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<mods:title id="C4CD3234D9F08A30769A046662AF4E0F">A taxonomic contribution to the genus Brachythele Ausserer, 1871 (Araneae, Nemesiidae) with a description of a new species from Rhodope Mountains, Bulgaria</mods:title>
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<mods:namePart id="B62511A1D718A19383CB01854685050D">Dimitrov, Dragomir</mods:namePart>
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<mods:namePart id="3C51E2BE65DF14EB92645D0A742856E6">Zonstein, Sergei</mods:namePart>
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<mods:affiliation id="FB736210C5A485323EDD5E1A4C1C3D3D">Steinhardt Museum of Natural History, Tel-Aviv University, Klauzner Str. 12, 69978 Tel Aviv, Israel. https: // orcid. org / 0000 - 0002 - 4515 - 0630</mods:affiliation>
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<treatment id="038309341135F35A969AFF4FFDB4FF49" ID-DOI="http://doi.org/10.5281/zenodo.6786071" ID-GBIF-Taxon="196373073" ID-Zenodo-Dep="6786071" LSID="urn:lsid:plazi:treatment:038309341135F35A969AFF4FFDB4FF49" httpUri="http://treatment.plazi.org/id/038309341135F35A969AFF4FFDB4FF49" lastPageId="4" lastPageNumber="587" pageId="3" pageNumber="586">
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<taxonomicName id="4C2AC3A11135F35D969AFF4FFDF9FF00" ID-CoL="N33Z" authority="Lazarov, 2005" authorityName="Lazarov" authorityYear="2005" box="[151,620,151,178]" class="Arachnida" family="Nemesiidae" genus="Brachythele" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="586" phylum="Arthropoda" rank="species" species="langourovi">
<emphasis id="B95E64301135F35D969AFF4FFDF9FF00" bold="true" box="[151,620,151,178]" pageId="3" pageNumber="586">
<emphasis id="B95E64301135F35D969AFF4FFE20FF03" bold="true" box="[151,437,151,178]" italics="true" pageId="3" pageNumber="586">Brachythele langourovi</emphasis>
<bibRefCitation id="EFBBC5D31135F35D97B0FF4FFDF9FF00" author="Lazarov, S. P." box="[445,620,151,177]" pageId="3" pageNumber="586" pagination="189 - 193" refId="ref3680" refString="Lazarov, S. P. (2005) A new spider species from Bulgaria, Brachythele langourovi sp. n. (Araneae, Nemesiidae). Revue Suisse de Zoologie, 112 (1), 189 - 193. https: // doi. org / 10.5962 / bhl. part. 80294" type="journal article" year="2005">Lazarov, 2005</bibRefCitation>
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<figureCitation id="1311A4A71135F35D969AFF63FE81FF64" box="[151,276,187,213]" captionStart="FIGURES 712" captionStartId="3.[151,264,1916,1940]" captionTargetBox="[151,1436,617,1893]" captionTargetId="figure-145@3.[151,1436,617,1893]" captionTargetPageId="3" captionText="FIGURES 712. Brachythele rhodopensis sp. n., holotype male, palp (79) and Brachythele langourovi, paratype, male, palp (1012). 7, 10 prolateral; 8, 11 ventral; 9, 12 retrolateral. Scale 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6786155" httpUri="https://zenodo.org/record/6786155/files/figure.png" pageId="3" pageNumber="586">Figs 1012</figureCitation>
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<paragraph id="8B95B8221135F35D969AFED8FDC4FEA6" blockId="3.[151,825,255,341]" box="[151,593,255,280]" pageId="3" pageNumber="586">
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<treatmentCitation id="0A8B9E331135F35D969AFED8FE7BFEA9" author="Lazarov, S. P." box="[151,494,255,280]" page="189" pageId="3" pageNumber="586" year="2005">
<taxonomicName id="4C2AC3A11135F35D969AFED8FE7BFEA9" ID-CoL="N33Z" authority="Lazarov, 2005: 189" authorityName="Lazarov" authorityPageNumber="189" authorityYear="2005" box="[151,494,255,280]" class="Arachnida" family="Nemesiidae" genus="Brachythele" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="586" phylum="Arthropoda" rank="species" species="langourovi">
<emphasis id="B95E64301135F35D969AFED8FEB4FEA9" box="[151,289,256,280]" italics="true" pageId="3" pageNumber="586">B. langourovi</emphasis>
<bibRefCitation id="EFBBC5D31135F35D9725FED8FE7BFEA9" author="Lazarov, S. P." box="[296,494,255,280]" pageId="3" pageNumber="586" pagination="189 - 193" refId="ref3680" refString="Lazarov, S. P. (2005) A new spider species from Bulgaria, Brachythele langourovi sp. n. (Araneae, Nemesiidae). Revue Suisse de Zoologie, 112 (1), 189 - 193. https: // doi. org / 10.5962 / bhl. part. 80294" type="journal article" year="2005">Lazarov, 2005: 189</bibRefCitation>
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, figs 15.
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<treatmentCitation id="0A8B9E331135F35D969AFEC7FE70FE87" author="Le Peru, B." box="[151,485,286,310]" page="88" pageId="3" pageNumber="586" year="2011">
<taxonomicName id="4C2AC3A11135F35D969AFEC7FE70FE87" ID-CoL="N33Z" authority=": Le Peru, 2011: 88" authorityName="Le" authorityYear="2011" box="[151,485,286,310]" class="Arachnida" family="Nemesiidae" genus="Brachythele" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="586" phylum="Arthropoda" rank="species" species="langourovi">
<emphasis id="B95E64301135F35D969AFEC7FEB4FE87" box="[151,289,286,310]" italics="true" pageId="3" pageNumber="586">B. langourovi</emphasis>
:
<bibRefCitation id="EFBBC5D31135F35D9722FEC7FE70FE87" author="Le Peru, B." box="[303,485,286,310]" pageId="3" pageNumber="586" pagination="1 - 522" refId="ref3782" refString="Le Peru, B. (2011) The spiders of Europe, a synthesis of data. Vol. 1. Atypidae to Theridiidae. Memoires de la Societe Linneenne de Lyon, 2, 1 - 522." type="journal article" year="2011">Le Peru, 2011: 88</bibRefCitation>
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, fig. 42.
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<treatmentCitation id="0A8B9E331135F35D969AFEE6FD72FEE4" author="Naumova, M. &amp; Hristovski, S. &amp; Hristov, G. H." box="[151,743,317,341]" page="505" pageId="3" pageNumber="586" year="2016">
<taxonomicName id="4C2AC3A11135F35D969AFEE6FD72FEE4" ID-CoL="N33Z" authority=": Naumova, Hristovski &amp; Hristov, 2016: 505" authorityName=": Naumova, Hristovski &amp; Hristov" authorityPageNumber="505" authorityYear="2016" box="[151,743,317,341]" class="Arachnida" family="Nemesiidae" genus="Brachythele" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="586" phylum="Arthropoda" rank="species" species="langourovi">
<emphasis id="B95E64301135F35D969AFEE6FEB4FEE4" box="[151,289,317,341]" italics="true" pageId="3" pageNumber="586">B. langourovi</emphasis>
:
<bibRefCitation id="EFBBC5D31135F35D9722FEE5FD72FEE4" author="Naumova, M. &amp; Hristovski, S. &amp; Hristov, G. H." box="[303,743,317,341]" pageId="3" pageNumber="586" pagination="503 - 511" refId="ref3822" refString="Naumova, M., Hristovski, S. &amp; Hristov, G. H. (2016) Spiders (Arachnida: Araneae) from Prespa National Park, Albania. Acta Zoologica Bulgarica, 68 (4), 503 - 511." type="journal article" year="2016">Naumova, Hristovski &amp; Hristov, 2016: 505</bibRefCitation>
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, fig 2B.
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<emphasis id="B95E64301135F35D969AFE5BFE16FE2C" bold="true" box="[151,387,387,413]" pageId="3" pageNumber="586">Material examined.</emphasis>
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,
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,
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,
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south of
<collectingMunicipality id="6BF122581135F35D922FFE5BFB0FFE2C" box="[1058,1178,387,413]" pageId="3" pageNumber="586">Kamenitsa</collectingMunicipality>
vill., deciduous forest (
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<emphasis id="B95E64301135F35D96ACFE70FEE4FE70" box="[161,369,423,450]" italics="true" pageId="3" pageNumber="586">Quercus coccifera</emphasis>
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), 5.04
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<collectingDate id="EFD0670A1135F35D97C6FE7FFDA8FE73" box="[459,573,423,450]" pageId="3" pageNumber="586" value="2002-05-09">9.05.2002</collectingDate>
</date>
,
<collectorName id="26DFDDF41135F35D9441FE7FFD5CFE73" box="[588,713,423,450]" pageId="3" pageNumber="586">S. Lazarov</collectorName>
leg. (
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)
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.
<materialsCitation id="3B42B27F1135F35D959DFE7FFDAFFE54" ID-GBIF-Occurrence="3821647306" collectionCode="SMF" country="Macedonia" location="North" municipality="Scopje" pageId="3" pageNumber="586" specimenCode="SMF 3482" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D2C73AB1135F35D959DFE7FFC77FE70" box="[912,994,423,449]" pageId="3" pageNumber="586" type="male">1 male</specimenCount>
,
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<collectingCountry id="F33DF8B21135F35D9230FE7FFB29FE70" box="[1085,1212,423,449]" name="Macedonia" pageId="3" pageNumber="586">Macedonia</collectingCountry>
,
<collectingMunicipality id="6BF122581135F35D92C7FE7FFA83FE70" box="[1226,1302,423,449]" pageId="3" pageNumber="586">Scopje</collectingMunicipality>
(Üsküb, as labelled), no other data (
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a)
</materialsCitation>
.
</paragraph>
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<emphasis id="B95E64301135F35D96CAFE37FE62FDB8" bold="true" box="[199,503,495,521]" pageId="3" pageNumber="586">Diagnosis (partial, male).</emphasis>
Males are morphologically similar to those of
<taxonomicName id="4C2AC3A11135F35D921FFE28FADFFDB8" authority="Zonstein, 2007" authorityName="Zonstein" authorityYear="2007" box="[1042,1354,495,522]" class="Arachnida" family="Nemesiidae" genus="Brachythele" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="586" phylum="Arthropoda" rank="species" species="bentzieni">
<emphasis id="B95E64301135F35D921FFE28FBB6FDB8" box="[1042,1059,496,521]" italics="true" pageId="3" pageNumber="586">B</emphasis>
.
<emphasis id="B95E64301135F35D923EFE37FB0DFDB8" box="[1075,1176,495,521]" italics="true" pageId="3" pageNumber="586">bentzieni</emphasis>
<bibRefCitation id="EFBBC5D31135F35D92ACFE37FADFFDB8" author="Zonstein, S. L." box="[1185,1354,495,521]" pageId="3" pageNumber="586" pagination="15 - 17" refId="ref4063" refString="Zonstein, S. L. (2007) A new species of the mygalomorph spider genus Brachythele Ausserer, 1871 (Aranei: Nemesiidae) from Greece. Arthropoda Selecta, 16, 15 - 17." type="journal article" year="2007">Zonstein, 2007</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2AC3A11135F35D9389FE28FE8AFD9C" authority="Dimitrov &amp; Zonstein, 2022" authorityName="Dimitrov &amp; Zonstein" authorityYear="2022" class="Arachnida" family="Nemesiidae" genus="Brachythele" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="586" phylum="Arthropoda" rank="species" species="rhodopensis" status="sp. nov.">
<emphasis id="B95E64301135F35D9389FE28FE8AFD9C" italics="true" pageId="3" pageNumber="586">B. rhodopensis</emphasis>
</taxonomicName>
<emphasis id="B95E64301135F35D972BFDCBFEF1FD9C" bold="true" box="[294,356,531,557]" pageId="3" pageNumber="586">
<taxonomicNameLabel id="A26DD94B1135F35D972BFDCBFEF1FD9C" box="[294,356,531,557]" pageId="3" pageNumber="586" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by having an embolic keel, but can be distinguished by the thin, curved embolic portion (
<figureCitation id="1311A4A71135F35D934CFDCBFF21FDE0" captionStart="FIGURES 712" captionStartId="3.[151,264,1916,1940]" captionTargetBox="[151,1436,617,1893]" captionTargetId="figure-145@3.[151,1436,617,1893]" captionTargetPageId="3" captionText="FIGURES 712. Brachythele rhodopensis sp. n., holotype male, palp (79) and Brachythele langourovi, paratype, male, palp (1012). 7, 10 prolateral; 8, 11 ventral; 9, 12 retrolateral. Scale 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6786155" httpUri="https://zenodo.org/record/6786155/files/figure.png" pageId="3" pageNumber="586">Figs 10, 12</figureCitation>
) vs. more massive in
<taxonomicName id="4C2AC3A11135F35D97A4FDE0FDC5FDE0" authority="Dimitrov &amp; Zonstein, 2022" authorityName="Dimitrov &amp; Zonstein" authorityYear="2022" box="[425,592,567,593]" class="Arachnida" family="Nemesiidae" genus="Brachythele" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="586" phylum="Arthropoda" rank="species" species="rhodopensis" status="sp. nov.">
<emphasis id="B95E64301135F35D97A4FDE0FDC5FDE0" box="[425,592,567,593]" italics="true" pageId="3" pageNumber="586">B. rhodopensis</emphasis>
</taxonomicName>
<emphasis id="B95E64301135F35D945AFDEFFD02FDE0" bold="true" box="[599,663,567,593]" pageId="3" pageNumber="586">
<taxonomicNameLabel id="A26DD94B1135F35D945AFDEFFD02FDE0" box="[599,663,567,593]" pageId="3" pageNumber="586" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="1311A4A71135F35D94A8FDEFFC9FFDE0" box="[677,778,567,593]" captionStart="FIGURES 712" captionStartId="3.[151,264,1916,1940]" captionTargetBox="[151,1436,617,1893]" captionTargetId="figure-145@3.[151,1436,617,1893]" captionTargetPageId="3" captionText="FIGURES 712. Brachythele rhodopensis sp. n., holotype male, palp (79) and Brachythele langourovi, paratype, male, palp (1012). 7, 10 prolateral; 8, 11 ventral; 9, 12 retrolateral. Scale 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6786155" httpUri="https://zenodo.org/record/6786155/files/figure.png" pageId="3" pageNumber="586">Figs 79</figureCitation>
,
<figureCitation id="1311A4A71135F35D951BFDEFFCC8FDE3" box="[790,861,567,594]" captionStart="FIGURES 1315" captionStartId="4.[151,264,959,983]" captionTargetBox="[151,1436,273,935]" captionTargetId="figure-52@4.[151,1436,272,935]" captionTargetPageId="4" captionText="FIGURES 1315. Brachythele rhodopensis sp. n., holotype male, palp. 13 prolateral; 14 ventral; 15 retrolateral. T—tegulum; E—embolus; K—embolic keel. Scale 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6786159" httpUri="https://zenodo.org/record/6786159/files/figure.png" pageId="3" pageNumber="586">1315</figureCitation>
) and straighter in
<taxonomicName id="4C2AC3A11135F35D9224FDE0FB38FDE0" authorityName="Zonstein" authorityYear="2007" box="[1065,1197,567,594]" class="Arachnida" family="Nemesiidae" genus="Brachythele" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="586" phylum="Arthropoda" rank="species" species="bentzieni">
<emphasis id="B95E64301135F35D9224FDE0FBAFFDE0" box="[1065,1082,568,593]" italics="true" pageId="3" pageNumber="586">B</emphasis>
.
<emphasis id="B95E64301135F35D9245FDEFFB38FDE0" box="[1096,1197,567,593]" italics="true" pageId="3" pageNumber="586">bentzieni</emphasis>
</taxonomicName>
(
<figureCitation id="1311A4A71135F35D92B1FDEFFAD5FDE0" box="[1212,1344,567,593]" captionStart="FIGURES 2627" captionStartId="7.[151,264,1657,1681]" captionTargetBox="[151,1427,781,1634]" captionTargetId="figure-286@7.[151,1436,777,1634]" captionTargetPageId="7" captionText="FIGURES 2627. Brachythele bentzieni, male palp. 26 ventral; 27 retrolateral. Scale 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6786167" httpUri="https://zenodo.org/record/6786167/files/figure.png" pageId="3" pageNumber="586">Figs 2627</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF55E8AA1135F35D969AF8A4FCFBF809" ID-DOI="http://doi.org/10.5281/zenodo.6786155" ID-Zenodo-Dep="6786155" httpUri="https://zenodo.org/record/6786155/files/figure.png" pageId="3" pageNumber="586" startId="3.[151,264,1916,1940]" targetBox="[151,1436,617,1893]" targetPageId="3">
<paragraph id="8B95B8221135F35D969AF8A4FCFBF809" blockId="3.[151,1437,1916,1977]" pageId="3" pageNumber="586">
<emphasis id="B95E64301135F35D969AF8A4FEDDF824" bold="true" box="[151,328,1916,1941]" pageId="3" pageNumber="586">FIGURES 712.</emphasis>
<taxonomicName id="4C2AC3A11135F35D9743F8A5FDDFF824" authority="Dimitrov &amp; Zonstein, 2022" authorityName="Dimitrov &amp; Zonstein" authorityYear="2022" box="[334,586,1917,1941]" class="Arachnida" family="Nemesiidae" genus="Brachythele" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="586" phylum="Arthropoda" rank="species" species="rhodopensis" status="sp. nov.">
<emphasis id="B95E64301135F35D9743F8A5FDDFF824" box="[334,586,1917,1941]" italics="true" pageId="3" pageNumber="586">Brachythele rhodopensis</emphasis>
</taxonomicName>
<emphasis id="B95E64301135F35D945FF8A5FD1EF824" bold="true" box="[594,651,1917,1941]" pageId="3" pageNumber="586">
<taxonomicNameLabel id="A26DD94B1135F35D945FF8A5FD1EF824" box="[594,651,1917,1941]" pageId="3" pageNumber="586" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype male, palp (79) and
<taxonomicName id="4C2AC3A11135F35D95DCF8A5FB54F824" authorityName=": Naumova, Hristovski &amp; Hristov" authorityYear="2016" box="[977,1217,1917,1941]" class="Arachnida" family="Nemesiidae" genus="Brachythele" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="586" phylum="Arthropoda" rank="species" species="langourovi">
<emphasis id="B95E64301135F35D95DCF8A5FB54F824" box="[977,1217,1917,1941]" italics="true" pageId="3" pageNumber="586">Brachythele langourovi</emphasis>
</taxonomicName>
, paratype, male, palp
<emphasis id="B95E64301135F35D969AF879FF7DF808" box="[151,232,1953,1977]" italics="true" pageId="3" pageNumber="586">(1012)</emphasis>
. 7, 10 prolateral; 8, 11 ventral; 9, 12 retrolateral. Scale 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C330EBA91132F35A96CAFF4EFDD1FF00" box="[199,580,150,177]" pageId="4" pageNumber="587" type="description">
<paragraph id="8B95B8221132F35A96CAFF4EFDD1FF00" blockId="4.[151,1436,150,248]" box="[199,580,150,177]" pageId="4" pageNumber="587">
<emphasis id="B95E64301132F35A96CAFF4EFECCFF01" bold="true" box="[199,345,150,176]" pageId="4" pageNumber="587">Description.</emphasis>
See
<bibRefCitation id="EFBBC5D31132F35A9782FF4FFDD5FF00" author="Lazarov, S. P." box="[399,576,150,177]" pageId="4" pageNumber="587" pagination="189 - 193" refId="ref3680" refString="Lazarov, S. P. (2005) A new spider species from Bulgaria, Brachythele langourovi sp. n. (Araneae, Nemesiidae). Revue Suisse de Zoologie, 112 (1), 189 - 193. https: // doi. org / 10.5962 / bhl. part. 80294" type="journal article" year="2005">Lazarov (2005)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C330EBA91132F35A96CAFF62FDB4FF49" pageId="4" pageNumber="587" type="distribution">
<paragraph id="8B95B8221132F35A96CAFF62FDB4FF49" blockId="4.[151,1436,150,248]" pageId="4" pageNumber="587">
<emphasis id="B95E64301132F35A96CAFF62FEF4FF65" bold="true" box="[199,353,186,212]" pageId="4" pageNumber="587">Distribution.</emphasis>
Previously known only from
<collectingCountry id="F33DF8B21132F35A94CFFF62FCB7FF65" box="[706,802,186,212]" name="Bulgaria" pageId="4" pageNumber="587">Bulgaria</collectingCountry>
(
<bibRefCitation id="EFBBC5D31132F35A9538FF62FBECFF65" author="World Spider Catalog" box="[821,1145,186,212]" pageId="4" pageNumber="587" refId="ref4101" refString="World Spider Catalog (2022) World Spider Catalog. Version 23.0. Natural History Museum Bern, Bern. Available from: http: // wsc. nmbe. ch (accessed 22 January 2022) https: // doi. org / 10.24436 / 2" type="book" year="2022">World Spider Catalog 2022</bibRefCitation>
), and here reported from North
<collectingCountry id="F33DF8B21132F35A96EDFF06FEC9FF49" box="[224,348,222,248]" name="Macedonia" pageId="4" pageNumber="587">Macedonia</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>