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<document id="BB4678A2B6713CA5AC8C9AA466A597DC" ID-CLB-Dataset="28431" ID-DOI="10.11646/zootaxa.4564.1.2" ID-GBIF-Dataset="d3a56aac-92aa-48f2-b9fe-9576a4c2a0ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="2588621" ID-ZooBank="27F70B5F-24DD-4C9E-BBA7-2B87476BD49C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552162070752" checkinUser="plazi" docAuthor="Gunter, Nicole L. &amp; Weir, Thomas A." docDate="2019" docId="03832713FFA38D7BA3DAFBC050F297A8" docLanguage="en" docName="zootaxa.4564.1.2.pdf" docOrigin="Zootaxa 4564 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepanus pygmaeus" docType="key" docVersion="9" lastPageNumber="50" masterDocId="FFBA5F6BFFAB8D72A34DFFB65569910A" masterDocTitle="Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group" masterLastPageNumber="80" masterPageNumber="41" pageNumber="49" updateTime="1698700752634" updateUser="ExternalLinkService">
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<mods:title id="94A53F3977EE9B5069937B415FDA9B13">Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group</mods:title>
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<mods:namePart id="62CD0397AAABA4385E8A90C0FD72E05F">Gunter, Nicole L.</mods:namePart>
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<treatment id="03832713FFA38D7BA3DAFBC050F297A8" ID-DOI="http://doi.org/10.5281/zenodo.5922792" ID-GBIF-Taxon="154366358" ID-Zenodo-Dep="5922792" LSID="urn:lsid:plazi:treatment:03832713FFA38D7BA3DAFBC050F297A8" httpUri="http://treatment.plazi.org/id/03832713FFA38D7BA3DAFBC050F297A8" lastPageId="9" lastPageNumber="50" pageId="8" pageNumber="49">
<subSubSection id="C330C58EFFA38D7AA3DAFBC05021944A" pageId="8" pageNumber="49" type="nomenclature">
<paragraph id="8B959605FFA38D7AA3DAFBC0578F959A" blockId="8.[151,742,1142,1168]" box="[151,742,1142,1168]" pageId="8" pageNumber="49">
<heading id="D0DD2169FFA38D7AA3DAFBC0578F959A" bold="true" box="[151,742,1142,1168]" fontSize="11" level="1" pageId="8" pageNumber="49" reason="1">
<emphasis id="B95E4A17FFA38D7AA3DAFBC0578F959A" bold="true" box="[151,742,1142,1168]" pageId="8" pageNumber="49">
Key to species
<taxonomicName id="4C2AED86FFA38D7AA203FBC0575C959A" ID-CoL="3T8VK" baseAuthorityName="Macleay" baseAuthorityYear="1888" box="[334,565,1142,1168]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFA38D7AA203FBC0575C959A" bold="true" box="[334,565,1142,1168]" italics="true" pageId="8" pageNumber="49">Lepanus pygmaeus</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
<paragraph id="8B959605FFA38D7AA3DAFB0B5021944A" blockId="8.[151,1437,1212,1344]" pageId="8" pageNumber="49">
Due to the subtle nature of some diagnostic characters, some specimens are difficult to diagnose. Images of some diagnostic characters are provided in
<figureCitation id="13118A80FFA38D7AA105FB6957AB95F2" box="[584,706,1247,1272]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStart-4="FIGURE 8" captionStart-5="FIGURE 9" captionStart-6="FIGURE 10" captionStart-7="FIGURE 11" captionStart-8="FIGURE 12" captionStartId-0="10.[151,250,1773,1795]" captionStartId-1="15.[151,250,1945,1967]" captionStartId-2="16.[151,250,1948,1970]" captionStartId-3="17.[151,250,1974,1996]" captionStartId-4="18.[151,250,1896,1918]" captionStartId-5="19.[151,250,1201,1223]" captionStartId-6="23.[151,250,1909,1931]" captionStartId-7="24.[151,250,1905,1927]" captionStartId-8="25.[151,250,1905,1927]" captionTargetBox-0="[153,1436,509,1747]" captionTargetBox-1="[245,1336,223,1911]" captionTargetBox-2="[235,1333,202,1834]" captionTargetBox-3="[497,1058,239,1909]" captionTargetBox-4="[190,1384,223,1858]" captionTargetBox-5="[205,1310,206,1167]" captionTargetBox-6="[167,1333,208,1895]" captionTargetBox-7="[195,1359,208,1854]" captionTargetBox-8="[231,1341,208,1744]" captionTargetId-0="figure@10.[151,1436,481,1759]" captionTargetId-1="figure@15.[233,1355,193,1926]" captionTargetId-2="figure@16.[213,1373,193,1926]" captionTargetId-3="figure@17.[403,1184,193,1953]" captionTargetId-4="figure@18.[173,1427,195,1873]" captionTargetId-5="figure@19.[194,1392,193,1180]" captionTargetId-6="figure@23.[164,1428,193,1895]" captionTargetId-7="figure@24.[194,1392,193,1889]" captionTargetId-8="figure@25.[230,1357,193,1890]" captionTargetPageId-0="10" captionTargetPageId-1="15" captionTargetPageId-2="16" captionTargetPageId-3="17" captionTargetPageId-4="18" captionTargetPageId-5="19" captionTargetPageId-6="23" captionTargetPageId-7="24" captionTargetPageId-8="25" captionText-0="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." captionText-1="FIGURE 5. Habitus of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil. All specimens in comparative scale. Scale bar 1 mm." captionText-2="FIGURE 6. Habitus of species within the Lepanus pygmaeus species group. A, Lepanus lingziae; B, Lepanus mckenziei; C, Lepanus menendezae; D, Lepanus pecki; E: Lepanus podocarp; F, Lepanus pungalina. All specimens in comparative scale. Scale bar 1 mm." captionText-3="FIGURE 7. Habitus of species within the Lepanus pygmaeus species group. A, Lepanus pygmaeus; B, Lepanus sauroni; C, Lepanus tozerensis. All specimens in comparative scale. Scale bar 1 mm." captionText-4="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." captionText-5="FIGURE 9. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus pecki; B, Lepanus podocarp; C: Lepanus pungalina; D, Lepanus pygmaeus; E, Lepanus sauroni; F, Lepanus tozerensis. All specimens in comparative scale. Scale bar 0.2 mm." captionText-6="FIGURE 10. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." captionText-7="FIGURE 11. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus lentil; B, Lepanus lingziae; C, Lepanus mckenziei; D, Lepanus menendezae; E, Lepanus pecki. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." captionText-8="FIGURE 12. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus podocarp; B, Lepanus pungalina; C, Lepanus pygmaeus; D, Lepanus sauroni; E: Lepanus tozerensis. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri-0="https://zenodo.org/record/2588629/files/figure.png" httpUri-1="https://zenodo.org/record/2588631/files/figure.png" httpUri-2="https://zenodo.org/record/2588633/files/figure.png" httpUri-3="https://zenodo.org/record/2588635/files/figure.png" httpUri-4="https://zenodo.org/record/2588637/files/figure.png" httpUri-5="https://zenodo.org/record/2588639/files/figure.png" httpUri-6="https://zenodo.org/record/2588641/files/figure.png" httpUri-7="https://zenodo.org/record/2588643/files/figure.png" httpUri-8="https://zenodo.org/record/2588645/files/figure.png" pageId="8" pageNumber="49">Figs. 4–12</figureCitation>
to help facilitate diagnoses. Given the small size of specimens (1.8–3.0 mm), a high-powered microscope with adequate lighting is recommended for identification. Furthermore, caution must be taken when interpreting setation on specimens as fine setae rub off and may appear absent.
</paragraph>
</subSubSection>
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<paragraph id="8B959605FFA38D7AA3DAFAD950F2948E" blockId="8.[151,1436,1391,2033]" box="[151,1435,1391,1412]" pageId="8" pageNumber="49">
<keyLead id="30DB3530FFA38D7AA3DAFAD950F2948E" box="[151,1435,1391,1412]" pageId="8" pageNumber="49">
1 Protibia with three teeth on outer edge, basal tooth sometimes very small (
<figureCitation id="13118A80FFA38D7AA0E4FAD95688948E" box="[937,993,1391,1412]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="8" pageNumber="49">Fig. 4</figureCitation>
I–J)................................ 2
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</paragraph>
<paragraph id="8B959605FFA38D7AA3DAFA3B50F294A8" blockId="8.[151,1436,1391,2033]" box="[151,1435,1421,1442]" pageId="8" pageNumber="49">
<keyLead id="30DB3530FFA38D7AA3DAFA3B50F294A8" box="[151,1435,1421,1442]" pageId="8" pageNumber="49">
- Protibia with two teeth on outer edge (
<figureCitation id="13118A80FFA38D7AA11CFA3B57E394A8" box="[593,650,1421,1442]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="8" pageNumber="49">Fig. 4</figureCitation>
G–H, K–L)....................................................... 3
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<keyStep id="30DE8EA0FFA38D7AA3DAFA1C50F2975D" pageId="8" pageNumber="49">
<paragraph id="8B959605FFA38D7AA3DAFA1C50F294F6" blockId="8.[151,1436,1391,2033]" pageId="8" pageNumber="49">
<keyLead id="30DB3530FFA38D7AA3DAFA1C50F294F6" pageId="8" pageNumber="49">
2 Pygidium with a tear-drop shaped depression extending under the usually straight upper edge, depression with an elongate tubercle (
<figureCitation id="13118A80FFA38D7AA276FA7C54EF94D5" box="[315,390,1482,1503]" captionStart="FIGURE 9" captionStartId="19.[151,250,1201,1223]" captionTargetBox="[205,1310,206,1167]" captionTargetId="figure@19.[194,1392,193,1180]" captionTargetPageId="19" captionText="FIGURE 9. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus pecki; B, Lepanus podocarp; C: Lepanus pungalina; D, Lepanus pygmaeus; E, Lepanus sauroni; F, Lepanus tozerensis. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588639/files/figure.png" pageId="8" pageNumber="49">Fig. 9E</figureCitation>
); male metatibiae with almost straight inner edge; female 6
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abdominal ventrite swollen in middle; aedeagus (
<figureCitation id="13118A80FFA38D7AA3A5FA51545094F6" box="[232,313,1511,1532]" captionStart="FIGURE 12" captionStartId="25.[151,250,1905,1927]" captionTargetBox="[231,1341,208,1744]" captionTargetId="figure@25.[230,1357,193,1890]" captionTargetPageId="25" captionText="FIGURE 12. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus podocarp; B, Lepanus pungalina; C, Lepanus pygmaeus; D, Lepanus sauroni; E: Lepanus tozerensis. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588645/files/figure.png" pageId="8" pageNumber="49">Fig.12D</figureCitation>
).........................................................
<taxonomicName id="4C2AED86FFA38D7AA0A9FA51507694F6" ID-CoL="3T8VN" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[996,1311,1511,1532]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="sauroni" status="sp. nov.">
<emphasis id="B95E4A17FFA38D7AA0A9FA5151E194F6" box="[996,1160,1511,1532]" italics="true" pageId="8" pageNumber="49">Lepanus sauroni</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFFA38D7AA664FA5150F294F6" box="[1321,1435,1511,1532]" pageId="8" pageNumber="49" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B959605FFA38D7AA3DAF9B250F2975D" blockId="8.[151,1436,1391,2033]" pageId="8" pageNumber="49">
<keyLead id="30DB3530FFA38D7AA3DAF9B250F2975D" pageId="8" pageNumber="49">
- Pygidium with an almost ovoid depression just extending under the usually angulate upper edge, depression with a round tubercle (
<figureCitation id="13118A80FFA38D7AA270F99254E49733" box="[317,397,1572,1593]" captionStart="FIGURE 9" captionStartId="19.[151,250,1201,1223]" captionTargetBox="[205,1310,206,1167]" captionTargetId="figure@19.[194,1392,193,1180]" captionTargetPageId="19" captionText="FIGURE 9. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus pecki; B, Lepanus podocarp; C: Lepanus pungalina; D, Lepanus pygmaeus; E, Lepanus sauroni; F, Lepanus tozerensis. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588639/files/figure.png" pageId="8" pageNumber="49">Fig. 9A</figureCitation>
); male metatibiae with expanded inner edge, creating a distinct flange (
<figureCitation id="13118A80FFA38D7AA71CF99251E29733" box="[1105,1163,1572,1593]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="8" pageNumber="49">Fig. 4</figureCitation>
D–E); female 6
<superScript id="7C5F3B4DFFA38D7AA669F99450469724" attach="left" box="[1316,1327,1570,1582]" fontSize="5" pageId="8" pageNumber="49">th</superScript>
abdominal ventrite not swollen; aedeagus as in
<figureCitation id="13118A80FFA38D7AA171F9F457E7975D" box="[572,654,1602,1623]" captionStart="FIGURE 11" captionStartId="24.[151,250,1905,1927]" captionTargetBox="[195,1359,208,1854]" captionTargetId="figure@24.[194,1392,193,1889]" captionTargetPageId="24" captionText="FIGURE 11. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus lentil; B, Lepanus lingziae; C, Lepanus mckenziei; D, Lepanus menendezae; E, Lepanus pecki. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588643/files/figure.png" pageId="8" pageNumber="49">Fig. 11E</figureCitation>
..............................
<taxonomicName id="4C2AED86FFA38D7AA0B3F9F45076975D" ID-CoL="3T8VC" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[1022,1311,1602,1623]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="pecki" status="sp. nov.">
<emphasis id="B95E4A17FFA38D7AA0B3F9F451E1975D" box="[1022,1160,1602,1623]" italics="true" pageId="8" pageNumber="49">Lepanus pecki</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFFA38D7AA664F9F450F2975D" box="[1321,1435,1602,1623]" pageId="8" pageNumber="49" rank="species">new species</taxonomicNameLabel>
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<paragraph id="8B959605FFA38D7AA3DAF9E950F2977E" blockId="8.[151,1436,1391,2033]" box="[151,1435,1631,1652]" pageId="8" pageNumber="49">
<keyLead id="30DB3530FFA38D7AA3DAF9E950F2977E" box="[151,1435,1631,1652]" pageId="8" pageNumber="49">
3 Front edge of protibia truncate (
<figureCitation id="13118A80FFA38D7AA15FF9E9570C977E" box="[530,613,1631,1652]" captionStart="FIGURE 3" captionStartId="7.[151,250,1911,1933]" captionTargetBox="[190,1318,415,1885]" captionTargetId="figure@7.[168,1467,409,1885]" captionTargetPageId="7" captionText="FIGURE 3. A, Dorsal aspect of key structures of Lepanus and other dung beetles; B, Ventral aspect of key structures of Lepanus and other dung beetles; C, key structures of a Lepanus head; D–F, aspect of the eye canthus relative to the eye: D, dividing the eye; E, nearly dividing the eye (separated by only a few eye facets); F, not dividing the eye; G–H, examples and key structures of Lepanus protibia: G, dorsal aspect of truncate protibia; H, dorsal aspect of deeply angulate protibia." httpUri="https://zenodo.org/record/2588627/files/figure.png" pageId="8" pageNumber="49">Figs. 3G</figureCitation>
,
<figureCitation id="13118A80FFA38D7AA122F9E95712977E" box="[623,635,1631,1652]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="8" pageNumber="49">4</figureCitation>
G–H); apical digit absent in females (
<figureCitation id="13118A80FFA38D7AA0AEF9E95158977E" box="[995,1073,1631,1652]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="8" pageNumber="49">Fig. 4G</figureCitation>
)............................. 4
</keyLead>
</paragraph>
<paragraph id="8B959605FFA38D7AA3DAF9CA50F2979B" blockId="8.[151,1436,1391,2033]" box="[151,1435,1660,1681]" pageId="8" pageNumber="49">
<keyLead id="30DB3530FFA38D7AA3DAF9CA50F2979B" box="[151,1435,1660,1681]" pageId="8" pageNumber="49">
- Front edge of protibia deeply angulate (
<figureCitation id="13118A80FFA38D7AA111F9CA57C0979B" box="[604,681,1660,1681]" captionStart="FIGURE 3" captionStartId="7.[151,250,1911,1933]" captionTargetBox="[190,1318,415,1885]" captionTargetId="figure@7.[168,1467,409,1885]" captionTargetPageId="7" captionText="FIGURE 3. A, Dorsal aspect of key structures of Lepanus and other dung beetles; B, Ventral aspect of key structures of Lepanus and other dung beetles; C, key structures of a Lepanus head; D–F, aspect of the eye canthus relative to the eye: D, dividing the eye; E, nearly dividing the eye (separated by only a few eye facets); F, not dividing the eye; G–H, examples and key structures of Lepanus protibia: G, dorsal aspect of truncate protibia; H, dorsal aspect of deeply angulate protibia." httpUri="https://zenodo.org/record/2588627/files/figure.png" pageId="8" pageNumber="49">Fig. 3H</figureCitation>
); apical digit present in females (as in
<figureCitation id="13118A80FFA38D7AA756F9CA5135979B" box="[1051,1116,1660,1681]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="8" pageNumber="49">
Fig.
<date id="FF94B0C5FFA38D7AA70AF9CA5135979B" box="[1095,1116,1660,1681]" pageId="8" pageNumber="49">4I</date>
</figureCitation>
)......................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DE8EA0FFA38D7AA3DAF92C50F29655" pageId="8" pageNumber="49">
<paragraph id="8B959605FFA38D7AA3DAF92C50F5960D" blockId="8.[151,1436,1391,2033]" pageId="8" pageNumber="49">
<keyLead id="30DB3530FFA38D7AA3DAF92C50F5960D" pageId="8" pageNumber="49">
4 Pygidial depression widely pear shaped with a somewhat triangular tubercle in the upper third, upper edge of pygidium narrowly angulate, depression just extending under the edge (
<figureCitation id="13118A80FFA38D7AA055F901560E97C6" box="[792,871,1719,1740]" captionStart="FIGURE 9" captionStartId="19.[151,250,1201,1223]" captionTargetBox="[205,1310,206,1167]" captionTargetId="figure@19.[194,1392,193,1180]" captionTargetPageId="19" captionText="FIGURE 9. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus pecki; B, Lepanus podocarp; C: Lepanus pungalina; D, Lepanus pygmaeus; E, Lepanus sauroni; F, Lepanus tozerensis. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588639/files/figure.png" pageId="8" pageNumber="49">Fig. 9D</figureCitation>
); punctures along basal margin of prothorax slightly elongate, differing from those on the disc; dorsal part of the eye moderate in size; aedeagus (
<figureCitation id="13118A80FFA38D7AA763F96251ED97E3" box="[1070,1156,1748,1769]" captionStart="FIGURE 12" captionStartId="25.[151,250,1905,1927]" captionTargetBox="[231,1341,208,1744]" captionTargetId="figure@25.[230,1357,193,1890]" captionTargetPageId="25" captionText="FIGURE 12. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus podocarp; B, Lepanus pungalina; C, Lepanus pygmaeus; D, Lepanus sauroni; E: Lepanus tozerensis. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588645/files/figure.png" pageId="8" pageNumber="49">Fig. 12C</figureCitation>
)..............................................................................................
<taxonomicName id="4C2AED86FFA38D7AA70CF94450F5960D" ID-CoL="3T8VK" authority="(Macleay, 1888)" baseAuthorityName="Macleay" baseAuthorityYear="1888" box="[1089,1436,1778,1799]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFA38D7AA70CF9445190960D" box="[1089,1273,1778,1799]" italics="true" pageId="8" pageNumber="49">Lepanus pygmaeus</emphasis>
(
<bibRefCitation id="EFBBEBF4FFA38D7AA648F94450FC960D" author="Macleay, W." box="[1285,1429,1778,1799]" pageId="8" pageNumber="49" pagination="897 - 904" refId="ref29625" refString="Macleay, W. (1888) The insects of King's Sound and its vicinity. II. The lamellicornes, Proceedings of the Linnean Society of New South Wales, 3, 897 - 904." type="journal article" year="1888">Macleay, 1888</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B959605FFA38D7AA3DAF8B950F29655" blockId="8.[151,1436,1391,2033]" pageId="8" pageNumber="49">
<keyLead id="30DB3530FFA38D7AA3DAF8B950F29655" pageId="8" pageNumber="49">
- Pygidial depression round with a round tubercle in the upper third, upper edge of pygidium broadly angulate, depression extending under the edge (
<figureCitation id="13118A80FFA38D7AA2ADF89A5740964B" box="[480,553,1836,1857]" captionStart="FIGURE 9" captionStartId="19.[151,250,1201,1223]" captionTargetBox="[205,1310,206,1167]" captionTargetId="figure@19.[194,1392,193,1180]" captionTargetPageId="19" captionText="FIGURE 9. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus pecki; B, Lepanus podocarp; C: Lepanus pungalina; D, Lepanus pygmaeus; E, Lepanus sauroni; F, Lepanus tozerensis. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588639/files/figure.png" pageId="8" pageNumber="49">Fig. 9F</figureCitation>
); punctures along basal margin of prothorax more round, similar to those on the disc; dorsal part of the eye narrower in size; aedeagus (
<figureCitation id="13118A80FFA38D7AA131F8FC57B89655" box="[636,721,1866,1887]" captionStart="FIGURE 12" captionStartId="25.[151,250,1905,1927]" captionTargetBox="[231,1341,208,1744]" captionTargetId="figure@25.[230,1357,193,1890]" captionTargetPageId="25" captionText="FIGURE 12. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus podocarp; B, Lepanus pungalina; C, Lepanus pygmaeus; D, Lepanus sauroni; E: Lepanus tozerensis. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588645/files/figure.png" pageId="8" pageNumber="49">Fig. 12E</figureCitation>
).....................
<taxonomicName id="4C2AED86FFA38D7AA080F8FC50769655" ID-CoL="3T8VR" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[973,1311,1866,1887]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="tozerensis" status="sp. nov.">
<emphasis id="B95E4A17FFA38D7AA080F8FC51E19655" box="[973,1160,1866,1887]" italics="true" pageId="8" pageNumber="49">Lepanus tozerensis</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFFA38D7AA664F8FC50F29655" box="[1321,1435,1866,1887]" pageId="8" pageNumber="49" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8B959605FFA38D7AA3DAF8D150F296BD" blockId="8.[151,1436,1391,2033]" pageId="8" pageNumber="49">
<keyLead id="30DB3530FFA38D7AA3DAF8D150F296BD" pageId="8" pageNumber="49">
<keyStep id="30DE8EA0FFA38D7AA3DAF8D150F296BD" pageId="8" pageNumber="49">
5 Pygidial depression narrow and vertically elongate, without a tubercle, occupying less than half the area of the pygidial disc (
<figureCitation id="13118A80FFA38D7AA3A5F83254599693" box="[232,304,1924,1945]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="8" pageNumber="49">Fig. 8F</figureCitation>
), basal carina of head complete, with a slight medial angle (
<figureCitation id="13118A80FFA38D7AA022F83256D29693" box="[879,955,1924,1945]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="8" pageNumber="49">Fig. 4B</figureCitation>
); aedeagus as in
<figureCitation id="13118A80FFA38D7AA710F83251D89693" box="[1117,1201,1924,1945]" captionStart="FIGURE 11" captionStartId="24.[151,250,1905,1927]" captionTargetBox="[195,1359,208,1854]" captionTargetId="figure@24.[194,1392,193,1889]" captionTargetPageId="24" captionText="FIGURE 11. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus lentil; B, Lepanus lingziae; C, Lepanus mckenziei; D, Lepanus menendezae; E, Lepanus pecki. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588643/files/figure.png" pageId="8" pageNumber="49">Fig. 11A</figureCitation>
.....................................................................................
<taxonomicName id="4C2AED86FFA38D7AA74DF814507696BD" ID-CoL="6P7XV" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[1024,1311,1954,1975]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="lentil" status="sp. nov.">
<emphasis id="B95E4A17FFA38D7AA74DF81451E196BD" box="[1024,1160,1954,1975]" italics="true" pageId="8" pageNumber="49">Lepanus lentil</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFFA38D7AA664F81450F296BD" box="[1321,1435,1954,1975]" pageId="8" pageNumber="49" rank="species">new species</taxonomicNameLabel>
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</keyLead>
</paragraph>
<paragraph id="8B959605FFA38D7BA3DAF80950F291ED" blockId="8.[151,1436,1391,2033]" lastBlockId="9.[151,1436,151,1698]" lastPageId="9" lastPageNumber="50" pageId="8" pageNumber="49">
<keyLead id="30DB3530FFA38D7BA3DAF80950F291ED" lastPageId="9" lastPageNumber="50" pageId="8" pageNumber="49">
- Pygidial depression of different shape, usually occupying at least one third to half the area of the pygidial disc (smaller and round in
<taxonomicName id="4C2AED86FFA38D7AA276F86A54A296FB" ID-CoL="3T8V3" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[315,459,2012,2033]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="menendezae">
<emphasis id="B95E4A17FFA38D7AA276F86A54A296FB" box="[315,459,2012,2033]" italics="true" pageId="8" pageNumber="49">L. menendezae</emphasis>
</taxonomicName>
), depression with or without a distinct tubercle; basal carina of head either obsolete, or interrupted medially............................................................................................. 6
<keyStep id="30DE8EA0FFA28D7BA3DAFF0350F291ED" pageId="9" pageNumber="50">
6 Pygidial depression a narrow or broad U-shape, without a distinct tubercle (
<figureCitation id="13118A80FFA28D7BA0F6FF03514691C0" box="[955,1071,181,202]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="9" pageNumber="50">Figs. 8A, G</figureCitation>
); basal carina of head absent or obsolete; basal margin of pronotum slightly angulate medially...................................................... 7
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B959605FFA28D7BA3DAFF5950F29035" blockId="9.[151,1436,151,1698]" pageId="9" pageNumber="50">
<keyLead id="30DB3530FFA28D7BA3DAFF5950F29035" pageId="9" pageNumber="50">
- Pygidial depression different, either round (
<emphasis id="B95E4A17FFA28D7BA1C1FF5957C6900E" box="[652,687,239,260]" italics="true" pageId="9" pageNumber="50">e.g.</emphasis>
,
<figureCitation id="13118A80FFA28D7BA1F0FF59564F900E" box="[701,806,239,260]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="9" pageNumber="50">Figs. 8C, I</figureCitation>
), ovoid (
<emphasis id="B95E4A17FFA28D7BA0CFFF5956CC900E" box="[898,933,239,260]" italics="true" pageId="9" pageNumber="50">e.g.</emphasis>
,
<figureCitation id="13118A80FFA28D7BA0FFFF595696900E" box="[946,1023,239,260]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="9" pageNumber="50">Fig. 8H</figureCitation>
), pear or tear-drop shaped (
<emphasis id="B95E4A17FFA28D7BA65CFF59505D900E" box="[1297,1332,239,260]" italics="true" pageId="9" pageNumber="50">e.g.</emphasis>
,
<figureCitation id="13118A80FFA28D7BA60FFF5950F1900E" box="[1346,1432,239,260]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="9" pageNumber="50">Figs. 8D</figureCitation>
,
<figureCitation id="13118A80FFA28D7BA3ACFEBB55849028" box="[225,237,269,290]" captionStart="FIGURE 9" captionStartId="19.[151,250,1201,1223]" captionTargetBox="[205,1310,206,1167]" captionTargetId="figure@19.[194,1392,193,1180]" captionTargetPageId="19" captionText="FIGURE 9. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus pecki; B, Lepanus podocarp; C: Lepanus pungalina; D, Lepanus pygmaeus; E, Lepanus sauroni; F, Lepanus tozerensis. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588639/files/figure.png" pageId="9" pageNumber="50">9</figureCitation>
B–C), with a distinct tubercle; basal carina of head present, medially interrupted (
<figureCitation id="13118A80FFA28D7BA0B9FEBB51289028" box="[1012,1089,269,290]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="9" pageNumber="50">Fig. 4A</figureCitation>
); basal margin of pronotum rounded medially............................................................................................. 8
</keyLead>
</paragraph>
<keyStep id="30DE8EA0FFA28D7BA3DAFEF150F2934D" pageId="9" pageNumber="50">
<paragraph id="8B959605FFA28D7BA3DAFEF150F290D8" blockId="9.[151,1436,151,1698]" pageId="9" pageNumber="50">
<keyLead id="30DB3530FFA28D7BA3DAFEF150F290D8" pageId="9" pageNumber="50">
7 Pygidial depression narrow U-shape, reticulate, occupying the medial third of the pygidial disc (
<figureCitation id="13118A80FFA28D7BA7CEFEF151BB9056" box="[1155,1234,327,348]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="9" pageNumber="50">Fig. 8A</figureCitation>
); surface of the head finely reticulate between clypeogenal suture and anterior edge, rest of head smooth (as in
<figureCitation id="13118A80FFA28D7BA778FED351E89070" box="[1077,1153,357,378]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="9" pageNumber="50">Fig. 4A</figureCitation>
); dorsal part of eye moderate, interocular space about 10 times the width of dorsal part of the eye; with a fine carina extending backwards from the mesometaventral suture (
<figureCitation id="13118A80FFA28D7BA2EDFE29548590BE" box="[416,492,415,436]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="9" pageNumber="50">Fig. 4C</figureCitation>
); elytral surface reticulate in apical half and at the sides, striae slightly impressed, striae 5–7 similar to striae 1–4; aedeagus as in
<figureCitation id="13118A80FFA28D7BA15EFE0B570190D8" box="[531,616,445,466]" captionStart="FIGURE 10" captionStartId="23.[151,250,1909,1931]" captionTargetBox="[167,1333,208,1895]" captionTargetId="figure@23.[164,1428,193,1895]" captionTargetPageId="23" captionText="FIGURE 10. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588641/files/figure.png" pageId="9" pageNumber="50">Fig. 10A</figureCitation>
..........................
<taxonomicName id="4C2AED86FFA28D7BA0E8FE0B507690D8" ID-CoL="6P7WG" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[933,1311,445,466]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="andersonorum" status="sp. nov.">
<emphasis id="B95E4A17FFA28D7BA0E8FE0B51E190D8" box="[933,1160,445,466]" italics="true" pageId="9" pageNumber="50">Lepanus andersonorum</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFFA28D7BA664FE0B50F290D8" box="[1321,1435,445,466]" pageId="9" pageNumber="50" rank="species">new species</taxonomicNameLabel>
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<paragraph id="8B959605FFA28D7BA3DAFE6C50F2934D" blockId="9.[151,1436,151,1698]" pageId="9" pageNumber="50">
<keyLead id="30DB3530FFA28D7BA3DAFE6C50F2934D" pageId="9" pageNumber="50">
- Pygidial depression broad U-shape, smooth, occupying at least medial half of the pygidial disc (
<figureCitation id="13118A80FFA28D7BA7D2FE6C518690E5" box="[1183,1263,474,495]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="9" pageNumber="50">Fig. 8G</figureCitation>
); surface of head entirely smooth; dorsal part of eye narrow in size, interocular space about 16 times the width of dorsal part of the eye; without a fine carina extending backwards from the meso-metaventral suture; elytral surface smooth, striae superficial, striae 5–7 weaker than striae 1–4; aedeagus as in
<figureCitation id="13118A80FFA28D7BA11BFD8457C3934D" box="[598,682,562,583]" captionStart="FIGURE 11" captionStartId="24.[151,250,1905,1927]" captionTargetBox="[195,1359,208,1854]" captionTargetId="figure@24.[194,1392,193,1889]" captionTargetPageId="24" captionText="FIGURE 11. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus lentil; B, Lepanus lingziae; C, Lepanus mckenziei; D, Lepanus menendezae; E, Lepanus pecki. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588643/files/figure.png" pageId="9" pageNumber="50">Fig. 11B</figureCitation>
..........................
<taxonomicName id="4C2AED86FFA28D7BA0A8FD845076934D" ID-CoL="6P7Y7" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[997,1311,562,583]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="lingziae" status="sp. nov.">
<emphasis id="B95E4A17FFA28D7BA0A8FD8451E1934D" box="[997,1160,562,583]" italics="true" pageId="9" pageNumber="50">Lepanus lingziae</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
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<keyLead id="30DB3530FFA28D7BA3DAFDF950F2936E" box="[151,1435,591,612]" pageId="9" pageNumber="50">
8 Pygidial depression pear or tear-drop shaped (
<figureCitation id="13118A80FFA28D7BA1D4FDF95786936E" box="[665,751,591,612]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="9" pageNumber="50">Figs. 8D</figureCitation>
,
<figureCitation id="13118A80FFA28D7BA1B4FDF9566C936E" box="[761,773,591,612]" captionStart="FIGURE 9" captionStartId="19.[151,250,1201,1223]" captionTargetBox="[205,1310,206,1167]" captionTargetId="figure@19.[194,1392,193,1180]" captionTargetPageId="19" captionText="FIGURE 9. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus pecki; B, Lepanus podocarp; C: Lepanus pungalina; D, Lepanus pygmaeus; E, Lepanus sauroni; F, Lepanus tozerensis. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588639/files/figure.png" pageId="9" pageNumber="50">9</figureCitation>
B–C).................................................. 9
</keyLead>
</paragraph>
<paragraph id="8B959605FFA28D7BA3DAFDDB50F29388" blockId="9.[151,1436,151,1698]" box="[151,1435,621,642]" pageId="9" pageNumber="50">
<keyLead id="30DB3530FFA28D7BA3DAFDDB50F29388" box="[151,1435,621,642]" pageId="9" pageNumber="50">
- Pygidial depression round or ovoid (
<figureCitation id="13118A80FFA28D7BA172FDDB57E99388" box="[575,640,621,642]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="9" pageNumber="50">Figs. 8</figureCitation>
B–C, E, H–I)..................................................... 11
</keyLead>
</paragraph>
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<keyStep id="30DE8EA0FFA28D7BA3DAFD3C50F29238" pageId="9" pageNumber="50">
<paragraph id="8B959605FFA28D7BA3DAFD3C50F293FD" blockId="9.[151,1436,151,1698]" pageId="9" pageNumber="50">
<keyLead id="30DB3530FFA28D7BA3DAFD3C50F293FD" pageId="9" pageNumber="50">
9 Pygidium with an evident secondary basal line across base of depression, pygidial depression tear-drop shaped, not reaching base, tubercle slightly elongate (
<figureCitation id="13118A80FFA28D7BA156FD11570E93B6" box="[539,615,679,700]" captionStart="FIGURE 9" captionStartId="19.[151,250,1201,1223]" captionTargetBox="[205,1310,206,1167]" captionTargetId="figure@19.[194,1392,193,1180]" captionTargetPageId="19" captionText="FIGURE 9. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus pecki; B, Lepanus podocarp; C: Lepanus pungalina; D, Lepanus pygmaeus; E, Lepanus sauroni; F, Lepanus tozerensis. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588639/files/figure.png" pageId="9" pageNumber="50">Fig. 9C</figureCitation>
); mesoventrite with discernible punctures at least at sides; surface of the head finely reticulate between clypeogenal suture and anterior edge (as in
<figureCitation id="13118A80FFA28D7BA07AFD7356ED93D0" box="[823,900,709,730]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="9" pageNumber="50">Fig. 4A</figureCitation>
); aedeagus as in
<figureCitation id="13118A80FFA28D7BA768FD73511093D0" box="[1061,1145,709,730]" captionStart="FIGURE 12" captionStartId="25.[151,250,1905,1927]" captionTargetBox="[231,1341,208,1744]" captionTargetId="figure@25.[230,1357,193,1890]" captionTargetPageId="25" captionText="FIGURE 12. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus podocarp; B, Lepanus pungalina; C, Lepanus pygmaeus; D, Lepanus sauroni; E: Lepanus tozerensis. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588645/files/figure.png" pageId="9" pageNumber="50">Fig. 12B</figureCitation>
......................................................................................
<taxonomicName id="4C2AED86FFA28D7BA082FD54507693FD" ID-CoL="3T8VJ" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[975,1311,738,759]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="pungalina" status="sp. nov.">
<emphasis id="B95E4A17FFA28D7BA082FD5451E193FD" box="[975,1160,738,759]" italics="true" pageId="9" pageNumber="50">Lepanus pungalina</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFFA28D7BA664FD5450F293FD" box="[1321,1435,738,759]" pageId="9" pageNumber="50" rank="species">new species</taxonomicNameLabel>
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<paragraph id="8B959605FFA28D7BA3DAFD4950F29238" blockId="9.[151,1436,151,1698]" pageId="9" pageNumber="50">
<keyLead id="30DB3530FFA28D7BA3DAFD4950F29238" pageId="9" pageNumber="50">
- Pygidium with secondary basal line not evident, merged with base, pygidial depression reaching base, tubercle elongate or an inverted V (
<figureCitation id="13118A80FFA28D7BA219FCAB54C09238" box="[340,425,797,818]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="9" pageNumber="50">Figs. 8D</figureCitation>
,
<figureCitation id="13118A80FFA28D7BA2FEFCAB54B89238" box="[435,465,797,818]" captionStart="FIGURE 9" captionStartId="19.[151,250,1201,1223]" captionTargetBox="[205,1310,206,1167]" captionTargetId="figure@19.[194,1392,193,1180]" captionTargetPageId="19" captionText="FIGURE 9. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus pecki; B, Lepanus podocarp; C: Lepanus pungalina; D, Lepanus pygmaeus; E, Lepanus sauroni; F, Lepanus tozerensis. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588639/files/figure.png" pageId="9" pageNumber="50">9B</figureCitation>
); mesoventrite with or without discernible punctures; surface of head entirely smooth.......... 10
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<paragraph id="8B959605FFA28D7BA3DAFC8C50F29280" blockId="9.[151,1436,151,1698]" pageId="9" pageNumber="50">
<keyLead id="30DB3530FFA28D7BA3DAFC8C50F29280" pageId="9" pageNumber="50">
10 Pygidial depression widely pear shaped, tubercle elongate (
<figureCitation id="13118A80FFA28D7BA067FC8C561E9245" box="[810,887,826,847]" captionStart="FIGURE 9" captionStartId="19.[151,250,1201,1223]" captionTargetBox="[205,1310,206,1167]" captionTargetId="figure@19.[194,1392,193,1180]" captionTargetPageId="19" captionText="FIGURE 9. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus pecki; B, Lepanus podocarp; C: Lepanus pungalina; D, Lepanus pygmaeus; E, Lepanus sauroni; F, Lepanus tozerensis. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588639/files/figure.png" pageId="9" pageNumber="50">Fig. 9B</figureCitation>
); elytral flange and groove relatively wide; colour dark brown to black with red humeri (
<figureCitation id="13118A80FFA28D7BA150FCE1570E9266" box="[541,615,855,876]" captionStart="FIGURE 6" captionStartId="16.[151,250,1948,1970]" captionTargetBox="[235,1333,202,1834]" captionTargetId="figure@16.[213,1373,193,1926]" captionTargetPageId="16" captionText="FIGURE 6. Habitus of species within the Lepanus pygmaeus species group. A, Lepanus lingziae; B, Lepanus mckenziei; C, Lepanus menendezae; D, Lepanus pecki; E: Lepanus podocarp; F, Lepanus pungalina. All specimens in comparative scale. Scale bar 1 mm." httpUri="https://zenodo.org/record/2588633/files/figure.png" pageId="9" pageNumber="50">Fig. 6E</figureCitation>
); mesoventrite with discernible punctures at least at sides; aedeagus as in
<figureCitation id="13118A80FFA28D7BA666FCE150169266" box="[1323,1407,855,876]" captionStart="FIGURE 12" captionStartId="25.[151,250,1905,1927]" captionTargetBox="[231,1341,208,1744]" captionTargetId="figure@25.[230,1357,193,1890]" captionTargetPageId="25" captionText="FIGURE 12. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus podocarp; B, Lepanus pungalina; C, Lepanus pygmaeus; D, Lepanus sauroni; E: Lepanus tozerensis. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588645/files/figure.png" pageId="9" pageNumber="50">Fig. 12A</figureCitation>
.................................................................
<taxonomicName id="4C2AED86FFA28D7BA099FCC350769280" ID-CoL="3T8VF" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[980,1311,885,906]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="podocarp" status="sp. nov.">
<emphasis id="B95E4A17FFA28D7BA099FCC351E19280" box="[980,1160,885,906]" italics="true" pageId="9" pageNumber="50">Lepanus podocarp</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFFA28D7BA664FCC350F29280" box="[1321,1435,885,906]" pageId="9" pageNumber="50" rank="species">new species</taxonomicNameLabel>
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</paragraph>
<paragraph id="8B959605FFA28D7BA3DAFC2450F292E8" blockId="9.[151,1436,151,1698]" pageId="9" pageNumber="50">
<keyLead id="30DB3530FFA28D7BA3DAFC2450F292E8" pageId="9" pageNumber="50">
- Pygidial depression narrowly tear-drop shaped, tubercle an inverted V (
<figureCitation id="13118A80FFA28D7BA0D1FC24568392AD" box="[924,1002,914,935]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="9" pageNumber="50">Fig. 8D</figureCitation>
); elytral flange and groove narrower; colour uniformly dark brown (
<figureCitation id="13118A80FFA28D7BA28FFC19576792CE" box="[450,526,943,964]" captionStart="FIGURE 5" captionStartId="15.[151,250,1945,1967]" captionTargetBox="[245,1336,223,1911]" captionTargetId="figure@15.[233,1355,193,1926]" captionTargetPageId="15" captionText="FIGURE 5. Habitus of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil. All specimens in comparative scale. Scale bar 1 mm." httpUri="https://zenodo.org/record/2588631/files/figure.png" pageId="9" pageNumber="50">Fig. 5D</figureCitation>
); mesoventrite virtually impunctate; aedeagus as in
<figureCitation id="13118A80FFA28D7BA74CFC19513F92CE" box="[1025,1110,943,964]" captionStart="FIGURE 10" captionStartId="23.[151,250,1909,1931]" captionTargetBox="[167,1333,208,1895]" captionTargetId="figure@23.[164,1428,193,1895]" captionTargetPageId="23" captionText="FIGURE 10. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588641/files/figure.png" pageId="9" pageNumber="50">Fig. 10D</figureCitation>
............................................................................................
<taxonomicName id="4C2AED86FFA28D7BA0A1FC7B507692E8" ID-CoL="6P89W" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[1004,1311,973,994]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="gubara" status="sp. nov.">
<emphasis id="B95E4A17FFA28D7BA0A1FC7B51E192E8" box="[1004,1160,973,994]" italics="true" pageId="9" pageNumber="50">Lepanus gubara</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFFA28D7BA664FC7B50F292E8" box="[1321,1435,973,994]" pageId="9" pageNumber="50" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8B959605FFA28D7BA3DAFC5C50F29530" blockId="9.[151,1436,151,1698]" pageId="9" pageNumber="50">
<keyLead id="30DB3530FFA28D7BA3DAFC5C50F29530" pageId="9" pageNumber="50">
11 Pygidial depression small and round, occupying less than half of the pygidial disc, separated from base, tubercle round and centrally located; secondary basal line evident (
<figureCitation id="13118A80FFA28D7BA1E5FBB157829516" box="[680,747,1031,1052]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="9" pageNumber="50">
Fig.
<date id="FF94B0C5FFA28D7BA198FBB157829516" box="[725,747,1031,1052]" pageId="9" pageNumber="50">8I</date>
</figureCitation>
); aedeagus as in
<figureCitation id="13118A80FFA28D7BA0C3FBB1568A9516" box="[910,995,1031,1052]" captionStart="FIGURE 11" captionStartId="24.[151,250,1905,1927]" captionTargetBox="[195,1359,208,1854]" captionTargetId="figure@24.[194,1392,193,1889]" captionTargetPageId="24" captionText="FIGURE 11. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus lentil; B, Lepanus lingziae; C, Lepanus mckenziei; D, Lepanus menendezae; E, Lepanus pecki. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588643/files/figure.png" pageId="9" pageNumber="50">Fig. 11D</figureCitation>
.................................................................................................
<taxonomicName id="4C2AED86FFA28D7BA0F6FB9350769530" ID-CoL="3T8V3" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[955,1311,1061,1082]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="menendezae" status="sp. nov.">
<emphasis id="B95E4A17FFA28D7BA0F6FB9351E19530" box="[955,1160,1061,1082]" italics="true" pageId="9" pageNumber="50">Lepanus menendezae</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFFA28D7BA664FB9350F29530" box="[1321,1435,1061,1082]" pageId="9" pageNumber="50" rank="species">new species</taxonomicNameLabel>
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</paragraph>
<paragraph id="8B959605FFA28D7BA3DAFBF450F2957E" blockId="9.[151,1436,151,1698]" pageId="9" pageNumber="50">
<keyLead id="30DB3530FFA28D7BA3DAFBF450F2957E" pageId="9" pageNumber="50">
- Pygidial depression larger, occupying about half of the pygidial disc, usually reaching the base (
<figureCitation id="13118A80FFA28D7BA733FBF451A9955D" box="[1150,1216,1090,1111]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="9" pageNumber="50">Figs. 8</figureCitation>
B–C, E, H); secondary basal line evident or not............................................................................... 12
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</paragraph>
</keyStep>
<keyStep id="30DE8EA0FFA28D7BA3DAFBCB50F29455" pageId="9" pageNumber="50">
<paragraph id="8B959605FFA28D7BA3DAFBCB50F2940D" blockId="9.[151,1436,151,1698]" pageId="9" pageNumber="50">
<keyLead id="30DB3530FFA28D7BA3DAFBCB50F2940D" pageId="9" pageNumber="50">
12 Pygidium with evident secondary basal line across base of depression, depression usually not reaching base, tubercle tubular with a dark tip (
<figureCitation id="13118A80FFA28D7BA2CEFB2C54B995A5" box="[387,464,1178,1199]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="9" pageNumber="50">Fig. 8E</figureCitation>
); mesoventrite with discernible punctures at least at sides and often reticulate; surface of the head finely reticulate between clypeogenal suture and anterior edge; elytral striae 6 and 7 appearing double, similar to striae 1–5; male metatibia with expanded inner edge, creating a small flange (
<figureCitation id="13118A80FFA28D7BA013FB6356CE95E0" box="[862,935,1237,1258]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="9" pageNumber="50">Fig. 4F</figureCitation>
); aedeagus as in
<figureCitation id="13118A80FFA28D7BA704FB6351F595E0" box="[1097,1180,1237,1258]" captionStart="FIGURE 10" captionStartId="23.[151,250,1909,1931]" captionTargetBox="[167,1333,208,1895]" captionTargetId="figure@23.[164,1428,193,1895]" captionTargetPageId="23" captionText="FIGURE 10. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588641/files/figure.png" pageId="9" pageNumber="50">Fig. 10E</figureCitation>
....................................................................................
<taxonomicName id="4C2AED86FFA28D7BA099FB445076940D" ID-CoL="6P7XX" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[980,1311,1266,1287]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="guthrieae" status="sp. nov.">
<emphasis id="B95E4A17FFA28D7BA099FB4451E1940D" box="[980,1160,1266,1287]" italics="true" pageId="9" pageNumber="50">Lepanus guthrieae</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFFA28D7BA664FB4450F2940D" box="[1321,1435,1266,1287]" pageId="9" pageNumber="50" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B959605FFA28D7BA3DAFAB950F29455" blockId="9.[151,1436,151,1698]" pageId="9" pageNumber="50">
<keyLead id="30DB3530FFA28D7BA3DAFAB950F29455" pageId="9" pageNumber="50">
- Pygidium with secondary basal line not evident, merged with base, upper edge of depression angulate, tubercle of various shapes (
<figureCitation id="13118A80FFA28D7BA260FA9B54079448" box="[301,366,1325,1346]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="9" pageNumber="50">Figs. 8</figureCitation>
B–C, H); mesoventrite virtually impunctate, not reticulate; surface of head usually entirely smooth; elytral striae 6 and 7 appearing single, different from striae 1–4 or 1–5; male metatibia with inner edge not expanded............... 13
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</paragraph>
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<keyStep id="30DE8EA0FFA28D7BA3DAFAD150F294F8" pageId="9" pageNumber="50">
<paragraph id="8B959605FFA28D7BA3DAFAD150F294BD" blockId="9.[151,1436,151,1698]" pageId="9" pageNumber="50">
<keyLead id="30DB3530FFA28D7BA3DAFAD150F294BD" pageId="9" pageNumber="50">
13 Apical tooth on protibiae with a slight notch on the inner edge (
<figureCitation id="13118A80FFA28D7BA02FFAD156D99476" box="[866,944,1383,1404]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="9" pageNumber="50">Fig. 4K</figureCitation>
) [often best viewed from below]; tubercle within pygidial depression low, round, or slightly elongate, same colour as depression (
<figureCitation id="13118A80FFA28D7BA0A9FA3351469490" box="[996,1071,1413,1434]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="9" pageNumber="50">Fig. 8B</figureCitation>
); eye canthus not dividing the eye (as in
<figureCitation id="13118A80FFA28D7BA3B4FA14542894BD" box="[249,321,1442,1463]" captionStart="FIGURE 3" captionStartId="7.[151,250,1911,1933]" captionTargetBox="[190,1318,415,1885]" captionTargetId="figure@7.[168,1467,409,1885]" captionTargetPageId="7" captionText="FIGURE 3. A, Dorsal aspect of key structures of Lepanus and other dung beetles; B, Ventral aspect of key structures of Lepanus and other dung beetles; C, key structures of a Lepanus head; D–F, aspect of the eye canthus relative to the eye: D, dividing the eye; E, nearly dividing the eye (separated by only a few eye facets); F, not dividing the eye; G–H, examples and key structures of Lepanus protibia: G, dorsal aspect of truncate protibia; H, dorsal aspect of deeply angulate protibia." httpUri="https://zenodo.org/record/2588627/files/figure.png" pageId="9" pageNumber="50">Fig. 3F</figureCitation>
); aedeagus as in
<figureCitation id="13118A80FFA28D7BA2A9FA14575194BD" box="[484,568,1442,1463]" captionStart="FIGURE 10" captionStartId="23.[151,250,1909,1931]" captionTargetBox="[167,1333,208,1895]" captionTargetId="figure@23.[164,1428,193,1895]" captionTargetPageId="23" captionText="FIGURE 10. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588641/files/figure.png" pageId="9" pageNumber="50">Fig. 10B</figureCitation>
..................................
<taxonomicName id="4C2AED86FFA28D7BA099FA14507694BD" ID-CoL="6P7WC" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[980,1311,1442,1463]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="crenidens" status="sp. nov.">
<emphasis id="B95E4A17FFA28D7BA099FA1451E194BD" box="[980,1160,1442,1463]" italics="true" pageId="9" pageNumber="50">Lepanus crenidens</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFFA28D7BA664FA1450F294BD" box="[1321,1435,1442,1463]" pageId="9" pageNumber="50" rank="species">new species</taxonomicNameLabel>
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</paragraph>
<paragraph id="8B959605FFA28D7BA3DAFA0950F294F8" blockId="9.[151,1436,151,1698]" pageId="9" pageNumber="50">
<keyLead id="30DB3530FFA28D7BA3DAFA0950F294F8" pageId="9" pageNumber="50">
- Apical tooth on protibiae without a notch on the inner edge; tubercle within pygidial depression with a darker tip, either shaped as an inverted V or slightly horizontal; eye canthus nearly dividing the eye (as in
<figureCitation id="13118A80FFA28D7BA0A1FA6B515D94F8" box="[1004,1076,1501,1522]" captionStart="FIGURE 3" captionStartId="7.[151,250,1911,1933]" captionTargetBox="[190,1318,415,1885]" captionTargetId="figure@7.[168,1467,409,1885]" captionTargetPageId="7" captionText="FIGURE 3. A, Dorsal aspect of key structures of Lepanus and other dung beetles; B, Ventral aspect of key structures of Lepanus and other dung beetles; C, key structures of a Lepanus head; D–F, aspect of the eye canthus relative to the eye: D, dividing the eye; E, nearly dividing the eye (separated by only a few eye facets); F, not dividing the eye; G–H, examples and key structures of Lepanus protibia: G, dorsal aspect of truncate protibia; H, dorsal aspect of deeply angulate protibia." httpUri="https://zenodo.org/record/2588627/files/figure.png" pageId="9" pageNumber="50">Fig. 3E</figureCitation>
)........................... 14
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</paragraph>
</keyStep>
<keyStep id="30DE8EA0FFA28D7BA3DAFA4C50F297A8" pageId="9" pageNumber="50">
<paragraph id="8B959605FFA28D7BA3DAFA4C50F29740" blockId="9.[151,1436,151,1698]" pageId="9" pageNumber="50">
<keyLead id="30DB3530FFA28D7BA3DAFA4C50F29740" pageId="9" pageNumber="50">
14 Pygidial depression with tubercle somewhat inverted V shaped (
<figureCitation id="13118A80FFA28D7BA000FA4C56F09705" box="[845,921,1530,1551]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="9" pageNumber="50">Fig. 8C</figureCitation>
); elytral striae 5–7 with discernible punctures at least in basal half; elytral flange and groove relatively wide; basal margin of pronotum with a distinct row of punctures (
<figureCitation id="13118A80FFA28D7BA60EF9A150E69726" box="[1347,1423,1559,1580]" captionStart="FIGURE 5" captionStartId="15.[151,250,1945,1967]" captionTargetBox="[245,1336,223,1911]" captionTargetId="figure@15.[233,1355,193,1926]" captionTargetPageId="15" captionText="FIGURE 5. Habitus of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil. All specimens in comparative scale. Scale bar 1 mm." httpUri="https://zenodo.org/record/2588631/files/figure.png" pageId="9" pageNumber="50">Fig. 5C</figureCitation>
); aedeagus as in
<figureCitation id="13118A80FFA28D7BA23CF98354AC9740" box="[369,453,1589,1610]" captionStart="FIGURE 10" captionStartId="23.[151,250,1909,1931]" captionTargetBox="[167,1333,208,1895]" captionTargetId="figure@23.[164,1428,193,1895]" captionTargetPageId="23" captionText="FIGURE 10. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588641/files/figure.png" pageId="9" pageNumber="50">Fig. 10C</figureCitation>
..............................................
<taxonomicName id="4C2AED86FFA28D7BA0A7F98350769740" ID-CoL="6P7VZ" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[1002,1311,1589,1610]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="feehani" status="sp. nov.">
<emphasis id="B95E4A17FFA28D7BA0A7F98351EE9740" box="[1002,1159,1589,1610]" italics="true" pageId="9" pageNumber="50">Lepanus feehani</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFFA28D7BA664F98350F29740" box="[1321,1435,1589,1610]" pageId="9" pageNumber="50" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B959605FFA28D7BA3DAF9E450F297A8" blockId="9.[151,1436,151,1698]" pageId="9" pageNumber="50">
<keyLead id="30DB3530FFA28D7BA3DAF9E450F297A8" pageId="9" pageNumber="50">
- Pygidial depression with tubercle slightly horizontal, not like an inverted V (
<figureCitation id="13118A80FFA28D7BA09EF9E45148976D" box="[979,1057,1618,1639]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="9" pageNumber="50">Fig. 8H</figureCitation>
); elytral striae 5–7 without discernible punctures; elytral flange and groove narrower; basal margin of pronotum without a distinct row of punctures (
<figureCitation id="13118A80FFA28D7BA7B2F9D95022978E" box="[1279,1355,1647,1668]" captionStart="FIGURE 6" captionStartId="16.[151,250,1948,1970]" captionTargetBox="[235,1333,202,1834]" captionTargetId="figure@16.[213,1373,193,1926]" captionTargetPageId="16" captionText="FIGURE 6. Habitus of species within the Lepanus pygmaeus species group. A, Lepanus lingziae; B, Lepanus mckenziei; C, Lepanus menendezae; D, Lepanus pecki; E: Lepanus podocarp; F, Lepanus pungalina. All specimens in comparative scale. Scale bar 1 mm." httpUri="https://zenodo.org/record/2588633/files/figure.png" pageId="9" pageNumber="50">Fig. 6B</figureCitation>
); aedeagus as in
<figureCitation id="13118A80FFA28D7BA276F93B54E797A8" box="[315,398,1677,1698]" captionStart="FIGURE 11" captionStartId="24.[151,250,1905,1927]" captionTargetBox="[195,1359,208,1854]" captionTargetId="figure@24.[194,1392,193,1889]" captionTargetPageId="24" captionText="FIGURE 11. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus lentil; B, Lepanus lingziae; C, Lepanus mckenziei; D, Lepanus menendezae; E, Lepanus pecki. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588643/files/figure.png" pageId="9" pageNumber="50">Fig. 11C</figureCitation>
................................................
<taxonomicName id="4C2AED86FFA28D7BA09FF93B507697A8" ID-CoL="3T8V2" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[978,1311,1677,1698]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="mckenziei" status="sp. nov.">
<emphasis id="B95E4A17FFA28D7BA09FF93B51EE97A8" box="[978,1159,1677,1698]" italics="true" pageId="9" pageNumber="50">Lepanus mckenziei</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFFA28D7BA664F93B50F297A8" box="[1321,1435,1677,1698]" pageId="9" pageNumber="50" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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