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<mods:title id="C1BAD31FEA59E5966C6FF9B91C855E98">A revision of the genus Heinzia Korge, 1971 (Coleoptera: Staphylinidae: Quediina), with description of a new species and its probable larva</mods:title>
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<taxonomicName id="7CFE4D2EE01FB16BC82DFD14D52608E6" ID-CoL="8WG9F" authority="Korge, 1971" authorityName="Korge" authorityYear="1971" box="[264,517,718,744]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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(
<figureCitation id="23C52A28E01FB16BCB30FD14D5AC08E6" box="[533,655,718,744]" captionStart-0="FIGURES 1 ­ 7" captionStart-1="FIGURES 8 ­ 13" captionStart-10="FIGURE 48" captionStart-2="FIGURES 14 ­ 19" captionStart-3="FIGURES 20 ­ 23" captionStart-4="FIGURES 24 ­ 27" captionStart-5="FIGURES 28 ­ 32" captionStart-6="FIGURES 33 ­ 35" captionStart-7="FIGURES 36 ­ 38" captionStart-8="FIGURES 39 ­ 45" captionStart-9="FIGURES 46 ­ 47" captionStartId-0="3.[264,382,1783,1807]" captionStartId-1="5.[264,383,1583,1607]" captionStartId-10="16.[264,367,1711,1735]" captionStartId-2="6.[264,383,1576,1600]" captionStartId-3="7.[264,383,1764,1788]" captionStartId-4="8.[264,383,1791,1815]" captionStartId-5="9.[264,383,1416,1440]" captionStartId-6="11.[264,383,1747,1771]" captionStartId-7="12.[264,383,1771,1795]" captionStartId-8="14.[264,383,1613,1637]" captionStartId-9="15.[264,383,1824,1848]" captionTargetBox-0="[289,1293,450,1752]" captionTargetBox-1="[287,1301,335,1549]" captionTargetBox-10="[324,1266,798,1690]" captionTargetBox-2="[286,1299,499,1546]" captionTargetBox-3="[300,1282,579,1733]" captionTargetBox-4="[281,1306,495,1759]" captionTargetBox-5="[324,1261,853,1386]" captionTargetBox-6="[315,1269,719,1731]" captionTargetBox-7="[288,1290,337,1740]" captionTargetBox-8="[273,1294,321,1581]" captionTargetBox-9="[337,1250,1177,1792]" captionTargetId-0="figure@3.[283,1304,440,1756]" captionTargetId-1="figure@5.[278,1308,329,1556]" captionTargetId-10="figure@16.[317,1270,793,1693]" captionTargetId-2="figure@6.[282,1306,489,1549]" captionTargetId-3="figure@7.[295,1292,569,1737]" captionTargetId-4="figure@8.[272,1315,489,1765]" captionTargetId-5="figure@9.[318,1269,849,1389]" captionTargetId-6="figure@11.[310,1278,713,1736]" captionTargetId-7="figure@12.[279,1307,329,1744]" captionTargetId-8="figure@14.[268,1319,315,1592]" captionTargetId-9="figure@15.[332,1255,1169,1797]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-10="16" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="11" captionTargetPageId-7="12" captionTargetPageId-8="14" captionTargetPageId-9="15" captionText-0="FIGURES 1 ­ 7. Mouthparts of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (paratype from Baribana cave). 1 – epipharynx; 2 – labrum; 3 – left mandible, dorsal view; 4 – right maxilla, ventral view; 5 – prementum; 6 – hypopharynx, left portion not shown; 7 – mentum. Scale bar 0.4 mm (1 ­ 5), 0.2 mm (6 ­ 7)." captionText-1="FIGURES 8 ­ 13. Details of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (8, 10, holotype; 12 ­ 13, paratype from Baribana cave) and H. variabilis Korge (9, Kalkanl Da {lar; 11, holotype). 8 ­ 9 – head; 10 ­ 11 – pronotum; 12 – left antenna; 13 – right metatarsus. Scale bar 1 mm (8 ­ 12), 0.4 mm (13)." captionText-10="FIGURE 48. Geographical distribution of Heinzia Korge. Solid circles – H. caucasica Gusarov &amp; Koval, sp. nov.; open circles – H. variabilis Korge." captionText-2="FIGURES 14 ­ 19. Details of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (14, 16, 18, paratype from Dolgaya cave) and H. variabilis Korge (15, 17, 19, Kalkanl Da {lar). 14 ­ 15 – male sternum 8; 16 ­ 17 – apex of mail sternum 8; 18 ­ 19 – male sternum 9. Scale bar 1 mm (14 ­ 15), 0.5 mm (16 ­ 19)." captionText-3="FIGURES 20 ­ 23. Aedeagus of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (20 ­ 21, paratype from Mezmay) and H. variabilis Korge (22 ­ 23, Kalkanl Da {lar). 20, 22 – ventral view; 21, 23, lateral view. Scale bar 0.4 mm." captionText-4="FIGURES 24 ­ 27. Details of aedeagus of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (24 ­ 25, holotype) and H. variabilis Korge (26 ­ 27, holotype). 24, 26 – median lobe, ventral view; 25, 27, paramere. Scale bar 0.2 mm." captionText-5="FIGURES 28 ­ 32. Apex of median lobe of aedeagus (lateral view) of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (28, paratypes from Mezmay; 29 ­ 30, Baribana cave; 31, Akhty) and H. variabilis Korge (32, Kalkanl Da {lar). Scale bar 0.2 mm." captionText-6="FIGURES 33 ­ 35. Details of female tergum 10 of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (33, 35, Baribana cave) and H. variabilis Korge (34, paratype). 33 ­ 34 – dorsal view; 35 – lateral view; LL – lateral lobe of female tergum 10; MP – medial process of female tergum 10. Scale bar 0.3 mm." captionText-7="FIGURES 36 ­ 38. Details of probable larva of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (Baribana cave). 36 – habitus, dorsal view; 37 – head, dorsal view, 38 – head, lateral view. Scale bar 5 mm (36), 0.8 mm (37 ­ 38)." captionText-8="FIGURES 39 ­ 45. Details of probable larva of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (Baribana cave). 39 – prementum; 40 – left mandible, dorsal view; 41 – left antenna; 42 – left maxilla, ventral view; 43 – protibiotarsus; 44 – apex of protibiotarsus; 45 – nasale. Scale bar 0.2 mm (39 ­ 43, 45), 0.1 mm (44)." captionText-9="FIGURES 46 ­ 47. Details of probable larva of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (Baribana cave). 46 ­ gula; 47 ­ pygopod and urogomphi. Scale bar 1 mm (46), 0.5 mm (47)." httpUri-0="https://zenodo.org/record/155769/files/figure.png" httpUri-1="https://zenodo.org/record/155770/files/figure.png" httpUri-10="https://zenodo.org/record/155780/files/figure.png" httpUri-2="https://zenodo.org/record/155772/files/figure.png" httpUri-3="https://zenodo.org/record/155773/files/figure.png" httpUri-4="https://zenodo.org/record/155774/files/figure.png" httpUri-5="https://zenodo.org/record/155775/files/figure.png" httpUri-6="https://zenodo.org/record/155776/files/figure.png" httpUri-7="https://zenodo.org/record/155777/files/figure.png" httpUri-8="https://zenodo.org/record/155778/files/figure.png" httpUri-9="https://zenodo.org/record/155779/files/figure.png" pageId="1" pageNumber="2">Figs. 1­48</figureCitation>
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<emphasis id="898AEABFE01FB16BC82DFCC3D678093E" box="[264,347,793,816]" italics="true" pageId="1" pageNumber="2">Heinzia</emphasis>
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: 33
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<taxonomicName id="7CFE4D2EE01FB16BC82DFCE6D56D095D" ID-CoL="8WG9F" authority="Smetana, 1977: 181" authorityName="Smetana" authorityPageNumber="181" authorityYear="1977" box="[264,590,828,851]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="898AEABFE01FB16BC82DFCE6D678095D" box="[264,347,828,851]" italics="true" pageId="1" pageNumber="2">Heinzia</emphasis>
: 
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: 181
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. 
<taxonomicName id="7CFE4D2EE01FB16BC82DFC85D5660978" ID-CoL="8WG9F" authority="Coiffait, 1978: 284" authorityName="Coiffait" authorityPageNumber="284" authorityYear="1978" box="[264,581,863,886]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="898AEABFE01FB16BC82DFC85D6780978" box="[264,347,863,886]" italics="true" pageId="1" pageNumber="2">Heinzia</emphasis>
: 
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<bibRefCitation id="DF6F4B5CE01FB16BC855FC85D52A0978" author="Coiffait" box="[368,521,863,886]" pageId="1" pageNumber="18" refString="Coiffait, H. (1978) Coleopteres Staphylinides de la Region Palearctique Occidentale. III. Sous famille Staphylininae, Tribu Quediini. Sous famille Paederinae, Tribu Pinophilini. Supplement a la Nouvelle Revue d'Entomologie, 8 (4), 1 - 364." type="journal article" year="1978">Coiffait, 1978</bibRefCitation>
: 284
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. 
<taxonomicName id="7CFE4D2EE01FB16BC82DFC5BD56D0996" ID-CoL="8WG9F" authority="Smetana, 1995: 131" authorityName="Smetana" authorityPageNumber="131" authorityYear="1995" box="[264,590,897,920]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="898AEABFE01FB16BC82DFC5BD6780996" box="[264,347,897,920]" italics="true" pageId="1" pageNumber="2">Heinzia</emphasis>
: 
<treatmentCitation id="3A5F10BCE01FB16BC855FC5BD56D0996" author="Smetana" box="[368,590,897,920]" page="131" pageId="1" pageNumber="2" year="1995">
<bibRefCitation id="DF6F4B5CE01FB16BC855FC5BD5310996" author="Smetana" box="[368,530,897,920]" pageId="1" pageNumber="19" refString="Smetana, A. (1995) Four new species of Strouhalium from Sichuan and Himalaya (Coleoptera: Staphylinidae: Quediina). Klapalekiana, 31, 131 - 139." type="journal article" year="1995">Smetana, 1995</bibRefCitation>
: 131
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</taxonomicName>
. 
<taxonomicName id="7CFE4D2EE01FB16BC82DFC7ED57509B5" ID-CoL="8WG9F" authority="Herman, 2001: 3054" authorityName="Herman" authorityPageNumber="3054" authorityYear="2001" box="[264,598,932,955]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="898AEABFE01FB16BC82DFC7ED67809B5" box="[264,347,932,955]" italics="true" pageId="1" pageNumber="2">Heinzia</emphasis>
: 
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: 3054
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<paragraph id="BB4136ADE01FB16BC82DFC34D68B0F7F" blockId="1.[264,1324,1006,1872]" pageId="1" pageNumber="2">
<emphasis id="898AEABFE01FB16BC82DFC34D6AB0E06" bold="true" box="[264,392,1006,1032]" pageId="1" pageNumber="2">Diagnosis.</emphasis>
<taxonomicName id="7CFE4D2EE01FB16BC8B0FC35D6D30E06" box="[405,496,1007,1032]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="898AEABFE01FB16BC8B0FC35D6D30E06" box="[405,496,1007,1032]" italics="true" pageId="1" pageNumber="2">Heinzia</emphasis>
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can be distinguished from other genera of the subtribe 
<taxonomicName id="7CFE4D2EE01FB16BCD54FC35D3FE0E07" box="[1137,1245,1007,1033]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="subTribe" subTribe="Quediina">Quediina</taxonomicName>
by the combination of the following characters: head on disc with dense punctation; antennae filiform with all segments longer than wide, first article shorter than second and third combined, penultimate article 1.3 times as long as wide (
<figureCitation id="23C52A28E01FB16BCAADFBBDD4C10E8F" box="[904,994,1127,1153]" captionStart="FIGURES 8 ­ 13" captionStartId="5.[264,383,1583,1607]" captionTargetBox="[287,1301,335,1549]" captionTargetId="figure@5.[278,1308,329,1556]" captionTargetPageId="5" captionText="FIGURES 8 ­ 13. Details of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (8, 10, holotype; 12 ­ 13, paratype from Baribana cave) and H. variabilis Korge (9, Kalkanl Da {lar; 11, holotype). 8 ­ 9 – head; 10 ­ 11 – pronotum; 12 – left antenna; 13 – right metatarsus. Scale bar 1 mm (8 ­ 12), 0.4 mm (13)." httpUri="https://zenodo.org/record/155770/files/figure.png" pageId="1" pageNumber="2">Fig. 12</figureCitation>
); segments of labial palpus glabrous, last segment cylindrical, 1.5 times as long as second (
<figureCitation id="23C52A28E01FB16BCADFFB55D3600EA7" box="[1018,1091,1167,1193]" captionStart="FIGURES 1 ­ 7" captionStartId="3.[264,382,1783,1807]" captionTargetBox="[289,1293,450,1752]" captionTargetId="figure@3.[283,1304,440,1756]" captionTargetPageId="3" captionText="FIGURES 1 ­ 7. Mouthparts of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (paratype from Baribana cave). 1 – epipharynx; 2 – labrum; 3 – left mandible, dorsal view; 4 – right maxilla, ventral view; 5 – prementum; 6 – hypopharynx, left portion not shown; 7 – mentum. Scale bar 0.4 mm (1 ­ 5), 0.2 mm (6 ­ 7)." httpUri="https://zenodo.org/record/155769/files/figure.png" pageId="1" pageNumber="2">Fig. 5</figureCitation>
); pronotum on disc with numerous scattered punctures (
<figureCitation id="23C52A28E01FB16BCB94FB6DD4190EDF" box="[689,826,1207,1233]" captionStart="FIGURES 8 ­ 13" captionStartId="5.[264,383,1583,1607]" captionTargetBox="[287,1301,335,1549]" captionTargetId="figure@5.[278,1308,329,1556]" captionTargetPageId="5" captionText="FIGURES 8 ­ 13. Details of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (8, 10, holotype; 12 ­ 13, paratype from Baribana cave) and H. variabilis Korge (9, Kalkanl Da {lar; 11, holotype). 8 ­ 9 – head; 10 ­ 11 – pronotum; 12 – left antenna; 13 – right metatarsus. Scale bar 1 mm (8 ­ 12), 0.4 mm (13)." httpUri="https://zenodo.org/record/155770/files/figure.png" pageId="1" pageNumber="2">Figs. 10­11</figureCitation>
), with 3 (in some specimens 2 or 4) punctures in dorsal row; prothoracic hypomera inflected and invisible in lateral view; female abdominal tergum 10 is split in two lateral lobes devoid of setae and bears sclerotized and setose medial process protruding posteriorly beyond posterior margin of the lateral lobes (
<figureCitation id="23C52A28E01FB16BC834FA8DD6B80F7F" box="[273,411,1367,1393]" captionStart="FIGURES 33 ­ 35" captionStartId="11.[264,383,1747,1771]" captionTargetBox="[315,1269,719,1731]" captionTargetId="figure@11.[310,1278,713,1736]" captionTargetPageId="11" captionText="FIGURES 33 ­ 35. Details of female tergum 10 of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (33, 35, Baribana cave) and H. variabilis Korge (34, paratype). 33 ­ 34 – dorsal view; 35 – lateral view; LL – lateral lobe of female tergum 10; MP – medial process of female tergum 10. Scale bar 0.3 mm." httpUri="https://zenodo.org/record/155776/files/figure.png" pageId="1" pageNumber="2">Figs. 33­35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F3E46526E01FB16BC81DFAA5D4EA0C36" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="BB4136ADE01FB16BC81DFAA5D5CC0FCF" blockId="1.[264,1324,1006,1872]" pageId="1" pageNumber="2">
<taxonomicName id="7CFE4D2EE01FB16BC81DFAA5D6B00F96" ID-CoL="8WG9F" box="[312,403,1407,1432]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="898AEABFE01FB16BC81DFAA5D6B00F96" box="[312,403,1407,1432]" italics="true" pageId="1" pageNumber="2">Heinzia</emphasis>
</taxonomicName>
is most closely related to 
<taxonomicName id="7CFE4D2EE01FB16BCBF2FAA5D3650F97" ID-CoL="8W3VQ" authority="Scheerpeltz, 1962" authorityName="Scheerpeltz" authorityYear="1962" box="[727,1094,1407,1433]" class="Insecta" family="Staphylinidae" genus="Strouhalium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="898AEABFE01FB16BCBF2FAA5D4440F96" box="[727,871,1407,1432]" italics="true" pageId="1" pageNumber="2">Strouhalium</emphasis>
<bibRefCitation id="DF6F4B5CE01FB16BCA54FAA5D3650F97" author="Scheerpeltz" box="[881,1094,1407,1433]" pageId="1" pageNumber="19" refString="Scheerpeltz, O. (1962) Eine neue Gattung und Art der Tribus Quediini (Col. Staphylinidae). Annalen des Naturhistorischen Museums in Wien, 65, 259 - 272." type="journal article" year="1962">Scheerpeltz, 1962</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="7CFE4D2EE01FB16BCDA0FAA5D6600FCF" authority="Hatch, 1957" authorityName="Hatch" authorityYear="1957" class="Insecta" family="Lymantriidae" genus="Beeria" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="898AEABFE01FB16BCDA0FAA5D3F00F96" box="[1157,1235,1407,1432]" italics="true" pageId="1" pageNumber="2">Beeria</emphasis>
<bibRefCitation id="DF6F4B5CE01FB16BCDF8FAA5D6600FCF" author="Hatch" pageId="1" pageNumber="18" refString="Hatch, M. H. (1957) The beetles of the Pacific Northwest. Part II: Staphyliniformia. Seattle: University of Washington Press, ix + 384 pp." type="book" year="1957">Hatch, 1957</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="DF6F4B5CE01FB16BC877FA7DD6C10FCF" author="Korge" box="[338,482,1447,1473]" pageId="1" pageNumber="19" refString="Korge, H. (1971) Beitrage zur Kenntnis der Koleopterenfauna Kleinasiens. Annotationes Zoologicae et Botanicae, 67, 1 - 68." type="journal article" year="1971">Korge 1971</bibRefCitation>
; 
<bibRefCitation id="DF6F4B5CE01FB16BC8CBFA7DD5B40FCF" author="Smetana" box="[494,663,1447,1473]" pageId="1" pageNumber="19" refString="Smetana, A. (1977) The Nearctic genus Beeria Hatch. Taxonomy, distribution and ecology (Coleoptera: Staphylinidae). Entomologica Scandinavica, 8, 177 - 190." type="journal article" year="1977">Smetana 1977</bibRefCitation>
, 
<bibRefCitation id="DF6F4B5CE01FB16BCB81FA7DD5C00FCF" author="Smetana" box="[676,739,1447,1473]" pageId="1" pageNumber="19" refString="Smetana, A. (1995) Four new species of Strouhalium from Sichuan and Himalaya (Coleoptera: Staphylinidae: Quediina). Klapalekiana, 31, 131 - 139." type="journal article" year="1995">1995</bibRefCitation>
).
</paragraph>
<paragraph id="BB4136ADE01FB16BC81DFA15D4EA0C36" blockId="1.[264,1324,1006,1872]" pageId="1" pageNumber="2">
<taxonomicName id="7CFE4D2EE01FB16BC81DFA15D6B00FE6" ID-CoL="8WG9F" box="[312,403,1487,1512]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="898AEABFE01FB16BC81DFA15D6B00FE6" box="[312,403,1487,1512]" italics="true" pageId="1" pageNumber="2">Heinzia</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="7CFE4D2EE01FB16BCB17FA15D5A30FE6" box="[562,640,1487,1512]" class="Insecta" family="Lymantriidae" genus="Beeria" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="898AEABFE01FB16BCB17FA15D5A30FE6" box="[562,640,1487,1512]" italics="true" pageId="1" pageNumber="2">Beeria</emphasis>
</taxonomicName>
in having prothoracic hypomera invisible in lateral view, numerous punctures in lateral portions of pronotum, protarsi equally dilated in both sexes, entire paramere, and modified female abdominal tergum 10.
</paragraph>
</subSubSection>
<subSubSection id="F3E46526E01FB168C81DF99DD6980F60" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="description">
<paragraph id="BB4136ADE01FB16BC81DF99DD4AB0C86" blockId="1.[264,1324,1006,1872]" pageId="1" pageNumber="2">
<taxonomicName id="7CFE4D2EE01FB16BC81DF99DD6B00C6E" box="[312,403,1607,1632]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="898AEABFE01FB16BC81DF99DD6B00C6E" box="[312,403,1607,1632]" italics="true" pageId="1" pageNumber="2">Heinzia</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="7CFE4D2EE01FB16BCB14F99DD5E20C6E" box="[561,705,1607,1632]" class="Insecta" family="Staphylinidae" genus="Strouhalium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="898AEABFE01FB16BCB14F99DD5E20C6E" box="[561,705,1607,1632]" italics="true" pageId="1" pageNumber="2">Strouhalium</emphasis>
</taxonomicName>
in having glabrous segments of labial palpus (
<bibRefCitation id="DF6F4B5CE01FB16BCDCEF99CD65F0C86" author="Smetana" pageId="1" pageNumber="19" refString="Smetana, A. (1995) Four new species of Strouhalium from Sichuan and Himalaya (Coleoptera: Staphylinidae: Quediina). Klapalekiana, 31, 131 - 139." type="journal article" year="1995">Smetana 1995</bibRefCitation>
) and modified female abdominal tergum 10.
</paragraph>
<paragraph id="BB4136ADE01FB16BC81DF94CD54B0D0E" blockId="1.[264,1324,1006,1872]" pageId="1" pageNumber="2">
<emphasis id="898AEABFE01FB16BC81DF94CD6F30CBE" bold="true" box="[312,464,1686,1712]" pageId="1" pageNumber="2">Description.</emphasis>
Length 
<quantity id="7C069B48E01FB16BCB1BF94CD5F90CBE" box="[574,730,1686,1712]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.15" metricValueMin="0.85" pageId="1" pageNumber="2" unit="mm" value="10.0" valueMax="11.5" valueMin="8.5">8.5­11.5 mm</quantity>
. Body from brownish black to black, legs brown with lighter tarsi and black inner surface of tibia, mouthparts and bases of antennal segments brown to brownish red.
</paragraph>
<paragraph id="BB4136ADE01FB168C81DF8D4D60408C8" blockId="1.[264,1324,1006,1872]" lastBlockId="2.[264,1324,284,1430]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
Head as long as wide, on disc with dense punctation except in anterior impunctate portion (
<figureCitation id="23C52A28E01FB16BC862F8ECD6910D5E" box="[327,434,1846,1872]" captionStart="FIGURES 8 ­ 13" captionStartId="5.[264,383,1583,1607]" captionTargetBox="[287,1301,335,1549]" captionTargetId="figure@5.[278,1308,329,1556]" captionTargetPageId="5" captionText="FIGURES 8 ­ 13. Details of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (8, 10, holotype; 12 ­ 13, paratype from Baribana cave) and H. variabilis Korge (9, Kalkanl Da {lar; 11, holotype). 8 ­ 9 – head; 10 ­ 11 – pronotum; 12 – left antenna; 13 – right metatarsus. Scale bar 1 mm (8 ­ 12), 0.4 mm (13)." httpUri="https://zenodo.org/record/155770/files/figure.png" pageId="1" pageNumber="2">Figs. 8­9</figureCitation>
), eyes slightly shorter than temples; infraorbital ridge absent; gular sutures separate. All antennal articles longer than wide, first article shorter than second and third combined, article 10 ­ 1.3 times as long as wide, last article 1.7­1.8 times as long as wide (
<figureCitation id="23C52A28E01CB168C834FEB6D64A0B88" box="[273,361,364,390]" captionStart="FIGURES 8 ­ 13" captionStartId="5.[264,383,1583,1607]" captionTargetBox="[287,1301,335,1549]" captionTargetId="figure@5.[278,1308,329,1556]" captionTargetPageId="5" captionText="FIGURES 8 ­ 13. Details of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (8, 10, holotype; 12 ­ 13, paratype from Baribana cave) and H. variabilis Korge (9, Kalkanl Da {lar; 11, holotype). 8 ­ 9 – head; 10 ­ 11 – pronotum; 12 – left antenna; 13 – right metatarsus. Scale bar 1 mm (8 ­ 12), 0.4 mm (13)." httpUri="https://zenodo.org/record/155770/files/figure.png" pageId="2" pageNumber="3">Fig. 12</figureCitation>
). Labrum strongly transverse, with deep medial emargination and strong setae at anterior margin (
<figureCitation id="23C52A28E01CB168C8E9FE4ED5370BA0" box="[460,532,404,430]" captionStart="FIGURES 1 ­ 7" captionStartId="3.[264,382,1783,1807]" captionTargetBox="[289,1293,450,1752]" captionTargetId="figure@3.[283,1304,440,1756]" captionTargetPageId="3" captionText="FIGURES 1 ­ 7. Mouthparts of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (paratype from Baribana cave). 1 – epipharynx; 2 – labrum; 3 – left mandible, dorsal view; 4 – right maxilla, ventral view; 5 – prementum; 6 – hypopharynx, left portion not shown; 7 – mentum. Scale bar 0.4 mm (1 ­ 5), 0.2 mm (6 ­ 7)." httpUri="https://zenodo.org/record/155769/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
). Adoral surface of labrum (epipharynx) densely covered with very fine setae, at anterior margin with dense row of long setae (
<figureCitation id="23C52A28E01CB168CAC7FE66D30A0BD8" box="[994,1065,444,470]" captionStart="FIGURES 1 ­ 7" captionStartId="3.[264,382,1783,1807]" captionTargetBox="[289,1293,450,1752]" captionTargetId="figure@3.[283,1304,440,1756]" captionTargetPageId="3" captionText="FIGURES 1 ­ 7. Mouthparts of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (paratype from Baribana cave). 1 – epipharynx; 2 – labrum; 3 – left mandible, dorsal view; 4 – right maxilla, ventral view; 5 – prementum; 6 – hypopharynx, left portion not shown; 7 – mentum. Scale bar 0.4 mm (1 ­ 5), 0.2 mm (6 ­ 7)." httpUri="https://zenodo.org/record/155769/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
). Mandibles slender with double tooth at inner margin (
<figureCitation id="23C52A28E01CB168CB82FE3ED5CC0BF0" box="[679,751,484,510]" captionStart="FIGURES 1 ­ 7" captionStartId="3.[264,382,1783,1807]" captionTargetBox="[289,1293,450,1752]" captionTargetId="figure@3.[283,1304,440,1756]" captionTargetPageId="3" captionText="FIGURES 1 ­ 7. Mouthparts of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (paratype from Baribana cave). 1 – epipharynx; 2 – labrum; 3 – left mandible, dorsal view; 4 – right maxilla, ventral view; 5 – prementum; 6 – hypopharynx, left portion not shown; 7 – mentum. Scale bar 0.4 mm (1 ­ 5), 0.2 mm (6 ­ 7)." httpUri="https://zenodo.org/record/155769/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
). Fourth segment of maxillary palpus fusiform, 5.5 times as long as wide, and 1.8 times as long as third (
<figureCitation id="23C52A28E01CB168CA80FDD6D4CD0828" box="[933,1006,524,550]" captionStart="FIGURES 1 ­ 7" captionStartId="3.[264,382,1783,1807]" captionTargetBox="[289,1293,450,1752]" captionTargetId="figure@3.[283,1304,440,1756]" captionTargetPageId="3" captionText="FIGURES 1 ­ 7. Mouthparts of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (paratype from Baribana cave). 1 – epipharynx; 2 – labrum; 3 – left mandible, dorsal view; 4 – right maxilla, ventral view; 5 – prementum; 6 – hypopharynx, left portion not shown; 7 – mentum. Scale bar 0.4 mm (1 ­ 5), 0.2 mm (6 ­ 7)." httpUri="https://zenodo.org/record/155769/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
). Ligula with apical emargination, labial palpus glabrous, third segment 1.5 times as long as second (
<figureCitation id="23C52A28E01CB168CD50FDEED39F0840" box="[1141,1212,564,590]" captionStart="FIGURES 1 ­ 7" captionStartId="3.[264,382,1783,1807]" captionTargetBox="[289,1293,450,1752]" captionTargetId="figure@3.[283,1304,440,1756]" captionTargetPageId="3" captionText="FIGURES 1 ­ 7. Mouthparts of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (paratype from Baribana cave). 1 – epipharynx; 2 – labrum; 3 – left mandible, dorsal view; 4 – right maxilla, ventral view; 5 – prementum; 6 – hypopharynx, left portion not shown; 7 – mentum. Scale bar 0.4 mm (1 ­ 5), 0.2 mm (6 ­ 7)." httpUri="https://zenodo.org/record/155769/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
). Lateral area of prementum with three pores and single spinose pore (
<figureCitation id="23C52A28E01CB168CAD2FD86D3600878" box="[1015,1091,604,630]" captionStart="FIGURES 1 ­ 7" captionStartId="3.[264,382,1783,1807]" captionTargetBox="[289,1293,450,1752]" captionTargetId="figure@3.[283,1304,440,1756]" captionTargetPageId="3" captionText="FIGURES 1 ­ 7. Mouthparts of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (paratype from Baribana cave). 1 – epipharynx; 2 – labrum; 3 – left mandible, dorsal view; 4 – right maxilla, ventral view; 5 – prementum; 6 – hypopharynx, left portion not shown; 7 – mentum. Scale bar 0.4 mm (1 ­ 5), 0.2 mm (6 ­ 7)." httpUri="https://zenodo.org/record/155769/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
). Hypopharyngeal lobes as in 
<figureCitation id="23C52A28E01CB168C8ACFD5ED6F30890" box="[393,464,644,670]" captionStart="FIGURES 1 ­ 7" captionStartId="3.[264,382,1783,1807]" captionTargetBox="[289,1293,450,1752]" captionTargetId="figure@3.[283,1304,440,1756]" captionTargetPageId="3" captionText="FIGURES 1 ­ 7. Mouthparts of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (paratype from Baribana cave). 1 – epipharynx; 2 – labrum; 3 – left mandible, dorsal view; 4 – right maxilla, ventral view; 5 – prementum; 6 – hypopharynx, left portion not shown; 7 – mentum. Scale bar 0.4 mm (1 ­ 5), 0.2 mm (6 ­ 7)." httpUri="https://zenodo.org/record/155769/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
. Mentum strongly transverse, with slightly concave anterior margin (
<figureCitation id="23C52A28E01CB168CDD9FD5ED63A08C8" captionStart="FIGURES 1 ­ 7" captionStartId="3.[264,382,1783,1807]" captionTargetBox="[289,1293,450,1752]" captionTargetId="figure@3.[283,1304,440,1756]" captionTargetPageId="3" captionText="FIGURES 1 ­ 7. Mouthparts of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (paratype from Baribana cave). 1 – epipharynx; 2 – labrum; 3 – left mandible, dorsal view; 4 – right maxilla, ventral view; 5 – prementum; 6 – hypopharynx, left portion not shown; 7 – mentum. Scale bar 0.4 mm (1 ­ 5), 0.2 mm (6 ­ 7)." httpUri="https://zenodo.org/record/155769/files/figure.png" pageId="2" pageNumber="3">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="BB4136ADE01CB168C81DFD0ED5390E08" blockId="2.[264,1324,284,1430]" pageId="2" pageNumber="3">
Pronotum on disc with dense scattered punctation (
<figureCitation id="23C52A28E01CB168CABEFD0ED30508E0" box="[923,1062,724,750]" captionStart="FIGURES 8 ­ 13" captionStartId="5.[264,383,1583,1607]" captionTargetBox="[287,1301,335,1549]" captionTargetId="figure@5.[278,1308,329,1556]" captionTargetPageId="5" captionText="FIGURES 8 ­ 13. Details of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (8, 10, holotype; 12 ­ 13, paratype from Baribana cave) and H. variabilis Korge (9, Kalkanl Da {lar; 11, holotype). 8 ­ 9 – head; 10 ­ 11 – pronotum; 12 – left antenna; 13 – right metatarsus. Scale bar 1 mm (8 ­ 12), 0.4 mm (13)." httpUri="https://zenodo.org/record/155770/files/figure.png" pageId="2" pageNumber="3">Figs. 10­11</figureCitation>
). Dorsal rows with 3 (in some specimens 2 or 4) punctures, hard to distinguish from scattered punctures. Prothoracic hypomera inflected, invisible in lateral view. Proepisterna large. Mesosternal process sharp, 0.4­0.5 times as long as mesocoxal cavity. Tarsi shorter than tibiae. All tibiae spinose. Tarsal formula 5­5­5, tarsal segments setose dorsally. Protarsal segments 1­ 4 dilated equally in both sexes. First segment of metatarsus as long as second and third combined, last segment much longer than first (
<figureCitation id="23C52A28E01CB168CA16FC1ED4AA09D0" box="[819,905,964,990]" captionStart="FIGURES 8 ­ 13" captionStartId="5.[264,383,1583,1607]" captionTargetBox="[287,1301,335,1549]" captionTargetId="figure@5.[278,1308,329,1556]" captionTargetPageId="5" captionText="FIGURES 8 ­ 13. Details of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (8, 10, holotype; 12 ­ 13, paratype from Baribana cave) and H. variabilis Korge (9, Kalkanl Da {lar; 11, holotype). 8 ­ 9 – head; 10 ­ 11 – pronotum; 12 – left antenna; 13 – right metatarsus. Scale bar 1 mm (8 ­ 12), 0.4 mm (13)." httpUri="https://zenodo.org/record/155770/files/figure.png" pageId="2" pageNumber="3">Fig. 13</figureCitation>
). Two long empodial setae present. Wings fully developed.
</paragraph>
<paragraph id="BB4136ADE01CB168C81DFBCED4F70EC0" blockId="2.[264,1324,284,1430]" pageId="2" pageNumber="3">
Abdominal tergum 7 with wide white palisade fringe. Male sternum 8 with medial emargination (
<figureCitation id="23C52A28E01CB168C897FBE6D51F0E58" box="[434,572,1084,1110]" captionStart="FIGURES 14 ­ 19" captionStartId="6.[264,383,1576,1600]" captionTargetBox="[286,1299,499,1546]" captionTargetId="figure@6.[282,1306,489,1549]" captionTargetPageId="6" captionText="FIGURES 14 ­ 19. Details of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (14, 16, 18, paratype from Dolgaya cave) and H. variabilis Korge (15, 17, 19, Kalkanl Da {lar). 14 ­ 15 – male sternum 8; 16 ­ 17 – apex of mail sternum 8; 18 ­ 19 – male sternum 9. Scale bar 1 mm (14 ­ 15), 0.5 mm (16 ­ 19)." httpUri="https://zenodo.org/record/155772/files/figure.png" pageId="2" pageNumber="3">Figs. 14­17</figureCitation>
). Female tergum 10 split into two broad lateral lobes (LL; 
<figureCitation id="23C52A28E01CB168CDD4FBE6D67B0E70" captionStart="FIGURES 33 ­ 35" captionStartId="11.[264,383,1747,1771]" captionTargetBox="[315,1269,719,1731]" captionTargetId="figure@11.[310,1278,713,1736]" captionTargetPageId="11" captionText="FIGURES 33 ­ 35. Details of female tergum 10 of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (33, 35, Baribana cave) and H. variabilis Korge (34, paratype). 33 ­ 34 – dorsal view; 35 – lateral view; LL – lateral lobe of female tergum 10; MP – medial process of female tergum 10. Scale bar 0.3 mm." httpUri="https://zenodo.org/record/155776/files/figure.png" pageId="2" pageNumber="3">Figs. 33, 35</figureCitation>
), the lobes poorly sclerotized and devoid of setae. Long sclerotized and setose medial process (MP; 
<figureCitation id="23C52A28E01CB168CB2CFB56D5BA0EA8" box="[521,665,1164,1190]" captionStart="FIGURES 33 ­ 35" captionStartId="11.[264,383,1747,1771]" captionTargetBox="[315,1269,719,1731]" captionTargetId="figure@11.[310,1278,713,1736]" captionTargetPageId="11" captionText="FIGURES 33 ­ 35. Details of female tergum 10 of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (33, 35, Baribana cave) and H. variabilis Korge (34, paratype). 33 ­ 34 – dorsal view; 35 – lateral view; LL – lateral lobe of female tergum 10; MP – medial process of female tergum 10. Scale bar 0.3 mm." httpUri="https://zenodo.org/record/155776/files/figure.png" pageId="2" pageNumber="3">Figs. 33, 35</figureCitation>
) attached at basis of tergum where the lobes meet and extended posteriad beyond margin of the lobes (
<figureCitation id="23C52A28E01CB168CA19FB6ED4E40EC0" box="[828,967,1204,1230]" captionStart="FIGURES 33 ­ 35" captionStartId="11.[264,383,1747,1771]" captionTargetBox="[315,1269,719,1731]" captionTargetId="figure@11.[310,1278,713,1736]" captionTargetPageId="11" captionText="FIGURES 33 ­ 35. Details of female tergum 10 of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (33, 35, Baribana cave) and H. variabilis Korge (34, paratype). 33 ­ 34 – dorsal view; 35 – lateral view; LL – lateral lobe of female tergum 10; MP – medial process of female tergum 10. Scale bar 0.3 mm." httpUri="https://zenodo.org/record/155776/files/figure.png" pageId="2" pageNumber="3">Figs. 33­35</figureCitation>
).
</paragraph>
<paragraph id="BB4136ADE01CB168C81DFB06D6980F60" blockId="2.[264,1324,284,1430]" pageId="2" pageNumber="3">
Median lobe of aedeagus straight (in lateral view) (
<figureCitation id="23C52A28E01CB168CAAAFB06D33D0EF8" box="[911,1054,1244,1270]" captionStart="FIGURES 20 ­ 23" captionStartId="7.[264,383,1764,1788]" captionTargetBox="[300,1282,579,1733]" captionTargetId="figure@7.[295,1292,569,1737]" captionTargetPageId="7" captionText="FIGURES 20 ­ 23. Aedeagus of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (20 ­ 21, paratype from Mezmay) and H. variabilis Korge (22 ­ 23, Kalkanl Da {lar). 20, 22 – ventral view; 21, 23, lateral view. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/155773/files/figure.png" pageId="2" pageNumber="3">Figs. 21, 23</figureCitation>
), with single subapical tooth on ventral surface (
<figureCitation id="23C52A28E01CB168CB0AFADED5E30F10" box="[559,704,1284,1310]" captionStart="FIGURES 24 ­ 27" captionStartId="8.[264,383,1791,1815]" captionTargetBox="[281,1306,495,1759]" captionTargetId="figure@8.[272,1315,489,1765]" captionTargetPageId="8" captionText="FIGURES 24 ­ 27. Details of aedeagus of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (24 ­ 25, holotype) and H. variabilis Korge (26 ­ 27, holotype). 24, 26 – median lobe, ventral view; 25, 27, paramere. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155774/files/figure.png" pageId="2" pageNumber="3">Figs. 24, 26</figureCitation>
). Paramere with 15­20 peg­like setae (
<figureCitation id="23C52A28E01CB168CDA8FADED23D0F10" box="[1165,1310,1284,1310]" captionStart="FIGURES 24 ­ 27" captionStartId="8.[264,383,1791,1815]" captionTargetBox="[281,1306,495,1759]" captionTargetId="figure@8.[272,1315,489,1765]" captionTargetPageId="8" captionText="FIGURES 24 ­ 27. Details of aedeagus of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (24 ­ 25, holotype) and H. variabilis Korge (26 ­ 27, holotype). 24, 26 – median lobe, ventral view; 25, 27, paramere. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155774/files/figure.png" pageId="2" pageNumber="3">Figs. 25, 27</figureCitation>
). Internal sac with single basal diverticulum (
<figureCitation id="23C52A28E01CB168CA2EFAF6D4B80F48" box="[779,923,1324,1350]" captionStart="FIGURES 20 ­ 23" captionStartId="7.[264,383,1764,1788]" captionTargetBox="[300,1282,579,1733]" captionTargetId="figure@7.[295,1292,569,1737]" captionTargetPageId="7" captionText="FIGURES 20 ­ 23. Aedeagus of Heinzia caucasica Gusarov &amp; Koval, sp. nov. (20 ­ 21, paratype from Mezmay) and H. variabilis Korge (22 ­ 23, Kalkanl Da {lar). 20, 22 – ventral view; 21, 23, lateral view. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/155773/files/figure.png" pageId="2" pageNumber="3">Figs. 21, 23</figureCitation>
). Aedeagus at rest with basal orifice facing left.
</paragraph>
</subSubSection>
<subSubSection id="F3E46526E01CB168C81DFAA1D3590F98" box="[312,1146,1403,1430]" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="BB4136ADE01CB168C81DFAA1D3590F98" blockId="2.[264,1324,284,1430]" box="[312,1146,1403,1430]" pageId="2" pageNumber="3">
<emphasis id="898AEABFE01CB168C81DFAA1D6F90F9B" bold="true" box="[312,474,1403,1429]" pageId="2" pageNumber="3">
<typeStatus id="6445880FE01CB168C81DFAA1D6560F9B" box="[312,373,1403,1429]" pageId="2" pageNumber="3">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="7CFE4D2EE01CB168C8CDFAA6D4740F98" authority="Korge, 1971" authorityName="Korge" authorityYear="1971" box="[488,855,1404,1430]" class="Insecta" family="Staphylinidae" genus="Heinzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="898AEABFE01CB168C8CDFAA6D59F0F9B" box="[488,700,1404,1429]" italics="true" pageId="2" pageNumber="3">Heinzia variabilis</emphasis>
<bibRefCitation id="DF6F4B5CE01CB168CBE6FAA6D4740F98" author="Korge" box="[707,855,1404,1430]" pageId="2" pageNumber="19" refString="Korge, H. (1971) Beitrage zur Kenntnis der Koleopterenfauna Kleinasiens. Annotationes Zoologicae et Botanicae, 67, 1 - 68." type="journal article" year="1971">Korge, 1971</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
</treatment>
</document>