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<document id="A01C5ED00CFF09232E4D82ECA83818EC" ID-CLB-Dataset="28431" ID-DOI="10.11646/zootaxa.4564.1.2" ID-GBIF-Dataset="d3a56aac-92aa-48f2-b9fe-9576a4c2a0ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="2588621" ID-ZooBank="27F70B5F-24DD-4C9E-BBA7-2B87476BD49C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552162070752" checkinUser="plazi" docAuthor="Gunter, Nicole L. &amp; Weir, Thomas A." docDate="2019" docId="03832713FFB18D69A3DAFE0151EC92C2" docLanguage="en" docName="zootaxa.4564.1.2.pdf" docOrigin="Zootaxa 4564 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepanus guthrieae Gunter &amp; Weir 2019, new species" docType="treatment" docUuid="6149EEFE-33BC-44A5-91B7-0EFE4B8EBCEF" docUuidSource="ZooBank" docVersion="9" lastPageNumber="68" masterDocId="FFBA5F6BFFAB8D72A34DFFB65569910A" masterDocTitle="Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group" masterLastPageNumber="80" masterPageNumber="41" pageNumber="67" updateTime="1698700752634" updateUser="ExternalLinkService">
<mods:mods id="6E7213282251AF4F847005C80D718F19" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="0CFA46CA3C6E9DC71D30D80906162818">Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group</mods:title>
</mods:titleInfo>
<mods:name id="B4EBFF71CB7811B20D16F874624D3E37" type="personal">
<mods:role id="342E67DD82B2E57317AF53D2CD2F7D55">
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<mods:namePart id="2A036D26C98BB2C27D1131A597F33C6A">Gunter, Nicole L.</mods:namePart>
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<mods:namePart id="380BC5A32241F708525F6E60528E468B">Weir, Thomas A.</mods:namePart>
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<mods:title id="0058C28A14E820D760FBC473EA2FBEEA">Zootaxa</mods:title>
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<mods:part id="2D184FFF7EF74DB642AECF027680D144">
<mods:date id="34F3B0C10CE6FFF18C921B72DFAA84C0">2019</mods:date>
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<mods:number id="F4CE0DDF56636D26D32F732098E0DF8F">2019-03-05</mods:number>
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<mods:number id="12A82CE5DA7B98424BE69F72607E86AD">4564</mods:number>
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<mods:classification id="45637B4374E45A6B67C9B2366CFB7037">journal article</mods:classification>
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<mods:identifier id="1133E41BC6CC2B19AD5132AF4D2D0B78" type="DOI">10.11646/zootaxa.4564.1.2</mods:identifier>
<mods:identifier id="A679FC44E3BC3EF8C89C9D506DC5463D" type="GBIF-Dataset">d3a56aac-92aa-48f2-b9fe-9576a4c2a0ec</mods:identifier>
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<treatment id="03832713FFB18D69A3DAFE0151EC92C2" ID-DOI="http://doi.org/10.5281/zenodo.5922810" ID-GBIF-Taxon="154366354" ID-Zenodo-Dep="5922810" LSID="urn:lsid:zoobank.org:act:6149EEFE-33BC-44A5-91B7-0EFE4B8EBCEF" httpUri="http://treatment.plazi.org/id/03832713FFB18D69A3DAFE0151EC92C2" lastPageId="27" lastPageNumber="68" pageId="26" pageNumber="67">
<subSubSection id="C330C58EFFB18D68A3DAFE0157B490D8" box="[151,733,439,466]" pageId="26" pageNumber="67" type="nomenclature">
<paragraph id="8B959605FFB18D68A3DAFE0157B490D8" blockId="26.[151,733,439,499]" box="[151,733,439,466]" pageId="26" pageNumber="67">
<heading id="D0DD2169FFB18D68A3DAFE0157B490D8" bold="true" box="[151,733,439,466]" fontSize="11" level="1" pageId="26" pageNumber="67" reason="1">
<emphasis id="B95E4A17FFB18D68A3DAFE0157B490D8" bold="true" box="[151,733,439,466]" pageId="26" pageNumber="67">
<taxonomicName id="4C2AED86FFB18D68A3DAFE01572890D8" ID-CoL="6P7XX" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[151,577,439,466]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="67" phylum="Arthropoda" rank="species" species="guthrieae" status="sp. nov.">
<emphasis id="B95E4A17FFB18D68A3DAFE01541090DB" bold="true" box="[151,377,439,465]" italics="true" pageId="26" pageNumber="67">Lepanus guthrieae</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFFB18D68A100FE0E57B490D8" box="[589,733,440,466]" pageId="26" pageNumber="67" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFB18D68A3DAFE6C549A90F9" box="[151,499,474,499]" pageId="26" pageNumber="67" type="description">
<paragraph id="8B959605FFB18D68A3DAFE6C549A90F9" blockId="26.[151,733,439,499]" box="[151,499,474,499]" pageId="26" pageNumber="67">
(
<figureCitation id="13118A80FFB18D68A3D2FE6C544590F9" box="[159,300,474,499]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="26" pageNumber="67">Figs. 4A, 4F</figureCitation>
,
<figureCitation id="13118A80FFB18D68A275FE6C543390F9" box="[312,346,474,499]" captionStart="FIGURE 5" captionStartId="15.[151,250,1945,1967]" captionTargetBox="[245,1336,223,1911]" captionTargetId="figure@15.[233,1355,193,1926]" captionTargetPageId="15" captionText="FIGURE 5. Habitus of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil. All specimens in comparative scale. Scale bar 1 mm." httpUri="https://zenodo.org/record/2588631/files/figure.png" pageId="26" pageNumber="67">5E</figureCitation>
,
<figureCitation id="13118A80FFB18D68A228FE6D54EF90F9" box="[357,390,475,499]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="26" pageNumber="67">8E</figureCitation>
,
<figureCitation id="13118A80FFB18D68A2DFFE6D54A890F9" box="[402,449,475,499]" captionStart="FIGURE 10" captionStartId="23.[151,250,1909,1931]" captionTargetBox="[167,1333,208,1895]" captionTargetId="figure@23.[164,1428,193,1895]" captionTargetPageId="23" captionText="FIGURE 10. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588641/files/figure.png" pageId="26" pageNumber="67">10E</figureCitation>
,
<figureCitation id="13118A80FFB18D68A280FE6C548390F9" box="[461,490,474,499]" captionStart="FIGURE 16" captionStartId="35.[151,250,1726,1748]" captionTargetBox="[151,1436,1037,1705]" captionTargetId="figure@35.[151,1436,1037,1705]" captionTargetPageId="35" captionText="FIGURE 16. Map of known records of Lepanus guthrieae including predicted distribution overlaid with predicted distribution boundary of L. pygmaeus (dashed line) taken from Fig. 13 map." httpUri="https://zenodo.org/record/2588653/files/figure.png" pageId="26" pageNumber="67">16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFB18D68A3DAFD9454D79491" pageId="26" pageNumber="67" type="materials_examined">
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<emphasis id="B95E4A17FFB18D68A3DAFD9454F79331" bold="true" box="[151,414,546,571]" pageId="26" pageNumber="67">
Type series.
<materialsCitation id="3B429C58FFB18D68A265FD9454F79331" ID-GBIF-Occurrence="2012745384" box="[296,414,546,571]" collectingDate="2005-08-23" collectingDateMax="2006-05-24" collectingDateMin="2005-08-23" collectionCode="CALM, PILBARA, SURVEY, ANIC, WAM" determinerName="T. A. Weir" latitude="-22.305832" location="Site" longLatPrecision="21" longitude="117.47972" pageId="26" pageNumber="67" specimenCode="TCMBW12" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="549128A7FFB18D68A265FD9454F09331" box="[296,409,546,571]" pageId="26" pageNumber="67" type="holotype">Holotype</typeStatus>
:
</materialsCitation>
</emphasis>
<specimenCount id="9D2C5D8CFFB18D68A2E8FD9554D39330" box="[421,442,547,570]" pageId="26" pageNumber="67" type="male">♂</specimenCount>
“W.A.:
<locationDeviation id="9A278F27FFB18D68A158FD9456509331" box="[533,825,546,571]" location="Tom Price" pageId="26" pageNumber="67">
<quantity id="4CD23BE0FFB18D68A158FD94573F9331" box="[533,598,546,571]" metricMagnitude="4" metricUnit="m" metricValue="5.3" pageId="26" pageNumber="67" unit="km" value="53.0">53km</quantity>
NNW of Tom Price
</locationDeviation>
,
<location id="8EF5C0DEFFB18D68A00EFD9456069331" LSID="urn:lsid:plazi:treatment:03832713FFB18D69A3DAFE0151EC92C2:8EF5C0DEFFB18D68A00EFD9456069331" box="[835,879,546,571]" latitude="-22.305832" longLatPrecision="21" longitude="117.47972" name="Site" pageId="26" pageNumber="67">Site</location>
<specimenCode id="DB8C3E7EFFB18D68A038FD9556979331" box="[885,1022,547,571]" collectionCode="TCMBW" pageId="26" pageNumber="67">TCMBW12</specimenCode>
, ethylene glycol pitfalls,
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,
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<collectingDate id="EFD0492DFFB18D68A279FDF154859355" box="[308,492,582,607]" pageId="26" pageNumber="67" value="2005-08-23">23 August 2005</collectingDate>
</date>
-
<date id="FF94B0C5FFB18D68A2B9FDF057F99355" box="[500,656,582,607]" pageId="26" pageNumber="67" value="2006-05-24">
<collectingDate id="EFD0492DFFB18D68A2B9FDF057F99355" box="[500,656,582,607]" pageId="26" pageNumber="67" value="2006-05-24">24 May 2006</collectingDate>
</date>
, CALM PILBARA SURVEY” / “ANIC Database No. 25-053803” / “
<taxonomicName id="4C2AED86FFB18D68A3ECFDDD546B9389" authorityName=", Matthews" authorityYear="1974" box="[161,258,619,643]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="79" phylum="Arthropoda" rank="genus">Lepanus</taxonomicName>
WA2 det.
<determinerName id="7E497AC1FFB18D68A233FDDC548C9389" box="[382,485,618,643]" pageId="26" pageNumber="67">T.A.Weir</determinerName>
2008” / “ANIC Image” in WAM.
<materialsCitation id="3B429C58FFB18D68A03CFDDC543F93C1" ID-GBIF-Occurrence="2012745366" collectingDate="2005-08-23" collectingDateMax="2006-05-24" collectingDateMin="2005-08-23" collectionCode="CALM, ANIC" collectorName="Pilbara Survey" latitude="-22.305832" location="Site" longLatPrecision="21" longitude="117.47972" pageId="26" pageNumber="67" specimenCode="TCMBW12" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia" typeStatus="paratype">
<emphasis id="B95E4A17FFB18D68A03CFDDC51899389" bold="true" box="[881,1248,618,643]" pageId="26" pageNumber="67">
<typeStatus id="549128A7FFB18D68A03CFDDC56849389" box="[881,1005,618,643]" pageId="26" pageNumber="67" type="paratype">Paratypes</typeStatus>
:
<collectingRegion id="49EE58E7FFB18D68A0B6FDDC51B29389" box="[1019,1243,618,643]" country="Australia" name="Western Australia" pageId="26" pageNumber="67">Western Australia</collectingRegion>
:
</emphasis>
<locationDeviation id="9A278F27FFB18D68A7A5FDDC546493AD" location="Tom Price" pageId="26" pageNumber="67">
<quantity id="4CD23BE0FFB18D68A7A5FDDC50589389" box="[1256,1329,618,643]" metricMagnitude="4" metricUnit="m" metricValue="5.3" pageId="26" pageNumber="67" unit="km" value="53.0">53 km</quantity>
NNW of Tom Price
</locationDeviation>
,
<location id="8EF5C0DEFFB18D68A25AFD38542A93AD" LSID="urn:lsid:plazi:treatment:03832713FFB18D69A3DAFE0151EC92C2:8EF5C0DEFFB18D68A25AFD38542A93AD" box="[279,323,654,679]" latitude="-22.305832" longLatPrecision="21" longitude="117.47972" name="Site" pageId="26" pageNumber="67" stateProvince="Western Australia">Site</location>
<specimenCode id="DB8C3E7EFFB18D68A207FD3954BB93AD" box="[330,466,655,679]" collectionCode="TCMBW" pageId="26" pageNumber="67">TCMBW12</specimenCode>
,
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,
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,
<date id="FF94B0C5FFB18D68A048FD3856E593AD" box="[773,908,654,679]" pageId="26" pageNumber="67" value="2005-08-23">
<collectingDate id="EFD0492DFFB18D68A048FD3856E593AD" box="[773,908,654,679]" pageId="26" pageNumber="67" value="2005-08-23">23.viii.2005</collectingDate>
</date>
–
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<collectingDate id="EFD0492DFFB18D68A0D6FD38516293AD" box="[923,1035,654,679]" pageId="26" pageNumber="67" value="2006-05-24">24.v.2006</collectingDate>
</date>
,
<collectionCode id="ED3B0EC0FFB18D68A75BFD39510E93AD" box="[1046,1127,655,679]" pageId="26" pageNumber="67">CALM</collectionCode>
<collectorName id="26DFF3D3FFB18D68A723FD38507B93AD" box="[1134,1298,654,679]" pageId="26" pageNumber="67">Pilbara Survey</collectorName>
(25-065526 ×
<specimenCount id="9D2C5D8CFFB18D68A3FDFD0555B593C0" box="[176,220,691,715]" pageId="26" pageNumber="67" type="female">1 ♀</specimenCount>
in
<collectionCode id="ED3B0EC0FFB18D68A248FD05542793C1" box="[261,334,691,715]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="26" pageNumber="67">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFB18D68A229FD04578393E5" ID-GBIF-Occurrence="2012745374" collectingDate="2005-08-31" collectingDateMax="2006-05-30" collectingDateMin="2005-08-31" collectionCode="CALM, ANIC" collectorName="Pilbara Survey" latitude="-22.607224" location="Site" longLatPrecision="21" longitude="117.935555" pageId="26" pageNumber="67" specimenCode="TCMBE10" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia" typeStatus="paratype">
<locationDeviation id="9A278F27FFB18D68A229FD0457FC93C1" box="[356,661,690,715]" location="Tom Price" pageId="26" pageNumber="67">
<quantity id="4CD23BE0FFB18D68A229FD0454AB93C1" box="[356,450,690,715]" metricMagnitude="4" metricUnit="m" metricValue="1.75" pageId="26" pageNumber="67" unit="km" value="17.5">17.5 km</quantity>
NE of Tom Price
</locationDeviation>
,
<location id="8EF5C0DEFFB18D68A1EFFD0457A793C1" LSID="urn:lsid:plazi:treatment:03832713FFB18D69A3DAFE0151EC92C2:8EF5C0DEFFB18D68A1EFFD0457A793C1" box="[674,718,690,715]" latitude="-22.607224" longLatPrecision="21" longitude="117.935555" name="Site" pageId="26" pageNumber="67" stateProvince="Western Australia">Site</location>
<specimenCode id="DB8C3E7EFFB18D68A19AFD05563F93C1" box="[727,854,691,715]" collectionCode="TCMBE" pageId="26" pageNumber="67">TCMBE10</specimenCode>
,
<geoCoordinate id="EE1EF0C2FFB18D68A029FD04568F93C1" box="[868,998,690,715]" direction="south" orientation="latitude" pageId="26" pageNumber="67" precision="15" value="-22.607224">22°36'26&quot;S</geoCoordinate>
,
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,
<date id="FF94B0C5FFB18D68A7DCFD04507E93C1" box="[1169,1303,690,715]" pageId="26" pageNumber="67" value="2005-08-31">
<collectingDate id="EFD0492DFFB18D68A7DCFD04507E93C1" box="[1169,1303,690,715]" pageId="26" pageNumber="67" value="2005-08-31">31.viii.2005</collectingDate>
</date>
–
<date id="FF94B0C5FFB18D68A66AFD0450FE93C1" box="[1319,1431,690,715]" pageId="26" pageNumber="67" value="2006-05-30">
<collectingDate id="EFD0492DFFB18D68A66AFD0450FE93C1" box="[1319,1431,690,715]" pageId="26" pageNumber="67" value="2006-05-30">30.v.2006</collectingDate>
</date>
,
<collectionCode id="ED3B0EC0FFB18D68A3DAFD61558193E5" box="[151,232,727,751]" pageId="26" pageNumber="67">CALM</collectionCode>
<collectorName id="26DFF3D3FFB18D68A3BFFD6054F293E5" box="[242,411,726,751]" pageId="26" pageNumber="67">Pilbara Survey</collectorName>
(25-053804
<date id="FF94B0C5FFB18D68A17EFD61572693E5" box="[563,591,727,751]" pageId="26" pageNumber="67">
x
<specimenCount id="9D2C5D8CFFB18D68A109FD61572693E5" box="[580,591,727,751]" pageId="26" pageNumber="67" type="female">1</specimenCount>
</date>
♀ in
<collectionCode id="ED3B0EC0FFB18D68A1D4FD61578B93E5" box="[665,738,727,751]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="26" pageNumber="67">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFB18D68A1B5FD61512D9219" ID-GBIF-Occurrence="2012745373" collectingDate="2004-05-08" collectingDateMax="2004-10-11" collectingDateMin="2004-05-08" collectionCode="CALM, ANIC" collectorName="Pilbara Survey" latitude="-21.394722" location="4 km N of Barowanna Hill" longLatPrecision="21" longitude="117.170555" pageId="26" pageNumber="67" specimenCount="2" specimenCount-male="2" stateProvince="Western Australia" typeStatus="paratype">
<location id="8EF5C0DEFFB18D68A1B5FD61515C93E5" LSID="urn:lsid:plazi:treatment:03832713FFB18D69A3DAFE0151EC92C2:8EF5C0DEFFB18D68A1B5FD61515C93E5" box="[760,1077,726,751]" latitude="-21.394722" longLatPrecision="21" longitude="117.170555" name="4 km N of Barowanna Hill" pageId="26" pageNumber="67" stateProvince="Western Australia">
<quantity id="4CD23BE0FFB18D68A1B5FD61565C93E5" box="[760,821,726,751]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="67" unit="km" value="4.0">4 km</quantity>
N of Barowanna Hill
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(Camp Curlewis),
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,
<geoCoordinate id="EE1EF0C2FFB18D68A3DAFD4D544E9219" box="[151,295,763,787]" direction="east" orientation="longitude" pageId="26" pageNumber="67" precision="15" value="117.170555">117°10'14&quot;E</geoCoordinate>
,
<date id="FF94B0C5FFB18D68A27BFD4D54B19219" box="[310,472,763,787]" pageId="26" pageNumber="67" value="2004-05-08" valueMax="2004-10-11" valueMin="2004-05-08">
<collectingDate id="EFD0492DFFB18D68A27BFD4D54B19219" box="[310,472,763,787]" pageId="26" pageNumber="67" value="2004-05-08" valueMax="2004-10-11" valueMin="2004-05-08">8.v–11.x.2004</collectingDate>
</date>
,
<collectionCode id="ED3B0EC0FFB18D68A2AAFD4D57519219" box="[487,568,763,787]" pageId="26" pageNumber="67">CALM</collectionCode>
<collectorName id="26DFF3D3FFB18D68A10FFD4C57839219" box="[578,746,762,787]" pageId="26" pageNumber="67">Pilbara Survey</collectorName>
(25-058206 ×
<specimenCount id="9D2C5D8CFFB18D68A0D1FD4D56A09218" box="[924,969,763,787]" pageId="26" pageNumber="67" type="male">2 ♂</specimenCount>
in
<collectionCode id="ED3B0EC0FFB18D68A0BEFD4D51559219" box="[1011,1084,763,787]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="26" pageNumber="67">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFB18D68A71FFD4D5199923D" ID-GBIF-Occurrence="2012745375" collectingDate="2003-11-25" collectingDateMax="2004-05-06" collectingDateMin="2003-11-25" collectionCode="CALM, ANIC" collectorName="Pilbara Survey" latitude="-21.876389" location="11 km SW of Coolawanyah" longLatPrecision="21" longitude="117.74139" pageId="26" pageNumber="67" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia" typeStatus="paratype">
<location id="8EF5C0DEFFB18D68A71FFD4D50F19219" LSID="urn:lsid:plazi:treatment:03832713FFB18D69A3DAFE0151EC92C2:8EF5C0DEFFB18D68A71FFD4D50F19219" box="[1106,1432,762,787]" latitude="-21.876389" longLatPrecision="21" longitude="117.74139" name="11 km SW of Coolawanyah" pageId="26" pageNumber="67" stateProvince="Western Australia">
<quantity id="4CD23BE0FFB18D68A71FFD4D51F59219" box="[1106,1180,762,787]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="26" pageNumber="67" unit="km" value="11.0">11 km</quantity>
SW of Coolawanyah
</location>
,
<geoCoordinate id="EE1EF0C2FFB18D68A3DAFCA85471923D" box="[151,280,798,823]" direction="south" orientation="latitude" pageId="26" pageNumber="67" precision="15" value="-21.876389">21°52'35&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFB18D68A26EFCA954DD923D" box="[291,436,799,823]" direction="east" orientation="longitude" pageId="26" pageNumber="67" precision="15" value="117.74139">117°44'29&quot;E</geoCoordinate>
,
<date id="FF94B0C5FFB18D68A2F3FCA8575C923D" box="[446,565,798,823]" pageId="26" pageNumber="67" value="2003-11-25">
<collectingDate id="EFD0492DFFB18D68A2F3FCA8575C923D" box="[446,565,798,823]" pageId="26" pageNumber="67" value="2003-11-25">25.xi.2003</collectingDate>
</date>
–
<date id="FF94B0C5FFB18D68A10BFCA857CE923D" box="[582,679,798,823]" pageId="26" pageNumber="67" value="2004-05-06">
<collectingDate id="EFD0492DFFB18D68A10BFCA857CE923D" box="[582,679,798,823]" pageId="26" pageNumber="67" value="2004-05-06">6.v.2004</collectingDate>
</date>
,
<collectionCode id="ED3B0EC0FFB18D68A1FFFCA9566A923D" box="[690,771,799,823]" pageId="26" pageNumber="67">CALM</collectionCode>
<collectorName id="26DFF3D3FFB18D68A044FCA856C7923D" box="[777,942,798,823]" pageId="26" pageNumber="67">Pilbara Survey</collectorName>
(25-058207 ×
<specimenCount id="9D2C5D8CFFB18D68A71EFCA95115923C" box="[1107,1148,799,823]" pageId="26" pageNumber="67" type="male">1 ♂</specimenCount>
in
<collectionCode id="ED3B0EC0FFB18D68A7D2FCA95181923D" box="[1183,1256,799,823]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="26" pageNumber="67">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFB18D68A7B6FCA9541A9275" ID-GBIF-Occurrence="2012745379" collectingDate="2003-11-21" collectingDateMax="2004-05-24" collectingDateMin="2003-11-21" collectionCode="CALM, WAM" collectorName="Pilbara Survey" latitude="-22.334723" location="1.8 km SSE of Mulga Downs Outcamp" longLatPrecision="21" longitude="118.988884" pageId="26" pageNumber="67" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Western Australia" typeStatus="paratype">
<location id="8EF5C0DEFFB18D68A7B6FCA954C09251" LSID="urn:lsid:plazi:treatment:03832713FFB18D69A3DAFE0151EC92C2:8EF5C0DEFFB18D68A7B6FCA954C09251" latitude="-22.334723" longLatPrecision="21" longitude="118.988884" name="1.8 km SSE of Mulga Downs Outcamp" pageId="26" pageNumber="67" stateProvince="Western Australia">
<quantity id="4CD23BE0FFB18D68A7B6FCA95021923D" box="[1275,1352,798,823]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="26" pageNumber="67" unit="km" value="1.8">1.8 km</quantity>
SSE of Mulga Downs Outcamp
</location>
,
<geoCoordinate id="EE1EF0C2FFB18D68A2F8FCF4575E9251" box="[437,567,834,859]" direction="south" orientation="latitude" pageId="26" pageNumber="67" precision="15" value="-22.334723">22°20'05&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFB18D68A109FCF457BD9251" box="[580,724,834,859]" direction="east" orientation="longitude" pageId="26" pageNumber="67" precision="15" value="118.988884">118°59'20&quot;E</geoCoordinate>
,
<date id="FF94B0C5FFB18D68A1ADFCF4563F9251" box="[736,854,834,859]" pageId="26" pageNumber="67" value="2003-11-21">
<collectingDate id="EFD0492DFFB18D68A1ADFCF4563F9251" box="[736,854,834,859]" pageId="26" pageNumber="67" value="2003-11-21">21.xi.2003</collectingDate>
</date>
–
<date id="FF94B0C5FFB18D68A02BFCF456BE9251" box="[870,983,834,859]" pageId="26" pageNumber="67" value="2004-05-24">
<collectingDate id="EFD0492DFFB18D68A02BFCF456BE9251" box="[870,983,834,859]" pageId="26" pageNumber="67" value="2004-05-24">24.v.2004</collectingDate>
</date>
,
<collectionCode id="ED3B0EC0FFB18D68A0A9FCF5515C9251" box="[996,1077,835,859]" pageId="26" pageNumber="67">CALM</collectionCode>
<collectorName id="26DFF3D3FFB18D68A770FCF4518A9251" box="[1085,1251,834,859]" pageId="26" pageNumber="67">Pilbara Survey</collectorName>
(25-058208 ×
<specimenCount id="9D2C5D8CFFB18D68A6C3FCF555C79274" pageId="26" pageNumber="67" type="male">1 ♂</specimenCount>
, ×
<specimenCount id="9D2C5D8CFFB18D68A39FFCD155959274" box="[210,252,871,895]" pageId="26" pageNumber="67" type="female">2 ♀</specimenCount>
in
<collectionCode id="ED3B0EC0FFB18D68A26CFCD154029275" box="[289,363,871,895]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="26" pageNumber="67" type="Museum">WAM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFB18D68A230FCD0518B9275" ID-GBIF-Occurrence="2012745386" box="[381,1250,870,895]" latitude="-16.633333" location="Cape Crawford" longLatPrecision="21" longitude="135.8" pageId="26" pageNumber="67" specimenCount="1" stateProvince="Northern Territory" typeStatus="paratype">
<emphasis id="B95E4A17FFB18D68A230FCD0570D9275" bold="true" box="[381,612,870,895]" pageId="26" pageNumber="67">
<collectingRegion id="49EE58E7FFB18D68A230FCD0570D9275" box="[381,612,870,895]" country="Australia" name="Northern Territory" pageId="26" pageNumber="67">Northern Territory</collectingRegion>
</emphasis>
:
<location id="8EF5C0DEFFB18D68A13EFCD1564B9275" LSID="urn:lsid:plazi:treatment:03832713FFB18D69A3DAFE0151EC92C2:8EF5C0DEFFB18D68A13EFCD1564B9275" box="[627,802,870,895]" latitude="-16.633333" longLatPrecision="21" longitude="135.8" name="Cape Crawford" pageId="26" pageNumber="67" stateProvince="Northern Territory">Cape Crawford</location>
,
<quantity id="4CD23BE0FFB18D68A063FCD156159275" box="[814,892,870,895]" metricMagnitude="3" metricUnit="m" metricValue="9.2" pageId="26" pageNumber="67" unit="km" value="9.2">9.2 km</quantity>
NE, [
<geoCoordinate id="EE1EF0C2FFB18D68A0F2FCD0512B9275" box="[959,1090,870,895]" direction="south" orientation="latitude" pageId="26" pageNumber="67" precision="15" value="-16.633333">16°38'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFB18D68A703FCD0518B9275" box="[1102,1250,870,895]" direction="east" orientation="longitude" pageId="26" pageNumber="67" precision="15" value="135.8">135°48'00&quot;E</geoCoordinate>
</materialsCitation>
<materialsCitation id="3B429C58FFB18D68A7AFFCD0542192E1" ID-GBIF-Occurrence="2012745376" collectingDate="2004-04-17" collectingDateMax="44-11-19" collectingDateMin="2004-04-17" collectionCode="ANIC, QM" collectorName="G. B. Monteith &amp; D. J. Cook &amp; Cape Crawford" latitude="-16.652777" longLatPrecision="21" longitude="135.76389" pageId="26" pageNumber="67" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Northern Territory" typeStatus="paratype">
],
<date id="FF94B0C5FFB18D68A7B8FCD050FE9275" box="[1269,1431,870,895]" pageId="26" pageNumber="67" value="2004-04-17" valueMax="2004-04-19" valueMin="2004-04-17">
<collectingDate id="EFD0492DFFB18D68A7B8FCD050FE9275" box="[1269,1431,870,895]" pageId="26" pageNumber="67" value="2004-04-17" valueMax="2004-04-19" valueMin="2004-04-17">17–19.iv.2004</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFB18D68A3DAFC3D545F92A9" box="[151,310,906,931]" pageId="26" pageNumber="67">G.B. Monteith</collectorName>
&amp;
<collectorName id="26DFF3D3FFB18D68A216FC3D54A592A9" box="[347,460,906,931]" pageId="26" pageNumber="67">D.J. Cook</collectorName>
(T
<date id="FF94B0C5FFB18D68A2A6FC3D572992A9" box="[491,576,907,931]" pageId="26" pageNumber="67" value="40-11-19">
<collectingDate id="EFD0492DFFB18D68A2A6FC3D572992A9" box="[491,576,907,931]" pageId="26" pageNumber="67" value="40-11-19">111940</collectingDate>
</date>
-T
<date id="FF94B0C5FFB18D68A114FC3D57C592A9" box="[601,684,907,931]" pageId="26" pageNumber="67" value="41-11-19">
<collectingDate id="EFD0492DFFB18D68A114FC3D57C592A9" box="[601,684,907,931]" pageId="26" pageNumber="67" value="41-11-19">111941</collectingDate>
</date>
1 ×
<specimenCount id="9D2C5D8CFFB18D68A192FC3D579F92A8" box="[735,758,907,930]" pageId="26" pageNumber="67" type="male">♂</specimenCount>
, 1 ×
<specimenCount id="9D2C5D8CFFB18D68A062FC3D562D92A8" box="[815,836,907,930]" pageId="26" pageNumber="67" type="female">♀</specimenCount>
in
<collectionCode id="ED3B0EC0FFB18D68A025FC3D56C592A9" box="[872,940,907,931]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="26" pageNumber="67">ANIC</collectionCode>
(also as 25-065473), T
<date id="FF94B0C5FFB18D68A7FCFC3D506D92A9" box="[1201,1284,907,931]" pageId="26" pageNumber="67" value="42-11-19">
<collectingDate id="EFD0492DFFB18D68A7FCFC3D506D92A9" box="[1201,1284,907,931]" pageId="26" pageNumber="67" value="42-11-19">111942</collectingDate>
</date>
-T
<date id="FF94B0C5FFB18D68A650FC3D501992A9" box="[1309,1392,907,931]" pageId="26" pageNumber="67" value="44-11-19">
<collectingDate id="EFD0492DFFB18D68A650FC3D501992A9" box="[1309,1392,907,931]" pageId="26" pageNumber="67" value="44-11-19">111944</collectingDate>
</date>
1 ×
<specimenCount id="9D2C5D8CFFB18D68A3DAFC1955C792CC" box="[151,174,943,966]" pageId="26" pageNumber="67" type="male">♂</specimenCount>
, 2 ×
<specimenCount id="9D2C5D8CFFB18D68A3A8FC19559392CC" box="[229,250,943,966]" pageId="26" pageNumber="67" type="female">♀</specimenCount>
in
<collectionCode id="ED3B0EC0FFB18D68A253FC19542692CD" box="[286,335,943,967]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="26" pageNumber="67" type="Museum">QM</collectionCode>
);
<collectorName id="26DFF3D3FFB18D68A22EFC19577892CD" box="[355,529,942,967]" pageId="26" pageNumber="67">Cape Crawford</collectorName>
,
<quantity id="4CD23BE0FFB18D68A150FC19570392CD" box="[541,618,942,967]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="26" pageNumber="67" unit="km" value="4.1">4.1 km</quantity>
NE,
<geoCoordinate id="EE1EF0C2FFB18D68A1E9FC18564F92CD" box="[676,806,942,967]" direction="south" orientation="latitude" pageId="26" pageNumber="67" precision="15" value="-16.652777">16°39'10&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFB18D68A07CFC1856AA92CD" box="[817,963,942,967]" direction="east" orientation="longitude" pageId="26" pageNumber="67" precision="15" value="135.76389">135°45'50&quot;E</geoCoordinate>
,
<date id="FF94B0C5FFB18D68A083FC18510792CD" box="[974,1134,942,967]" pageId="26" pageNumber="67" value="2004-04-17" valueMax="2004-04-19" valueMin="2004-04-17">
<collectingDate id="EFD0492DFFB18D68A083FC18510792CD" box="[974,1134,942,967]" pageId="26" pageNumber="67" value="2004-04-17" valueMax="2004-04-19" valueMin="2004-04-17">17–19.iv.2004</collectingDate>
</date>
(T
<date id="FF94B0C5FFB18D68A7C0FC18518892CD" box="[1165,1249,942,967]" pageId="26" pageNumber="67" value="36-11-19">
<collectingDate id="EFD0492DFFB18D68A7C0FC18518892CD" box="[1165,1249,942,967]" pageId="26" pageNumber="67" value="36-11-19">111936</collectingDate>
</date>
-T
<date id="FF94B0C5FFB18D68A7B7FC18502792CD" box="[1274,1358,942,967]" pageId="26" pageNumber="67" value="39-11-19">
<collectingDate id="EFD0492DFFB18D68A7B7FC18502792CD" box="[1274,1358,942,967]" pageId="26" pageNumber="67" value="39-11-19">111939</collectingDate>
</date>
3 ×
<specimenCount id="9D2C5D8CFFB18D68A6CDFC1950FE92CC" box="[1408,1431,943,966]" pageId="26" pageNumber="67" type="male">♂</specimenCount>
, 1 ×
<specimenCount id="9D2C5D8CFFB18D68A381FC65558892E0" box="[204,225,979,1002]" pageId="26" pageNumber="67" type="female">♀</specimenCount>
in
<collectionCode id="ED3B0EC0FFB18D68A243FC65542992E1" box="[270,320,979,1003]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="26" pageNumber="67" type="Museum">QM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFB18D68A215FC6450FE92E1" ID-GBIF-Occurrence="2012745370" box="[344,1431,978,1003]" collectingDate="2004-04-16" collectingDateMax="31-11-19" collectingDateMin="2004-04-16" collectionCode="QM" latitude="-19.993612" location="Kiama Creek" longLatPrecision="21" longitude="137.76721" municipality="Barkly Highway" pageId="26" pageNumber="67" specimenCount="1" specimenCount-male="1" stateProvince="Northern Territory" typeStatus="paratype">
<collectingMunicipality id="6BF10C7FFFB18D68A215FC64577192E1" box="[344,536,978,1003]" pageId="26" pageNumber="67">Barkly Highway</collectingMunicipality>
,
<location id="8EF5C0DEFFB18D68A16AFC6457AD92E1" LSID="urn:lsid:plazi:treatment:03832713FFB18D69A3DAFE0151EC92C2:8EF5C0DEFFB18D68A16AFC6457AD92E1" box="[551,708,978,1003]" latitude="-19.993612" longLatPrecision="21" longitude="137.76721" municipality="Barkly Highway" name="Kiama Creek" pageId="26" pageNumber="67" stateProvince="Northern Territory">Kiama Creek</location>
,
<geoCoordinate id="EE1EF0C2FFB18D68A19EFC64563D92E1" box="[723,852,978,1003]" direction="south" orientation="latitude" pageId="26" pageNumber="67" precision="15" value="-19.993612">19°59'37&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFB18D68A028FC64568192E1" box="[869,1000,978,1003]" direction="east" orientation="longitude" pageId="26" pageNumber="67" precision="15" value="137.76721">137°46'2&quot;E</geoCoordinate>
,
<date id="FF94B0C5FFB18D68A0B5FC6451F192E1" box="[1016,1176,978,1003]" pageId="26" pageNumber="67" value="2004-04-16" valueMax="2004-04-21" valueMin="2004-04-16">
<collectingDate id="EFD0492DFFB18D68A0B5FC6451F192E1" box="[1016,1176,978,1003]" pageId="26" pageNumber="67" value="2004-04-16" valueMax="2004-04-21" valueMin="2004-04-16">16–21.iv.2004</collectingDate>
</date>
(T
<quantity id="4CD23BE0FFB18D68A7F0FC65505B92E1" box="[1213,1330,978,1003]" metricMagnitude="3" metricUnit="m" metricValue="2.8430473999999997" pageId="26" pageNumber="67" unit="in" value="111931.0">
<date id="FF94B0C5FFB18D68A7F0FC65507892E1" box="[1213,1297,979,1003]" pageId="26" pageNumber="67" value="31-11-19">
<collectingDate id="EFD0492DFFB18D68A7F0FC65507892E1" box="[1213,1297,979,1003]" pageId="26" pageNumber="67" value="31-11-19">111931</collectingDate>
</date>
in
</quantity>
<collectionCode id="ED3B0EC0FFB18D68A670FC65500392E1" box="[1341,1386,979,1003]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="26" pageNumber="67" type="Museum">QM</collectionCode>
<specimenCount id="9D2C5D8CFFB18D68A63BFC6550E492E1" box="[1398,1421,979,1003]" pageId="26" pageNumber="67" type="male">♂</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFB18D68A3DAFC40560C9505" ID-GBIF-Occurrence="2012745381" box="[151,869,1014,1039]" latitude="-19.1" location="159 km S Normanton" longLatPrecision="21" longitude="141.06667" pageId="26" pageNumber="67" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<emphasis id="B95E4A17FFB18D68A3DAFC40544D9505" bold="true" box="[151,292,1014,1039]" pageId="26" pageNumber="67">
<collectingRegion id="49EE58E7FFB18D68A3DAFC40544D9505" box="[151,292,1014,1039]" country="Australia" name="Queensland" pageId="26" pageNumber="67">Queensland</collectingRegion>
</emphasis>
:
<location id="8EF5C0DEFFB18D68A279FC4057479504" LSID="urn:lsid:plazi:treatment:03832713FFB18D69A3DAFE0151EC92C2:8EF5C0DEFFB18D68A279FC4057479504" box="[308,558,1014,1039]" latitude="-19.1" longLatPrecision="21" longitude="141.06667" name="159 km S Normanton" pageId="26" pageNumber="67" stateProvince="Queensland">
<quantity id="4CD23BE0FFB18D68A279FC4054E29505" box="[308,395,1014,1039]" metricMagnitude="5" metricUnit="m" metricValue="1.59" pageId="26" pageNumber="67" unit="km" value="159.0">159 km</quantity>
S Normanton
</location>
, [
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,
<geoCoordinate id="EE1EF0C2FFB18D68A19FFC41560C9505" box="[722,869,1015,1039]" direction="east" orientation="longitude" pageId="26" pageNumber="67" precision="15" value="141.06667">141°04'00&quot;E</geoCoordinate>
</materialsCitation>
<materialsCitation id="3B429C58FFB18D68A028FC4157B2955D" ID-GBIF-Occurrence="2012745362" collectingDate="1976-03-11" collectingDateMax="34-03-26" collectingDateMin="1976-03-11" collectionCode="QDPC, CMNC, QM" collectorName="R. I. Storey &amp; Project" latitude="-20.9775" longLatPrecision="21" longitude="141.80112" pageId="26" pageNumber="67" specimenCode="CMNEN-00030249" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Queensland" typeStatus="paratype">
],
<date id="FF94B0C5FFB18D68A034FC4051779505" box="[889,1054,1014,1039]" pageId="26" pageNumber="67" value="1976-03-11" valueMax="1976-03-13" valueMin="1976-03-11">
<collectingDate id="EFD0492DFFB18D68A034FC4051779505" box="[889,1054,1014,1039]" pageId="26" pageNumber="67" value="1976-03-11" valueMax="1976-03-13" valueMin="1976-03-11">11–13.iii.1976</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFB18D68A766FC4151CE9505" box="[1067,1191,1015,1039]" pageId="26" pageNumber="67">R.I. Storey</collectorName>
(25-
<date id="FF94B0C5FFB18D68A794FC4050589505" box="[1241,1329,1014,1039]" pageId="26" pageNumber="67" value="34-03-26">
<collectingDate id="EFD0492DFFB18D68A794FC4050589505" box="[1241,1329,1014,1039]" pageId="26" pageNumber="67" value="34-03-26">032634</collectingDate>
</date>
×
<specimenCount id="9D2C5D8CFFB18D68A61FFC4150149504" box="[1362,1405,1015,1039]" pageId="26" pageNumber="67" type="female">1 ♀</specimenCount>
in
<collectionCode id="ED3B0EC0FFB18D68A3DAFBAD558C9539" box="[151,229,1051,1075]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34157" lsid="urn:lsid:biocol.org:col:34157" name="Queensland Primary Industries Insect Collection" pageId="26" pageNumber="67" type="Other type of research institution/biorepository">QDPC</collectionCode>
,
<specimenCode id="DB8C3E7EFFB18D68A3B6FBAD54B09539" box="[251,473,1051,1075]" collectionCode="CMNEN-" pageId="26" pageNumber="67">CMNEN-00030249</specimenCode>
in
<collectionCode id="ED3B0EC0FFB18D68A15EFBAD570F9539" box="[531,614,1051,1075]" pageId="26" pageNumber="67">CMNC</collectionCode>
<specimenCount id="9D2C5D8CFFB18D68A135FBAD57E59539" box="[632,652,1051,1075]" pageId="26" pageNumber="67" type="female">♀</specimenCount>
);
<collectingMethod id="526BEE12FFB18D68A1FDFBAC51769538" box="[688,1055,1050,1075]" pageId="26" pageNumber="67">Toorak Research Station Trap</collectingMethod>
1, NHT
<collectorName id="26DFF3D3FFB18D68A7E9FBAC519F9539" box="[1188,1270,1050,1075]" pageId="26" pageNumber="67">Project</collectorName>
, [
<geoCoordinate id="EE1EF0C2FFB18D68A659FBAC50FE9539" box="[1300,1431,1050,1075]" direction="south" orientation="latitude" pageId="26" pageNumber="67" precision="15" value="-20.9775">20°58'39&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFB18D68A3DAFB895442955D" box="[151,299,1087,1111]" direction="east" orientation="longitude" pageId="26" pageNumber="67" precision="15" value="141.80112">141°48'04&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFB18D68A273FB8854D9955D" box="[318,432,1086,1111]" pageId="26" pageNumber="67" value="2002-02-10">
<collectingDate id="EFD0492DFFB18D68A273FB8854D9955D" box="[318,432,1086,1111]" pageId="26" pageNumber="67" value="2002-02-10">10.ii.2002</collectingDate>
</date>
(T
<quantity id="4CD23BE0FFB18D68A29CFB89572C955D" box="[465,581,1086,1111]" metricMagnitude="3" metricUnit="m" metricValue="3.5311841999999998" pageId="26" pageNumber="67" unit="in" value="139023.0">139023 in</quantity>
×
<specimenCount id="9D2C5D8CFFB18D68A123FB8957F0955C" box="[622,665,1087,1111]" pageId="26" pageNumber="67" type="male">1 ♂</specimenCount>
<collectionCode id="ED3B0EC0FFB18D68A1EFFB8957BD955D" box="[674,724,1087,1111]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="26" pageNumber="67" type="Museum">QM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFB18D68A1A5FB8856159571" ID-GBIF-Occurrence="2012745385" collectingDate="2007-02-11" collectingDateMax="2007-02-12" collectingDateMin="2007-02-11" collectionCode="QM" collectorName="G. B. Monteith" latitude="-19.815832" location="Fletcher Creek Rest Area" longLatPrecision="21" longitude="146.0539" pageId="26" pageNumber="67" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<location id="8EF5C0DEFFB18D68A1A5FB885164955D" LSID="urn:lsid:plazi:treatment:03832713FFB18D69A3DAFE0151EC92C2:8EF5C0DEFFB18D68A1A5FB885164955D" box="[744,1037,1086,1111]" latitude="-19.815832" longLatPrecision="21" longitude="146.0539" name="Fletcher Creek Rest Area" pageId="26" pageNumber="67" stateProvince="Queensland">Fletcher Creek Rest Area</location>
,14895, [
<geoCoordinate id="EE1EF0C2FFB18D68A721FB885199955D" box="[1132,1264,1086,1111]" direction="south" orientation="latitude" pageId="26" pageNumber="67" precision="15" value="-19.815832">19°48'57&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFB18D68A7B0FB8850F8955D" box="[1277,1425,1086,1111]" direction="east" orientation="longitude" pageId="26" pageNumber="67" precision="15" value="146.0539">146°03'14&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFB18D68A3DAFBD4545D9571" box="[151,308,1122,1147]" pageId="26" pageNumber="67" value="2007-02-11" valueMax="2007-02-12" valueMin="2007-02-11">
<collectingDate id="EFD0492DFFB18D68A3DAFBD4545D9571" box="[151,308,1122,1147]" pageId="26" pageNumber="67" value="2007-02-11" valueMax="2007-02-12" valueMin="2007-02-11">11–12.ii.2007</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFB18D68A20BFBD554879571" box="[326,494,1122,1147]" pageId="26" pageNumber="67">G.B. Monteith</collectorName>
, dung pit fall (T
<quantity id="4CD23BE0FFB18D68A1F3FBD4565E9571" box="[702,823,1122,1147]" metricMagnitude="3" metricUnit="m" metricValue="3.656711" pageId="26" pageNumber="67" unit="in" value="143965.0">143965 in</quantity>
<collectionCode id="ED3B0EC0FFB18D68A008FBD5561D9571" box="[837,884,1123,1147]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="26" pageNumber="67" type="Museum">QM</collectionCode>
)
</materialsCitation>
<specimenCount id="9D2C5D8CFFB18D68A0C4FBD556C89570" box="[905,929,1123,1146]" pageId="26" pageNumber="67" type="male">♂</specimenCount>
; Beames Brook
<materialsCitation id="3B429C58FFB18D68A723FBD454C795C9" ID-GBIF-Occurrence="2012745368" collectingDate="1972-05-20" collectionCode="QM" collectorName="G. B. &amp; S. R. Monteith" latitude="-17.962778" location="Burketown" longLatPrecision="21" longitude="139.50555" pageId="26" pageNumber="67" specimenCode="UQIC 71247, ANIC 25-058521" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="paratype">
<quantity id="4CD23BE0FFB18D68A723FBD451DC9571" box="[1134,1205,1122,1147]" metricMagnitude="4" metricUnit="m" metricValue="2.414016" pageId="26" pageNumber="67" unit="mi" value="15.0">15 mi</quantity>
SW of
<location id="8EF5C0DEFFB18D68A66DFBD450F49571" LSID="urn:lsid:plazi:treatment:03832713FFB18D69A3DAFE0151EC92C2:8EF5C0DEFFB18D68A66DFBD450F49571" box="[1312,1437,1122,1147]" latitude="-17.962778" longLatPrecision="21" longitude="139.50555" name="Burketown" pageId="26" pageNumber="67" stateProvince="Queensland">Burketown</location>
[
<geoCoordinate id="EE1EF0C2FFB18D68A3D3FB30544E9595" box="[158,295,1158,1183]" direction="south" orientation="latitude" pageId="26" pageNumber="67" precision="15" value="-17.962778">17°57’46”S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFB18D68A27EFB3154A59595" box="[307,460,1159,1183]" direction="east" orientation="longitude" pageId="26" pageNumber="67" precision="15" value="139.50555">139°30’20”E</geoCoordinate>
],
<date id="FF94B0C5FFB18D68A292FB3157399595" box="[479,592,1159,1183]" pageId="26" pageNumber="67" value="1972-05-20">
<collectingDate id="EFD0492DFFB18D68A292FB3157399595" box="[479,592,1159,1183]" pageId="26" pageNumber="67" value="1972-05-20">20.v.1972</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFB18D68A111FB31563C9595" box="[604,853,1158,1183]" pageId="26" pageNumber="67">G.B. &amp; S.R. Monteith</collectorName>
, mercury vapour lamp (
<specimenCode id="DB8C3E7EFFB18D68A72CFB3151919595" box="[1121,1272,1159,1183]" collectionCode="UQIC" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34515" lsid="urn:lsid:biocol.org:col:34515" name="University of Queensland Insect Collection" pageId="26" pageNumber="67">UQIC 71247</specimenCode>
, also as
<specimenCode id="DB8C3E7EFFB18D68A615FB31547995C9" collectionCode="ANIC" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="26" pageNumber="67">ANIC 25-058521</specimenCode>
×
<specimenCount id="9D2C5D8CFFB18D68A263FB1D543895C9" box="[302,337,1195,1219]" pageId="26" pageNumber="67" type="female">1♀</specimenCount>
in
<collectionCode id="ED3B0EC0FFB18D68A238FB1D54CF95C9" box="[373,422,1195,1219]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="26" pageNumber="67" type="Museum">QM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B959605FFB18D68A38AFB7854D79491" blockId="26.[151,1437,546,2011]" pageId="26" pageNumber="67">
<emphasis id="B95E4A17FFB18D68A38AFB78576995ED" bold="true" box="[199,512,1230,1255]" pageId="26" pageNumber="67">Other material examined.</emphasis>
Additional specimens from three localities of the CALM survey not included in the
<typeStatus id="549128A7FFB18D68A38FFB45544B9401" box="[194,290,1267,1291]" pageId="26" pageNumber="67" type="paratype">paratype</typeStatus>
series above were also examined.
<materialsCitation id="3B429C58FFB18D68A1F8FB45566E9425" ID-GBIF-Occurrence="2012745361" collectingDate="2004-05-08" collectingDateMax="2004-10-11" collectingDateMin="2004-05-08" collectionCode="CALM" latitude="-21.394722" location="4 km N of Barowanna Hill" longLatPrecision="21" longitude="117.170555" pageId="26" pageNumber="67" specimenCount="1" stateProvince="Western Australia">
<collectingRegion id="49EE58E7FFB18D68A1F8FB4556ED9401" box="[693,900,1266,1291]" country="Australia" name="Western Australia" pageId="26" pageNumber="67">Western Australia</collectingRegion>
:
<location id="8EF5C0DEFFB18D68A0DCFB4551A19401" LSID="urn:lsid:plazi:treatment:03832713FFB18D69A3DAFE0151EC92C2:8EF5C0DEFFB18D68A0DCFB4551A19401" box="[913,1224,1266,1291]" latitude="-21.394722" longLatPrecision="21" longitude="117.170555" name="4 km N of Barowanna Hill" pageId="26" pageNumber="67" stateProvince="Western Australia">
<quantity id="4CD23BE0FFB18D68A0DCFB4556A59401" box="[913,972,1266,1291]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="67" unit="km" value="4.0">4 km</quantity>
N of Barowanna Hill
</location>
(Camp Curlewis),
<geoCoordinate id="EE1EF0C2FFB18D68A3DAFAA154709425" box="[151,281,1303,1327]" direction="south" orientation="latitude" pageId="26" pageNumber="67" precision="15" value="-21.394722">21°23'41&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFB18D68A265FAA154D19425" box="[296,440,1303,1327]" direction="east" orientation="longitude" pageId="26" pageNumber="67" precision="15" value="117.170555">117°10'14&quot;E</geoCoordinate>
,
<date id="FF94B0C5FFB18D68A28AFAA157409425" box="[455,553,1303,1327]" pageId="26" pageNumber="67" value="2004-05-08">
<collectingDate id="EFD0492DFFB18D68A28AFAA157409425" box="[455,553,1303,1327]" pageId="26" pageNumber="67" value="2004-05-08">8.v.2004</collectingDate>
</date>
–
<date id="FF94B0C5FFB18D68A174FAA157CE9425" box="[569,679,1303,1327]" pageId="26" pageNumber="67" value="2004-10-11">
<collectingDate id="EFD0492DFFB18D68A174FAA157CE9425" box="[569,679,1303,1327]" pageId="26" pageNumber="67" value="2004-10-11">11.x.2004</collectingDate>
</date>
,
<collectionCode id="ED3B0EC0FFB18D68A1FBFAA1566E9425" box="[694,775,1303,1327]" pageId="26" pageNumber="67">CALM</collectionCode>
</materialsCitation>
<materialsCitation id="3B429C58FFB18D68A05FFAA051939425" ID-GBIF-Occurrence="2012745364" box="[786,1274,1302,1327]" collectingDate="1999-10" collectionCode="ANIC" location="Pilbara Survey" pageId="26" pageNumber="67" specimenCount="1" stateProvince="Western Australia">
<location id="8EF5C0DEFFB18D68A05FFAA056D29425" LSID="urn:lsid:plazi:treatment:03832713FFB18D69A3DAFE0151EC92C2:8EF5C0DEFFB18D68A05FFAA056D29425" box="[786,955,1302,1327]" name="Pilbara Survey" pageId="26" pageNumber="67" stateProvince="Western Australia">Pilbara Survey</location>
(25-058205
<date id="FF94B0C5FFB18D68A71EFAA151149425" box="[1107,1149,1303,1327]" pageId="26" pageNumber="67" value="1999-10">
<collectingDate id="EFD0492DFFB18D68A71EFAA151149425" box="[1107,1149,1303,1327]" pageId="26" pageNumber="67" value="1999-10">
x
<quantity id="4CD23BE0FFB18D68A72CFAA151149425" box="[1121,1149,1303,1327]" metricMagnitude="0" metricUnit="m" metricValue="2.5146" pageId="26" pageNumber="67" unit="in" value="99.0">99</quantity>
</collectingDate>
</date>
in
<collectionCode id="ED3B0EC0FFB18D68A7E5FAA151989425" box="[1192,1265,1303,1327]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="26" pageNumber="67">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFB18D68A644FAA056F99459" ID-GBIF-Occurrence="2012745369" collectingDate="2003-11-24" collectingDateMax="2004-05-07" collectingDateMin="2003-11-24" collectionCode="CALM" latitude="-21.6425" location="Millstream" longLatPrecision="21" longitude="117.03694" pageId="26" pageNumber="67" specimenCount="1" stateProvince="Western Australia">
<quantity id="4CD23BE0FFB18D68A644FAA0502C9425" box="[1289,1349,1302,1327]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="26" pageNumber="67" unit="km" value="6.0">6 km</quantity>
SW of
<location id="8EF5C0DEFFB18D68A3DAFA8C547C9459" LSID="urn:lsid:plazi:treatment:03832713FFB18D69A3DAFE0151EC92C2:8EF5C0DEFFB18D68A3DAFA8C547C9459" box="[151,277,1338,1363]" latitude="-21.6425" longLatPrecision="21" longitude="117.03694" name="Millstream" pageId="26" pageNumber="67" stateProvince="Western Australia">Millstream</location>
,
<geoCoordinate id="EE1EF0C2FFB18D68A26DFA8D54CB9459" box="[288,418,1339,1363]" direction="south" orientation="latitude" pageId="26" pageNumber="67" precision="15" value="-21.6425">21°38'33&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFB18D68A2E3FA8D57579459" box="[430,574,1339,1363]" direction="east" orientation="longitude" pageId="26" pageNumber="67" precision="15" value="117.03694">117°02'13&quot;E</geoCoordinate>
,
<date id="FF94B0C5FFB18D68A107FA8C57AB9459" box="[586,706,1338,1363]" pageId="26" pageNumber="67" value="2003-11-24">
<collectingDate id="EFD0492DFFB18D68A107FA8C57AB9459" box="[586,706,1338,1363]" pageId="26" pageNumber="67" value="2003-11-24">24.xi.2003</collectingDate>
</date>
–
<date id="FF94B0C5FFB18D68A19FFA8C565B9459" box="[722,818,1338,1363]" pageId="26" pageNumber="67" value="2004-05-07">
<collectingDate id="EFD0492DFFB18D68A19FFA8C565B9459" box="[722,818,1338,1363]" pageId="26" pageNumber="67" value="2004-05-07">7.v.2004</collectingDate>
</date>
,
<collectionCode id="ED3B0EC0FFB18D68A072FA8D56F99459" box="[831,912,1339,1363]" pageId="26" pageNumber="67">CALM</collectionCode>
</materialsCitation>
<materialsCitation id="3B429C58FFB18D68A0DAFA8C50049459" ID-GBIF-Occurrence="2012745358" box="[919,1389,1338,1363]" collectingDate="1934-10" collectionCode="ANIC" location="Pilbara Survey" pageId="26" pageNumber="67" specimenCount="1" stateProvince="Western Australia">
<location id="8EF5C0DEFFB18D68A0DAFA8C51549459" LSID="urn:lsid:plazi:treatment:03832713FFB18D69A3DAFE0151EC92C2:8EF5C0DEFFB18D68A0DAFA8C51549459" box="[919,1085,1338,1363]" name="Pilbara Survey" pageId="26" pageNumber="67" stateProvince="Western Australia">Pilbara Survey</location>
(25-058211
<date id="FF94B0C5FFB18D68A780FA8D519E9459" box="[1229,1271,1339,1363]" pageId="26" pageNumber="67" value="1934-10">
<collectingDate id="EFD0492DFFB18D68A780FA8D519E9459" box="[1229,1271,1339,1363]" pageId="26" pageNumber="67" value="1934-10">
x
<quantity id="4CD23BE0FFB18D68A796FA8D519E9459" box="[1243,1271,1339,1363]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" pageId="26" pageNumber="67" unit="in" value="34.0">34</quantity>
</collectingDate>
</date>
in
<collectionCode id="ED3B0EC0FFB18D68A651FA8D500C9459" box="[1308,1381,1339,1363]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="26" pageNumber="67">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFB18D68A634FA8C5149947D" ID-GBIF-Occurrence="2012745367" collectingDate="2003-11-25" collectingDateMax="2004-05-06" collectingDateMin="2003-11-25" collectionCode="CALM" latitude="-21.864723" location="Coolawanyah" longLatPrecision="21" longitude="117.78694" pageId="26" pageNumber="67" specimenCount="1" stateProvince="Western Australia">
<quantity id="4CD23BE0FFB18D68A634FA8C55D2947D" metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="26" pageNumber="67" unit="km" value="6.5">6.5 km</quantity>
SSW
<location id="8EF5C0DEFFB18D68A248FAE854CB947D" LSID="urn:lsid:plazi:treatment:03832713FFB18D69A3DAFE0151EC92C2:8EF5C0DEFFB18D68A248FAE854CB947D" box="[261,418,1374,1399]" latitude="-21.864723" longLatPrecision="21" longitude="117.78694" name="Coolawanyah" pageId="26" pageNumber="67" stateProvince="Western Australia">Coolawanyah</location>
,
<geoCoordinate id="EE1EF0C2FFB18D68A2E3FAE85759947D" box="[430,560,1374,1399]" direction="south" orientation="latitude" pageId="26" pageNumber="67" precision="15" value="-21.864723">21°51'53&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFB18D68A170FAE957A5947D" box="[573,716,1375,1399]" direction="east" orientation="longitude" pageId="26" pageNumber="67" precision="15" value="117.78694">117°47'13&quot;E</geoCoordinate>
,
<date id="FF94B0C5FFB18D68A194FAE85638947D" box="[729,849,1374,1399]" pageId="26" pageNumber="67" value="2003-11-25">
<collectingDate id="EFD0492DFFB18D68A194FAE85638947D" box="[729,849,1374,1399]" pageId="26" pageNumber="67" value="2003-11-25">25.xi.2003</collectingDate>
</date>
–
<date id="FF94B0C5FFB18D68A02CFAE856AB947D" box="[865,962,1374,1399]" pageId="26" pageNumber="67" value="2004-05-06">
<collectingDate id="EFD0492DFFB18D68A02CFAE856AB947D" box="[865,962,1374,1399]" pageId="26" pageNumber="67" value="2004-05-06">6.v.2004</collectingDate>
</date>
,
<collectionCode id="ED3B0EC0FFB18D68A082FAE95149947D" box="[975,1056,1375,1399]" pageId="26" pageNumber="67">CALM</collectionCode>
</materialsCitation>
<materialsCitation id="3B429C58FFB18D68A765FAE854D39491" ID-GBIF-Occurrence="2012745357" collectingDate="x 1" collectionCode="ANIC" location="Pilbara Survey" pageId="26" pageNumber="67" specimenCount="1" stateProvince="Western Australia">
<location id="8EF5C0DEFFB18D68A765FAE851A6947D" LSID="urn:lsid:plazi:treatment:03832713FFB18D69A3DAFE0151EC92C2:8EF5C0DEFFB18D68A765FAE851A6947D" box="[1064,1231,1374,1399]" name="Pilbara Survey" pageId="26" pageNumber="67" stateProvince="Western Australia">Pilbara Survey</location>
(25-058209
<date id="FF94B0C5FFB18D68A62FFAE95017947D" box="[1378,1406,1375,1399]" pageId="26" pageNumber="67">
<collectingDate id="EFD0492DFFB18D68A62FFAE95017947D" box="[1378,1406,1375,1399]" pageId="26" pageNumber="67">
x
<quantity id="4CD23BE0FFB18D68A63EFAE95017947D" box="[1395,1406,1375,1399]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="26" pageNumber="67" unit="in" value="1.0">1</quantity>
</collectingDate>
</date>
in
<collectionCode id="ED3B0EC0FFB18D68A3DAFA3555B49491" box="[151,221,1411,1435]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="26" pageNumber="67">ANIC</collectionCode>
) (partial specimen)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFB18D69A38AFA1051A3904A" lastPageId="27" lastPageNumber="68" pageId="26" pageNumber="67" type="description">
<paragraph id="8B959605FFB18D68A38AFA1050FF94B5" blockId="26.[151,1437,546,2011]" box="[199,1430,1446,1471]" pageId="26" pageNumber="67">
<emphasis id="B95E4A17FFB18D68A38AFA10543094B5" bold="true" box="[199,345,1446,1471]" pageId="26" pageNumber="67">Description.</emphasis>
Bicoloured; pronotum and elytra light brown, head black with brown edges. Antennal club grey.
</paragraph>
<paragraph id="8B959605FFB18D68A38AFA7C519B94E8" blockId="26.[151,1437,546,2011]" box="[199,1266,1482,1507]" pageId="26" pageNumber="67">
Total length: 2.4–3.0 mm.
<typeStatus id="549128A7FFB18D68A2BDFA7C573194E9" box="[496,600,1482,1507]" pageId="26" pageNumber="67" type="holotype">Holotype</typeStatus>
measurements: Total length
<quantity id="4CD23BE0FFB18D68A0D1FA7D569D94E8" box="[924,1012,1483,1507]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="26" pageNumber="67" unit="mm" value="2.8">2.8 mm</quantity>
, elytral width
<quantity id="4CD23BE0FFB18D68A7D8FA7D518794E8" box="[1173,1262,1483,1507]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="26" pageNumber="67" unit="mm" value="1.7">1.7 mm</quantity>
.
</paragraph>
<paragraph id="8B959605FFB18D68A38AFA5851F09779" blockId="26.[151,1437,546,2011]" pageId="26" pageNumber="67">Male: Head: U-shaped between clypeal teeth, which are upturned. Margin slightly angulate at clypeogenal suture. Genal angle weakly defined. Basal carina present but interrupted medially. Head reticulate between clypeal suture and anterior edge. Dorsal part of eyes narrow in size, separated by interocular space approximately 17–18 times eye width (35: 2); eye canthus not dividing eye. Head 1.25 × wider than long (43:34).</paragraph>
<paragraph id="8B959605FFB18D68A38AF9C8578297B1" blockId="26.[151,1437,546,2011]" pageId="26" pageNumber="67">Prothorax: Hypomeral surface finely reticulate with light punctation. Hypomeral striae half length of hypomeron. Pronotum 1.6 × wider than long (74:46).</paragraph>
<paragraph id="8B959605FFB18D68A38AF97051259609" blockId="26.[151,1437,546,2011]" pageId="26" pageNumber="67">Elytra: Surface smooth, intervals with extremely fine setae extending backwards. Striae double with very fine punctate edges. All striae on disc equal width. Elytra 1.15 × wider than long (87:75).</paragraph>
<paragraph id="8B959605FFB18D68A38AF8B954DC9665" blockId="26.[151,1437,546,2011]" pageId="26" pageNumber="67">Legs: Front edge of protibiae deeply angulate; apical digit elongate and sharp; 2 teeth on outer edge. Inner edge of metatibiae gently curved and with a slight flange. Metatarsi with 5th tarsomere longest. Tarsal claws intermediate and toothed.</paragraph>
<paragraph id="8B959605FFB18D68A38AF8CC504896D1" blockId="26.[151,1437,546,2011]" pageId="26" pageNumber="67">Abdomen: Pygidium with a variably round to ovoid shaped depression, containing a pillar-shaped tubercle. Depression occupies approximately half size of disc and extends under the concave upper edge of pygidium. Abdominal surface is very finely reticulate right across. Segment 6 finely punctate and finely reticulate.</paragraph>
<paragraph id="8B959605FFB08D69A38AFF2154A69017" blockId="27.[151,1436,151,968]" pageId="27" pageNumber="68">Pterothorax: Medial lobe of metaventrite finely punctate and broadly margined between mesocoxae. Lateral lobe of metaventrite punctate and finely reticulate. Meso-metaventral suture very slightly arched. Mesoventrite with slight notch anteriorly. Mesoventrite finely reticulate and punctate at least at sides. Mesepimeron and metanepisternum reticulate.</paragraph>
<paragraph id="8B959605FFB08D69A38AFE9151A3904A" blockId="27.[151,1436,151,968]" box="[199,1226,295,320]" pageId="27" pageNumber="68">
Female: As per the
<taxonomicName id="4C2AED86FFB08D69A2EFFE9F5759904A" authorityName=": Matthews" authorityYear="1974" box="[418,560,296,320]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFB08D69A2EFFE9F54D8904A" box="[418,433,297,320]" italics="true" pageId="27" pageNumber="68">L</emphasis>
.
<emphasis id="B95E4A17FFB08D69A28DFE9F5759904A" box="[448,560,297,320]" italics="true" pageId="27" pageNumber="68">pygmaeus</emphasis>
</taxonomicName>
species group and pygidial depression slightly more ovoid.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFB08D69A38AFEFA518C9082" pageId="27" pageNumber="68" type="etymology">
<paragraph id="8B959605FFB08D69A38AFEFA518C9082" blockId="27.[151,1436,151,968]" pageId="27" pageNumber="68">
<emphasis id="B95E4A17FFB08D69A38AFEFA5427906F" bold="true" box="[199,334,332,357]" pageId="27" pageNumber="68">Etymology.</emphasis>
Named in honour of Nadine Guthrie, entomologist with the
<collectingRegion id="49EE58E7FFB08D69A753FEFB5182906F" box="[1054,1259,332,357]" country="Australia" name="Western Australia" pageId="27" pageNumber="68">Western Australia</collectingRegion>
Department of Environment and Conservation, who collected much of the material for the CALM Pilbara survey.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFB08D69A38AFE225411938F" pageId="27" pageNumber="68" type="distribution">
<paragraph id="8B959605FFB08D69A38AFE225411938F" blockId="27.[151,1436,151,968]" pageId="27" pageNumber="68">
<emphasis id="B95E4A17FFB08D69A38AFE22540990A7" bold="true" box="[199,352,404,429]" pageId="27" pageNumber="68">Distribution.</emphasis>
A widespread species known from the Western Australian PIL bioregion,
<collectingRegion id="49EE58E7FFB08D69A784FE2250F590A7" box="[1225,1436,404,429]" country="Australia" name="Northern Territory" pageId="27" pageNumber="68">Northern Territory</collectingRegion>
GFU bioregion, and
<collectingRegion id="49EE58E7FFB08D69A2CBFE01576390DA" box="[390,522,439,464]" country="Australia" name="Queensland" pageId="27" pageNumber="68">Queensland</collectingRegion>
GUP and EIU bioregions (
<figureCitation id="13118A80FFB08D69A005FE0156F790DA" box="[840,926,439,464]" captionStart="FIGURE 16" captionStartId="35.[151,250,1726,1748]" captionTargetBox="[151,1436,1037,1705]" captionTargetId="figure@35.[151,1436,1037,1705]" captionTargetPageId="35" captionText="FIGURE 16. Map of known records of Lepanus guthrieae including predicted distribution overlaid with predicted distribution boundary of L. pygmaeus (dashed line) taken from Fig. 13 map." httpUri="https://zenodo.org/record/2588653/files/figure.png" pageId="27" pageNumber="68">Fig. 16</figureCitation>
). Given the disjunct distribution and limited sampling in much of northern
<collectingCountry id="F33DD695FFB08D69A2BAFE6A570990FF" box="[503,608,476,501]" name="Australia" pageId="27" pageNumber="68">Australia</collectingCountry>
, this species is likely to occur in numerous other IBRA regions such as those predicted by BIOCLIM. Our BIOCLIM model predicted
<taxonomicName id="4C2AED86FFB08D69A028FDB756999312" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[869,1008,511,536]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="species" species="guthrieae">
<emphasis id="B95E4A17FFB08D69A028FDB756999312" box="[869,1008,511,536]" italics="true" pageId="27" pageNumber="68">L. guthrieae</emphasis>
</taxonomicName>
may also occur in Carnarvon (
<collectingCountry id="F33DD695FFB08D69A61DFDB650E69312" box="[1360,1423,512,536]" name="Central African Republic" pageId="27" pageNumber="68">CAR</collectingCountry>
), Gascoyne (GAS), Little Sandy Desert (LSD), Great Sandy Desert (GSD), DAL, CEK, OVP, STU, GUC, CEA, ARC, Tanami (TAN), Davenport Murchison Ranges (DMR), Mitchell Grass Downs (MGD), MII, DEU, BBN, and CYP IBRA regions.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFB08D69A38AFD3951EC92C2" pageId="27" pageNumber="68" type="discussion">
<paragraph id="8B959605FFB08D69A38AFD3951AE9232" blockId="27.[151,1436,151,968]" pageId="27" pageNumber="68">
<emphasis id="B95E4A17FFB08D69A38AFD39542693A2" bold="true" box="[199,335,655,680]" pageId="27" pageNumber="68">Comments.</emphasis>
<taxonomicName id="4C2AED86FFB08D69A210FD27575C93A2" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[349,565,655,680]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="species" species="guthrieae">
<emphasis id="B95E4A17FFB08D69A210FD27575C93A2" box="[349,565,655,680]" italics="true" pageId="27" pageNumber="68">Lepanus guthrieae</emphasis>
</taxonomicName>
has previously been referred to as “
<taxonomicName id="4C2AED86FFB08D69A0B1FD27513293A2" authorityName=", Matthews" authorityYear="1974" box="[1020,1115,657,680]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FFB08D69A0B1FD27513293A2" box="[1020,1115,657,680]" italics="true" pageId="27" pageNumber="68">Lepanus</emphasis>
</taxonomicName>
WA2” under the informal nomenclature sensu
<bibRefCitation id="EFBBEBF4FFB08D69A2CDFD03570A93C6" author="Yeates, D. &amp; Gunter, N. L. &amp; Weir, T. &amp; Cameron, S. L." box="[384,611,692,717]" pageId="27" pageNumber="68" pagination="92 - 99" refId="ref30201" refString="Yeates, D. K, Gunter, N. L., Weir, T. &amp; Cameron, S. L. (2011) Developing a benchmark invertebrate group for biodiversity modelling. In: West, J. (Ed.), 21 st Century Taxonomy: accelerating research and discovery of Australia's biodiversity, Taxonomy Research and Identification Network Final Report- March, 2011, pp. 92 - 99. Available from: https: // www. environment. gov. au / system / files / pages / 56 c 08 f 6 f- 7 bf 7 - 4594 - 8175 - b 4 c 143110156 / files / hub-taxonomy. pdf (accessed 12 December 2018)" type="journal article" year="2011">
Yeates
<emphasis id="B95E4A17FFB08D69A29EFD03576E93C7" box="[467,519,692,717]" italics="true" pageId="27" pageNumber="68">et al</emphasis>
. (2011)
</bibRefCitation>
. A specimen (COL 2465) from Cape Crawford,
<collectingRegion id="49EE58E7FFB08D69A7D8FD02500E93C7" box="[1173,1383,692,717]" country="Australia" name="Northern Territory" pageId="27" pageNumber="68">Northern Territory</collectingRegion>
was sequenced in the study of
<bibRefCitation id="EFBBEBF4FFB08D69A2F1FD6E57F293FA" author="Gunter, N. L. &amp; Monteith, G. B. &amp; Cameron, S. L. &amp; Weir, T." box="[444,667,727,752]" pageId="27" pageNumber="68" pagination="1 - 27" refId="ref29280" refString="Gunter, N. L., Monteith, G. B., Cameron, S. L. &amp; Weir, T. A (2018) Evidence from Australian mesic zone dung beetles support Gondwanan vicariance and Mesozoic origin of the Scarabaeinae (Coleoptera: Scarabaeidae). Invertebrate Systematics and Evolution, 1 - 27. [Advance online publication] https: // doi. org / 10.1163 / 1876312 X- 00002171" type="book chapter" year="2018">
Gunter
<emphasis id="B95E4A17FFB08D69A15DFD6F572193FA" box="[528,584,727,752]" italics="true" pageId="27" pageNumber="68">et al.</emphasis>
(2018)
</bibRefCitation>
is designated as a
<typeStatus id="549128A7FFB08D69A020FD6E56A793FA" box="[877,974,728,752]" pageId="27" pageNumber="68" type="paratype">paratype</typeStatus>
here (ANIC) and was recovered as sister to
<taxonomicName id="4C2AED86FFB08D69A3F8FD4B542F921F" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[181,326,764,789]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="species" species="pungalina">
<emphasis id="B95E4A17FFB08D69A3F8FD4B542F921F" box="[181,326,764,789]" italics="true" pageId="27" pageNumber="68">L. pungalina</emphasis>
</taxonomicName>
within a member of the monsoonal clade of
<taxonomicName id="4C2AED86FFB08D69A004FD4B56C1921E" authorityName=", Matthews" authorityYear="1974" box="[841,936,765,788]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FFB08D69A004FD4B56C1921E" box="[841,936,765,788]" italics="true" pageId="27" pageNumber="68">Lepanus</emphasis>
</taxonomicName>
, which contains species in the
<taxonomicName id="4C2AED86FFB08D69A640FD4B50F5921E" authorityName=": Matthews" authorityYear="1974" box="[1293,1436,765,788]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFB08D69A640FD4B50F5921E" box="[1293,1436,765,788]" italics="true" pageId="27" pageNumber="68">L. pygmaeus</emphasis>
</taxonomicName>
species group. All specimens of
<taxonomicName id="4C2AED86FFB08D69A14EFC9757E59232" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[515,652,799,824]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="species" species="guthrieae">
<emphasis id="B95E4A17FFB08D69A14EFC9757E59232" box="[515,652,799,824]" italics="true" pageId="27" pageNumber="68">L. guthrieae</emphasis>
</taxonomicName>
have been collected in pitfall traps in open forests.
</paragraph>
<paragraph id="8B959605FFB08D69A38AFCF251EC92C2" blockId="27.[151,1436,151,968]" pageId="27" pageNumber="68">
A specimen from Beames Brook,
<quantity id="4CD23BE0FFB08D69A12EFCF257DB9257" box="[611,690,836,861]" metricMagnitude="4" metricUnit="m" metricValue="2.414016" pageId="27" pageNumber="68" unit="mi" value="15.0">15 mi.</quantity>
SW Burketown (25-058521 in QM) listed as
<taxonomicName id="4C2AED86FFB08D69A7A8FCF35433928A" authority="in Matthews (1974)" authorityName="in Matthews" authorityYear="1974" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFB08D69A7A8FCF350109256" box="[1253,1401,837,860]" italics="true" pageId="27" pageNumber="68">L. pygmaeus</emphasis>
in
<bibRefCitation id="EFBBEBF4FFB08D69A3DAFCD15433928A" author="Matthews, E. G." box="[151,346,871,896]" pageId="27" pageNumber="68" pagination="1 - 211" refId="ref29693" refString="Matthews, E. G. (1974) A revision of the scarabaeine dung beetles of Australia. II. Tribe Scarabaeini, Australian Journal of Zoology, Supplementary Series, 24, 1 - 211." type="journal article" year="1974">Matthews (1974)</bibRefCitation>
</taxonomicName>
revision is most similar to
<taxonomicName id="4C2AED86FFB08D69A1D7FCDF564C928A" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[666,805,871,896]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="species" species="guthrieae">
<emphasis id="B95E4A17FFB08D69A1D7FCDF564C928A" box="[666,805,871,896]" italics="true" pageId="27" pageNumber="68">L. guthrieae</emphasis>
</taxonomicName>
, although this specimen is a teneral female it is easily differentiated from
<taxonomicName id="4C2AED86FFB08D69A237FC3B576592AE" authorityName="in Matthews" authorityYear="1974" box="[378,524,909,932]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFB08D69A237FC3B576592AE" box="[378,524,909,932]" italics="true" pageId="27" pageNumber="68">L. pygmaeus</emphasis>
</taxonomicName>
on the basis of the apical edge of the protibia being angulate (truncate in
<taxonomicName id="4C2AED86FFB08D69A6C8FC3B546E92C2" authorityName="in Matthews" authorityYear="1974" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFB08D69A6C8FC3B546E92C2" italics="true" pageId="27" pageNumber="68">L. pygmaeus</emphasis>
</taxonomicName>
) and the almost ovoid pygidial depression (distinct pear shape in
<taxonomicName id="4C2AED86FFB08D69A0AAFC07511C92C2" authorityName="in Matthews" authorityYear="1974" box="[999,1141,945,968]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="68" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFB08D69A0AAFC07511C92C2" box="[999,1141,945,968]" italics="true" pageId="27" pageNumber="68">L. pygmaeus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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