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<document id="A55EDB5613F99ACBAF63767B58EFF82B" ID-CLB-Dataset="8225" ID-DOI="10.11646/zootaxa.4860.1.3" ID-GBIF-Dataset="a333c0b8-de3f-4f3b-a9b9-05453686efc4" ID-ISSN="1175-5326" ID-Zenodo-Dep="4413508" ID-ZooBank="38F1E2A5-5DE0-4B95-959A-6347BC593AF0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609700513105" checkinUser="plazi" docAuthor="Mielke, Carlos G. C., Grehan, John R. &amp; Monzón-Sierra, José" docDate="2020" docId="0383282C7E25FFE94390FC5B474019BB" docLanguage="en" docName="zootaxa.4860.1.3.pdf" docOrigin="Zootaxa 4860 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Schausiana maishei Mielke &amp; Grehan &amp; Monzón-Sierra 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="75" masterDocId="FFBA50547E23FFE14307FFAD431F1D1A" masterDocTitle="Taxonomic revision of Schausiana Viette with two new species from Guatemala and notes on biogeography and correlated tectonics (Lepidoptera: Hepialidae)" masterLastPageNumber="91" masterPageNumber="67" pageNumber="73" updateTime="1698851864825" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="7A72F89CBC9CBB3AF317787120F403C8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E5779150C641E77F643348A2DB25EF69">
<mods:title id="D976881C39293B2A6C5D108F0F3E8010">Taxonomic revision of Schausiana Viette with two new species from Guatemala and notes on biogeography and correlated tectonics (Lepidoptera: Hepialidae)</mods:title>
</mods:titleInfo>
<mods:name id="0FD2856844AC7563D28AFF58575AF039" type="personal">
<mods:role id="5275070CA5A7D53E52174D104C13FF8D">
<mods:roleTerm id="4399DFEE5B35C8AE46AD1FD7D7E643A9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2354BD97F799C4087F22D3EC72DA7D96">Mielke, Carlos G. C.</mods:namePart>
<mods:affiliation id="1C24A445C0B7FB73CB3FDD4F1558B068">Caixa postal 1206, 84.145 - 000 Carambeí, Paraná, Brazil.</mods:affiliation>
</mods:name>
<mods:name id="BF1F203724BCCFC1ABA0EBD8D67111B9" type="personal">
<mods:role id="1F7A413840C2A09F860874E9B70D90E4">
<mods:roleTerm id="6C7553B058975E19E3DD481410BB9E4B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0003B0AC27F5A3CB8BB55E657C200FFC">Grehan, John R.</mods:namePart>
<mods:affiliation id="97A6ECB7E6D66054BA1FA61EC8387D7D">Research Associate, McGuire Center for Lepidoptera &amp; Biodiversity, Gainesville, FL 32611, USA.</mods:affiliation>
</mods:name>
<mods:name id="D45B2371B3A70BD759072720C8EF30B0" type="personal">
<mods:role id="2D8A1C33D78738A3D8757E4858BBD5EC">
<mods:roleTerm id="3A628A3B01E6D94B3674587DE63F69DC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9408B9282A19EFC7D76E48364C2E7FBB">Monzón-Sierra, José</mods:namePart>
<mods:nameIdentifier id="9A818CC03EC979750B3F0AB37EE3DEF1" type="ORCID">0000-0002-0733-1895</mods:nameIdentifier>
<mods:affiliation id="1551D9492C4353F57ED11439D06DA336">Centro de Estudios Ambientales y Biodiversidad, Universidad del Valle de Guatemala, 18 Avenida 11 - 95 Zona 15, Vista Hermosa III, Guatemala, Guatemala, C. A. jmonzon @ uvg. edu. gt; https: // orcid. org / 0000 - 0002 - 0733 - 1895</mods:affiliation>
<mods:nameIdentifier id="001CBFF60C1B8755650604FC29067EB7" type="email">jmonzon@uvg.edu.gt</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="D2D5E92581ACD9FCD53D7A96973379ED">text</mods:typeOfResource>
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<mods:titleInfo id="94E07500B97559B7A3C521ACCD1ADB5F">
<mods:title id="4780760C59D8EA63222440ACD3C07CB9">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0BFB134B8E90FEB8527D5BFB14B67E1F">
<mods:date id="18043553C3BE96D026AFACCFD2D1CE25">2020</mods:date>
<mods:detail id="58F021C52FEF063E42FB9F1BB2B91685" type="pubDate">
<mods:number id="60D7549748A4119940DF649C93EBC83A">2020-10-09</mods:number>
</mods:detail>
<mods:detail id="F091463C2D8123720DCBD4875834BBDD" type="volume">
<mods:number id="EF5A80BD910ADF783913D7EF74190833">4860</mods:number>
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<mods:number id="9BF6FF60C9ED915390CB38033E78056F">1</mods:number>
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<mods:start id="02FC7B0C50BAE17D855E115E72857652">67</mods:start>
<mods:end id="DE61043A69F611370B78E6312EEFB673">91</mods:end>
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<mods:classification id="7ABE969E8199E6BB6F287B24FE00E320">journal article</mods:classification>
<mods:identifier id="D7E6162805180142B0CF1C67532CEA63" type="CLB-Dataset">8225</mods:identifier>
<mods:identifier id="FBD758CAE4FF56148225FCC87766EFE4" type="DOI">10.11646/zootaxa.4860.1.3</mods:identifier>
<mods:identifier id="5CF9EB4ABA22E7900F783E0C53AAB802" type="GBIF-Dataset">a333c0b8-de3f-4f3b-a9b9-05453686efc4</mods:identifier>
<mods:identifier id="F0ADC0BE39A60C59350A8B2B25FD2657" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EEB0D38E246DA0346FCCB5343D23F04D" type="Zenodo-Dep">4413508</mods:identifier>
<mods:identifier id="211D14F317C9587BD3C93E15647248C8" type="ZooBank">38F1E2A5-5DE0-4B95-959A-6347BC593AF0</mods:identifier>
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<treatment id="0383282C7E25FFE94390FC5B474019BB" ID-DOI="http://doi.org/10.5281/zenodo.4413536" ID-GBIF-Taxon="177324807" ID-Zenodo-Dep="4413536" LSID="urn:lsid:plazi:treatment:0383282C7E25FFE94390FC5B474019BB" httpUri="http://treatment.plazi.org/id/0383282C7E25FFE94390FC5B474019BB" lastPageId="8" lastPageNumber="75" pageId="6" pageNumber="73">
<subSubSection id="C330CAB17E25FFE74390FC5B42EF190B" box="[151,496,1014,1041]" pageId="6" pageNumber="73" type="nomenclature">
<paragraph id="8B95993A7E25FFE74390FC5B42EF190B" blockId="6.[151,496,1014,1077]" box="[151,496,1014,1041]" pageId="6" pageNumber="73">
<heading id="D0DD2E567E25FFE74390FC5B42EF190B" bold="true" box="[151,496,1014,1041]" fontSize="11" level="1" pageId="6" pageNumber="73" reason="1">
<emphasis id="B95E45287E25FFE74390FC5B42EF190B" bold="true" box="[151,496,1014,1041]" pageId="6" pageNumber="73">
<taxonomicName id="4C2AE2B97E25FFE74390FC5B4294190B" authority="Mielke &amp; Grehan &amp; Monzón-Sierra, 2020" authorityName="Mielke &amp; Grehan &amp; Monzón-Sierra" authorityYear="2020" box="[151,395,1014,1041]" class="Insecta" family="Hepialidae" genus="Schausiana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="73" phylum="Arthropoda" rank="species" species="maishei" status="sp. nov.">
<emphasis id="B95E45287E25FFE74390FC5B4294190B" bold="true" box="[151,395,1014,1041]" italics="true" pageId="6" pageNumber="73">Schausiana maishei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26DF8537E25FFE74295FC5A42EF190B" box="[402,496,1015,1041]" pageId="6" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C330CAB17E25FFE74390FBB642AD192F" box="[151,434,1050,1077]" pageId="6" pageNumber="73" type="description">
<paragraph id="8B95993A7E25FFE74390FBB642AD192F" blockId="6.[151,496,1014,1077]" box="[151,434,1050,1077]" pageId="6" pageNumber="73">
(
<figureCitation id="131185BF7E25FFE74398FBB643F8192F" box="[159,231,1051,1077]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1864,1888]" captionTargetBox="[156,1431,194,1830]" captionTargetId="figure-26@4.[156,1432,181,1841]" captionTargetPageId="4" captionText="FIGURES 1–5. Schausiana spp. FW areas and specialized scales. Areas: 1, S. trojesa ♀. Specialized scales: 2, S. trojesa ♀; 3, S. maishei, sp. nov. ♀; 4, S. chalciope, sp. nov. ♀; 5. S. pharus ♀. Arrows indicate the position of such scales." figureDoi="http://doi.org/10.5281/zenodo.4413510" httpUri="https://zenodo.org/record/4413510/files/figure.png" pageId="6" pageNumber="73">Figs 3</figureCitation>
,
<figureCitation id="131185BF7E25FFE743F3FBB64232192F" box="[244,301,1051,1077]" captionStart="FIGURES 9–11" captionStartId="7.[151,264,1945,1969]" captionTargetBox="[232,1410,193,1899]" captionTargetId="figure-20@7.[153,1434,181,1921]" captionTargetPageId="7" captionText="FIGURES 9–11. Schausiana maishei, sp. nov. 9, ♂ PT (CGCM 32.071): 9a, dorsal view; 9b, ventral view (CGCM). 10, ♀ PT (CGCM 36.646): 10a, dorsal view; 10b, ventral view (CGCM). 11, ♀ HT: dorsal view (UVG). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4413514" httpUri="https://zenodo.org/record/4413514/files/figure.png" pageId="6" pageNumber="73">9–11</figureCitation>
,
<figureCitation id="131185BF7E25FFE7423EFBB64248192F" box="[313,343,1051,1077]" captionStart="FIGURES 22–27" captionStartId="14.[151,264,1900,1924]" captionTargetBox="[159,1418,181,1870]" captionTargetId="figure-26@14.[158,1429,181,1877]" captionTargetPageId="14" captionText="FIGURES 22–27. Schausiana spp. labial palpi, legs and wing venation. 22–23, labial palpi of S. trojesa: 22, ♀ (Morelos, Cuernavaca (UNAM)); 23, ♂ (Morelos, Tepoztlán (UNAM)). Unscaled. 24–25, legs: 24, S. maishei, sp. nov. (CGCM 23.236); 25, S. pharus (CGCM 7.110). Scale bar = 5 mm. 26–27, wing venation of S. trojesa: 26, ♂ (arrows indicating 1A incomplete and 2A+3A); 27, ♀ (arrow indicating CuP mesally incomplete)." figureDoi="http://doi.org/10.5281/zenodo.4413522" httpUri="https://zenodo.org/record/4413522/files/figure.png" pageId="6" pageNumber="73">24</figureCitation>
,
<figureCitation id="131185BF7E25FFE74264FBB6429E192F" box="[355,385,1051,1077]" captionStart="FIGURES 28–33" captionStartId="15.[151,264,1922,1946]" captionTargetBox="[179,1396,633,1892]" captionTargetId="figure-214@15.[167,1420,633,1898]" captionTargetPageId="15" captionText="FIGURES 28–33. Schausiana spp. abdomen (upper side, tergites; lower side, sternites). 28–29, S. trojesa: 28, ♂ (CGCM 39417); 29, ♀ (CGCM 33.184). 30, S. maishei, sp. nov., ♂ PT (CGCM 23.236). 31, S. chalciope, sp. nov., ♀ PT (CGCM 41.258). 32–33, S. pharus: 32, ♂ (CGCM 7.110); 33, ♀ (CGCM 5.797). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4413524" httpUri="https://zenodo.org/record/4413524/files/figure.png" pageId="6" pageNumber="73">30</figureCitation>
,
<figureCitation id="131185BF7E25FFE7428AFBB742B5192F" box="[397,426,1050,1077]" captionStart="FIGURES 34–36" captionStartId="16.[151,264,1795,1819]" captionTargetBox="[264,1317,196,1761]" captionTargetId="figure-26@16.[242,1345,181,1771]" captionTargetPageId="16" captionText="FIGURES 34–36. Schausiana spp. ♂ genitalia. 34, S. trojesa (CGCM 39.417): 34a, ventral view; 34b, ventral view with base of valva exposed (arrow indicates the hook at the base of the valva); 34c, dorsal view (internal; arrow indicating the mesal ridge of fultura inferior); 34d, lateral view. 35, S. maishei, sp. nov., ♂ PT (CGCM 23.236): 35a, ventral view (arrow indicates the hook at the base of the valva); 35b, ventral view with pseudotegumen moved upwards (arrow indicating the mesal ridge of fultura inferior); 35c, dorsal view (internal); 35d, lateral view. 36, S. pharus (CGCM 7.110): 36a, ventral view (arrow indicates the hook at the base of the valva); 36b, dorsal view (internal; arrow indicating the mesal ridge of fultura inferior); 36c, lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4413526" httpUri="https://zenodo.org/record/4413526/files/figure.png" pageId="6" pageNumber="73">35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C330CAB17E25FFE74390FBCE461218DB" pageId="6" pageNumber="73" type="materials_examined">
<paragraph id="8B95993A7E25FFE74390FBCE474419DF" blockId="6.[151,1437,1122,2014]" pageId="6" pageNumber="73">
<emphasis id="B95E45287E25FFE74390FBCE425B1966" bold="true" box="[151,324,1122,1149]" pageId="6" pageNumber="73">Type material</emphasis>
.
<materialsCitation id="3B4293677E25FFE74250FBCE468C19BB" ID-GBIF-Occurrence="3034649321" collectingDate="2017-04-22" collectorName=" Monzon &amp; Naumann" country="Guatemala" elevation="2600" latitude="14.518525" location="Finca El Pilar" longLatPrecision="1" longitude="-90.691246" municipality="San Cristobal El Bajo" pageId="6" pageNumber="73" specimenCount="1" specimenCount-female="1" stateProvince="Sacatepequez" typeStatus="holotype">
<typeStatus id="549127987E25FFE74250FBCE42A01967" box="[343,447,1123,1149]" pageId="6" pageNumber="73" type="holotype">Holotype</typeStatus>
<specimenCount id="9D2C52B37E25FFE742CAFBCE42FD1967" box="[461,482,1123,1149]" pageId="6" pageNumber="73" type="female">♀</specimenCount>
(
<figureCitation id="131185BF7E25FFE742F0FBCE41501967" box="[503,591,1123,1149]" captionStart="FIGURES 9–11" captionStartId="7.[151,264,1945,1969]" captionTargetBox="[232,1410,193,1899]" captionTargetId="figure-20@7.[153,1434,181,1921]" captionTargetPageId="7" captionText="FIGURES 9–11. Schausiana maishei, sp. nov. 9, ♂ PT (CGCM 32.071): 9a, dorsal view; 9b, ventral view (CGCM). 10, ♀ PT (CGCM 36.646): 10a, dorsal view; 10b, ventral view (CGCM). 11, ♀ HT: dorsal view (UVG). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4413514" httpUri="https://zenodo.org/record/4413514/files/figure.png" pageId="6" pageNumber="73">Fig. 11</figureCitation>
):
<collectingCountry id="F33DD9AA7E25FFE7416DFBCE40041967" box="[618,795,1123,1149]" name="Guatemala" pageId="6" pageNumber="73">GUATEMALA</collectingCountry>
,
<collectingRegion id="49EE57D87E25FFE7402BFBCE40D81967" box="[812,967,1123,1149]" country="Guatemala" name="Sacatepequez" pageId="6" pageNumber="73">Sacatepéquez</collectingRegion>
,
<collectorName id="26DFFCEC7E25FFE740DFFBCE47FF1967" box="[984,1248,1123,1149]" pageId="6" pageNumber="73">
<collectingMunicipality id="6BF103407E25FFE740DFFBCE47FF1967" box="[984,1248,1123,1149]" pageId="6" pageNumber="73">San Cristobal El Bajo</collectingMunicipality>
</collectorName>
,
<collectorName id="26DFFCEC7E25FFE747F6FBCE46871967" box="[1265,1432,1123,1149]" pageId="6" pageNumber="73">
<location id="8EF5CFE17E25FFE747F6FBCE46871967" LSID="urn:lsid:plazi:treatment:0383282C7E25FFE94390FC5B474019BB:8EF5CFE17E25FFE747F6FBCE46871967" box="[1265,1432,1123,1149]" country="Guatemala" latitude="14.518525" longLatPrecision="1" longitude="-90.691246" municipality="San Cristobal El Bajo" name="Finca El Pilar" pageId="6" pageNumber="73" stateProvince="Sacatepequez">Finca El Pilar</location>
</collectorName>
, Cerro Cucurucho.
<quantity id="4CD234DF7E25FFE7426BFB2A42A219BB" box="[364,445,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="6" pageNumber="73" unit="m" value="2600.0">
<elevation id="00077E097E25FFE7426BFB2A42A219BB" box="[364,445,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="6" pageNumber="73" unit="m" value="2600.0">2600m</elevation>
</quantity>
.
<date id="FF94BFFA7E25FFE742CDFB2A417819BB" box="[458,615,1159,1185]" pageId="6" pageNumber="73" value="2017-04-22">
<collectingDate id="EFD046127E25FFE742CDFB2A417819BB" box="[458,615,1159,1185]" pageId="6" pageNumber="73" value="2017-04-22">22 abril 2017</collectingDate>
</date>
,
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<geoCoordinate id="EE1EFFFD7E25FFE741F2FB2A406619BB" box="[757,889,1159,1185]" degrees="90.691247" direction="west" orientation="longitude" pageId="6" pageNumber="73" precision="1" value="-90.691246">-90.691247</geoCoordinate>
,
<collectorName id="26DFFCEC7E25FFE74081FB2A470319BB" box="[902,1052,1159,1185]" pageId="6" pageNumber="73">Col. Monzón</collectorName>
y
<collectorName id="26DFFCEC7E25FFE7473AFB2A47B519BB" box="[1085,1194,1159,1185]" pageId="6" pageNumber="73">Naumann</collectorName>
[
<emphasis id="B95E45287E25FFE747BAFB2A47C019BB" box="[1213,1247,1159,1185]" italics="true" pageId="6" pageNumber="73">leg</emphasis>
.] / JMS 10.939
</materialsCitation>
/
<materialsCitation id="3B4293677E25FFE74390FB06474819DF" ID-GBIF-Occurrence="3034649307" box="[151,1111,1195,1221]" collectionCode="UVG" country="Guatemala" pageId="6" pageNumber="73" specimenCount="1" stateProvince="Sacatepequez" typeStatus="holotype">
<typeStatus id="549127987E25FFE74390FB06425F19DF" box="[151,320,1195,1221]" pageId="6" pageNumber="73" type="holotype">HOLOTYPUS</typeStatus>
,
<taxonomicName id="4C2AE2B97E25FFE7424BFB06409C19DF" authority="C. Mielke, Grehan &amp; Monzon" authorityName="C. Mielke, Grehan &amp; Monzon" box="[332,899,1195,1221]" class="Insecta" family="Hepialidae" genus="Schausiana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="73" phylum="Arthropoda" rank="species" species="maishei">
<emphasis id="B95E45287E25FFE7424BFB06413419DF" box="[332,555,1195,1221]" italics="true" pageId="6" pageNumber="73">Schausiana maishei</emphasis>
C. Mielke, Grehan &amp; Monzón
</taxonomicName>
des. 2020/ (
<collectionCode id="ED3B01FF7E25FFE7470AFB06474F19DF" box="[1037,1104,1195,1221]" country="Guatemala" httpUri="http://grbio.org/cool/vx15-6kjy" name="Universidad del Valle" pageId="6" pageNumber="73">UVG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B95993A7E25FFE743C0FB62461218DB" blockId="6.[151,1437,1122,2014]" pageId="6" pageNumber="73">
<materialsCitation id="3B4293677E25FFE743C0FB62401019F2" ID-GBIF-Occurrence="3034649306" box="[199,783,1230,1257]" collectorName="All" country="Guatemala" location="Paratypes" pageId="6" pageNumber="73" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Paratypes" typeStatus="paratype">
<emphasis id="B95E45287E25FFE743C0FB62422019F3" bold="true" box="[199,319,1231,1257]" pageId="6" pageNumber="73">
<collectingRegion id="49EE57D87E25FFE743C0FB62422019F3" box="[199,319,1231,1257]" name="Paratypes" pageId="6" pageNumber="73">
<typeStatus id="549127987E25FFE743C0FB62422019F3" box="[199,319,1231,1257]" pageId="6" pageNumber="73" type="paratype">Paratypes</typeStatus>
</collectingRegion>
</emphasis>
: (in total
<specimenCount id="9D2C52B37E25FFE742B7FB6242C219F3" box="[432,477,1231,1257]" pageId="6" pageNumber="73" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D2C52B37E25FFE742EDFB63410B19F3" box="[490,532,1230,1257]" pageId="6" pageNumber="73" type="female">5 ♀</specimenCount>
).
<collectorName id="26DFFCEC7E25FFE7412BFB62414F19F3" box="[556,592,1231,1257]" pageId="6" pageNumber="73">All</collectorName>
<emphasis id="B95E45287E25FFE7415FFB63401019F2" bold="true" box="[600,783,1230,1256]" pageId="6" pageNumber="73">
<collectingCountry id="F33DD9AA7E25FFE7415FFB63401019F2" box="[600,783,1230,1256]" name="Guatemala" pageId="6" pageNumber="73">GUATEMALA</collectingCountry>
</emphasis>
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.
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<subSubSection id="C330CAB17E25FFE743C0FA6741341B13" pageId="6" pageNumber="73" type="diagnosis">
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<emphasis id="B95E45287E25FFE743C0FA67422518FE" bold="true" box="[199,314,1482,1508]" pageId="6" pageNumber="73">Diagnosis</emphasis>
. Immediately distinguished from all other
<taxonomicName id="4C2AE2B97E25FFE74027FA6640BE18FF" authorityName="Viette" authorityYear="1949" box="[800,929,1483,1509]" class="Insecta" family="Hepialidae" genus="Schausiana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="73" phylum="Arthropoda" rank="genus">
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by the crenulated margins of the FW and its dorsal olivish-brown ground colour.
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(
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a–b, 24, 30, 35a–d).
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. Frons and vertex dark olivish-brown. Antenna with ~26 antenomeres.
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<paragraph id="8B95993A7E25FFE943C0F9F6472D1C07" blockId="6.[151,1437,1122,2014]" lastBlockId="8.[151,1437,151,1185]" lastPageId="8" lastPageNumber="75" pageId="6" pageNumber="73">
<emphasis id="B95E45287E25FFE743C0F9F642071B6F" box="[199,280,1627,1653]" italics="true" pageId="6" pageNumber="73">Thorax</emphasis>
. Dorsally pro- and mesothorax concolourous with head with scattered long and light piliform scales, metathorax anteriorly as the former, posteriorly with scales dark yellow distally, ventrally dark olivish-brown; metathorax pale yellow with long piliform scales. Legs (
<figureCitation id="131185BF7E25FFE7401AF90E406C1BA7" box="[797,883,1699,1725]" captionStart="FIGURES 22–27" captionStartId="14.[151,264,1900,1924]" captionTargetBox="[159,1418,181,1870]" captionTargetId="figure-26@14.[158,1429,181,1877]" captionTargetPageId="14" captionText="FIGURES 22–27. Schausiana spp. labial palpi, legs and wing venation. 22–23, labial palpi of S. trojesa: 22, ♀ (Morelos, Cuernavaca (UNAM)); 23, ♂ (Morelos, Tepoztlán (UNAM)). Unscaled. 24–25, legs: 24, S. maishei, sp. nov. (CGCM 23.236); 25, S. pharus (CGCM 7.110). Scale bar = 5 mm. 26–27, wing venation of S. trojesa: 26, ♂ (arrows indicating 1A incomplete and 2A+3A); 27, ♀ (arrow indicating CuP mesally incomplete)." figureDoi="http://doi.org/10.5281/zenodo.4413522" httpUri="https://zenodo.org/record/4413522/files/figure.png" pageId="6" pageNumber="73">Fig. 24</figureCitation>
): epiphysis absent; metatibia hypertrophied with inflated thin-walled metatibial gland. FW length:
<quantity id="4CD234DF7E25FFE741C8F96B40531BFB" box="[719,844,1734,1761]" metricMagnitude="-2" metricUnit="m" metricValue="5.9" metricValueMax="6.1" metricValueMin="5.7" pageId="6" pageNumber="73" unit="mm" value="59.0" valueMax="61.0" valueMin="57.0">57–61 mm</quantity>
, width:
<quantity id="4CD234DF7E25FFE740ADF96A473A1BFB" box="[938,1061,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="6" pageNumber="73" unit="mm" value="25.0" valueMax="27.0" valueMin="23.0">23–27 mm</quantity>
(ratio ~2.4), wingspan:
<quantity id="4CD234DF7E25FFE7463CF96A43D91A1F" metricMagnitude="-1" metricUnit="m" metricValue="1.165" metricValueMax="1.2" metricValueMin="1.13" pageId="6" pageNumber="73" unit="mm" value="116.5" valueMax="120.0" valueMin="113.0">113–120 mm</quantity>
; elongated, tornus indistinct; costal margin convex, apex pronounced and acute, outer margin crenulate, slightly concave from apex to Rs4, inner margin convex. DFW ground colour olivish-brown with numerous dark brown dots and longitudinal and transverse black stripes throughout the wing; anal area with two irregular and detached black spots; antemedial area marked by a dark olivish-brown U-band with the distal arm extending from the stigma to between Cu and A veins; postdiscal, premarginal bands and marginal spots concolorous with the U-shaped band; postdiscal band broken between Rs3-Rs4 and M
<subScript id="17AE9B7F7E25FFE7419AF81C41BA1AA5" attach="left" box="[669,677,1969,1983]" fontSize="6" pageId="6" pageNumber="73">2</subScript>
-M
<subScript id="17AE9B7F7E25FFE741CFF81C41CF1AA5" attach="left" box="[712,720,1969,1983]" fontSize="6" pageId="6" pageNumber="73">3</subScript>
;premarginal band oblique(sometimes connected to the postdiscal band posteriorly); marginal area distally lighter with darker diamond shaped spots at the insertion of the veins at the wing margin; stigma silver and subtriangular; evenly spaced white specialized scales on the veins, long and piliform closer to wing base, spine-like nearer wing margin. DHW outer margin slightly crenulate, ground colour greyishbrown, costal margin distally brown with some darker spots, marginal band narrow, yellow, as for veins distally. VFW and VHW greyish-brown with apex and veins distally coloured dark yellow.
</paragraph>
<caption id="DF55C9B27E24FFE64390F83446121ACF" ID-DOI="http://doi.org/10.5281/zenodo.4413514" ID-Zenodo-Dep="4413514" httpUri="https://zenodo.org/record/4413514/files/figure.png" pageId="7" pageNumber="74" startId="7.[151,264,1945,1969]" targetBox="[232,1410,193,1899]" targetPageId="7">
<paragraph id="8B95993A7E24FFE64390F83446121ACF" blockId="7.[151,1436,1945,2005]" pageId="7" pageNumber="74">
<emphasis id="B95E45287E24FFE64390F834425F1AAB" bold="true" box="[151,320,1945,1969]" pageId="7" pageNumber="74">FIGURES 9–11</emphasis>
.
<taxonomicName id="4C2AE2B97E24FFE6424BF83441091AAB" authority="Mielke &amp; Grehan &amp; Monzón-Sierra, 2020" authorityName="Mielke &amp; Grehan &amp; Monzón-Sierra" authorityYear="2020" box="[332,534,1945,1969]" class="Insecta" family="Hepialidae" genus="Schausiana" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="74" phylum="Arthropoda" rank="species" species="maishei" status="sp. nov.">
<emphasis id="B95E45287E24FFE6424BF83441091AAB" box="[332,534,1945,1969]" italics="true" pageId="7" pageNumber="74">Schausiana maishei</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26DF8537E24FFE64124F834416A1AAB" box="[547,629,1945,1969]" pageId="7" pageNumber="74" rank="species">
<emphasis id="B95E45287E24FFE64124F83441241AAB" bold="true" box="[547,571,1945,1969]" pageId="7" pageNumber="74">sp</emphasis>
.
<emphasis id="B95E45287E24FFE64140F83441711AAB" bold="true" box="[583,622,1945,1969]" pageId="7" pageNumber="74">nov</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B95E45287E24FFE6417CF83441971AAB" bold="true" box="[635,648,1945,1969]" pageId="7" pageNumber="74">9</emphasis>
, ♂ PT (CGCM 32.071): 9a, dorsal view; 9b, ventral view (CGCM).
<emphasis id="B95E45287E24FFE64647F83446461AAB" bold="true" box="[1344,1369,1945,1969]" pageId="7" pageNumber="74">10</emphasis>
, ♀ PT (CGCM 36.646): 10a, dorsal view; 10b, ventral view (CGCM).
<emphasis id="B95E45287E24FFE64018F81040281ACF" bold="true" box="[799,823,1981,2005]" pageId="7" pageNumber="74">11</emphasis>
, ♀ HT: dorsal view (UVG). Scale bar = 1 cm.
</paragraph>
</caption>
<paragraph id="8B95993A7E2BFFE943C0FE8A411B1CCB" blockId="8.[151,1437,151,1185]" pageId="8" pageNumber="75">
<emphasis id="B95E45287E2BFFE943C0FE8A422E1C5B" box="[199,305,295,321]" italics="true" pageId="8" pageNumber="75">Abdomen</emphasis>
(
<figureCitation id="131185BF7E2BFFE94246FE8A42891C5B" box="[321,406,295,321]" captionStart="FIGURES 28–33" captionStartId="15.[151,264,1922,1946]" captionTargetBox="[179,1396,633,1892]" captionTargetId="figure-214@15.[167,1420,633,1898]" captionTargetPageId="15" captionText="FIGURES 28–33. Schausiana spp. abdomen (upper side, tergites; lower side, sternites). 28–29, S. trojesa: 28, ♂ (CGCM 39417); 29, ♀ (CGCM 33.184). 30, S. maishei, sp. nov., ♂ PT (CGCM 23.236). 31, S. chalciope, sp. nov., ♀ PT (CGCM 41.258). 32–33, S. pharus: 32, ♂ (CGCM 7.110); 33, ♀ (CGCM 5.797). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4413524" httpUri="https://zenodo.org/record/4413524/files/figure.png" pageId="8" pageNumber="75">Fig. 30</figureCitation>
). Segments I and II dorsally and ventrally concolorous with the DHW, segments III to VIII dorsally dark-brown with some dark blue reflecting scales, lighter posteriorly on each segment, ventrally pale yellow. Tergite II with slightly concave lateral ridge near margin, sternite II elongate with all corners projected and the lateral and posterior margins concave; tergite VIII relatively tiny, trapezoidal, only slightly sclerotized, sternite VIII as a narrow trapezoidal bar.
</paragraph>
<paragraph id="8B95993A7E2BFFE943C0FE7641341FEB" blockId="8.[151,1437,151,1185]" pageId="8" pageNumber="75">
<emphasis id="B95E45287E2BFFE943C0FE7642731CEF" box="[199,364,475,501]" italics="true" pageId="8" pageNumber="75">Male genitalia</emphasis>
(
<figureCitation id="131185BF7E2BFFE9427CFE7642CE1CEF" box="[379,465,474,501]" captionStart="FIGURES 34–36" captionStartId="16.[151,264,1795,1819]" captionTargetBox="[264,1317,196,1761]" captionTargetId="figure-26@16.[242,1345,181,1771]" captionTargetPageId="16" captionText="FIGURES 34–36. Schausiana spp. ♂ genitalia. 34, S. trojesa (CGCM 39.417): 34a, ventral view; 34b, ventral view with base of valva exposed (arrow indicates the hook at the base of the valva); 34c, dorsal view (internal; arrow indicating the mesal ridge of fultura inferior); 34d, lateral view. 35, S. maishei, sp. nov., ♂ PT (CGCM 23.236): 35a, ventral view (arrow indicates the hook at the base of the valva); 35b, ventral view with pseudotegumen moved upwards (arrow indicating the mesal ridge of fultura inferior); 35c, dorsal view (internal); 35d, lateral view. 36, S. pharus (CGCM 7.110): 36a, ventral view (arrow indicates the hook at the base of the valva); 36b, dorsal view (internal; arrow indicating the mesal ridge of fultura inferior); 36c, lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4413526" httpUri="https://zenodo.org/record/4413526/files/figure.png" pageId="8" pageNumber="75">Figs 35</figureCitation>
a–d). Tegumen as a narrow rectangular bar fused to pseudotegumen. Saccus U–shaped with the mesal portion of the posterior margin slightly projected ventrally as a shelf along the apodemal suture. Tergal lobe well marked, fused to the pseudotegumen, each side globular, occupying about 1/4 of the posterior margin of the pseudotegumen. Pseudotegumen forming an irregular plate, increasingly sclerotized ventrally with a ridge parallel to the anterior and posterior margins; edge of anogenital region serrate; antero-ventrally slightly projected and heavily sclerotized. Fultura inferior rectangular with rounded edges and a longitudinal medial ridge. Fultura superior reduced and triangular. Valva distally short, lobate and setose, sacculus narrowing anteriorly terminating in prominent medially angled spine.
</paragraph>
<paragraph id="8B95993A7E2BFFE943C0FD5642E01E0F" blockId="8.[151,1437,151,1185]" box="[199,511,763,789]" pageId="8" pageNumber="75">
<emphasis id="B95E45287E2BFFE943C0FD5642011E0F" bold="true" box="[199,286,763,789]" pageId="8" pageNumber="75">Female</emphasis>
(
<figureCitation id="131185BF7E2BFFE9422AFD56426A1E0F" box="[301,373,763,789]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1864,1888]" captionTargetBox="[156,1431,194,1830]" captionTargetId="figure-26@4.[156,1432,181,1841]" captionTargetPageId="4" captionText="FIGURES 1–5. Schausiana spp. FW areas and specialized scales. Areas: 1, S. trojesa ♀. Specialized scales: 2, S. trojesa ♀; 3, S. maishei, sp. nov. ♀; 4, S. chalciope, sp. nov. ♀; 5. S. pharus ♀. Arrows indicate the position of such scales." figureDoi="http://doi.org/10.5281/zenodo.4413510" httpUri="https://zenodo.org/record/4413510/files/figure.png" pageId="8" pageNumber="75">Figs 3</figureCitation>
,
<figureCitation id="131185BF7E2BFFE94285FD5642821E0F" box="[386,413,763,789]" captionStart="FIGURES 9–11" captionStartId="7.[151,264,1945,1969]" captionTargetBox="[232,1410,193,1899]" captionTargetId="figure-20@7.[153,1434,181,1921]" captionTargetPageId="7" captionText="FIGURES 9–11. Schausiana maishei, sp. nov. 9, ♂ PT (CGCM 32.071): 9a, dorsal view; 9b, ventral view (CGCM). 10, ♀ PT (CGCM 36.646): 10a, dorsal view; 10b, ventral view (CGCM). 11, ♀ HT: dorsal view (UVG). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4413514" httpUri="https://zenodo.org/record/4413514/files/figure.png" pageId="8" pageNumber="75">10</figureCitation>
a–b, 11).
</paragraph>
<paragraph id="8B95993A7E2BFFE943C0FCB242BC1E23" blockId="8.[151,1437,151,1185]" box="[199,419,799,825]" pageId="8" pageNumber="75">
<emphasis id="B95E45287E2BFFE943C0FCB2421B1E23" box="[199,260,799,825]" italics="true" pageId="8" pageNumber="75">Head</emphasis>
. As for the
<specimenCount id="9D2C52B37E2BFFE94280FCB242801E23" box="[391,415,799,825]" pageId="8" pageNumber="75" type="male">♂</specimenCount>
.
</paragraph>
<paragraph id="8B95993A7E2BFFE943C0FCEE478A1E47" blockId="8.[151,1437,151,1185]" box="[199,1173,834,861]" pageId="8" pageNumber="75">
<emphasis id="B95E45287E2BFFE943C0FCEE42071E47" box="[199,280,835,861]" italics="true" pageId="8" pageNumber="75">Thorax</emphasis>
. FW length:
<quantity id="4CD234DF7E2BFFE942ACFCEE41391E47" box="[427,550,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" metricValueMax="8.2" metricValueMin="4.6" pageId="8" pageNumber="75" unit="mm" value="64.0" valueMax="82.0" valueMin="46.0">46–82 mm</quantity>
, width:
<quantity id="4CD234DF7E2BFFE94187FCEE41E61E47" box="[640,761,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.3" metricValueMin="1.7" pageId="8" pageNumber="75" unit="mm" value="25.0" valueMax="33.0" valueMin="17.0">17–33 mm</quantity>
(ratio ~2.4), wingspan:
<quantity id="4CD234DF7E2BFFE94700FCEF478E1E47" box="[1031,1169,834,861]" metricMagnitude="-1" metricUnit="m" metricValue="1.28" metricValueMax="1.59" metricValueMin="0.97" pageId="8" pageNumber="75" unit="mm" value="128.0" valueMax="159.0" valueMin="97.0">97–159 mm</quantity>
.
</paragraph>
<paragraph id="8B95993A7E2BFFE943C0FCCA42CF1E9B" blockId="8.[151,1437,151,1185]" box="[199,464,871,897]" pageId="8" pageNumber="75">
<emphasis id="B95E45287E2BFFE943C0FCCA422E1E9B" box="[199,305,871,897]" italics="true" pageId="8" pageNumber="75">Abdomen</emphasis>
. As for the
<specimenCount id="9D2C52B37E2BFFE942B3FCCA42D31E9B" box="[436,460,871,897]" pageId="8" pageNumber="75" type="male">♂</specimenCount>
.
</paragraph>
<paragraph id="8B95993A7E2BFFE943C0FC26412C1EBF" blockId="8.[151,1437,151,1185]" box="[199,563,907,933]" pageId="8" pageNumber="75">
<emphasis id="B95E45287E2BFFE943C0FC2642991EBF" box="[199,390,907,933]" italics="true" pageId="8" pageNumber="75">Female genitalia</emphasis>
. Not dissected.
</paragraph>
<paragraph id="8B95993A7E2BFFE943C0FC03474E1EF7" blockId="8.[151,1437,151,1185]" pageId="8" pageNumber="75">
<emphasis id="B95E45287E2BFFE943C0FC0342E01ED2" bold="true" box="[199,511,942,968]" pageId="8" pageNumber="75">Geographical distribution</emphasis>
.
<emphasis id="B95E45287E2BFFE9410AFC0241EB1ED3" box="[525,756,943,969]" italics="true" pageId="8" pageNumber="75">
<taxonomicName id="4C2AE2B97E2BFFE9410AFC0241EF1ED3" authority="Mielke &amp; Grehan &amp; Monzón-Sierra, 2020" authorityName="Mielke &amp; Grehan &amp; Monzón-Sierra" authorityYear="2020" box="[525,752,943,969]" class="Insecta" family="Hepialidae" genus="Schausiana" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="75" phylum="Arthropoda" rank="species" species="maishei" status="sp. nov.">Schausiana maishei</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A26DF8537E2BFFE941FBFC0240481ED3" box="[764,855,943,969]" pageId="8" pageNumber="75" rank="species">
<emphasis id="B95E45287E2BFFE941FBFC0240091ED3" bold="true" box="[764,790,943,969]" pageId="8" pageNumber="75">sp</emphasis>
.
<emphasis id="B95E45287E2BFFE94022FC02404E1ED3" bold="true" box="[805,849,943,969]" pageId="8" pageNumber="75">nov</emphasis>
.
</taxonomicNameLabel>
, is known from four localities in the vicinity of the Motagua-Polochic Fracture Zone in
<collectingCountry id="F33DD9AA7E2BFFE94128FC7E41B71EF7" box="[559,680,979,1005]" name="Guatemala" pageId="8" pageNumber="75">Guatemala</collectingCountry>
at altitudes around
<quantity id="4CD234DF7E2BFFE94082FC7F40FE1EF7" box="[901,993,978,1005]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="75" unit="m" value="2500.0">2,500 m</quantity>
(
<figureCitation id="131185BF7E2BFFE940F7FC7E475B1EF7" box="[1008,1092,979,1005]" captionStart="FIGURE 48" captionStartId="18.[151,250,1727,1751]" captionTargetBox="[227,1436,921,1704]" captionTargetId="figure-316@18.[151,1436,921,1704]" captionTargetPageId="18" captionText="FIGURE 48. Distribution of Schausiana spp. within southern North and Central America Eastern boundary of Guerrero terrane shown as blue dashed line." figureDoi="http://doi.org/10.5281/zenodo.4413530" httpUri="https://zenodo.org/record/4413530/files/figure.png" pageId="8" pageNumber="75">Fig. 48</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C330CAB17E2BFFE943C0FC5A42CC190B" box="[199,467,1015,1041]" pageId="8" pageNumber="75" type="biology_ecology">
<paragraph id="8B95993A7E2BFFE943C0FC5A42CC190B" blockId="8.[151,1437,151,1185]" box="[199,467,1015,1041]" pageId="8" pageNumber="75">
<emphasis id="B95E45287E2BFFE943C0FC5A4250190B" bold="true" box="[199,335,1015,1041]" pageId="8" pageNumber="75">Host plants</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C330CAB17E2BFFE943C0FBB641621943" pageId="8" pageNumber="75" type="etymology">
<paragraph id="8B95993A7E2BFFE943C0FBB641621943" blockId="8.[151,1437,151,1185]" pageId="8" pageNumber="75">
<emphasis id="B95E45287E2BFFE943C0FBB64257192F" bold="true" box="[199,328,1051,1077]" pageId="8" pageNumber="75">Etymology</emphasis>
. The specific epithet is dedicated to Maishe Dickman for his friendship and sustained support through his field expeditions in
<collectingCountry id="F33DD9AA7E2BFFE942FBFB9241661943" box="[508,633,1087,1113]" name="Guatemala" pageId="8" pageNumber="75">Guatemala</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C330CAB17E2BFFE943C0FBCE474019BB" pageId="8" pageNumber="75" type="discussion">
<paragraph id="8B95993A7E2BFFE943C0FBCE474019BB" blockId="8.[151,1437,151,1185]" pageId="8" pageNumber="75">
<emphasis id="B95E45287E2BFFE943C0FBCE422B1967" bold="true" box="[199,308,1123,1149]" pageId="8" pageNumber="75">Remarks</emphasis>
. Similarities between
<emphasis id="B95E45287E2BFFE9412AFBCE400C1967" box="[557,787,1123,1149]" italics="true" pageId="8" pageNumber="75">
<taxonomicName id="4C2AE2B97E2BFFE9412AFBCE40101967" authority="Mielke &amp; Grehan &amp; Monzón-Sierra, 2020" authorityName="Mielke &amp; Grehan &amp; Monzón-Sierra" authorityYear="2020" box="[557,783,1123,1149]" class="Insecta" family="Hepialidae" genus="Schausiana" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="75" phylum="Arthropoda" rank="species" species="maishei" status="sp. nov.">Schausiana maishei</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A26DF8537E2BFFE9401DFBCE406C1967" box="[794,883,1123,1149]" pageId="8" pageNumber="75" rank="species">
<emphasis id="B95E45287E2BFFE9401DFBCE402B1967" bold="true" box="[794,820,1123,1149]" pageId="8" pageNumber="75">sp</emphasis>
.
<emphasis id="B95E45287E2BFFE94045FBCE40711967" bold="true" box="[834,878,1123,1149]" pageId="8" pageNumber="75">nov</emphasis>
.
</taxonomicNameLabel>
, and the previous species in wing ornamentation, scales on the thorax and DFW may suggest a close relationship between these species.
</paragraph>
</subSubSection>
</treatment>
</document>
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