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<mods:title id="F32F2869742B92D4FF6FC3D177F4C8D1">Taxonomic revision of Eunota togata (LaFerté-Sénectère, 1841) (Coleoptera: Cicindelidae) in North America with a new subspecies from western Texas and New Mexico, United States</mods:title>
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<paragraph id="8B956772FF80CB12BEA0FB9D2E32FBB4" blockId="13.[192,819,1045,1074]" box="[192,819,1045,1074]" pageId="13" pageNumber="12">
<heading id="D0DDD01EFF80CB12BEA0FB9D2E32FBB4" box="[192,819,1045,1074]" fontSize="12" level="2" pageId="13" pageNumber="12" reason="2">
<emphasis id="B95EBB60FF80CB12BEA0FB9D2E32FBB4" bold="true" box="[192,819,1045,1074]" pageId="13" pageNumber="12">
<taxonomicName id="4C2A1CF1FF80CB12BEA0FB9D2FA1FBB7" authority="(Willis, 1967)" authorityName="Acciavatti" authorityYear="2021" baseAuthorityName="Willis" baseAuthorityYear="1967" box="[192,672,1045,1074]" class="Insecta" family="Carabidae" genus="Eunota" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="subSpecies" species="togata" status="stat. nov." subSpecies="latilabris">
<emphasis id="B95EBB60FF80CB12BEA0FB9D2CEBFBB7" bold="true" box="[192,490,1045,1074]" italics="true" pageId="13" pageNumber="12">Eunota togata latilabris</emphasis>
(
<bibRefCitation id="EFBB1A83FF80CB12BF99FB9E2F96FBB7" author="Willis HL" box="[505,663,1045,1074]" pageId="13" pageNumber="12" pagination="145 - 313" refId="ref19135" refString="Willis HL. 1967. Bionomics and zoogeography of tiger beetles of saline habitats in the central United States (Coleoptera: Cicindelidae). The University of Kansas Science Bulletin 67 (5): 145 - 313." type="journal article" year="1967">Willis, 1967</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A26D061BFF80CB12BCCDFB9E2E32FBB4" box="[685,819,1045,1073]" pageId="13" pageNumber="12">new status</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33034F9FF80CB12BE80FBB42CDAFBF3" pageId="13" pageNumber="12" type="reference_group">
<paragraph id="8B956772FF80CB12BE80FBB42CDAFBF3" blockId="13.[224,1442,1087,1481]" pageId="13" pageNumber="12">
<treatmentCitationGroup id="AB3A405CFF80CB12BE80FBB42CDAFBF3" pageId="13" pageNumber="12">
<treatmentCitation id="0A8B4163FF80CB12BE80FBB42F96FBDD" author="Horn W." box="[224,663,1087,1112]" page="17" pageId="13" pageNumber="12" year="1897">
<taxonomicName id="4C2A1CF1FF80CB12BE80FBB42F96FBDD" ID-CoL="V8MB" authority="Horn 1897: 17" authorityName="Horn" authorityPageNumber="17" authorityYear="1897" baseAuthorityName="Horn" baseAuthorityYear="1897" box="[224,663,1087,1112]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="variety" species="togata" variety="apicalis">
<emphasis id="B95EBB60FF80CB12BE80FBB42C83FBD2" box="[224,386,1087,1111]" italics="true" pageId="13" pageNumber="12">Cicindela togata</emphasis>
var.
<emphasis id="B95EBB60FF80CB12BFD2FBB42CFDFBD2" box="[434,508,1087,1111]" italics="true" pageId="13" pageNumber="12">apicalis</emphasis>
<bibRefCitation id="EFBB1A83FF80CB12BC63FBCB2F96FBDD" author="Horn W." box="[515,663,1088,1112]" pageId="13" pageNumber="12" pagination="17 - 20" refId="ref18073" refString="Horn W. 1897. Drei neue Cicindelen und uber Neolaphra Bedel. Entomologische Nachrichten 23 (2): 17 - 20." type="journal article" year="1897">Horn 1897: 17</bibRefCitation>
</taxonomicName>
</treatmentCitation>
: [(
<treatmentCitation id="0A8B4163FF80CB12BCCDFBCB290BFBD2" author="Bousquet Y &amp; Larochelle A." box="[685,1034,1087,1112]" page="66" pageId="13" pageNumber="12" year="1993">
<bibRefCitation id="EFBB1A83FF80CB12BCCDFBCB290BFBD2" author="Bousquet Y &amp; Larochelle A." box="[685,1034,1087,1112]" pageId="13" pageNumber="12" refId="ref17535" refString="Bousquet Y, Larochelle A. 1993. Catalogue of the Geadephaga (Coleoptera: Trachypachidae, Rhysodidae, Carabidae including Cicindelini) of America North of Mexico. Memoirs of the Entomological Society of Canada, 167. 397 p." type="book" year="1993">Bousquet and Larochelle 1993: 66</bibRefCitation>
</treatmentCitation>
) as primary homonym of
<taxonomicName id="4C2A1CF1FF80CB12BB75FBB42CD3FBF3" ID-CoL="8L9NY" authority="Chaudoir, 1843" authorityName="Chaudoir" authorityYear="1843" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="B95EBB60FF80CB12BB75FBB42C2AFBF3" italics="true" pageId="13" pageNumber="12">Cicindela apicalis</emphasis>
<bibRefCitation id="EFBB1A83FF80CB12BF51FBD52CD3FBF3" author="Chaudoir" box="[305,466,1118,1142]" pageId="13" pageNumber="12" pagination="671 - 791" refId="ref17717" refString="Chaudoir M de. 1843. Carabiques nouveaux. Bulletin de la Societe Imperiale des Naturalistes de Moscou 16: 671 - 791." type="journal article" year="1843">Chaudoir, 1843</bibRefCitation>
</taxonomicName>
].
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</paragraph>
</subSubSection>
<subSubSection id="C33034F9FF80CB12BF60FBF52838FAC8" pageId="13" pageNumber="12" type="materials_examined">
<paragraph id="8B956772FF80CB12BF60FBF52838FAC8" blockId="13.[224,1442,1087,1481]" pageId="13" pageNumber="12">
<emphasis id="B95EBB60FF80CB12BF60FBF52C6FFB10" box="[256,366,1150,1173]" italics="true" pageId="13" pageNumber="12">Type status</emphasis>
.
<typeStatus id="5491D9D0FF80CB12BF1AFBF62CD5FB10" box="[378,468,1149,1173]" pageId="13" pageNumber="12" type="syntype">Syntypes</typeStatus>
(
<specimenCount id="9D2CACFBFF80CB12BF84FBF62F03FB12" box="[484,514,1149,1175]" pageId="13" pageNumber="12" type="female">1♀</specimenCount>
, other
<specimenCount id="9D2CACFBFF80CB12BC2AFBF52F5CFB12" box="[586,605,1150,1175]" pageId="13" pageNumber="12" type="male"></specimenCount>
<specimenCount id="9D2CACFBFF80CB12BC3DFBF52F71FB12" box="[605,624,1150,1175]" pageId="13" pageNumber="12" type="female"></specimenCount>
not enumerated) (
<bibRefCitation id="EFBB1A83FF80CB12BD4DFBF62EAEFB10" author="Dobler H." box="[813,943,1149,1173]" pageId="13" pageNumber="12" pagination="355 - 419" refId="ref17741" refString="Dobler H. 1973. Katalog der in den Sammlungen des ehemaligen Deutschen Entomologischen Institutes aufbewahrten Typen - IX, Coleoptera, Cicindelidae. Beitrage Entomologie 23: 355 - 419." type="journal article" year="1973">Döbler 1973</bibRefCitation>
;
<bibRefCitation id="EFBB1A83FF80CB12BDD8FBF62950FB10" author="Bousquet Y." box="[952,1105,1149,1173]" pageId="13" pageNumber="12" pagination="1 - 1722" refId="ref17475" refString="Bousquet Y. 2012. Catalogue of the Geadephaga of Canada and the United States. ZooKeys 245: 1 - 1722." type="journal article" year="2012">Bousquet 2012</bibRefCitation>
) [unexamined; concept based on specimens from Kackley, Kansas, collected by Knaus at
<collectionCode id="ED3BFFB7FF80CB12BD59FB172E8DFB31" box="[825,908,1180,1204]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="13" pageNumber="12" type="Museum">CMNH</collectionCode>
].
<emphasis id="B95EBB60FF80CB12BDFCFB172939FB31" box="[924,1080,1180,1204]" italics="true" pageId="13" pageNumber="12">Type depository</emphasis>
.
<collectionCode id="ED3BFFB7FF80CB12BA24FB172978FB31" box="[1092,1145,1180,1204]" pageId="13" pageNumber="12">SDEI</collectionCode>
[formerly
<collectionCode id="ED3BFFB7FF80CB12BA88FB172812FB36" box="[1256,1299,1180,1203]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="13" pageNumber="12" type="Other type of research institution/biorepository">DEI</collectionCode>
, Eberswalde],
<collectingCountry id="F33D27E2FF80CB12BF61FB302C61FB56" box="[257,352,1211,1235]" name="Germany" pageId="13" pageNumber="12">Germany</collectingCountry>
(
<bibRefCitation id="EFBB1A83FF80CB12BF10FB312CF5FB56" author="Dobler H." box="[368,500,1210,1235]" pageId="13" pageNumber="12" pagination="355 - 419" refId="ref17741" refString="Dobler H. 1973. Katalog der in den Sammlungen des ehemaligen Deutschen Entomologischen Institutes aufbewahrten Typen - IX, Coleoptera, Cicindelidae. Beitrage Entomologie 23: 355 - 419." type="journal article" year="1973">Döbler 1973</bibRefCitation>
;
<bibRefCitation id="EFBB1A83FF80CB12BC60FB302F9FFB56" author="Bousquet Y." box="[512,670,1211,1235]" pageId="13" pageNumber="12" pagination="1 - 1722" refId="ref17475" refString="Bousquet Y. 2012. Catalogue of the Geadephaga of Canada and the United States. ZooKeys 245: 1 - 1722." type="journal article" year="2012">Bousquet 2012</bibRefCitation>
).
<emphasis id="B95EBB60FF80CB12BCCFFB302E2FFB57" box="[687,814,1210,1234]" italics="true" pageId="13" pageNumber="12">Type locality</emphasis>
. Kakley (sic), Kansas [Kackley, Republic County]; Nebraska [in original description].
<emphasis id="B95EBB60FF80CB12BC64FB512FB0FB74" box="[516,689,1241,1265]" italics="true" pageId="13" pageNumber="12">Type locality note</emphasis>
: This taxon type locality (
<figureCitation id="13117BF7FF80CB12BDDBFB522905FB74" box="[955,1028,1241,1265]" captionStart="Figures 4245" captionStartId="15.[192,279,1129,1155]" captionTargetBox="[204,1430,221,1097]" captionTargetId="figure-9@15.[194,1442,213,1107]" captionTargetPageId="15" captionText="Figures 4245. Eunota togata described taxa type localities. 42) E. t. togata, TL: Port Isabel, Cameron Co., Texas; coastal saltwater estuaries. 43) E. t. latilabris, TL: Kackley, Republic Co., Kansas; salt marsh and brine pond on Saint Johns Creek. 44) E. t. globicollis, TL: Englewood, Clark Co., Kansas; saline banks of Cimarron River. 45) E. t. fascinans, TL: Santa Rosa, Guadalupe Co., New Mexico; saline soils from freshwater ponds above Pecos River." figureDoi="http://doi.org/10.5281/zenodo.5041468" httpUri="https://zenodo.org/record/5041468/files/figure.png" pageId="13" pageNumber="12">Fig. 43</figureCitation>
,
<figureCitation id="13117BF7FF80CB12BA6FFB522928FB74" box="[1039,1065,1241,1265]" captionStart="Figure 50" captionStartId="19.[193,269,1833,1859]" captionTargetBox="[192,1441,952,1802]" captionTargetId="figure-393@19.[189,1444,949,1804]" captionTargetPageId="19" captionText="Figure 50. Eunota togata geographical distribution in the United States from study specimens. = E. t. togata;" figureDoi="http://doi.org/10.5281/zenodo.5041472" httpUri="https://zenodo.org/record/5041472/files/figure.png" pageId="13" pageNumber="12">50</figureCitation>
) is herein fixed at a site within a salt marsh surrounding a small pond southwest of Kackley, Republic County, Kansas, with these GPS coordinates:
<geoCoordinate id="EE1E01B5FF80CB12BB2DFB73289FFA95" box="[1357,1438,1272,1296]" degrees="39.6897" direction="north" orientation="latitude" pageId="13" pageNumber="12" precision="5" value="39.6897">39.6897</geoCoordinate>
,
<geoCoordinate id="EE1E01B5FF80CB12BE9FFA9C2C5FFAAA" box="[255,350,1303,1327]" degrees="97.9281" direction="west" orientation="longitude" pageId="13" pageNumber="12" precision="5" value="-97.9281">97.9281</geoCoordinate>
,
<specimenCount id="9D2CACFBFF80CB12BF0AFA9D2CABFAAB" box="[362,426,1302,1326]" pageId="13" pageNumber="12" type="generic">
<quantity id="4CD2CA97FF80CB12BF0AFA9D2CA7FAAB" box="[362,422,1302,1326]" metricMagnitude="2" metricUnit="m" metricValue="4.26" pageId="13" pageNumber="12" unit="m" value="426.0">426m</quantity>
.
</specimenCount>
This salt marsh is along Saint Johns Creek within Jamestown, Kansas, Wildlife Management Area.
<emphasis id="B95EBB60FF80CB12BF62FABE2C4DFAC8" box="[258,332,1333,1357]" italics="true" pageId="13" pageNumber="12">Habitat</emphasis>
: Small openings with bare, salt encrusted soils around small pond within salt marsh at type locality.
</paragraph>
</subSubSection>
<subSubSection id="C33034F9FF80CB12BE80FADF2866FA4D" pageId="13" pageNumber="12" type="reference_group">
<paragraph id="8B956772FF80CB12BE80FADF283FFAE9" blockId="13.[224,1442,1087,1481]" box="[224,1342,1364,1388]" pageId="13" pageNumber="12">
<treatmentCitationGroup id="AB3A405CFF80CB12BE80FADF283FFAE9" box="[224,1342,1364,1388]" pageId="13" pageNumber="12">
<taxonomicName id="4C2A1CF1FF80CB12BE80FADF2F5FFAE9" authority="Willis, 1967" authorityName="Willis" authorityYear="1967" box="[224,606,1364,1388]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="subSpecies" species="togata" subSpecies="latilabris">
<emphasis id="B95EBB60FF80CB12BE80FADF2CDEFAE9" box="[224,479,1364,1388]" italics="true" pageId="13" pageNumber="12">Cicindela togata latilabris</emphasis>
<bibRefCitation id="EFBB1A83FF80CB12BF84FADF2F5FFAE9" author="Willis HL" box="[484,606,1364,1388]" pageId="13" pageNumber="12" pagination="145 - 313" refId="ref19135" refString="Willis HL. 1967. Bionomics and zoogeography of tiger beetles of saline habitats in the central United States (Coleoptera: Cicindelidae). The University of Kansas Science Bulletin 67 (5): 145 - 313." type="journal article" year="1967">Willis, 1967</bibRefCitation>
</taxonomicName>
: [(
<treatmentCitation id="0A8B4163FF80CB12BC14FADF2E1DFAE9" author="Willis HL" box="[628,796,1364,1388]" page="286" pageId="13" pageNumber="12" year="1967">
<bibRefCitation id="EFBB1A83FF80CB12BC14FADF2E1DFAE9" author="Willis HL" box="[628,796,1364,1388]" pageId="13" pageNumber="12" pagination="145 - 313" refId="ref19135" refString="Willis HL. 1967. Bionomics and zoogeography of tiger beetles of saline habitats in the central United States (Coleoptera: Cicindelidae). The University of Kansas Science Bulletin 67 (5): 145 - 313." type="journal article" year="1967">Willis 1967: 286</bibRefCitation>
</treatmentCitation>
) replacement name for
<taxonomicName id="4C2A1CF1FF80CB12BA6CFADF2835FAE9" ID-CoL="V8MB" authority="Horn, 1897" authorityName="Horn" authorityYear="1897" baseAuthorityName="Horn" baseAuthorityYear="1897" box="[1036,1332,1364,1388]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="subSpecies" species="togata" subSpecies="apicalis">
<emphasis id="B95EBB60FF80CB12BA6CFADF29B7FAE9" box="[1036,1206,1364,1388]" italics="true" pageId="13" pageNumber="12">C. togata apicalis</emphasis>
<bibRefCitation id="EFBB1A83FF80CB12BADCFADF2835FAE9" author="Horn W." box="[1212,1332,1364,1388]" pageId="13" pageNumber="12" pagination="17 - 20" refId="ref18073" refString="Horn W. 1897. Drei neue Cicindelen und uber Neolaphra Bedel. Entomologische Nachrichten 23 (2): 17 - 20." type="journal article" year="1897">Horn, 1897</bibRefCitation>
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B956772FF80CB12BE80FAF82C93FA2F" blockId="13.[224,1442,1087,1481]" pageId="13" pageNumber="12">
<treatmentCitationGroup id="AB3A405CFF80CB12BE80FAF82C93FA2F" pageId="13" pageNumber="12">
<taxonomicName id="4C2A1CF1FF80CB12BE80FAF82F65FA0E" authority="Willis, 1967" authorityName="Willis" authorityYear="1967" box="[224,612,1395,1419]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="subSpecies" species="togata" subSpecies="latilabris">
<emphasis id="B95EBB60FF80CB12BE80FAF82CE0FA0E" box="[224,481,1395,1419]" italics="true" pageId="13" pageNumber="12">Cicindela togata latilabris</emphasis>
<bibRefCitation id="EFBB1A83FF80CB12BF88FAF82F65FA0E" author="Willis HL" box="[488,612,1395,1419]" pageId="13" pageNumber="12" pagination="145 - 313" refId="ref19135" refString="Willis HL. 1967. Bionomics and zoogeography of tiger beetles of saline habitats in the central United States (Coleoptera: Cicindelidae). The University of Kansas Science Bulletin 67 (5): 145 - 313." type="journal article" year="1967">Willis, 1967</bibRefCitation>
</taxonomicName>
: [(
<treatmentCitation id="0A8B4163FF80CB12BC1BFAF82EDBFA0E" author="Bousquet Y &amp; Larochelle A." box="[635,986,1395,1419]" page="66" pageId="13" pageNumber="12" year="1993">
<bibRefCitation id="EFBB1A83FF80CB12BC1BFAF82EDBFA0E" author="Bousquet Y &amp; Larochelle A." box="[635,986,1395,1419]" pageId="13" pageNumber="12" refId="ref17535" refString="Bousquet Y, Larochelle A. 1993. Catalogue of the Geadephaga (Coleoptera: Trachypachidae, Rhysodidae, Carabidae including Cicindelini) of America North of Mexico. Memoirs of the Entomological Society of Canada, 167. 397 p." type="book" year="1993">Bousquet and Larochelle 1993: 66</bibRefCitation>
</treatmentCitation>
) assigned as synonym of
<taxonomicName id="4C2A1CF1FF80CB12BABFFAF82C8AFA2F" authority="(Casey, 1913)" baseAuthorityName="Casey" baseAuthorityYear="1913" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="subSpecies" species="togata" subSpecies="globicollis">
<emphasis id="B95EBB60FF80CB12BABFFAF828A0FA0E" box="[1247,1441,1395,1419]" italics="true" pageId="13" pageNumber="12">C. togata globicollis</emphasis>
(
<bibRefCitation id="EFBB1A83FF80CB12BF68FA192C84FA2F" author="Casey TL" box="[264,389,1426,1450]" pageId="13" pageNumber="12" refId="ref17628" refString="Casey TL. 1913. Memoirs on the Coleoptera. IV. The New Era Printing Company; Lancaster, PA. 400 p." type="book" year="1913">Casey, 1913</bibRefCitation>
)
</taxonomicName>
]
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B956772FF80CB12BE80FA3A2866FA4D" blockId="13.[224,1442,1087,1481]" box="[224,1383,1456,1481]" pageId="13" pageNumber="12">
<taxonomicName id="4C2A1CF1FF80CB12BE80FA3A2F5BFA4D" authority="(Willis, 1967)" authorityName="Acciavatti" authorityYear="2021" baseAuthorityName="Willis" baseAuthorityYear="1967" box="[224,602,1456,1480]" class="Insecta" family="Carabidae" genus="Eunota" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="subSpecies" species="togata" subSpecies="latilabris">
<emphasis id="B95EBB60FF80CB12BE80FA3A2CC8FA4D" box="[224,457,1456,1480]" italics="true" pageId="13" pageNumber="12">Eunota togata latilabris</emphasis>
(
<bibRefCitation id="EFBB1A83FF80CB12BFB6FA3B2F52FA4D" author="Willis HL" box="[470,595,1456,1480]" pageId="13" pageNumber="12" pagination="145 - 313" refId="ref19135" refString="Willis HL. 1967. Bionomics and zoogeography of tiger beetles of saline habitats in the central United States (Coleoptera: Cicindelidae). The University of Kansas Science Bulletin 67 (5): 145 - 313." type="journal article" year="1967">Willis, 1967</bibRefCitation>
)
</taxonomicName>
: [(
<bibRefCitation id="EFBB1A83FF80CB12BC10FA3B2E0CFA4D" author="Bousquet Y." box="[624,781,1456,1480]" pageId="13" pageNumber="12" pagination="1 - 1722" refId="ref17475" refString="Bousquet Y. 2012. Catalogue of the Geadephaga of Canada and the United States. ZooKeys 245: 1 - 1722." type="journal article" year="2012">Bousquet 2012</bibRefCitation>
) assigned as synonym of
<taxonomicName id="4C2A1CF1FF80CB12BA6CFA3A285CFA4D" ID-CoL="7H5LW" authority="(Casey, 1913)" baseAuthorityName="Casey" baseAuthorityYear="1913" box="[1036,1373,1456,1480]" class="Insecta" family="Carabidae" genus="Eunota" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="subSpecies" species="togata" subSpecies="globicollis">
<emphasis id="B95EBB60FF80CB12BA6CFA3A29CBFA4D" box="[1036,1226,1456,1480]" italics="true" pageId="13" pageNumber="12">E. togata globicollis</emphasis>
(
<bibRefCitation id="EFBB1A83FF80CB12BAB7FA3B2854FA4D" author="Casey TL" box="[1239,1365,1456,1480]" pageId="13" pageNumber="12" refId="ref17628" refString="Casey TL. 1913. Memoirs on the Coleoptera. IV. The New Era Printing Company; Lancaster, PA. 400 p." type="book" year="1913">Casey, 1913</bibRefCitation>
)
</taxonomicName>
].
</paragraph>
</subSubSection>
</treatment>
</document>