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<document id="CF6EDC86036858B4DB534D75F7F58930" ID-CLB-Dataset="269966" ID-DOI="10.5281/zenodo.5337424" ID-GBIF-Dataset="7c26f212-ab38-4e92-b461-cdb7dc073af5" ID-ISSN="0253-116X" ID-Zenodo-Dep="5337424" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1630178635768" checkinUser="carolina" docAuthor="Puthz, V." docDate="2012" docId="038313247C4BFFCEFF01FAE73EB6E210" docLanguage="en" docName="LinzerbiolBeitr.2012.44.2.1431-1475.pdf" docOrigin="Linzer biologische Beiträge 44 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Stenus perodiosus Puthz 2012, nov.sp." docType="treatment" docVersion="8" lastPageNumber="1448" masterDocId="FFBA6B5C7C5BFFDFFFB3FFE33D00E112" masterDocTitle="On the Stenus LATREILLE 1797 from Taiwan with non-spotted elytra (Coleoptera, Staphylinidae) (327 Contribution to the Knowledge of Steninae)" masterLastPageNumber="1475" masterPageNumber="1431" pageNumber="1447" updateTime="1699176368675" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="401A727BF1BC92CC6021029918863337">On the Stenus LATREILLE 1797 from Taiwan with non-spotted elytra (Coleoptera, Staphylinidae) (327 Contribution to the Knowledge of Steninae)</mods:title>
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<mods:namePart id="426178FB814EFC25F17A9C3BA477DF4B">Puthz, V.</mods:namePart>
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<mods:title id="823568DBA13F7885E4D93B90DB71F1B9">Linzer biologische Beiträge</mods:title>
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<mods:date id="2465369DC96C38CAA7FEDD5488879AB5">2012</mods:date>
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<mods:number id="F31F68B31BDF376BA711C4579B0F732D">2012-12-28</mods:number>
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<mods:number id="E7FC74100BCF813ED4BF707162423926">44</mods:number>
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<mods:identifier id="B4325443A6AE989318925F8F40910F26" type="DOI">10.5281/zenodo.5337424</mods:identifier>
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<mods:identifier id="A404612F233F74C34CAC3D2239271933" type="Zenodo-Dep">5337424</mods:identifier>
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<treatment id="038313247C4BFFCEFF01FAE73EB6E210" ID-DOI="http://doi.org/10.5281/zenodo.8392890" ID-GBIF-Taxon="213626800" ID-Zenodo-Dep="8392890" LSID="urn:lsid:plazi:treatment:038313247C4BFFCEFF01FAE73EB6E210" httpUri="http://treatment.plazi.org/id/038313247C4BFFCEFF01FAE73EB6E210" lastPageId="17" lastPageNumber="1448" pageId="16" pageNumber="1447">
<subSubSection id="C330F1B97C4BFFCFFF01FAE73F71E408" box="[178,625,1284,1306]" pageId="16" pageNumber="1447" type="nomenclature">
<paragraph id="8B95A2327C4BFFCFFF01FAE73F71E408" blockId="16.[178,625,1284,1306]" box="[178,625,1284,1306]" pageId="16" pageNumber="1447">
<heading id="D0DD155E7C4BFFCFFF01FAE73F71E408" box="[178,625,1284,1306]" fontSize="8" level="2" pageId="16" pageNumber="1447" reason="2">
<emphasis id="B95E7E207C4BFFCFFF01FAE73F71E408" bold="true" box="[178,625,1284,1306]" pageId="16" pageNumber="1447">
<taxonomicName id="4C2AD9B17C4BFFCFFF01FAE73C6FE408" ID-CoL="52B4G" authority="Puthz, 2012" authorityName="Puthz" authorityYear="2012" box="[178,367,1284,1306]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1447" phylum="Arthropoda" rank="species" species="perodiosus" status="sp. nov.">
<emphasis id="B95E7E207C4BFFCFFF01FAE73C6FE408" bold="true" box="[178,367,1284,1306]" italics="true" pageId="16" pageNumber="1447">Stenus perodiosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26DC35B7C4BFFCFFEC6FAE73CC2E408" box="[373,450,1284,1306]" pageId="16" pageNumber="1447" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="1311BEB77C4BFFCFFE7CFAE73F1EE408" box="[463,542,1284,1306]" captionStart="Figs 1-16" captionStartId="38.[178,218,1324,1344]" captionTargetBox="[178,1082,220,1305]" captionTargetId="figure-8@38.[178,1085,674,834]" captionTargetPageId="38" captionText="Figs 1-16: Sternite 8 of male (1-8, 13, 14) and of female (9, 10), posterior portion of sternite 9 of male (11, 12), anterior portion of aedeagus (15, 16) of Stenus hirtellus SHARP (1), S. piliferus MOTSCHULSKY (2), S. perpastus nov.sp. (3, PT), S. perpinguis nov.sp. (4, PT), S. perpropinquus nov.sp. (5, PT), S. persimplex nov.sp. (6, PT), S. perturbator nov.sp. (7, PT), S. pervilis nov.sp. (8, PT), S. persculptus nov.sp. (9, PT), S. perspicabilis nov.sp. (10, PT), S. sugayai nov.sp. (11, PT), S. perodiosus nov.sp. (12, PT), S. ascendor (13, 15, PT), S. hanami HROMÁDKA (14, 16, HT). – Scale bar = 0.1mm (1= 2-10; 11=12; 13=14; 15=16)." figureDoi="http://doi.org/10.5281/zenodo.5337426" httpUri="https://zenodo.org/record/5337426/files/figure.png" pageId="16" pageNumber="1447">Figs 12</figureCitation>
,
<figureCitation id="1311BEB77C4BFFCFFD99FAE73F45E408" box="[554,581,1284,1306]" captionStart="Figs 30-41" captionStartId="40.[178,218,1177,1197]" captionTargetBox="[179,1083,220,1157]" captionTargetId="figure-8@40.[178,1083,682,843]" captionTargetPageId="40" captionText="Figs 30-41: Ventral aspect of aedeagus (30-34, 37-41) outline of apical portion of aedeagus (35, 36) of Stenus persubtilis nov.sp. (30, T 146, PT), S. pervilis nov.sp. (31, T 12, PT), S. perversor nov.sp. (32, T 59, PT), S. permodestus nov.sp. (33, T 179, PT), S. perplicatus nov.sp. (34-36: Shouchia, PTT), S. persimplex nov.sp. (37, T 177, PT), S. perodiosus nov.sp. (38, T 85, PT), S. perrarus nov.sp. (39, T 5, PT), S. permolestus nov.sp. (40, T 119, PT), S. pertenuis nov.sp. (41, T 160, PT). – Scale bar = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.5337430" httpUri="https://zenodo.org/record/5337430/files/figure.png" pageId="16" pageNumber="1447">38</figureCitation>
,
<figureCitation id="1311BEB77C4BFFCFFDE3FAE73F6BE408" box="[592,619,1284,1306]" captionStart="Figs 55-69" captionStartId="42.[178,218,1301,1321]" captionTargetBox="[178,1085,219,1281]" captionTargetId="figure-149@42.[178,1085,219,1281]" captionTargetPageId="42" captionText="Figs 55-69: Spermathecae of PTT of Stenus pervenustus nov.sp. (55, T 138), S. persculptus nov.sp. (56, T 107), S. persculpturatus nov.sp. (57, T 88), S. pervilis nov.sp. (58, T 120), S. sugayai nov.sp. (59, Anmashan), S. persubtilis nov.sp. (60, T 199), S. perplicatus nov.sp. (61, Meifeng), S. perturbator nov.sp. (62, T 187), S. persimplex nov.sp. (63, T 177), S. perrarus nov.sp. (64, Fushan), S. perspicabilis nov.sp. (65, T 144), S. perpinguis nov.sp. (66, T 29), S. perodiosus nov.sp. (67, T 84), S. perpunctus nov.sp. (68, T 72), S. perversor nov.sp. (69, T 59)." figureDoi="http://doi.org/10.5281/zenodo.5337434" httpUri="https://zenodo.org/record/5337434/files/figure.png" pageId="16" pageNumber="1447">67</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C330F1B97C4BFFCFFF01FAD73CB5E4BA" pageId="16" pageNumber="1447" type="materials_examined">
<paragraph id="8B95A2327C4BFFCFFF01FAD73CB5E4BA" blockId="16.[178,1086,1332,1448]" pageId="16" pageNumber="1447">
Type material:
<materialsCitation id="3B42A86F7C4BFFCFFE17FAD73F1CE45A" ID-GBIF-Occurrence="4417845408" box="[420,540,1332,1352]" collectingDate="1991-05-17" collectionCode="Smetana" collectorName="A. Smetana" country="Taiwan" elevation="3375" location="Yushan National Park" pageId="16" pageNumber="1447" specimenCount="1" stateProvince="Nantou" typeStatus="holotype">
<typeStatus id="54911C907C4BFFCFFE17FAD73CF5E45A" box="[420,501,1332,1352]" pageId="16" pageNumber="1447" type="holotype">Holotype</typeStatus>
()
</materialsCitation>
<materialsCitation id="3B42A86F7C4BFFCFFD93FAD73E94E472" ID-GBIF-Occurrence="4417845523" collectingDate="1991-05-17" collectionCode="Smetana" collectorName="A. Smetana" country="Taiwan" elevation="3375" location="Yushan National Park" pageId="16" pageNumber="1447" specimenCount="36" stateProvince="Nantou" typeStatus="paratype">
and 17, 19 -
<typeStatus id="54911C907C4BFFCFFD79FAD73E22E45A" box="[714,802,1332,1352]" pageId="16" pageNumber="1447" type="paratype">paratypes</typeStatus>
:
<collectingCountry id="F33DE2A27C4BFFCFFC99FAD73E6FE45A" box="[810,879,1332,1352]" name="Taiwan" pageId="16" pageNumber="1447">Taiwan</collectingCountry>
:
<collectingRegion id="49EE6CD07C4BFFCFFCC5FAD73EF2E45A" box="[886,1010,1332,1352]" country="Taiwan" name="Nantou" pageId="16" pageNumber="1447">Nantou Hsien</collectingRegion>
:
<location id="8EF5F4E97C4BFFCFFC49FAD73C5AE472" LSID="urn:lsid:plazi:treatment:038313247C4BFFCEFF01FAE73EB6E210:8EF5F4E97C4BFFCFFC49FAD73C5AE472" country="Taiwan" name="Yushan National Park" pageId="16" pageNumber="1447" stateProvince="Nantou">Yushan National Park</location>
,
<locationDeviation id="9A27BB107C4BFFCFFED0FAAF3F34E472" box="[355,564,1356,1376]" location="Pai-Yun Hut" pageId="16" pageNumber="1447">
<quantity id="4CD20FD77C4BFFCFFED0FAAF3CA1E472" box="[355,417,1356,1376]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="16" pageNumber="1447" unit="km" value="1.8">1.8 km</quantity>
W Pai-Yun Hut
</locationDeviation>
,
<quantity id="4CD20FD77C4BFFCFFD8EFAAF3F7CE472" box="[573,636,1356,1376]" metricMagnitude="3" metricUnit="m" metricValue="3.375" pageId="16" pageNumber="1447" unit="m" value="3375.0">
<elevation id="000745017C4BFFCFFD8EFAAF3F7CE472" box="[573,636,1356,1376]" metricMagnitude="3" metricUnit="m" metricValue="3.375" pageId="16" pageNumber="1447" unit="m" value="3375.0">3375m</elevation>
</quantity>
,
<date id="FF9484F27C4BFFCFFD35FAAF3FE3E472" box="[646,739,1356,1376]" pageId="16" pageNumber="1447" value="1991-05-17">
<collectingDate id="EFD07D1A7C4BFFCFFD35FAAF3FE3E472" box="[646,739,1356,1376]" pageId="16" pageNumber="1447" value="1991-05-17">17.V.1991</collectingDate>
</date>
,
<collectorName id="26DFC7E47C4BFFCFFD5EFAAF3E54E472" box="[749,852,1356,1376]" pageId="16" pageNumber="1447">A. Smetana</collectorName>
(T 85)
</materialsCitation>
.
<materialsCitation id="3B42A86F7C4BFFCFFC2DFAAF3E4CE46A" ID-GBIF-Occurrence="4417845496" collectingDate="1991-05-16" collectionCode="cN, cT, cP" collectorName="A. Smetana" country="Taiwan" elevation="3350" location="Pai-Yun Hut" pageId="16" pageNumber="1447" specimenCount="20" stateProvince="Nantou" typeStatus="paratype">
<typeStatus id="54911C907C4BFFCFFC2DFAAF3EF6E472" box="[926,1014,1356,1376]" pageId="16" pageNumber="1447" type="paratype">Paratypes</typeStatus>
: 11, 9: ibidem,
<quantity id="4CD20FD77C4BFFCFFED0FA873C92E46A" box="[355,402,1380,1400]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="1447" unit="km" value="2.0">2 km</quantity>
W
<location id="8EF5F4E97C4BFFCFFE00FA873F25E46A" LSID="urn:lsid:plazi:treatment:038313247C4BFFCEFF01FAE73EB6E210:8EF5F4E97C4BFFCFFE00FA873F25E46A" box="[435,549,1380,1400]" name="Pai-Yun Hut" pageId="16" pageNumber="1447">Pai-Yun Hut</location>
,
<quantity id="4CD20FD77C4BFFCFFD9CFA873F6EE46A" box="[559,622,1380,1400]" metricMagnitude="3" metricUnit="m" metricValue="3.35" pageId="16" pageNumber="1447" unit="m" value="3350.0">
<elevation id="000745017C4BFFCFFD9CFA873F6EE46A" box="[559,622,1380,1400]" metricMagnitude="3" metricUnit="m" metricValue="3.35" pageId="16" pageNumber="1447" unit="m" value="3350.0">3350m</elevation>
</quantity>
,
<date id="FF9484F27C4BFFCFFDCBFA873FD5E46A" box="[632,725,1380,1400]" pageId="16" pageNumber="1447" value="1991-05-16">
<collectingDate id="EFD07D1A7C4BFFCFFDCBFA873FD5E46A" box="[632,725,1380,1400]" pageId="16" pageNumber="1447" value="1991-05-16">16.V.1991</collectingDate>
</date>
, idem (T 84)
</materialsCitation>
;
<materialsCitation id="3B42A86F7C4BFFCFFCE5FA873CB1E4BA" ID-GBIF-Occurrence="4417845483" collectingDate="1991-05-13" collectionCode="cN, cT, cP" collectorName="Smetana" country="Taiwan" elevation="2700" location="Clif" municipality="Yushan National Park" pageId="16" pageNumber="1447" specimenCount="2" stateProvince="Nantou" typeStatus="paratype">
1, 1:
<collectingMunicipality id="6BF138487C4BFFCFFC1AFA873C09E482" pageId="16" pageNumber="1447">Yushan National Park</collectingMunicipality>
, Mun-Li
<location id="8EF5F4E97C4BFFCFFEEFFA9F3C80E482" LSID="urn:lsid:plazi:treatment:038313247C4BFFCEFF01FAE73EB6E210:8EF5F4E97C4BFFCFFEEFFA9F3C80E482" box="[348,384,1404,1424]" municipality="Yushan National Park" name="Clif" pageId="16" pageNumber="1447">Clif</location>
,
<quantity id="4CD20FD77C4BFFCFFE3AFA9F3CC9E482" box="[393,457,1404,1424]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="16" pageNumber="1447" unit="m" value="2700.0">
<elevation id="000745017C4BFFCFFE3AFA9F3CC9E482" box="[393,457,1404,1424]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="16" pageNumber="1447" unit="m" value="2700.0">2700m</elevation>
</quantity>
,
<date id="FF9484F27C4BFFCFFE62FA9F3F2FE482" box="[465,559,1404,1424]" pageId="16" pageNumber="1447" value="1991-05-13">
<collectingDate id="EFD07D1A7C4BFFCFFE62FA9F3F2FE482" box="[465,559,1404,1424]" pageId="16" pageNumber="1447" value="1991-05-13">13.V.1991</collectingDate>
</date>
, idem (T 79).- HT and PTT in coll. Smetana (Ottawa), PTT also in cN, cT and in cP
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C330F1B97C4BFFCEFF01FA543E1EE372" lastPageId="17" lastPageNumber="1448" pageId="16" pageNumber="1447" type="description">
<paragraph id="8B95A2327C4BFFCFFF01FA543FDDE4DE" blockId="16.[178,733,1462,1484]" box="[178,733,1462,1484]" pageId="16" pageNumber="1447">
Description: About as in
<taxonomicName id="4C2AD9B17C4BFFCFFDBBFA543F8DE4DE" authority="Puthz, 2012" authorityName="Puthz" authorityYear="2012" box="[520,653,1463,1484]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1447" phylum="Arthropoda" rank="species" species="persubtilis" status="sp. nov.">
<emphasis id="B95E7E207C4BFFCFFDBBFA543F8DE4DE" box="[520,653,1463,1484]" italics="true" pageId="16" pageNumber="1447">S. persubtilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26DC35B7C4BFFCFFD20FA543FDDE4DE" box="[659,733,1463,1484]" pageId="16" pageNumber="1447" rank="species">nov.sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="8B95A2327C4BFFCFFF01FA3F3F6CE4E0" blockId="16.[178,620,1500,1522]" box="[178,620,1500,1522]" pageId="16" pageNumber="1447">
Length:
<quantity id="4CD20FD77C4BFFCFFEB4FA3F3C76E4E0" box="[263,374,1500,1522]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="16" pageNumber="1447" unit="mm" value="3.0" valueMax="3.5" valueMin="2.5">2.5-3.5mm</quantity>
(forebody:
<quantity id="4CD20FD77C4BFFCFFE5DFA3F3F60E4E0" box="[494,608,1500,1522]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="16" pageNumber="1447" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4-1.6mm</quantity>
).
</paragraph>
<paragraph id="8B95A2327C4BFFCFFF01F9E13EBBE705" blockId="16.[178,955,1537,1559]" box="[178,955,1537,1559]" pageId="16" pageNumber="1447">PM of the HT: HW: 29; DE: 15; PW: 21; PL: 21; EW: 26.5; EL: 23; SL: 17.</paragraph>
<paragraph id="8B95A2327C4BFFCEFF01F9C43F84E091" blockId="16.[178,1084,1574,1596]" lastBlockId="17.[178,1086,219,387]" lastPageId="17" lastPageNumber="1448" pageId="16" pageNumber="1447">Male: Metasternum shallowly impressed, moderately coarsely and moderately densely punctate, interstices often as large as half diameter of punctures. Anterior sternites simple, sternite 7 finer and denser punctate and pubescent posteromedially. Sternite 8 with a very shallow apical emargination (LS: ED= 50:2). Sternite 9 with comparatively short apicolateral teeth (fig. 12). Aedeagus (fig. 38), apical portion triangularly narrowed, apex acute, expulsion hooks connected medially; parameres slightly longer than median lobe, with about 8-9 apical setae.</paragraph>
<paragraph id="8B95A2327C4AFFCEFF01FE703F79E0F1" blockId="17.[178,1085,402,483]" pageId="17" pageNumber="1448">Female: Sternite 8 at posterior margin slightly projecting, apex moderately broadly rounded. Valvifer with a comparatively short apicolateral tooth. Spermatheca (fig. 67), infundibulum more than 3x as long as broad.</paragraph>
<paragraph id="8B95A2327C4AFFCEFF01FE113E1EE372" blockId="17.[178,1085,498,608]" pageId="17" pageNumber="1448">
About as
<taxonomicName id="4C2AD9B17C4AFFCEFEA4FE103C9EE315" authorityName="Puthz" authorityYear="2012" box="[279,414,498,520]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1448" phylum="Arthropoda" rank="species" species="persubtilis">
<emphasis id="B95E7E207C4AFFCEFEA4FE103C9EE315" box="[279,414,498,520]" italics="true" pageId="17" pageNumber="1448">S. persubtilis</emphasis>
</taxonomicName>
, but the elytra with deep longitudinal impressions in lateral half and near suture; median portion of frons distinctly exceeding the level of medial eye margins; longitudinal impression of pronotum deep, broadly impunctate and deeply reticulate, dull; abdominal punctation throughout very dense.
</paragraph>
</subSubSection>
<subSubSection id="C330F1B97C4AFFCEFF01FD8C3C20E3D2" pageId="17" pageNumber="1448" type="discussion">
<paragraph id="8B95A2327C4AFFCEFF01FD8C3C20E3D2" blockId="17.[178,1085,623,704]" pageId="17" pageNumber="1448">Comparative note: This new species resembles many similar species with deep elytral impressions, from which it should be distinguished by examining the sexual characters.</paragraph>
</subSubSection>
<subSubSection id="C330F1B97C4AFFCEFF01FD333EB6E210" pageId="17" pageNumber="1448" type="etymology">
<paragraph id="8B95A2327C4AFFCEFF01FD333EB6E210" blockId="17.[178,1085,719,770]" pageId="17" pageNumber="1448">
Etymology: Because of its great similarity with other species of the group and the difficulty to identify this species I name it &quot;
<emphasis id="B95E7E207C4AFFCEFDD8FD0F3FDAE213" box="[619,730,748,769]" italics="true" pageId="17" pageNumber="1448">perodiosus</emphasis>
&quot; (Lat.= very odious).
</paragraph>
</subSubSection>
</treatment>
</document>
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