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<mods:title id="43378FC7F6AFD01651A915EE8722E275">Open-holed trapdoor spiders of the genus Teyl (Mygalomorphae: Nemesiidae Anamini) from Western Australias Pilbara bioregion: a new species and expanded phylogenetic assessment</mods:title>
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Family
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Nemesiidae
<bibRefCitation id="EFBB6EDD267DFFAFFE64B0E1637B1EB0" author="Simon, E." box="[397,551,225,251]" pageId="6" pageNumber="355" pagination="169 - 220" refId="ref8044" refString="Simon, E. (1889) Arachnides. In: Voyage de M. E. Simon au Venezuela (decembre 1887 - avril 1888). 4 e Memoire. Annales de la Soci e t e Entomologique de France, Series 6, 9, 169 - 220." type="journal article" year="1889">Simon, 1889</bibRefCitation>
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Tribe Anamini
<bibRefCitation id="EFBB6EDD267DFFAFFEBCB12960B31F08" author="Simon, E." box="[341,495,297,323]" pageId="6" pageNumber="355" pagination="169 - 220" refId="ref8044" refString="Simon, E. (1889) Arachnides. In: Voyage de M. E. Simon au Venezuela (decembre 1887 - avril 1888). 4 e Memoire. Annales de la Soci e t e Entomologique de France, Series 6, 9, 169 - 220." type="journal article" year="1889">Simon, 1889</bibRefCitation>
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Anamae
<bibRefCitation id="EFBB6EDD267DFFAFFF05B16D60F81FCD" author="Simon, E." box="[236,420,365,390]" pageId="6" pageNumber="355" pagination="169 - 220" refId="ref8044" refString="Simon, E. (1889) Arachnides. In: Voyage de M. E. Simon au Venezuela (decembre 1887 - avril 1888). 4 e Memoire. Annales de la Soci e t e Entomologique de France, Series 6, 9, 169 - 220." type="journal article" year="1889">Simon, 1889: 178</bibRefCitation>
.
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genus
<taxonomicName id="4C2A68AF267DFFAFFDCEB16E62561FCD" authority="L. Koch, 1873" authorityName="L. Koch" authorityYear="1873" box="[551,778,365,390]" class="Arachnida" family="Nemesiidae" genus="Aname" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="355" phylum="Arthropoda" rank="genus">
<emphasis id="B95ECF3E267DFFAFFDCEB16E63311FCE" box="[551,621,366,389]" italics="true" pageId="6" pageNumber="355">Aname</emphasis>
L.
<bibRefCitation id="EFBB6EDD267DFFAFFD79B16D62561FCD" author="Koch, L." box="[656,778,365,390]" pageId="6" pageNumber="355" refId="ref7198" refString="Koch, L. (1873) Die Arachniden Australiens, Nach der Natur Beschrieben und Abgebildet. Bauer &amp; Raspe, Nurnberg, 718 pp." type="book" year="1873">Koch, 1873</bibRefCitation>
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.
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<emphasis id="B95ECF3E267DFFAFFF7EB1B460571F85" bold="true" box="[151,267,436,462]" pageId="6" pageNumber="355">Remarks.</emphasis>
The tribe Anamini is an unambiguously monophyletic assemblage of nine genera, characterized by a strongly developed posterior maxillary heel bearing a field of cuspules (
<figureCitation id="13110FA9267DFFAFFC29B1D8655F1FB9" box="[960,1027,472,498]" captionStart="FIGURES 413" captionStartId="10.[151,264,1902,1927]" captionTargetBox="[164,1423,166,1871]" captionTargetId="figure@10.[164,1423,166,1877]" captionTargetPageId="10" captionText="FIGURES 413. Teyl heuretes sp. nov., male holotype (WAM T91918) from Weeli Wolli Creek region, Hamersley Range (Western Australia; PIL), somatic morphology: 45, carapace and abdomen, dorsal view; 6, cephalothorax, lateral view; 7, eyes, dorsal view; 8, mouthparts, ventral view; 910, cephalothorax and abdomen, ventral view; 11, leg I, prolateral view; 12, leg I tibia, prolateral view; 13, leg I tibia, retrolateral view. Scale bars = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3460200" httpUri="https://zenodo.org/record/3460200/files/figure.png" pageId="6" pageNumber="355">Fig. 8</figureCitation>
), in combination with an absence of tarsal claw tufts, and two rows of teeth on the paired claws (both family-level characters; see
<bibRefCitation id="EFBB6EDD267DFFAFFB5DB1FD64621C5D" author="Raven, R. J." box="[1204,1342,507,535]" pageId="6" pageNumber="355" pagination="1 - 180" refId="ref7757" refString="Raven, R. J. (1985) The spider infraorder Mygalomorphae (Araneae): cladistics and systematics. Bulletin of the American Museum of Natural History, 182, 1 - 180." type="journal article" year="1985">Raven 1985</bibRefCitation>
;
<bibRefCitation id="EFBB6EDD267DFFAFFAA3B1FD60481C71" author="Harvey, M. S. &amp; Hillyer, M. J. &amp; Main, B. Y. &amp; Moulds, T. A. &amp; Raven, R. J. &amp; Rix, M. G. &amp; Vink, C. J. &amp; Huey, J. A." pageId="6" pageNumber="355" pagination="407 - 452" refId="ref6824" refString="Harvey, M. S., Hillyer, M. J., Main, B. Y., Moulds, T. A., Raven, R. J., Rix, M. G., Vink, C. J. &amp; Huey, J. A. (2018) Phylogenetic relationships of the Australasian open-holed trapdoor spiders (Araneae: Mygalomorphae: Nemesiidae: Anaminae): multilocus molecular analyses resolve the generic classification of a highly diverse fauna. Zoological Journal of the Linnean Society, 184, 407 - 452. https: // doi. org / 10.1093 / zoolinnean / zlx 111" type="journal article" year="2018">
Harvey
<emphasis id="B95ECF3E267DFFAFFF7EB221618C1C71" box="[151,208,544,570]" italics="true" pageId="6" pageNumber="355">et al.</emphasis>
2018
</bibRefCitation>
). The burrows of these spiders are almost exclusively open, and often include a secondary escape shaft which branches off from the burrow proper (hence the other commonly applied name of wishbone spiders; see
<bibRefCitation id="EFBB6EDD267DFFAFFF7EB268604A1CC9" author="Main, B. Y." box="[151,278,616,642]" pageId="6" pageNumber="355" pagination="599 - 606" refId="ref7551" refString="Main, B. Y. (1993) From flood avoidance to foraging: adaptive shifts in trapdoor spider behaviour. Memoirs of the Queensland Museum, 33, 599 - 606." type="journal article" year="1993">Main 1993</bibRefCitation>
). The tribe is distributed Australia-wide, and the Australian fauna is remarkably diverse, with 100200 new species (and likely many more) still to be described (
<bibRefCitation id="EFBB6EDD267DFFAFFCF3B28C657E1CED" author="Castalanelli, M. A. &amp; Teale, R. &amp; Rix, M. G. &amp; Kennington, W. J. &amp; Harvey, M. S." box="[794,1058,652,678]" pageId="6" pageNumber="355" pagination="375 - 385" refId="ref6576" refString="Castalanelli, M. A., Teale, R., Rix, M. G., Kennington, W. J. &amp; Harvey, M. S. (2014) Barcoding of mygalomorph spiders (Araneae: Mygalomorphae) in the Pilbara bioregion of Western Australia reveals a highly diverse biota. Invertebrate Systematics, 28, 375 - 385. https: // doi. org / 10.1071 / IS 13058" type="journal article" year="2014">
Castalanelli
<emphasis id="B95ECF3E267DFFAFFC4EB28D62831CED" box="[935,991,652,678]" italics="true" pageId="6" pageNumber="355">et al.</emphasis>
2014
</bibRefCitation>
;
<bibRefCitation id="EFBB6EDD267DFFAFFBC5B28D65A31CED" author="Harvey, M. S. &amp; Hillyer, M. J. &amp; Main, B. Y. &amp; Moulds, T. A. &amp; Raven, R. J. &amp; Rix, M. G. &amp; Vink, C. J. &amp; Huey, J. A." box="[1068,1279,652,679]" pageId="6" pageNumber="355" pagination="407 - 452" refId="ref6824" refString="Harvey, M. S., Hillyer, M. J., Main, B. Y., Moulds, T. A., Raven, R. J., Rix, M. G., Vink, C. J. &amp; Huey, J. A. (2018) Phylogenetic relationships of the Australasian open-holed trapdoor spiders (Araneae: Mygalomorphae: Nemesiidae: Anaminae): multilocus molecular analyses resolve the generic classification of a highly diverse fauna. Zoological Journal of the Linnean Society, 184, 407 - 452. https: // doi. org / 10.1093 / zoolinnean / zlx 111" type="journal article" year="2018">
Harvey
<emphasis id="B95ECF3E267DFFAFFB6FB28D65EB1CED" box="[1158,1207,652,678]" italics="true" pageId="6" pageNumber="355">et al</emphasis>
. 2018
</bibRefCitation>
; MSH, MGR, JAH, unpublished data).
</paragraph>
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