280 lines
35 KiB
XML
280 lines
35 KiB
XML
<document id="B7E3A299DDC90C745F66A3755C65B1D3" ID-DOI="10.11646/zootaxa.5375.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="10170174" ID-ZooBank="30782E67-2E2C-4B63-9809-85C3945187AE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700559471735" checkinUser="plazi" docAuthor="Inoue, Shota & Nomura, Shûhei" docDate="2023" docId="03861C35FFDDFFFFFF69813DFC2BFCA8" docLanguage="en" docName="zootaxa.5375.1.4.pdf" docOrigin="Zootaxa 5375 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5375.1.4/52314" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tyrodes amamianus Inoue & Nomura 2023, sp. nov." docType="treatment" docUuid="6653A617-8647-4433-B713-4A287B488467" docUuidSource="ZooBank" docVersion="2" lastPageNumber="91" masterDocId="FFBF644DFFD8FFF7FFFE846AFFACFFFB" masterDocTitle="New species and record of Tyrodes Raffray (Coleoptera: Staphylinidae: Pselaphinae) from Japan" masterLastPageNumber="92" masterPageNumber="83" pageNumber="88" updateTime="1700561800485" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="4FE82C7B144CF22344F6BE38461FD265">
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<mods:title id="7FCAB86AC8C8FA4AEA67A5510323C1A7">New species and record of Tyrodes Raffray (Coleoptera: Staphylinidae: Pselaphinae) from Japan</mods:title>
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<mods:name id="435A625CE7CA531393F2F1E6D1660644" type="personal">
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<mods:roleTerm id="BD1BEA6F1B72A8B9B8195803C8C588F4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8C34F708FD476445384E7FB75D8F3B08">Inoue, Shota</mods:namePart>
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<mods:nameIdentifier id="E2962DC8C99EAD77A13628FE68F86998" type="ORCID">0000-0002-7005-6531</mods:nameIdentifier>
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<mods:affiliation id="01672EF714850412DFDDCBE2C10D9BD2">University Education Center, Tokyo Metropolitan University, 1 - 1 Minami-Osawa, Hachioji, Tokyo, 192 - 0397 Japan shota 666. shigure @ gmail. com; https: // orcid. org / 0000 - 0002 - 7005 - 6531 & The Kyushu University Museum, Fukuoka, 812 - 8581 Japan.</mods:affiliation>
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<mods:nameIdentifier id="16B1473AA102CBA9C1ADCDC636909B86" type="email">shota666.shigure@gmail.com</mods:nameIdentifier>
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<mods:name id="F861CC799F7A97529870AD60E7D39B92" type="personal">
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</mods:role>
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<mods:namePart id="8F4BAD31310E3EBBC599B70B2D0A0E6B">Nomura, Shûhei</mods:namePart>
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<mods:nameIdentifier id="A9814E7972609D028E3001C46B779394" type="ORCID">0000-0003-1086-5078</mods:nameIdentifier>
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<mods:affiliation id="52DF2939A3560109F01E9B46DC773F03">Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1, Amakubo, Tsukuba-shi, Ibaraki, 305 - 0005 Japan nomura @ kahaku. go. jp; https: // orcid. org / 0000 - 0003 - 1086 - 5078</mods:affiliation>
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<mods:nameIdentifier id="DF24C54DC324465E10FE5C67C898779E" type="email">nomura@kahaku.go.jp</mods:nameIdentifier>
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<mods:typeOfResource id="04B213DB8A52346DF46D828451511EBC">text</mods:typeOfResource>
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<mods:title id="7C21AA2293C67696123BDF85323CB7F6">Zootaxa</mods:title>
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<mods:part id="26D667C5B2024A5CA08141828F0A7B69">
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<mods:date id="80F14815E56DBFB38178C6114A44E9B6">2023</mods:date>
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<mods:number id="1EC5FAECBE196B573D49FA64F428A9E5">2023-11-21</mods:number>
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<mods:number id="1BF674789FEC04F65E4D1D8FC05924CA">5375</mods:number>
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<mods:url id="78A4DD31D0EBBD2FFE72A529FCE1F663">https://www.mapress.com/zt/article/download/zootaxa.5375.1.4/52314</mods:url>
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<mods:classification id="CDB795E45F4F1D91CB1EE50FA3EE293D">journal article</mods:classification>
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<treatment id="03861C35FFDDFFFFFF69813DFC2BFCA8" LSID="urn:lsid:zoobank.org:act:6653A617-8647-4433-B713-4A287B488467" httpUri="http://treatment.plazi.org/id/03861C35FFDDFFFFFF69813DFC2BFCA8" lastPageId="8" lastPageNumber="91" pageId="5" pageNumber="88">
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<subSubSection id="C335FEA8FFDDFFF2FF69813DFE5CFA89" box="[151,496,1367,1394]" pageId="5" pageNumber="88" type="nomenclature">
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<paragraph id="8B90AD23FFDDFFF2FF69813DFE5CFA89" blockId="5.[151,814,1367,1466]" box="[151,496,1367,1394]" pageId="5" pageNumber="88">
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<heading id="D0D81A4FFFDDFFF2FF69813DFE5CFA89" bold="true" box="[151,496,1367,1394]" fontSize="11" level="1" pageId="5" pageNumber="88" reason="1">
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<emphasis id="B95B7131FFDDFFF2FF69813DFE5CFA89" bold="true" box="[151,496,1367,1394]" pageId="5" pageNumber="88">
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<taxonomicName id="4C2FD6A0FFDDFFF2FF69813DFE20FA89" authority="Inoue & Nomura, 2023" authorityName="Inoue & Nomura" authorityYear="2023" box="[151,396,1367,1394]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="species" species="amamianus" status="sp. nov.">
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<emphasis id="B95B7131FFDDFFF2FF69813DFE20FA89" bold="true" box="[151,396,1367,1394]" italics="true" pageId="5" pageNumber="88">Tyrodes amamianus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A268CC4AFFDDFFF2FE6D8132FE5CFA89" box="[403,496,1368,1394]" pageId="5" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C335FEA8FFDDFFF2FF698116FC82FA41" pageId="5" pageNumber="88" type="description">
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<paragraph id="8B90AD23FFDDFFF2FF698116FE91FA6D" blockId="5.[151,814,1367,1466]" box="[151,317,1404,1431]" pageId="5" pageNumber="88">
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(
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<figureCitation id="1314B1A6FFDDFFF2FF618116FF44FA6C" box="[159,232,1404,1431]" captionStart="FIGURE 5" captionStartId="6.[151,250,1710,1734]" captionTargetBox="[158,1429,198,1681]" captionTargetId="figure-27@6.[151,1436,181,1686]" captionTargetPageId="6" captionText="FIGURE 5. Morphology of Tyrodes amamianus sp. nov., male. A. Dorsal habitus. B. Ventral habitus. C. Antennomere 1. D. Metaventral projection. E. Lateral side of abdomen. F. Tergite VIII. G. Sternite VIII. H. Sternite IX. Scale bars: 1.0 mm in A, B; 0.5 mm in E; 0.2 mm in D; 0.1 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.10170201" httpUri="https://zenodo.org/record/10170201/files/figure.png" pageId="5" pageNumber="88">Figs 5</figureCitation>
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,
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<figureCitation id="1314B1A6FFDDFFF2FF0A8116FE99FA6D" box="[244,309,1404,1430]" captionStart="FIGURE 6" captionStartId="7.[151,250,1433,1457]" captionTargetBox="[154,1432,184,1406]" captionTargetId="figure-19@7.[151,1435,181,1409]" captionTargetPageId="7" captionText="FIGURE 6. Aedeagus of Tyrodes segrex Kurbatov (A–C) and Tyrodes amamianus sp. nov. (D–F). A, D. Ventral view. B, E. Lateral view. C, F. Dorsal view. Scale bar: 0.1 mm in A–F." figureDoi="http://doi.org/10.5281/zenodo.10170207" httpUri="https://zenodo.org/record/10170207/files/figure.png" pageId="5" pageNumber="88">6D–F</figureCitation>
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)
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</paragraph>
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<paragraph id="8B90AD23FFDDFFF2FF6981CAFC82FA41" blockId="5.[151,814,1367,1466]" box="[151,814,1440,1466]" pageId="5" pageNumber="88">[Japanese common name: Amami-Nisekoke-arizukamushi]</paragraph>
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</subSubSection>
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<subSubSection id="C335FEA8FFDDFFF2FF698182FAB4F9DD" pageId="5" pageNumber="88" type="materials_examined">
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<paragraph id="8B90AD23FFDDFFF2FF698182FAB4F9DD" blockId="5.[151,1437,1512,2043]" pageId="5" pageNumber="88">
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<emphasis id="B95B7131FFDDFFF2FF698182FEECF9F9" bold="true" box="[151,320,1512,1538]" pageId="5" pageNumber="88">Type material</emphasis>
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.
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<materialsCitation id="3B47A77EFFDDFFF2FEAE8182FAB4F9DD" collectingDate="1991-04-02" collectorName="Is. Amami-oshima & T. Ueno" country="Japan" determinerName="S. Nomura" location="Mt. Yuwandake" pageId="5" pageNumber="88" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="B95B7131FFDDFFF2FEAE8182FE54F9F9" bold="true" box="[336,504,1512,1538]" pageId="5" pageNumber="88">
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||
<typeStatus id="54941381FFDDFFF2FEAE8182FE5FF9F9" box="[336,499,1512,1538]" pageId="5" pageNumber="88" type="holotype">HOLOTYPE</typeStatus>
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:
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</emphasis>
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³, “
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<collectingCountry id="F338EDB3FFDDFFF2FDD18182FD27F9F9" box="[559,651,1512,1538]" name="Japan" pageId="5" pageNumber="88">JAPAN</collectingCountry>
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: RYUKYUS /
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<collectorName id="26DAC8F5FFDDFFF2FCC98182FBABF9F9" box="[823,1031,1512,1538]" pageId="5" pageNumber="88">Is. Amami-oshima</collectorName>
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/
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||
<location id="8EF0FBF8FFDDFFF2FBDF8182FB78F9F9" LSID="urn:lsid:plazi:treatment:03861C35FFDDFFFFFF69813DFC2BFCA8:8EF0FBF8FFDDFFF2FBDF8182FB78F9F9" box="[1057,1236,1512,1538]" country="Japan" name="Mt. Yuwandake" pageId="5" pageNumber="88">Mt. Yuwandake</location>
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/
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||
<date id="FF918BE3FFDDFFF2FB138182FAC1F9F9" box="[1261,1389,1512,1538]" pageId="5" pageNumber="88" value="1991-04-02">
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||
<collectingDate id="EFD5720BFFDDFFF2FB138182FAC1F9F9" box="[1261,1389,1512,1538]" pageId="5" pageNumber="88" value="1991-04-02">2. IV. 1991</collectingDate>
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||
</date>
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||
/
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||
<collectorName id="26DAC8F5FFDDFFF2FA788182FF78F9DD" pageId="5" pageNumber="88">T. Ueno</collectorName>
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leg. // FAUNA JAPONICA /
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<taxonomicName id="4C2FD6A0FFDDFFF2FDDF8266FD12F9DD" box="[545,702,1548,1574]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
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,
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<taxonomicName id="4C2FD6A0FFDDFFF2FD378266FCE3F9DD" box="[713,847,1548,1574]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="subFamily" subFamily="Pselaphinae">Pselaphinae</taxonomicName>
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||
//
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<taxonomicName id="4C2FD6A0FFDDFFF2FC928266FC53F9DD" box="[876,1023,1548,1574]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="species" species="undefined-2">
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<emphasis id="B95B7131FFDDFFF2FC928266FC6FF9DD" box="[876,963,1548,1574]" italics="true" pageId="5" pageNumber="88">Tyrodes</emphasis>
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sp. 2
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</taxonomicName>
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||
//
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<determinerName id="7E4C41E7FFDDFFF2FBE38266FB3AF9DD" box="[1053,1174,1548,1574]" pageId="5" pageNumber="88">S. Nomura</determinerName>
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det., 2021”
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C335FEA8FFDDFFF2FF39825AFCADF969" pageId="5" pageNumber="88" type="diagnosis">
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||
<paragraph id="8B90AD23FFDDFFF2FF39825AFCADF969" blockId="5.[151,1437,1512,2043]" pageId="5" pageNumber="88">
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<emphasis id="B95B7131FFDDFFF2FF39825AFEEEF9B1" bold="true" box="[199,322,1584,1610]" pageId="5" pageNumber="88">Diagnosis.</emphasis>
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This species is easily distinguished from its congener by the elongate antennomeres 9–10, abdominal sternite IV with densely setose patch at posterolateral in concavity and endophallus composed of two sclerites which are long and basally twisted one and apically bifid one.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C335FEA8FFDDFFFFFF3982F6FE32FE0B" lastPageId="8" lastPageNumber="91" pageId="5" pageNumber="88" type="description">
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||
<paragraph id="8B90AD23FFDDFFF2FF3982F6FB67F94D" blockId="5.[151,1437,1512,2043]" box="[199,1227,1692,1719]" pageId="5" pageNumber="88">
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<emphasis id="B95B7131FFDDFFF2FF3982F6FEFDF94D" bold="true" box="[199,337,1692,1718]" pageId="5" pageNumber="88">Description</emphasis>
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. Male (
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<figureCitation id="1314B1A6FFDDFFF2FE5682F6FE41F94C" box="[424,493,1692,1719]" captionStart="FIGURE 5" captionStartId="6.[151,250,1710,1734]" captionTargetBox="[158,1429,198,1681]" captionTargetId="figure-27@6.[151,1436,181,1686]" captionTargetPageId="6" captionText="FIGURE 5. Morphology of Tyrodes amamianus sp. nov., male. A. Dorsal habitus. B. Ventral habitus. C. Antennomere 1. D. Metaventral projection. E. Lateral side of abdomen. F. Tergite VIII. G. Sternite VIII. H. Sternite IX. Scale bars: 1.0 mm in A, B; 0.5 mm in E; 0.2 mm in D; 0.1 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.10170201" httpUri="https://zenodo.org/record/10170201/files/figure.png" pageId="5" pageNumber="88">Fig. 5</figureCitation>
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). BL
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<quantity id="4CD700C6FFDDFFF2FDD282F6FD3FF94D" box="[556,659,1692,1718]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" pageId="5" pageNumber="88" unit="mm" value="1.92">1.92 mm</quantity>
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. Dorsal surface with dense, moderately long setae.
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</paragraph>
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||
<paragraph id="8B90AD23FFDDFFF1FF3982AAFB3FF801" blockId="5.[151,1437,1512,2043]" lastBlockId="6.[151,1436,1875,2043]" lastPageId="6" lastPageNumber="89" pageId="5" pageNumber="88">
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Head slightly wider than long (HL/HW 0.87), HL
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||
<quantity id="4CD700C6FFDDFFF2FD0982AAFCF0F921" box="[759,860,1728,1754]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="5" pageNumber="88" unit="mm" value="0.34">0.34 mm</quantity>
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, HW
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<quantity id="4CD700C6FFDDFFF2FC6282AAFBAEF921" box="[924,1026,1728,1754]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="5" pageNumber="88" unit="mm" value="0.39">0.39 mm</quantity>
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, with both antennal tubercles closely situated each other, forming short frontal rostrum; frontal rostrum narrow, about half as long as HL, 1/3 as wide as HW, with few punctures, with small, nude frontal fovea at middle; posterolateral clypeal region strongly concave, its edge demarcated, angularly projected laterally and visible in dorsal view; vertex finely punctate, with a pair of setose fovea; eyes large, prominent; postocular margin rounded, with tufts of dense setae. Antennae each long,
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<quantity id="4CD700C6FFDDFFF2FAE3833AFAD3F891" box="[1309,1407,1872,1898]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" pageId="5" pageNumber="88" unit="mm" value="1.21">1.21 mm</quantity>
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in length; antennomeres 1 each longer than wide,
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<quantity id="4CD700C6FFDDFFF2FD63831EFD52F875" box="[669,766,1908,1934]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="88" unit="mm" value="0.12">0.12 mm</quantity>
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in length, with triangular projection on external side of basal part (
|
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<figureCitation id="1314B1A6FFDDFFF2FF2C83F2FE87F848" box="[210,299,1944,1971]" captionStart="FIGURE 5" captionStartId="6.[151,250,1710,1734]" captionTargetBox="[158,1429,198,1681]" captionTargetId="figure-27@6.[151,1436,181,1686]" captionTargetPageId="6" captionText="FIGURE 5. Morphology of Tyrodes amamianus sp. nov., male. A. Dorsal habitus. B. Ventral habitus. C. Antennomere 1. D. Metaventral projection. E. Lateral side of abdomen. F. Tergite VIII. G. Sternite VIII. H. Sternite IX. Scale bars: 1.0 mm in A, B; 0.5 mm in E; 0.2 mm in D; 0.1 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.10170201" httpUri="https://zenodo.org/record/10170201/files/figure.png" pageId="5" pageNumber="88">Fig. 5C</figureCitation>
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); relative length of each antennomere to antennomeres 1 as follows: 1.0: 0.72: 0.72: 0.63: 0.63: 0.55: 0.55: 0.55: 1.28: 1.28: 2.28; antennomeres 2 each longer than wide; antennomeres 3 each slightly widened towards apex, with posterior part 2/3 as wide as anterior part; antennomeres 3–5 each longer than wide; antennomeres 6–8 each as long as wide; antennomeres 9–11 each enlarged to form club, longer than wide, length/width ratio of 9–11 as follows: 1.47: 1.36: 1.89; antennomeres 11 each largest, subconical, about twice times longer than wide. Maxillary palpi with palpomeres 2 each narrowed and pedunculate at basal half, gradually widened apicad; palpomeres 3 narrowed and pedunculate at basal third, gradually widened apicad; palpomeres 4 each narrowed and pedunculate at base, widened apically, widest at middle, then narrowed apically, with apical appendage.
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</paragraph>
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<caption id="DF50FDABFFDEFFF1FF6982C4FD21F8F5" ID-DOI="http://doi.org/10.5281/zenodo.10170201" ID-Zenodo-Dep="10170201" httpUri="https://zenodo.org/record/10170201/files/figure.png" pageId="6" pageNumber="89" startId="6.[151,250,1710,1734]" targetBox="[158,1429,198,1681]" targetPageId="6" targetType="figure">
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<paragraph id="8B90AD23FFDEFFF1FF6982C4FD21F8F5" blockId="6.[151,1436,1710,1806]" pageId="6" pageNumber="89">
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||
<emphasis id="B95B7131FFDEFFF1FF6982C4FEA2F93D" bold="true" box="[151,270,1710,1734]" pageId="6" pageNumber="89">FIGURE 5</emphasis>
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||
. Morphology of
|
||
<taxonomicName id="4C2FD6A0FFDEFFF1FE4182C4FD26F93D" authority="Inoue & Nomura, 2023" authorityName="Inoue & Nomura" authorityYear="2023" box="[447,650,1710,1734]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="species" species="amamianus" status="sp. nov.">
|
||
<emphasis id="B95B7131FFDEFFF1FE4182C4FD26F93D" box="[447,650,1710,1734]" italics="true" pageId="6" pageNumber="89">Tyrodes amamianus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95B7131FFDEFFF1FD6F82C4FD4FF93D" bold="true" box="[657,739,1710,1734]" pageId="6" pageNumber="89">
|
||
<taxonomicNameLabel id="A268CC4AFFDEFFF1FD6F82C4FD4FF93D" box="[657,739,1710,1734]" pageId="6" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male.
|
||
<emphasis id="B95B7131FFDEFFF1FCD182C4FCE4F93D" bold="true" box="[815,840,1710,1734]" pageId="6" pageNumber="89">A.</emphasis>
|
||
Dorsal habitus.
|
||
<emphasis id="B95B7131FFDEFFF1FC0F82C4FBA4F93D" bold="true" box="[1009,1032,1710,1734]" pageId="6" pageNumber="89">B.</emphasis>
|
||
Ventral habitus.
|
||
<emphasis id="B95B7131FFDEFFF1FB4682C4FB7CF93D" bold="true" box="[1208,1232,1710,1734]" pageId="6" pageNumber="89">C.</emphasis>
|
||
Antennomere 1.
|
||
<emphasis id="B95B7131FFDEFFF1FA7A82C4FA30F93D" bold="true" box="[1412,1436,1710,1734]" pageId="6" pageNumber="89">D.</emphasis>
|
||
Metaventral projection.
|
||
<emphasis id="B95B7131FFDEFFF1FE7382B8FE08F911" bold="true" box="[397,420,1746,1770]" pageId="6" pageNumber="89">E.</emphasis>
|
||
Lateral side of abdomen.
|
||
<emphasis id="B95B7131FFDEFFF1FD4E82B8FD6FF911" bold="true" box="[688,707,1746,1770]" pageId="6" pageNumber="89">F.</emphasis>
|
||
Tergite VIII.
|
||
<emphasis id="B95B7131FFDEFFF1FCAE82B8FCC7F911" bold="true" box="[848,875,1746,1770]" pageId="6" pageNumber="89">G.</emphasis>
|
||
Sternite VIII.
|
||
<emphasis id="B95B7131FFDEFFF1FBFE82B8FBB6F911" bold="true" box="[1024,1050,1746,1770]" pageId="6" pageNumber="89">H.</emphasis>
|
||
Sternite IX. Scale bars: 1.0 mm in A, B; 0.5 mm in E; 0.2 mm in D; 0.1 mm in C, F–H.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF50FDABFFDFFFF0FF6981F3FD46FA2E" ID-DOI="http://doi.org/10.5281/zenodo.10170207" ID-Zenodo-Dep="10170207" httpUri="https://zenodo.org/record/10170207/files/figure.png" pageId="7" pageNumber="90" startId="7.[151,250,1433,1457]" targetBox="[154,1432,184,1406]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8B90AD23FFDFFFF0FF6981F3FD46FA2E" blockId="7.[151,1436,1433,1493]" pageId="7" pageNumber="90">
|
||
<emphasis id="B95B7131FFDFFFF0FF6981F3FEA3FA4A" bold="true" box="[151,271,1433,1457]" pageId="7" pageNumber="90">FIGURE 6</emphasis>
|
||
. Aedeagus of
|
||
<taxonomicName id="4C2FD6A0FFDFFFF0FE5A81F3FD0FFA4A" authority="Kurbatov" authorityName="Kurbatov" authorityYear="1990" box="[420,675,1433,1457]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="segrex">
|
||
<emphasis id="B95B7131FFDFFFF0FE5A81F3FD97FA4A" box="[420,571,1433,1457]" italics="true" pageId="7" pageNumber="90">Tyrodes segrex</emphasis>
|
||
Kurbatov
|
||
</taxonomicName>
|
||
(A–C) and
|
||
<taxonomicName id="4C2FD6A0FFDFFFF0FCDE81F3FC40FA4A" authority="Inoue & Nomura, 2023" authorityName="Inoue & Nomura" authorityYear="2023" box="[800,1004,1433,1457]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="amamianus" status="sp. nov.">
|
||
<emphasis id="B95B7131FFDFFFF0FCDE81F3FC40FA4A" box="[800,1004,1433,1457]" italics="true" pageId="7" pageNumber="90">Tyrodes amamianus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A268CC4AFFDFFFF0FC0D81F0FBE8FA4A" box="[1011,1092,1434,1457]" pageId="7" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(D–F).
|
||
<emphasis id="B95B7131FFDFFFF0FB6981F3FB7CFA4A" bold="true" box="[1175,1232,1433,1457]" pageId="7" pageNumber="90">A, D.</emphasis>
|
||
Ventral view.
|
||
<emphasis id="B95B7131FFDFFFF0FA9881F3FA30FA4A" bold="true" box="[1382,1436,1433,1457]" pageId="7" pageNumber="90">B, E.</emphasis>
|
||
Lateral view.
|
||
<emphasis id="B95B7131FFDFFFF0FEDC81D7FEFFFA2E" bold="true" box="[290,339,1469,1493]" pageId="7" pageNumber="90">C, F.</emphasis>
|
||
Dorsal view. Scale bar: 0.1 mm in A–F.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B90AD23FFDFFFF0FF39824CFDECF859" blockId="7.[151,1437,1574,2026]" pageId="7" pageNumber="90">
|
||
Pronotum as long as wide (PL/PW 1.00), PL
|
||
<quantity id="4CD700C6FFDFFFF0FD46824CFCB0F9BB" box="[696,796,1574,1600]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="7" pageNumber="90" unit="mm" value="0.42">0.42 mm</quantity>
|
||
, PW
|
||
<quantity id="4CD700C6FFDFFFF0FCA8824CFC17F9BB" box="[854,955,1574,1600]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="7" pageNumber="90" unit="mm" value="0.42">0.42 mm</quantity>
|
||
, widest at apical third, with anterior margin shorter than posterior margin, finely punctate, with transverse antebasal sulcus connecting small median setose fovea and large lateral setose foveae. Elytra nearly trapezoidal, wider than long (EL/EW 0.73), EL
|
||
<quantity id="4CD700C6FFDFFFF0FB6D8207FB54F97C" box="[1171,1272,1645,1671]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="90" unit="mm" value="0.6">0.60 mm</quantity>
|
||
, EW
|
||
<quantity id="4CD700C6FFDFFFF0FACD8207FA34F97C" box="[1331,1432,1645,1671]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="7" pageNumber="90" unit="mm" value="0.81">0.81 mm</quantity>
|
||
, longer than pronotum, finely punctate; each elytron with two basal setose foveae; discal stria each shallow, extend to elytral midpoint. Mesoventrite with setose median mesoventral foveae small, with lateral mesoventral foveae setose, with lateral mesocoxal foveae setose; metaventrite finely punctate, polished, with median metaventral foveae setose, with setose small projection at middle (
|
||
<figureCitation id="1314B1A6FFDFFFF0FD598291FCADF8EE" box="[679,769,1786,1813]" captionStart="FIGURE 5" captionStartId="6.[151,250,1710,1734]" captionTargetBox="[158,1429,198,1681]" captionTargetId="figure-27@6.[151,1436,181,1686]" captionTargetPageId="6" captionText="FIGURE 5. Morphology of Tyrodes amamianus sp. nov., male. A. Dorsal habitus. B. Ventral habitus. C. Antennomere 1. D. Metaventral projection. E. Lateral side of abdomen. F. Tergite VIII. G. Sternite VIII. H. Sternite IX. Scale bars: 1.0 mm in A, B; 0.5 mm in E; 0.2 mm in D; 0.1 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.10170201" httpUri="https://zenodo.org/record/10170201/files/figure.png" pageId="7" pageNumber="90">Fig. 5D</figureCitation>
|
||
); posterior margin of metaventrite with looped notch. Legs with coxae lacking spines; with femora each broadest at middle; profemora each simple; protibiae and metatibiae each nearly straight; mesotibiae each weakly arcuate at apical third; tarsi each elongate, about half as long as each tibia, with metatarsi longest; tarsomeres 1 each very short; tarsomere 2 and 3 each elongate; 3 each slightly longer than 2, with symmetrical tarsal claws.
|
||
</paragraph>
|
||
<paragraph id="8B90AD23FFDFFFFFFF3983C6FC43FEC5" blockId="7.[151,1437,1574,2026]" lastBlockId="8.[151,1437,150,852]" lastPageId="8" lastPageNumber="91" pageId="7" pageNumber="90">
|
||
Abdomen wider than long (AL/AW 0.69), AL
|
||
<quantity id="4CD700C6FFDFFFF0FD2883C6FC91F83D" box="[726,829,1964,1991]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="7" pageNumber="90" unit="mm" value="0.57">0.57 mm</quantity>
|
||
, AW
|
||
<quantity id="4CD700C6FFDFFFF0FC8283C6FC4FF83D" box="[892,995,1964,1990]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="7" pageNumber="90" unit="mm" value="0.82">0.82 mm</quantity>
|
||
, finely punctate, with tergite IV longer than V, with shallow basal sulcus; IV–VIII each with a pair of basolateral setose foveae; IV–VII each with a pair of well-developed paratergites; tergite VIII (
|
||
<figureCitation id="1314B1A6FFD0FFFFFD9B84FDFD1AFF4A" box="[613,694,150,177]" captionStart="FIGURE 5" captionStartId="6.[151,250,1710,1734]" captionTargetBox="[158,1429,198,1681]" captionTargetId="figure-27@6.[151,1436,181,1686]" captionTargetPageId="6" captionText="FIGURE 5. Morphology of Tyrodes amamianus sp. nov., male. A. Dorsal habitus. B. Ventral habitus. C. Antennomere 1. D. Metaventral projection. E. Lateral side of abdomen. F. Tergite VIII. G. Sternite VIII. H. Sternite IX. Scale bars: 1.0 mm in A, B; 0.5 mm in E; 0.2 mm in D; 0.1 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.10170201" httpUri="https://zenodo.org/record/10170201/files/figure.png" pageId="8" pageNumber="91">Fig. 5F</figureCitation>
|
||
) semicircular, transverse; sternite IV with shallow basal transverse sulcus; posterolateral part of sternite IV shallowly concave and with densely setose patch on medial side of its shallow concavity, anterolateral part of sternite V strongly concave, its concavity glabrous (
|
||
<figureCitation id="1314B1A6FFD0FFFFFB7684B7FB72FF03" box="[1160,1246,221,248]" captionStart="FIGURE 5" captionStartId="6.[151,250,1710,1734]" captionTargetBox="[158,1429,198,1681]" captionTargetId="figure-27@6.[151,1436,181,1686]" captionTargetPageId="6" captionText="FIGURE 5. Morphology of Tyrodes amamianus sp. nov., male. A. Dorsal habitus. B. Ventral habitus. C. Antennomere 1. D. Metaventral projection. E. Lateral side of abdomen. F. Tergite VIII. G. Sternite VIII. H. Sternite IX. Scale bars: 1.0 mm in A, B; 0.5 mm in E; 0.2 mm in D; 0.1 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.10170201" httpUri="https://zenodo.org/record/10170201/files/figure.png" pageId="8" pageNumber="91">Fig. 5E</figureCitation>
|
||
); sternite IV–VII flat at middle; sternite VIII (
|
||
<figureCitation id="1314B1A6FFD0FFFFFE2C856BFD85FEE7" box="[466,553,257,284]" captionStart="FIGURE 5" captionStartId="6.[151,250,1710,1734]" captionTargetBox="[158,1429,198,1681]" captionTargetId="figure-27@6.[151,1436,181,1686]" captionTargetPageId="6" captionText="FIGURE 5. Morphology of Tyrodes amamianus sp. nov., male. A. Dorsal habitus. B. Ventral habitus. C. Antennomere 1. D. Metaventral projection. E. Lateral side of abdomen. F. Tergite VIII. G. Sternite VIII. H. Sternite IX. Scale bars: 1.0 mm in A, B; 0.5 mm in E; 0.2 mm in D; 0.1 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.10170201" httpUri="https://zenodo.org/record/10170201/files/figure.png" pageId="8" pageNumber="91">Fig. 5G</figureCitation>
|
||
) transverse, emarginate at middle of apical margin; sternite IX (
|
||
<figureCitation id="1314B1A6FFD0FFFFFB0C856BFAE6FEE7" box="[1266,1354,257,284]" captionStart="FIGURE 5" captionStartId="6.[151,250,1710,1734]" captionTargetBox="[158,1429,198,1681]" captionTargetId="figure-27@6.[151,1436,181,1686]" captionTargetPageId="6" captionText="FIGURE 5. Morphology of Tyrodes amamianus sp. nov., male. A. Dorsal habitus. B. Ventral habitus. C. Antennomere 1. D. Metaventral projection. E. Lateral side of abdomen. F. Tergite VIII. G. Sternite VIII. H. Sternite IX. Scale bars: 1.0 mm in A, B; 0.5 mm in E; 0.2 mm in D; 0.1 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.10170201" httpUri="https://zenodo.org/record/10170201/files/figure.png" pageId="8" pageNumber="91">Fig. 5H</figureCitation>
|
||
) small, rounded, apical margin with sparce setae, with membranous plates at lateral.
|
||
</paragraph>
|
||
<paragraph id="8B90AD23FFD0FFFFFF398522FA8AFE37" blockId="8.[151,1437,150,852]" pageId="8" pageNumber="91">
|
||
Aedeagus (
|
||
<figureCitation id="1314B1A6FFD0FFFFFEB98522FE13FE99" box="[327,447,328,354]" captionStart="FIGURE 6" captionStartId="7.[151,250,1433,1457]" captionTargetBox="[154,1432,184,1406]" captionTargetId="figure-19@7.[151,1435,181,1409]" captionTargetPageId="7" captionText="FIGURE 6. Aedeagus of Tyrodes segrex Kurbatov (A–C) and Tyrodes amamianus sp. nov. (D–F). A, D. Ventral view. B, E. Lateral view. C, F. Dorsal view. Scale bar: 0.1 mm in A–F." figureDoi="http://doi.org/10.5281/zenodo.10170207" httpUri="https://zenodo.org/record/10170207/files/figure.png" pageId="8" pageNumber="91">Fig. 6D–F</figureCitation>
|
||
)
|
||
<quantity id="4CD700C6FFD0FFFFFE2E8522FD9AFE98" box="[464,566,328,355]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="8" pageNumber="91" unit="mm" value="0.32">0.32 mm</quantity>
|
||
in length, asymmetrical, sclerotized; median lobe with broad basal capsule; apical half narrowed apically, bended laterally in dorso-ventral view; apical orifice broad; parameres reduced, to form apically truncated plate; endophallus composed of long, basally twisted sclerite and short, apically bifid sclerite; base of both sclerites membranous, and base of short, apically bifid sclerite gradually obscured.
|
||
</paragraph>
|
||
<paragraph id="8B90AD23FFD0FFFFFF3985BCFE32FE0B" blockId="8.[151,1437,150,852]" box="[199,414,470,496]" pageId="8" pageNumber="91">Female. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C335FEA8FFD0FFFFFF398593FC0AFDE8" box="[199,934,505,534]" pageId="8" pageNumber="91" type="distribution">
|
||
<paragraph id="8B90AD23FFD0FFFFFF398593FC0AFDE8" blockId="8.[151,1437,150,852]" box="[199,934,505,534]" pageId="8" pageNumber="91">
|
||
<emphasis id="B95B7131FFD0FFFFFF398593FEF5FDE8" bold="true" box="[199,345,505,531]" pageId="8" pageNumber="91">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F338EDB3FFD0FFFFFE998590FE0BFDEF" box="[359,423,506,532]" name="Japan" pageId="8" pageNumber="91">Japan</collectingCountry>
|
||
(Amami-Ôshima Is., Kagoshima-ken,
|
||
<collectingCountry id="F338EDB3FFD0FFFFFCA28593FC32FDE8" box="[860,926,505,531]" name="Japan" pageId="8" pageNumber="91">Japan</collectingCountry>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C335FEA8FFD0FFFFFF398677FC2BFCA8" pageId="8" pageNumber="91" type="etymology">
|
||
<paragraph id="8B90AD23FFD0FFFFFF398677FBD6FDCC" blockId="8.[151,1437,150,852]" box="[199,1146,541,569]" pageId="8" pageNumber="91">
|
||
<emphasis id="B95B7131FFD0FFFFFF398677FEE2FDCC" bold="true" box="[199,334,541,567]" pageId="8" pageNumber="91">Etymology.</emphasis>
|
||
The new species is named after its
|
||
<typeStatus id="54941381FFD0FFFFFD228677FCA0FDCC" box="[732,780,541,567]" pageId="8" pageNumber="91">type</typeStatus>
|
||
locality, Amami-Ôshima
|
||
<collectingCountry id="F338EDB3FFD0FFFFFBD08677FBDAFDCC" box="[1070,1142,541,567]" name="Iceland" pageId="8" pageNumber="91">Island</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B90AD23FFD0FFFFFF39862AFB44FD3E" blockId="8.[151,1437,150,852]" pageId="8" pageNumber="91">
|
||
<emphasis id="B95B7131FFD0FFFFFF39862AFE1DFDA1" bold="true" box="[199,433,576,602]" pageId="8" pageNumber="91">Comparative notes.</emphasis>
|
||
As in the original descriptions, the antennomeres 9–10 are relatively quadrate or slightly shorter than wide in all
|
||
<taxonomicName id="4C2FD6A0FFD0FFFFFE54860EFDF7FD85" box="[426,603,612,638]" pageId="8" pageNumber="91">
|
||
<emphasis id="B95B7131FFD0FFFFFE54860EFDADFD85" box="[426,513,612,638]" italics="true" pageId="8" pageNumber="91">Tyrodes</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
except for the new species and
|
||
<taxonomicName id="4C2FD6A0FFD0FFFFFC33860EFAF3FD85" authority="Zhang & Yin 2023" authorityName="Zhang & Yin" authorityYear="2023" box="[973,1375,612,638]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="91" phylum="Arthropoda" rank="species" species="tibialis">
|
||
<emphasis id="B95B7131FFD0FFFFFC33860EFBD7FD85" box="[973,1147,612,638]" italics="true" pageId="8" pageNumber="91">Tyrodes tibialis</emphasis>
|
||
<bibRefCitation id="EFBED0D2FFD0FFFFFB7A860EFAF3FD85" author="Zhang, Y. - Q. & Yin, Z. - W." box="[1156,1375,612,638]" pageId="8" pageNumber="91" pagination="596 - 599" refId="ref5190" refString="Zhang, Y. - Q. & Yin, Z. - W. (2023) A new species of Tyrodes Raffray (Coleoptera: Staphylinidae: Pselaphinae) from China. Zootaxa, 5293 (3), 596 - 599. https: // doi. org / 10.11646 / zootaxa. 5293.3.11" type="journal article" year="2023">Zhang & Yin 2023</bibRefCitation>
|
||
</taxonomicName>
|
||
. This new species shares the elongate antennomeres 9–10 with
|
||
<taxonomicName id="4C2FD6A0FFD0FFFFFCE386E2FC2BFD5A" authorityName="Zhang & Yin" authorityYear="2023" box="[797,903,647,673]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="91" phylum="Arthropoda" rank="species" species="tibialis">
|
||
<emphasis id="B95B7131FFD0FFFFFCE386E2FC2BFD5A" box="[797,903,647,673]" italics="true" pageId="8" pageNumber="91">T. tibialis</emphasis>
|
||
</taxonomicName>
|
||
but the new species is distinguished by lacking a lateral expansion of the aedeagal median lobe and lacking the small projection of each metatibia.
|
||
</paragraph>
|
||
<paragraph id="8B90AD23FFD0FFFFFF3986A4FC2BFCA8" blockId="8.[151,1437,150,852]" pageId="8" pageNumber="91">
|
||
Among Japanese species, the new species is separated from
|
||
<taxonomicName id="4C2FD6A0FFD0FFFFFC6A86A5FC50FD13" authorityName="Kurbatov" authorityYear="1990" box="[916,1020,719,744]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="91" phylum="Arthropoda" rank="species" species="segrex">
|
||
<emphasis id="B95B7131FFD0FFFFFC6A86A5FC50FD13" box="[916,1020,719,744]" italics="true" pageId="8" pageNumber="91">T. segrex</emphasis>
|
||
</taxonomicName>
|
||
by a combination of the following characteristics: antennomeres 9 and 10 each longer than wide, metafemora simple, sternite IV with densely setose posterolateral patch in cavity; sternite IV–VII flat at middle (sternite IV–VII shallowly impress at middle with tufts of setae in
|
||
<taxonomicName id="4C2FD6A0FFD0FFFFFEEF8750FED9FCA8" authorityName="Kurbatov" authorityYear="1990" box="[273,373,826,851]" class="Insecta" family="Staphylinidae" genus="Tyrodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="91" phylum="Arthropoda" rank="species" species="segrex">
|
||
<emphasis id="B95B7131FFD0FFFFFEEF8750FED9FCA8" box="[273,373,826,851]" italics="true" pageId="8" pageNumber="91">T. segrex</emphasis>
|
||
</taxonomicName>
|
||
), and large and elongate aedeagal endophallus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |