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<document id="C1E17BC1308629ADF028D211158920E5" ID-CLB-Dataset="21808" ID-DOI="10.5852/ejt.2014.100" ID-GBIF-Dataset="5329bfc4-61b2-48f5-a96a-caf4ba31039b" ID-ISSN="2118-9773" ID-Zenodo-Dep="3860450" ID-ZooBank="B3E6C489-6D96-4AF5-A33D-EE8329A9321B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1590595554426" checkinUser="carolina" docAuthor="Enghoff, Henrik" docDate="2014" docId="334F2769C146FFA51CD6F346FE4EFC89" docLanguage="en" docName="ejt-2014-100.pdf.imf" docOrigin="European Journal of Taxonomy 100" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Chaleponcus ibis Enghoff 2014, sp. nov." docType="treatment" docUuid="E6764255-8F2F-4B2D-8F3C-26F40FA01EB0" docUuidSource="ZooBank" docVersion="7" lastPageNumber="46" masterDocId="CF765F11C16DFF881E57F137FFA0FFA3" masterDocTitle="A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="75" masterPageNumber="1" pageNumber="44" updateTime="1698841637397" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="66ED89828AB503A622801DDC6250FDF8">A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<mods:namePart id="A5CAF563B358E30C90D5BAD232B51A3C">Enghoff, Henrik</mods:namePart>
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<mods:title id="8FD217F86C40DB2C32651B97F33B26CC">European Journal of Taxonomy</mods:title>
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<mods:date id="AA6FA00CC998A62F9A8F71443BF9D8BC">2014</mods:date>
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<mods:number id="75D4928927BFBBC1019A55F9C0DC158D">2014-10-24</mods:number>
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<treatment id="334F2769C146FFA51CD6F346FE4EFC89" ID-DOI="http://doi.org/10.5281/zenodo.3861215" ID-GBIF-Taxon="164362784" ID-Zenodo-Dep="3861215" LSID="urn:lsid:zoobank.org:act:E6764255-8F2F-4B2D-8F3C-26F40FA01EB0" httpUri="http://treatment.plazi.org/id/334F2769C146FFA51CD6F346FE4EFC89" lastPageId="45" lastPageNumber="46" pageId="43" pageNumber="44">
<subSubSection id="F3FCC5F4C146FFA31CD6F346FC13FD2F" box="[641,947,625,652]" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="BB59967FC146FFA31CD6F346FC13FD2F" blockId="43.[373,1215,625,722]" box="[641,947,625,652]" pageId="43" pageNumber="44">
<heading id="E0112113C146FFA31CD6F346FC13FD2F" box="[641,947,625,652]" centered="true" fontSize="11" level="2" pageId="43" pageNumber="44" reason="2">
<taxonomicName id="7CE6EDFCC146FFA31CD6F346FCF1FD2F" ID-CoL="TH6P" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[641,849,625,652]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="ibis" status="sp. nov.">
<emphasis id="89924A6DC146FFA31CD6F346FCF1FD2F" bold="true" box="[641,849,625,652]" italics="true" pageId="43" pageNumber="44">Chaleponcus ibis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C146FFA31D0FF345FC13FD2F" box="[856,947,626,652]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C146FFA31F22F3A3FCF4FD71" pageId="43" pageNumber="44" type="description">
<paragraph id="BB59967FC146FFA31F22F3A3FB1FFD0C" blockId="43.[373,1215,625,722]" box="[373,1215,660,687]" pageId="43" pageNumber="44">
<uri id="CF779A7DC146FFA31F22F3A3FB1FFD0C" box="[373,1215,660,687]" pageId="43" pageNumber="44">
urn:lsid:zoobank.org:act:
<uuid id="CF40ACAAC146FFA31CC1F3A3FB1FFD0C" box="[662,1215,660,687]" pageId="43" pageNumber="44">E6764255-8F2F-4B2D-8F3C-26F40FA01EB0</uuid>
</uri>
</paragraph>
<paragraph id="BB59967FC146FFA31C88F38FFCF4FD71" blockId="43.[373,1215,625,722]" box="[735,852,696,722]" pageId="43" pageNumber="44">
<figureCitation id="23DD8AFAC146FFA31C88F38FFC8AFD71" box="[735,810,696,722]" captionStart="Fig" captionStartId="15.[187,230,1816,1842]" captionTargetBox="[247,1356,507,1717]" captionTargetId="figure@15.[200,1385,488,1723]" captionTargetPageId="15" captionText="Fig. 9. Chaleponcus spp., outline of coxite of left gonopod, posterior view. (The transverse plane of the gonopod is usually slightly at angles with the exact transverse plane of the millipede, so it may be necessary to tilt the gonopod slightly in order to obtain the view shown on the drawings.) The stippled lines indicate the place where the telopodital basomere emerges from the coxal cavity. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3860472" httpUri="https://zenodo.org/record/3860472/files/figure.png" pageId="43" pageNumber="44">Figs 9</figureCitation>
,
<figureCitation id="23DD8AFAC146FFA31D60F38FFCF4FD71" box="[823,852,696,722]" captionStart="Fig" captionStartId="44.[189,232,1893,1919]" captionTargetBox="[190,1398,501,1853]" captionTargetId="figure@44.[189,1398,501,1853]" captionTargetPageId="44" captionText="Fig. 22. Chaleponcus ibis sp. nov., left gonopod. A. Posterior view. B. Anterior view. C. Telopodite, mesal-anterior view. al = anterior lobe of telomere, al = accessory lamella?, cu = cucullus, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior lobe of telomere, ps = proximal spine of solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860498" httpUri="https://zenodo.org/record/3860498/files/figure.png" pageId="43" pageNumber="44">22</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C146FFA31EEAF3CEFACFFC2A" pageId="43" pageNumber="44" type="diagnosis">
<paragraph id="BB59967FC146FFA31EEAF3CEFE96FCB0" blockId="43.[189,310,761,787]" box="[189,310,761,787]" pageId="43" pageNumber="44">
<heading id="E0112113C146FFA31EEAF3CEFE96FCB0" bold="true" box="[189,310,761,787]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="89924A6DC146FFA31EEAF3CEFE96FCB0" bold="true" box="[189,310,761,787]" pageId="43" pageNumber="44">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC146FFA31EEAF21FFACFFC2A" blockId="43.[189,1398,808,905]" pageId="43" pageNumber="44">
Large. Gonopod coxa (
<figureCitation id="23DD8AFAC146FFA31F80F21FFD81FCE1" box="[471,545,808,834]" captionStart="Fig" captionStartId="15.[187,230,1816,1842]" captionTargetBox="[247,1356,507,1717]" captionTargetId="figure@15.[200,1385,488,1723]" captionTargetPageId="15" captionText="Fig. 9. Chaleponcus spp., outline of coxite of left gonopod, posterior view. (The transverse plane of the gonopod is usually slightly at angles with the exact transverse plane of the millipede, so it may be necessary to tilt the gonopod slightly in order to obtain the view shown on the drawings.) The stippled lines indicate the place where the telopodital basomere emerges from the coxal cavity. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3860472" httpUri="https://zenodo.org/record/3860472/files/figure.png" pageId="43" pageNumber="44">Fig. 9</figureCitation>
) without a lateral process; metaplical shelf-spine very long. Telomere with a rugged anterior lamella and posterior lamella ending in short, parallel, elongate lobes. Claws of walking legs surrounded by palisade of setae, a character shared only with
<taxonomicName id="7CE6EDFCC146FFA31A6AF258FAADFC2B" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1085,1293,878,905]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="circumvallatus" status="sp. nov.">
<emphasis id="89924A6DC146FFA31A6AF258FAADFC2B" box="[1085,1293,878,905]" italics="true" pageId="43" pageNumber="44">C. circumvallatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C146FFA31B43F258FACFFC2A" box="[1300,1391,879,905]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C146FFA31EEAF287FA8DFC5B" box="[189,1325,944,1017]" pageId="43" pageNumber="44" type="etymology">
<paragraph id="BB59967FC146FFA31EEAF287FEE4FC69" blockId="43.[189,324,944,970]" box="[189,324,944,970]" pageId="43" pageNumber="44">
<heading id="E0112113C146FFA31EEAF287FEE4FC69" bold="true" box="[189,324,944,970]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="89924A6DC146FFA31EEAF287FEE4FC69" bold="true" box="[189,324,944,970]" pageId="43" pageNumber="44">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC146FFA31EEAF2E9FA8DFC5B" blockId="43.[189,1325,990,1017]" box="[189,1325,990,1017]" pageId="43" pageNumber="44">The name is a noun in apposition and refers to the somewhat ibis bill-like metaplical shelf-spine.</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C146FFA31EEAF528FC20FAE1" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="BB59967FC146FFA31EEAF528FDB7FB99" blockId="43.[189,535,1055,1082]" box="[189,535,1055,1082]" pageId="43" pageNumber="44">
<emphasis id="89924A6DC146FFA31EEAF528FE2DFB9A" bold="true" box="[189,397,1055,1081]" pageId="43" pageNumber="44">Material studied</emphasis>
(total:
<specimenCount id="ADE05DF6C146FFA31FB5F517FDAFFB99" box="[482,527,1056,1082]" pageId="43" pageNumber="44" type="male">1 ♂</specimenCount>
)
</paragraph>
<materialsCitation id="0B8E9C22C146FFA31EBDF579FAC4FB0C" ID-GBIF-Occurrence="2626747302" collectingDate="1994-07" collectingDateMax="1994-11-31" collectingDateMin="1994-07" collectionCode="ZMUC" collectorName="D. Moyer" country="Tanzania" county="Udzungwa Mts" latitude="-7.713611" location="Udekwa village" longLatPrecision="21" longitude="36.420834" municipality="West Kilombero" pageId="43" pageNumber="44" specimenCount="1" specimenCount-male="1" stateProvince="Iringa Region" typeStatus="holotype">
<paragraph id="BB59967FC146FFA31EBDF579FEFCFBCB" blockId="43.[189,1398,1102,1199]" box="[234,348,1102,1128]" pageId="43" pageNumber="44">
<emphasis id="89924A6DC146FFA31EBDF579FEFCFBCB" bold="true" box="[234,348,1102,1128]" pageId="43" pageNumber="44">
<typeStatus id="645D28DDC146FFA31EBDF579FEFCFBCB" box="[234,348,1102,1128]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="BB59967FC146FFA31EEAF546FAC4FB0C" blockId="43.[189,1398,1102,1199]" pageId="43" pageNumber="44">
<collectingCountry id="C3F1D6EFC146FFA31EEAF546FEF7FB28" box="[189,343,1137,1163]" name="Tanzania" pageId="43" pageNumber="44">TANZANIA</collectingCountry>
:
<specimenCount id="ADE05DF6C146FFA31F34F545FEDBFB2F" box="[355,379,1138,1164]" pageId="43" pageNumber="44" type="male"></specimenCount>
,
<collectingRegion id="7922589DC146FFA31FDEF546FD8FFB28" box="[393,559,1137,1163]" country="Tanzania" name="Iringa" pageId="43" pageNumber="44">Iringa Region</collectingRegion>
,
<collectingRegion id="7922589DC146FFA31C6BF546FD46FB28" box="[572,742,1137,1163]" country="Tanzania" name="Iringa" pageId="43" pageNumber="44">Iringa District</collectingRegion>
,
<collectingCounty id="5238EEF3C146FFA31CA4F546FC0AFB2F" box="[755,938,1137,1164]" pageId="43" pageNumber="44">Udzungwa Mts</collectingCounty>
,
<collectingMunicipality id="5B3D0C05C146FFA31DE0F545FBD9FB28" box="[951,1145,1137,1164]" pageId="43" pageNumber="44">West Kilombero</collectingMunicipality>
FR,
<location id="BE39C0A4C146FFA31AE2F546FAD2FB28" LSID="urn:lsid:plazi:treatment:334F2769C146FFA51CD6F346FE4EFC89:BE39C0A4C146FFA31AE2F546FAD2FB28" box="[1205,1394,1137,1163]" country="Tanzania" county="Udzungwa Mts" latitude="-7.713611" longLatPrecision="21" longitude="36.420834" municipality="West Kilombero" name="Udekwa village" pageId="43" pageNumber="44" stateProvince="Iringa Region">Udekwa village</location>
,
<location id="BE39C0A4C146FFA31EEAF5A3FE22FB0C" LSID="urn:lsid:plazi:treatment:334F2769C146FFA51CD6F346FE4EFC89:BE39C0A4C146FFA31EEAF5A3FE22FB0C" box="[189,386,1172,1199]" country="Tanzania" county="Udzungwa Mts" latitude="-7.713611" longLatPrecision="21" longitude="36.420834" municipality="West Kilombero" name="Nyambanito Mt." pageId="43" pageNumber="44" stateProvince="Iringa Region">Nyambanito Mt.</location>
,
<location id="BE39C0A4C146FFA31FD9F5A3FD91FB0C" LSID="urn:lsid:plazi:treatment:334F2769C146FFA51CD6F346FE4EFC89:BE39C0A4C146FFA31FD9F5A3FD91FB0C" box="[398,561,1172,1199]" country="Tanzania" county="Udzungwa Mts" latitude="-7.713611" longLatPrecision="21" longitude="36.420834" municipality="West Kilombero" name="Ukami Forest" pageId="43" pageNumber="44" stateProvince="Iringa Region">Ukami Forest</location>
,
<geoCoordinate id="DED2F0B8C146FFA31C6AF5A2FD78FB0C" box="[573,728,1173,1199]" degrees="07" direction="south" minutes="42" orientation="latitude" pageId="43" pageNumber="44" precision="15" seconds="49" value="-7.713611">07° 4249” S</geoCoordinate>
,
<geoCoordinate id="DED2F0B8C146FFA31CB2F5A3FCDBFB0C" box="[741,891,1172,1199]" degrees="36" direction="east" minutes="25" orientation="longitude" pageId="43" pageNumber="44" precision="15" seconds="15" value="36.420834">36°2515” E</geoCoordinate>
,
<date id="CF58B0BFC146FFA31DD0F5A3FB9DFB0C" box="[903,1085,1172,1199]" pageId="43" pageNumber="44" value="1994-07" valueMax="1994-11-31" valueMin="1994-07-00">
<collectingDate id="DF1C4957C146FFA31DD0F5A3FB9DFB0C" box="[903,1085,1172,1199]" pageId="43" pageNumber="44" value="1994-07" valueMax="1994-11-31" valueMin="1994-07-00">Jul.Nov. 1994</collectingDate>
</date>
.
<collectorName id="1613F3A9C146FFA31A1FF5A2FB1AFB0C" box="[1096,1210,1173,1199]" pageId="43" pageNumber="44">D. Moyer</collectorName>
leg. (
<collectionCode id="DDF70EBAC146FFA31AABF5A2FAF8FB0C" box="[1276,1368,1173,1199]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="43" pageNumber="44">ZMUC</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="BB59967FC146FFA31EEAF5E1FEFDFB53" blockId="43.[189,349,1238,1264]" box="[189,349,1238,1264]" pageId="43" pageNumber="44">
<heading id="E0112113C146FFA31EEAF5E1FEFDFB53" bold="true" box="[189,349,1238,1264]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="89924A6DC146FFA31EEAF5E1FEFDFB53" bold="true" box="[189,349,1238,1264]" pageId="43" pageNumber="44">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC146FFA31EEAF432FC20FAE1" blockId="43.[189,1398,1284,1346]" pageId="43" pageNumber="44">
<collectingCountry id="C3F1D6EFC146FFA31EEAF432FEF7FABC" box="[189,343,1285,1311]" name="Tanzania" pageId="43" pageNumber="44">TANZANIA</collectingCountry>
:
<collectingRegion id="7922589DC146FFA31F31F433FDB0FABD" box="[358,528,1284,1310]" country="Tanzania" name="Iringa" pageId="43" pageNumber="44">Iringa Region</collectingRegion>
,
<collectingRegion id="7922589DC146FFA31C76F433FD6EFABD" box="[545,718,1284,1310]" country="Tanzania" name="Iringa" pageId="43" pageNumber="44">Iringa District</collectingRegion>
, Udzungwa Mts, West Kilombero
<collectionCode id="DDF70EBAC146FFA31A2DF432FB01FABC" box="[1146,1185,1285,1311]" pageId="43" pageNumber="44">FR</collectionCode>
, Udekwa village, Nyambanito Mt., Ukami Forest,
<geoCoordinate id="DED2F0B8C146FFA31C6AF41FFD78FAE1" box="[573,728,1320,1346]" degrees="07" direction="south" minutes="42" orientation="latitude" pageId="43" pageNumber="44" precision="15" seconds="49" value="-7.713611">07° 4249” S</geoCoordinate>
,
<geoCoordinate id="DED2F0B8C146FFA31CB2F410FCDCFAE1" box="[741,892,1319,1346]" degrees="36" direction="east" minutes="25" orientation="longitude" pageId="43" pageNumber="44" precision="15" seconds="15" value="36.420834">36°2515” E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C146FFA41EEAF45EFBA9FE76" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="description">
<paragraph id="BB59967FC146FFA31EEAF45EFE02FA27" blockId="43.[189,418,1385,1412]" box="[189,418,1385,1412]" pageId="43" pageNumber="44">
<emphasis id="89924A6DC146FFA31EEAF45EFEEEFA20" bold="true" box="[189,334,1385,1411]" pageId="43" pageNumber="44">Description</emphasis>
(male)
</paragraph>
<paragraph id="BB59967FC146FFA31EEAF4AFFB91FA11" blockId="43.[189,1073,1431,1458]" box="[189,1073,1431,1458]" pageId="43" pageNumber="44">
DIAMETER.
<quantity id="7C1E3B9AC146FFA31F6BF4A0FE39FA11" box="[316,409,1431,1458]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="43" pageNumber="44" unit="mm" value="3.5">3.5 mm</quantity>
, 54 podous rings (the largest of all species in the group).
</paragraph>
<paragraph id="BB59967FC146FFA31EEAF4EEFB34F9B4" blockId="43.[189,1398,1497,1559]" pageId="43" pageNumber="44">
COLOUR. After 20 years in alcohol ground colour greyish, posterior part of metazonites yellow-brownish; traces of a broad dorsal
<emphasis id="89924A6DC146FFA31F81F4CBFDACF9B5" box="[470,524,1532,1558]" italics="true" pageId="43" pageNumber="44">dark</emphasis>
stripe (most other species have traces of a light stripe).
</paragraph>
<paragraph id="BB59967FC146FFA31EEAF709FC8FF9D8" blockId="43.[189,1398,1597,1659]" pageId="43" pageNumber="44">ANAL VALVES. Each with a strong, apparently broken dorsal and smaller, but well-developed ventral spine. Margin raised, setiferous tubercles on ravelins.</paragraph>
<paragraph id="BB59967FC146FFA31EEAF794FAF3F91F" blockId="43.[189,1363,1698,1725]" box="[189,1363,1698,1725]" pageId="43" pageNumber="44">
LIMBUS. With triangular lobes; lobes
<emphasis id="89924A6DC146FFA31C3CF794FD27F91F" box="[619,647,1699,1724]" italics="true" pageId="43" pageNumber="44">ca</emphasis>
. as long as wide, margin micro-dentate, outer surface striate.
</paragraph>
<paragraph id="BB59967FC146FFA31EEAF7D3FC43F882" blockId="43.[189,1398,1764,1826]" pageId="43" pageNumber="44">
TARSAL SETATION. Unusual: claws surrounded and almost hidden by numerous stiff setae, accessory claw much longer than claw (as in
<taxonomicName id="7CE6EDFCC146FFA31C4FF630FD47F882" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[536,743,1799,1825]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="circumvallatus" status="sp. nov.">
<emphasis id="89924A6DC146FFA31C4FF630FD47F882" box="[536,743,1799,1825]" italics="true" pageId="43" pageNumber="44">C. circumvallatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C146FFA31CB8F630FCE9F882" box="[751,841,1799,1825]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
,
<figureCitation id="23DD8AFAC146FFA31D0FF630FC75F882" box="[856,981,1799,1826]" captionStart="Fig" captionStartId="12.[187,230,760,786]" captionTargetBox="[314,1268,265,748]" captionTargetId="figure@12.[314,1268,265,748]" captionTargetPageId="12" captionText="Fig. 5. Chaleponcus spp., tip of tarsus and claw of walking legs. — A. C. mwanihanensis sp. nov. as an example of normal setation. BC. C. circumvallatus sp. nov. with the claw surrounded by a palisade of setae. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860462" httpUri="https://zenodo.org/record/3860462/files/figure.png" pageId="43" pageNumber="44">Fig. 5B, C</figureCitation>
).
</paragraph>
<paragraph id="BB59967FC146FFA41EEAF67FFAD6FE85" blockId="43.[189,1399,1864,1996]" lastBlockId="44.[189,1398,267,470]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
GONOPOD COXA (
<figureCitation id="23DD8AFAC146FFA31F2FF67FFE62F8C1" box="[376,450,1864,1890]" captionStart="Fig" captionStartId="15.[187,230,1816,1842]" captionTargetBox="[247,1356,507,1717]" captionTargetId="figure@15.[200,1385,488,1723]" captionTargetPageId="15" captionText="Fig. 9. Chaleponcus spp., outline of coxite of left gonopod, posterior view. (The transverse plane of the gonopod is usually slightly at angles with the exact transverse plane of the millipede, so it may be necessary to tilt the gonopod slightly in order to obtain the view shown on the drawings.) The stippled lines indicate the place where the telopodital basomere emerges from the coxal cavity. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3860472" httpUri="https://zenodo.org/record/3860472/files/figure.png" pageId="43" pageNumber="44">Figs 9</figureCitation>
,
<figureCitation id="23DD8AFAC146FFA31F99F67FFE4FF8C1" box="[462,495,1864,1890]" captionStart="Fig" captionStartId="44.[189,232,1893,1919]" captionTargetBox="[190,1398,501,1853]" captionTargetId="figure@44.[189,1398,501,1853]" captionTargetPageId="44" captionText="Fig. 22. Chaleponcus ibis sp. nov., left gonopod. A. Posterior view. B. Anterior view. C. Telopodite, mesal-anterior view. al = anterior lobe of telomere, al = accessory lamella?, cu = cucullus, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior lobe of telomere, ps = proximal spine of solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860498" httpUri="https://zenodo.org/record/3860498/files/figure.png" pageId="43" pageNumber="44">22</figureCitation>
). Stout,
<emphasis id="89924A6DC146FFA31C19F67EFDD1F8C1" box="[590,625,1865,1890]" italics="true" pageId="43" pageNumber="44">ca.</emphasis>
3 × as long as wide. Lateral margin irregularly convex, continuing in smooth curve in apical margin; cucullus (
<emphasis id="89924A6DC146FFA31C85F65BFD4EF826" box="[722,750,1900,1925]" italics="true" pageId="43" pageNumber="44">cu</emphasis>
) hence regularly rounded. Metaplica posterio-mesally with triangular process (
<emphasis id="89924A6DC146FFA31FB5F6A7FDA6F80A" box="[482,518,1936,1961]" italics="true" pageId="43" pageNumber="44">mp</emphasis>
). Metaplical shelf (
<emphasis id="89924A6DC146FFA31CA2F6A7FCB6F80A" box="[757,790,1936,1961]" italics="true" pageId="43" pageNumber="44">ms</emphasis>
) relatively small, giving rise to very long, slightly curved metaplical shelf-spine (
<emphasis id="89924A6DC146FFA31C70F684FDF3F86F" box="[551,595,1971,1996]" italics="true" pageId="43" pageNumber="44">mss</emphasis>
). GONOPOD TELOPODITE (
<figureCitation id="23DD8AFAC141FFA41F9EF03CFD81FE86" box="[457,545,267,294]" captionStart="Fig" captionStartId="44.[189,232,1893,1919]" captionTargetBox="[190,1398,501,1853]" captionTargetId="figure@44.[189,1398,501,1853]" captionTargetPageId="44" captionText="Fig. 22. Chaleponcus ibis sp. nov., left gonopod. A. Posterior view. B. Anterior view. C. Telopodite, mesal-anterior view. al = anterior lobe of telomere, al = accessory lamella?, cu = cucullus, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior lobe of telomere, ps = proximal spine of solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860498" httpUri="https://zenodo.org/record/3860498/files/figure.png" pageId="44" pageNumber="45">Fig. 22</figureCitation>
). Solenomere with a very long, straight, mesad proximal spine (
<emphasis id="89924A6DC141FFA41B1CF03BFAC5FE86" box="[1355,1381,268,293]" italics="true" pageId="44" pageNumber="45">ps</emphasis>
).
</paragraph>
<paragraph id="BB59967FC141FFA41EEAF019FCDAFEEB" blockId="44.[189,1398,267,470]" box="[189,890,302,329]" pageId="44" pageNumber="45">Telomere distally with two branches, both curving anteriad:</paragraph>
<paragraph id="BB59967FC141FFA41EEAF065FEF8FE10" blockId="44.[189,1398,267,470]" pageId="44" pageNumber="45">
• an irregular shaped anterior lamella (
<emphasis id="89924A6DC141FFA41C91F065FD47FECF" box="[710,743,338,364]" italics="true" pageId="44" pageNumber="45">al)</emphasis>
with coarsely dentate proximal part and a smooth, subrectangular distal part (which maybe corresponds to the accessory lobe,
<emphasis id="89924A6DC141FFA41A0DF042FB24FE2C" box="[1114,1156,373,399]" italics="true" pageId="44" pageNumber="45">al¸</emphasis>
in
<taxonomicName id="7CE6EDFCC141FFA41AFFF042FAD6FE2C" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1192,1398,373,399]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="circumvallatus" status="sp. nov.">
<emphasis id="89924A6DC141FFA41AFFF042FAD6FE2C" box="[1192,1398,373,399]" italics="true" pageId="44" pageNumber="45">C. circumvallatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C141FFA41EBAF0AEFEEBFE10" box="[237,331,409,435]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
),
</paragraph>
<paragraph id="BB59967FC141FFA41EEAF08BFBA9FE76" blockId="44.[189,1398,267,470]" box="[189,1033,443,470]" pageId="44" pageNumber="45">
• a posterior lamella (
<emphasis id="89924A6DC141FFA41F8FF08CFE4FFE76" box="[472,495,443,469]" italics="true" pageId="44" pageNumber="45">pl</emphasis>
) ending in two short, parallel, elongate lobes.
</paragraph>
</subSubSection>
<caption id="EF99C6F7C141FFA41EEAF652FCB6F84A" ID-DOI="http://doi.org/10.5281/zenodo.3860498" ID-Zenodo-Dep="3860498" httpUri="https://zenodo.org/record/3860498/files/figure.png" pageId="44" pageNumber="45" startId="44.[189,232,1893,1919]" targetBox="[190,1398,501,1853]" targetPageId="44">
<paragraph id="BB59967FC141FFA41EEAF652FCB6F84A" blockId="44.[189,1398,1893,2025]" pageId="44" pageNumber="45">
<emphasis id="89924A6DC141FFA41EEAF652FEBBF8DC" bold="true" box="[189,283,1893,1919]" pageId="44" pageNumber="45">Fig. 22.</emphasis>
<taxonomicName id="7CE6EDFCC141FFA41F72F652FE52F8DC" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[293,498,1893,1919]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="ibis" status="sp. nov.">
<emphasis id="89924A6DC141FFA41F72F652FE52F8DC" box="[293,498,1893,1919]" italics="true" pageId="44" pageNumber="45">Chaleponcus ibis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C141FFA41FABF651FDFCF823" box="[508,604,1894,1920]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
, left gonopod.
<emphasis id="89924A6DC141FFA41D45F652FC87F8DC" bold="true" box="[786,807,1893,1919]" pageId="44" pageNumber="45">A</emphasis>
. Posterior view.
<emphasis id="89924A6DC141FFA41DA3F652FBA8F8DC" bold="true" box="[1012,1032,1893,1919]" pageId="44" pageNumber="45">B</emphasis>
. Anterior view.
<emphasis id="89924A6DC141FFA41A9BF652FB41F8DC" bold="true" box="[1228,1249,1893,1919]" pageId="44" pageNumber="45">C</emphasis>
. Telopodite, mesal-anterior view.
<emphasis id="89924A6DC141FFA41FE4F6BFFE6AF801" box="[435,458,1928,1954]" italics="true" pageId="44" pageNumber="45">al</emphasis>
= anterior lobe of telomere,
<emphasis id="89924A6DC141FFA41D4DF6BFFC9BF801" box="[794,827,1928,1954]" italics="true" pageId="44" pageNumber="45">al</emphasis>
= accessory lamella?,
<emphasis id="89924A6DC141FFA41A13F6BEFBC0F801" box="[1092,1120,1929,1954]" italics="true" pageId="44" pageNumber="45">cu</emphasis>
= cucullus,
<emphasis id="89924A6DC141FFA41AB8F6BEFAB3F801" box="[1263,1299,1929,1954]" italics="true" pageId="44" pageNumber="45">mp</emphasis>
= mesal metaplical process,
<emphasis id="89924A6DC141FFA41FF2F69BFE66F866" box="[421,454,1964,1989]" italics="true" pageId="44" pageNumber="45">ms</emphasis>
= metaplical shelf,
<emphasis id="89924A6DC141FFA41CF9F69BFD7AF866" box="[686,730,1964,1989]" italics="true" pageId="44" pageNumber="45">mss</emphasis>
= metaplical shelf-spine,
<emphasis id="89924A6DC141FFA41A5DF69CFB81F866" box="[1034,1057,1963,1989]" italics="true" pageId="44" pageNumber="45">pl</emphasis>
= posterior lobe of telomere,
<emphasis id="89924A6DC141FFA41EEAF6E7FF77F84A" box="[189,215,2000,2025]" italics="true" pageId="44" pageNumber="45">ps</emphasis>
= proximal spine of solenomere. Scales 0.1 mm.
</paragraph>
</caption>
<subSubSection id="F3FCC5F4C140FFA51EEAF03CFAF6FEF0" pageId="45" pageNumber="46" type="multiple">
<paragraph id="BB59967FC140FFA51EEAF03CFE4EFE86" blockId="45.[189,494,267,293]" box="[189,494,267,293]" pageId="45" pageNumber="46">
<heading id="E0112113C140FFA51EEAF03CFE4EFE86" bold="true" box="[189,494,267,293]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="89924A6DC140FFA51EEAF03CFE4EFE86" bold="true" box="[189,494,267,293]" pageId="45" pageNumber="46">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC140FFA51EEAF00DFAF6FEF0" blockId="45.[189,1366,313,340]" box="[189,1366,313,340]" pageId="45" pageNumber="46">Known only from Nyambanito Mt. in West Kilombero Scarp FR. No altitude or habitat information.</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C140FFA51EEAF04CFE4EFC89" pageId="45" pageNumber="46" type="discussion">
<paragraph id="BB59967FC140FFA51EEAF04CFE3EFE36" blockId="45.[189,414,379,405]" box="[189,414,379,405]" pageId="45" pageNumber="46">
<heading id="E0112113C140FFA51EEAF04CFE3EFE36" bold="true" box="[189,414,379,405]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="89924A6DC140FFA51EEAF04CFE3EFE36" bold="true" box="[189,414,379,405]" pageId="45" pageNumber="46">Coexisting species</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC140FFA51EEAF09DFF49FDA9" blockId="45.[189,1399,425,522]" pageId="45" pageNumber="46">
<taxonomicName id="7CE6EDFCC140FFA51EEAF09DFEF3FE60" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[189,339,425,452]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="basiliscus" status="sp. nov.">
<emphasis id="89924A6DC140FFA51EEAF09DFEF3FE60" box="[189,339,425,452]" italics="true" pageId="45" pageNumber="46">C. basiliscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C140FFA51F0EF09DFE13FE67" box="[345,435,426,452]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
was found in the same sample as
<taxonomicName id="7CE6EDFCC140FFA51D69F09DFC2AFE60" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[830,906,425,452]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="ibis" status="sp. nov.">
<emphasis id="89924A6DC140FFA51D69F09DFC2AFE60" box="[830,906,425,452]" italics="true" pageId="45" pageNumber="46">C. ibis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C140FFA51DC6F09DFC4BFE67" box="[913,1003,426,452]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
In addition,
<taxonomicName id="7CE6EDFCC140FFA51A28F09DFAEEFE60" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1151,1358,425,452]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="circumvallatus" status="sp. nov.">
<emphasis id="89924A6DC140FFA51A28F09DFAEEFE60" box="[1151,1358,425,452]" italics="true" pageId="45" pageNumber="46">C. circumvallatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C140FFA51B02F09DFF4FFE44" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="7CE6EDFCC140FFA51EAAF0FAFE28FE44" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[253,392,461,487]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="gracilior" status="sp. nov.">
<emphasis id="89924A6DC140FFA51EAAF0FAFE28FE44" box="[253,392,461,487]" italics="true" pageId="45" pageNumber="46">C. gracilior</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C140FFA51FC7F0FAFE4BFE44" box="[400,491,461,487]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="7CE6EDFCC140FFA51FADF0FAFDFAFE45" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[506,602,461,487]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="netus" status="sp. nov.">
<emphasis id="89924A6DC140FFA51FADF0FAFDFAFE45" box="[506,602,461,487]" italics="true" pageId="45" pageNumber="46">C. netus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C140FFA51C35F0FAFD1DFE44" box="[610,701,461,487]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="7CE6EDFCC140FFA51CA1F0FAFCF7FE45" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[758,855,461,487]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="tintin" status="sp. nov.">
<emphasis id="89924A6DC140FFA51CA1F0FAFCF7FE45" box="[758,855,461,487]" italics="true" pageId="45" pageNumber="46">C. tintin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C140FFA51D09F0FAFC19FE44" box="[862,953,461,487]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
were found in West Kilombero Scarp FR.
</paragraph>
<paragraph id="BB59967FC140FFA51EEAF306FEA3FDE8" blockId="45.[189,259,561,587]" box="[189,259,561,587]" pageId="45" pageNumber="46">
<heading id="E0112113C140FFA51EEAF306FEA3FDE8" bold="true" box="[189,259,561,587]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="89924A6DC140FFA51EEAF306FEA3FDE8" bold="true" box="[189,259,561,587]" pageId="45" pageNumber="46">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC140FFA51EEAF357FE4EFC89" blockId="45.[189,1399,608,810]" pageId="45" pageNumber="46">
Resembles the almost similar-sized
<taxonomicName id="7CE6EDFCC140FFA51C3BF357FC9DFDD9" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[620,829,608,634]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="circumvallatus" status="sp. nov.">
<emphasis id="89924A6DC140FFA51C3BF357FC9DFDD9" box="[620,829,608,634]" italics="true" pageId="45" pageNumber="46">C. circumvallatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C140FFA51D10F357FC04FDD9" box="[839,932,608,634]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
both in the unusual tarsal setation and in gonopod structure.
<taxonomicName id="7CE6EDFCC140FFA51F9FF3B4FDB7FD3E" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[456,535,643,669]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="ibis" status="sp. nov.">
<emphasis id="89924A6DC140FFA51F9FF3B4FDB7FD3E" box="[456,535,643,669]" italics="true" pageId="45" pageNumber="46">C. ibis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C140FFA51C75F3B3FDDFFD3D" box="[546,639,644,670]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="7CE6EDFCC140FFA51D74F3B4FC55FD3E" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[803,1013,643,669]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="circumvallatus" status="sp. nov.">
<emphasis id="89924A6DC140FFA51D74F3B4FC55FD3E" box="[803,1013,643,669]" italics="true" pageId="45" pageNumber="46">C. circumvallatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C140FFA51A57F3B3FBC0FD3D" box="[1024,1120,644,670]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
, i.a., in having distinct lobes on the limbus, in having the metaplical shelf-spine (
<emphasis id="89924A6DC140FFA51D2AF390FC09FD63" box="[893,937,679,704]" italics="true" pageId="45" pageNumber="46">mss</emphasis>
) much longer, in having the proximal part of the anterior telomeral lobe (
<emphasis id="89924A6DC140FFA51C38F3FEFD26FD40" box="[623,646,713,739]" italics="true" pageId="45" pageNumber="46">al</emphasis>
) coarsely dentate, and in having two relatively short, parallel elongate lobes on the posterior telomeral lamella, as opposed to the two very long, diverging spines in
<taxonomicName id="7CE6EDFCC140FFA51EEAF227FE2CFC89" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[189,396,784,810]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="circumvallatus" status="sp. nov.">
<emphasis id="89924A6DC140FFA51EEAF227FE2CFC89" box="[189,396,784,810]" italics="true" pageId="45" pageNumber="46">C. circumvallatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C140FFA51FC3F227FE4EFC89" box="[404,494,784,810]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>