber="17" type="description">
<paragraph id="8B90160F57389060FF71FBF1FB16FABB" blockId="16.[151,1437,618,1759]" pageId="16" pageNumber="17">
<emphasis id="B95BCA1D57389060FF71FBF1FDA3FBBF" bold="true" box="[199,546,1086,1111]" pageId="16" pageNumber="17">
Description. Male (
<typeStatus id="5494A8AD57389060FE1CFBF1FD94FBBF" box="[426,533,1086,1111]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
).
</emphasis>
Total length: 2.43. Carapace yellow, 1.45 long, 1.01 wide, 0.78 high (
<figureCitation id="13140A8A57389060FA98FBF1FA11FBBF" box="[1326,1424,1086,1111]" captionStart="FIGURES 17" captionStartId="28.[151,264,1660,1682]" captionTargetBox="[340,1244,413,1635]" captionTargetId="figure@28.[340,1247,411,1639]" captionTargetPageId="28" captionText="FIGURES 17 a – i. A. guatavita sp. nov., a) male holotype (ANDES – IN 3155). A. pae sp. nov., b) male holotype (ICN – Ar 6576). A. quimbaya sp. nov., c) male holotype (ICN – Ar 7746). A. quinini sp. nov., d) male holotype (ICN – Ar 8266); e) female paratype (ICN – Ar 8267). A. sie sp. nov., f) male holotype (ICN – Ar 8254); g) female paratype (ICN – Ar 8255). A. sikuani sp. nov., h) male holotype (ICN – Ar 6558); i) female paratype (ICN – Ar 7185). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828240/files/figure.png" pageId="16" pageNumber="17">Fig. 17h</figureCitation>
). OC black, 0.61 long. Anterior eye row 1.14 wide and posterior 0.91 wide. Sternum yellow, 0.54 long, 0.44 wide. Labium yellow, 0.10 long, 0.20 wide. Chelicerae yellow with black proximal side, with three retromarginal and two promarginal teeth. Palp brown with distal femur and patella with long white hairs, long embolic lamella (EL), and a bifurcated RTA, with its ventral and dorsal lobes with the same shape (
<figureCitation id="13140A8A57389060FC57FB01FBE5FB0F" box="[993,1124,1230,1255]" captionStart="FIGURES 10" captionStartId="17.[151,264,1888,1910]" captionTargetBox="[175,1413,209,1862]" captionTargetId="figure@17.[163,1427,193,1871]" captionTargetPageId="17" captionText="FIGURES 10 a – f. A. sikuani sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. A. somondoco sp. nov., e) epigyne, cleared, ventral view; f) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, PED = process on emblic disc, ED = embolic disc, EF = embolic filament, EL = embolic lamella, fd = fertilization duct, po = epigynal pocket, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, sp = spermatheca, TL = tegular lobe. Scale bars: 0.25 mm (a – b, e – f), 0.20 mm (c – d)." httpUri="https://zenodo.org/record/828226/files/figure.png" pageId="16" pageNumber="17">Figs 10a–b</figureCitation>
,
<figureCitation id="13140A8A57389060FBC7FB01FB3AFB0F" box="[1137,1211,1230,1255]" captionStart="FIGURES 22" captionStartId="33.[151,264,1900,1922]" captionTargetBox="[195,1386,193,1874]" captionTargetId="figure@33.[162,1426,193,1883]" captionTargetPageId="33" captionText="FIGURES 22 a – k. Amphidraus spp, left male palp. A. sikuani sp. nov., a) ventral view; b) same, retrolateral view. A. sotairensis sp. nov., c) ventral view; d) same, retrolateral view. A. sua sp. nov., e) ventral view; f) same, retrolateral view. A. tisquesusa sp. nov., g) ventral view; h) same, retrolateral view. A. tanimuca sp. nov., i) ventral view of the bulb and cymbium; j) palp without cymbium to show the RTA, retrolateral view; k) same, prolateral view; l) same, ventral view." httpUri="https://zenodo.org/record/828250/files/figure.png" pageId="16" pageNumber="17">22a–b</figureCitation>
). Legs 1342, I dark brown, II–IV yellow. Leg macrosetae: femur, I–II, IV d 1 di; III d 2 di; patella, III–IV r 1 me; tibia, I
<date id="FF9130CF57389060FB4EFB3CFAB2FAE3" box="[1272,1331,1267,1291]" pageId="16" pageNumber="17">v 2-2</date>
-2; II
<date id="FF9130CF57389060FAC6FB3CFF26FAC7" pageId="16" pageNumber="17">v 1- 2</date>
-2, p 1 di; III–IV
<date id="FF9130CF57389060FED3FAD8FE09FAC7" box="[357,392,1303,1327]" pageId="16" pageNumber="17">v 1</date>
pr, p 1-0-1, r 1-0-1; metatarsus, I
<date id="FF9130CF57389060FCB4FAD8FCBEFAC7" box="[770,831,1303,1327]" pageId="16" pageNumber="17">v 2-2</date>
; II
<date id="FF9130CF57389060FCD2FAD8FC21FAC7" box="[868,928,1303,1327]" pageId="16" pageNumber="17">v 2-2</date>
, p 1 di; III
<date id="FF9130CF57389060FBAAFAD8FBC0FAC7" box="[1052,1089,1303,1327]" pageId="16" pageNumber="17">v 2</date>
-0-1, p 1-0-2, r 1-0-2; IV
<date id="FF9130CF57389060FAEFFAD8FAF9FAC7" box="[1369,1400,1303,1327]" pageId="16" pageNumber="17">v 1</date>
-0- 1, p 1-0-2, r 1-0-2. Abdomen white with an uneven dark brown pattern of marks (
<figureCitation id="13140A8A57389060FB9EFAF5FB0AFABB" box="[1064,1163,1338,1363]" captionStart="FIGURES 17" captionStartId="28.[151,264,1660,1682]" captionTargetBox="[340,1244,413,1635]" captionTargetId="figure@28.[340,1247,411,1639]" captionTargetPageId="28" captionText="FIGURES 17 a – i. A. guatavita sp. nov., a) male holotype (ANDES – IN 3155). A. pae sp. nov., b) male holotype (ICN – Ar 6576). A. quimbaya sp. nov., c) male holotype (ICN – Ar 7746). A. quinini sp. nov., d) male holotype (ICN – Ar 8266); e) female paratype (ICN – Ar 8267). A. sie sp. nov., f) male holotype (ICN – Ar 8254); g) female paratype (ICN – Ar 8255). A. sikuani sp. nov., h) male holotype (ICN – Ar 6558); i) female paratype (ICN – Ar 7185). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828240/files/figure.png" pageId="16" pageNumber="17">Fig. 17h</figureCitation>
).
</paragraph>
<paragraph id="8B90160F57389060FF71FA91FB81F99B" blockId="16.[151,1437,618,1759]" pageId="16" pageNumber="17">
<emphasis id="B95BCA1D57389060FF71FA91FE2FFA9F" bold="true" box="[199,430,1374,1399]" pageId="16" pageNumber="17">
Female (
<typeStatus id="5494A8AD57389060FE87FA91FE20FA9F" box="[305,417,1374,1399]" pageId="16" pageNumber="17" type="paratype">paratype</typeStatus>
).
</emphasis>
Total length 2.50. Carapace yellow with some posterior black marks, 1.26 long, 0.91 wide, 0.60 high (
<figureCitation id="13140A8A57389060FEE8FA4DFE3DFA73" box="[350,444,1410,1435]" captionStart="FIGURES 17" captionStartId="28.[151,264,1660,1682]" captionTargetBox="[340,1244,413,1635]" captionTargetId="figure@28.[340,1247,411,1639]" captionTargetPageId="28" captionText="FIGURES 17 a – i. A. guatavita sp. nov., a) male holotype (ANDES – IN 3155). A. pae sp. nov., b) male holotype (ICN – Ar 6576). A. quimbaya sp. nov., c) male holotype (ICN – Ar 7746). A. quinini sp. nov., d) male holotype (ICN – Ar 8266); e) female paratype (ICN – Ar 8267). A. sie sp. nov., f) male holotype (ICN – Ar 8254); g) female paratype (ICN – Ar 8255). A. sikuani sp. nov., h) male holotype (ICN – Ar 6558); i) female paratype (ICN – Ar 7185). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828240/files/figure.png" pageId="16" pageNumber="17">
Fig.
<date id="FF9130CF57389060FE23FA4DFE3DFA73" box="[405,444,1410,1435]" pageId="16" pageNumber="17" value="1917-01">17i</date>
</figureCitation>
). OC black, 0.47 long. Anterior eye row 0.86 wide and posterior 0.78 wide. Sternum yellow, 0.48 long, 0.40 wide. Labium yellow, 0.15 long, 0.18 wide. Chelicerae yellow, with five tiny retromarginal teeth, and one promarginal tooth. Legs 4312, all yellow. Leg macrosetae: femur, II–III d 1 di; patellae, III–IV r 1 me; tibiae, I
<date id="FF9130CF57389060FE95FA20FEE1F9EF" box="[291,352,1519,1543]" pageId="16" pageNumber="17">v 2-2</date>
-2; II
<date id="FF9130CF57389060FE2AFA20FE54F9EF" box="[412,469,1519,1543]" pageId="16" pageNumber="17">v 1-2</date>
-2, p 1 di; III–IV
<date id="FF9130CF57389060FD20FA20FD38F9EF" box="[662,697,1519,1543]" pageId="16" pageNumber="17">v 1</date>
me, p 1-1, r 1-1; metatarsus, I
<date id="FF9130CF57389060FBA6FA20FBCCF9EF" box="[1040,1101,1519,1543]" pageId="16" pageNumber="17">v 2-2</date>
; II
<date id="FF9130CF57389060FBC7FA20FB2CF9EF" box="[1137,1197,1519,1543]" pageId="16" pageNumber="17">v 2-2</date>
, p 1 di; III
<date id="FF9130CF57389060FA9FFA20FAE5F9EF" box="[1321,1380,1519,1543]" pageId="16" pageNumber="17">v 2-1</date>
, p 1- 0-2, r 1-0-2; IV
<date id="FF9130CF57389060FEF8F9DCFEF3F9C3" box="[334,370,1555,1579]" pageId="16" pageNumber="17">v 1</date>
pr, p 1-0-2, r 1-0-2. Abdomen gray, with a dark brown reticulated pattern of marks (
<figureCitation id="13140A8A57389060FA85F9DDFA0EF9C3" box="[1331,1423,1554,1579]" captionStart="FIGURES 17" captionStartId="28.[151,264,1660,1682]" captionTargetBox="[340,1244,413,1635]" captionTargetId="figure@28.[340,1247,411,1639]" captionTargetPageId="28" captionText="FIGURES 17 a – i. A. guatavita sp. nov., a) male holotype (ANDES – IN 3155). A. pae sp. nov., b) male holotype (ICN – Ar 6576). A. quimbaya sp. nov., c) male holotype (ICN – Ar 7746). A. quinini sp. nov., d) male holotype (ICN – Ar 8266); e) female paratype (ICN – Ar 8267). A. sie sp. nov., f) male holotype (ICN – Ar 8254); g) female paratype (ICN – Ar 8255). A. sikuani sp. nov., h) male holotype (ICN – Ar 6558); i) female paratype (ICN – Ar 7185). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828240/files/figure.png" pageId="16" pageNumber="17">
Fig.
<date id="FF9130CF57389060FADEF9DDFA0EF9C3" box="[1384,1423,1554,1579]" pageId="16" pageNumber="17" value="1917-01">17i</date>
</figureCitation>
). Epigyne (
<figureCitation id="13140A8A57389060FEB3F9F9FE09F9A7" box="[261,392,1590,1615]" captionStart="FIGURES 10" captionStartId="17.[151,264,1888,1910]" captionTargetBox="[175,1413,209,1862]" captionTargetId="figure@17.[163,1427,193,1871]" captionTargetPageId="17" captionText="FIGURES 10 a – f. A. sikuani sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. A. somondoco sp. nov., e) epigyne, cleared, ventral view; f) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, PED = process on emblic disc, ED = embolic disc, EF = embolic filament, EL = embolic lamella, fd = fertilization duct, po = epigynal pocket, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, sp = spermatheca, TL = tegular lobe. Scale bars: 0.25 mm (a – b, e – f), 0.20 mm (c – d)." httpUri="https://zenodo.org/record/828226/files/figure.png" pageId="16" pageNumber="17">Figs 10c–d</figureCitation>
,
<figureCitation id="13140A8A57389060FE23F9F9FE7DF9A7" box="[405,508,1590,1615]" captionStart="FIGURES 25" captionStartId="36.[151,264,1889,1911]" captionTargetBox="[185,1412,194,1864]" captionTargetId="figure@36.[151,1436,193,1867]" captionTargetPageId="36" captionText="FIGURES 25 a – l. Amphidraus spp, epigyne. A. sikuani sp. nov., a) ventral view; g) disposition of the internal structures. A. somondoco sp. nov., b) ventral view; h) disposition of the internal structures. A. sua sp. nov., c) ventral view; i) disposition of the internal structures. A. tundama sp. nov., d) ventral view; j) disposition of the internal structures. A. zaque sp. nov., e) ventral view; k) disposition of the internal structures. A. zipa sp. nov., f) ventral view; l) disposition of the internal structures." httpUri="https://zenodo.org/record/828256/files/figure.png" pageId="16" pageNumber="17">
25a,
<quantity id="4CD7BBEA57389060FE78F9F9FE7DF9A7" box="[462,508,1590,1615]" metricMagnitude="-2" metricUnit="kg" metricValue="2.5" pageId="16" pageNumber="17" unit="g" value="25.0">25g</quantity>
</figureCitation>
), with anterior copulatory openings, short copulatory ducts and posterior slender spermathecae, with a rounded pocket in a posteriorly extended epigynal plate.
</paragraph>
</subSubSection>
<subSubSection id="C335458457389060FF71F9B1FBF0F953" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8B90160F57389060FF71F9B1FBF0F953" blockId="16.[151,1437,618,1759]" pageId="16" pageNumber="17">
<emphasis id="B95BCA1D57389060FF71F9B1FECEF97F" bold="true" box="[199,335,1662,1687]" pageId="16" pageNumber="17">Comments.</emphasis>
The
<typeStatus id="5494A8AD57389060FE39F9B1FE70F97F" box="[399,497,1662,1687]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
male was collected in a yellow tray trap, and the female
<typeStatus id="5494A8AD57389060FB21F9B0FB76F97F" box="[1175,1271,1663,1687]" pageId="16" pageNumber="17" type="paratype">paratype</typeStatus>
was collected manually in a rotten trunk in the soil, in a secondary lowland semi wet forest (
<figureCitation id="13140A8A57389060FBB7F96DFBE5F953" box="[1025,1124,1698,1723]" captionStart="FIGURES 26" captionStartId="37.[151,264,1181,1203]" captionTargetBox="[151,1436,193,1159]" captionTargetId="figure@37.[151,1436,193,1159]" captionTargetPageId="37" captionText="FIGURES 26 a – d. Different altitudinal ecosystems where some of the species of Amphidraus here described were found in Colombia. a) Páramo ecosystem, at Las Lagunas Encantadas Natural Reserve, Duitama, Colombia, habitat of A. tundama sp. nov. (~ 3800 m); b) Andean wet forest at Cueva de los Guácharos National Natural Park, Huila, Colombia, habitat of A. guaitipan sp. nov. (~ 2000 m); c) Andean Oak forest (Quercus humboldtii) at Quinini Natural Reserve, Cundinamarca, Colombia, habitat of A. quinini sp. nov. (~ 1700 m); d) Secondary lowland semi wet forest on the outskirts of the city of Villavicencio, Meta, Colombia, habitat of A. sikuani sp. nov. (~ 500 m)." httpUri="https://zenodo.org/record/828258/files/figure.png" pageId="16" pageNumber="17">Fig. 26d</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C335458457389060FF71F909FB09F936" box="[199,1160,1734,1759]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8B90160F57389060FF71F909FB09F936" blockId="16.[151,1437,618,1759]" box="[199,1160,1734,1759]" pageId="16" pageNumber="17">
<emphasis id="B95BCA1D57389060FF71F909FEE1F937" bold="true" box="[199,352,1734,1759]" pageId="16" pageNumber="17">Distribution.</emphasis>
<collectingCountry id="F338569F57389060FED1F909FE54F937" box="[359,469,1734,1759]" name="Colombia" pageId="16" pageNumber="17">Colombia</collectingCountry>
(
<collectingRegion id="49EBD8ED57389060FE52F908FDA0F937" box="[484,545,1735,1759]" country="Colombia" name="Meta" pageId="16" pageNumber="17">Meta</collectingRegion>
) (
<figureCitation id="13140A8A57389060FD8EF909FD0DF937" box="[568,652,1734,1759]" captionStart="FIGURE 27" captionStartId="38.[151,250,1317,1339]" captionTargetBox="[138,1418,182,1273]" captionTargetId="figure@38.[137,1422,179,1282]" captionTargetPageId="38" captionText="FIGURE 27. Distribution records of the species of the genus Amphidraus Simon from Colombia treated here. Symbols with a dot inside represent species that have distributions in the Cundiboyacense high-Andean plateau, in the Boyacá and Cundinamarca departments." httpUri="https://zenodo.org/record/828260/files/figure.png" pageId="16" pageNumber="17">Fig. 27</figureCitation>
). Known altitudinal distribution:
<quantity id="4CD7BBEA57389060FBB4F909FB05F936" box="[1026,1156,1734,1759]" metricMagnitude="2" metricUnit="m" metricValue="4.14" metricValueMax="4.529999999999999" metricValueMin="3.75" pageId="16" pageNumber="17" unit="m" value="414.0" valueMax="453.0" valueMin="375.0">
375–
<specimenCount id="9D29DD8657389060FB8CF909FB05F936" box="[1082,1156,1734,1759]" pageId="16" pageNumber="17" type="generic">453 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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