150 lines
19 KiB
XML
150 lines
19 KiB
XML
<document id="305CE6A14839637378239020C986132C" ID-CLB-Dataset="37244" ID-DOI="10.11646/zootaxa.4226.4.9" ID-GBIF-Dataset="2fc811b7-a188-45a3-8283-928707d193ee" ID-ISSN="1175-5326" ID-Zenodo-Dep="265181" ID-ZooBank="570DC764-EF4E-4F29-9220-0A048767998C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1485855655015" checkinUser="GgServerImporter" docAuthor="Makarkin, Vladimir N." docDate="2017" docId="03AC5871FFD52546FF72FB724AB358F3" docLanguage="en" docName="data.5A19B6D72489CE0684E5E86DC52D17D7.cached" docOrigin="Zootaxa 4226 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Xenoberotha angustialata Makarkin, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="597" masterDocId="FF952009FFD42545FFE5FFEC4E7A5D42" masterDocTitle="An interesting new genus of Berothinae (Neuroptera: Berothidae) from the early Eocene Green River Formation, Colorado" masterLastPageNumber="600" masterPageNumber="594" pageNumber="595" updateTime="1698436549968" updateUser="ExternalLinkService">
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<mods:mods id="8954DEA6D9BD5CC138D6F5C41E7CEAA7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E91E74608DF648DF913CDE94892611D3">
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<mods:title id="D58CE64898903C1009BEBB9D06D76CDC">An interesting new genus of Berothinae (Neuroptera: Berothidae) from the early Eocene Green River Formation, Colorado</mods:title>
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</mods:titleInfo>
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<mods:name id="DE5A6D984248E9A31AF3B6CECED529DA" type="personal">
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<mods:role id="5E36E8BF226064EFB5A246102885D9CA">
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<mods:roleTerm id="8500377367162064B93ACBBCFBBC1739">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="77A8C0C37BD9FF433A307BB253F3FEE2">Makarkin, Vladimir N.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="AE6F631CB23287D95A2EB75D899F4BAC">text</mods:typeOfResource>
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<mods:titleInfo id="A77CD5E2D18671A10E669CFE27FACA29">
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<mods:title id="745CE97887EB70B3DDCE93C1471CEFA8">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="760A87965B6D806CE41710EAA38806E3">
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<mods:date id="6902BF59845100DE8D1B4537EE84F9F3">2017</mods:date>
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<mods:detail id="DE1ACC16BC28627EFDEE8A8CF92C18EE" type="volume">
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<mods:number id="15A31583E32FD8D7C184C577AD19401B">4226</mods:number>
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</mods:detail>
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<mods:detail id="17685D13B17A697D5EB0E605353FCA65" type="issue">
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<mods:number id="1690E18609333080A38D9788BE4F60B1">4</mods:number>
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</mods:detail>
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<mods:start id="6A61A1360F2E96C5D669F4F894F136E0">594</mods:start>
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<mods:end id="BC7FAC4D791F220B1FC38856F9EBD272">600</mods:end>
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</mods:part>
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<mods:classification id="A03C80D25F6701A85CE58FF1FDC573EB">journal article</mods:classification>
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<mods:identifier id="D7FF99AC90F99CDD259709B243ADBE46" type="CLB-Dataset">37244</mods:identifier>
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<mods:identifier id="5E1692DC94952912D29B4E497A15DCCA" type="DOI">10.11646/zootaxa.4226.4.9</mods:identifier>
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<mods:identifier id="72F856B832AE5C1255FD2148DB87FA69" type="GBIF-Dataset">2fc811b7-a188-45a3-8283-928707d193ee</mods:identifier>
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<mods:identifier id="4805EEA64B7C898992612280932258CE" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="03D271B5869E381B4F95358BA84A3F3E" type="Zenodo-Dep">265181</mods:identifier>
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</mods:mods>
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<treatment id="03AC5871FFD52546FF72FB724AB358F3" ID-DOI="http://doi.org/10.5281/zenodo.6028972" ID-GBIF-Taxon="127628128" ID-Zenodo-Dep="6028972" LSID="urn:lsid:plazi:treatment:03AC5871FFD52546FF72FB724AB358F3" httpUri="http://treatment.plazi.org/id/03AC5871FFD52546FF72FB724AB358F3" lastPageId="3" lastPageNumber="597" pageId="1" pageNumber="595">
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<subSubSection id="C31FBAECFFD52544FF72FB724E835998" pageId="1" pageNumber="595" type="nomenclature">
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<paragraph id="8BBAE967FFD52544FF72FB724C4F59FA" blockId="1.[151,565,1182,1242]" box="[151,565,1182,1208]" pageId="1" pageNumber="595">
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<heading id="D0F25E0BFFD52544FF72FB724C4F59FA" bold="true" box="[151,565,1182,1208]" fontSize="11" level="1" pageId="1" pageNumber="595" reason="1">
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<emphasis id="B9713575FFD52544FF72FB724C4F59FA" bold="true" box="[151,565,1182,1208]" pageId="1" pageNumber="595">
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<taxonomicName id="4C0592E4FFD52544FF72FB724FB559FA" ID-CoL="5C9GS" box="[151,463,1182,1208]" class="Insecta" family="Berothidae" genus="Xenoberotha" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="595" phylum="Arthropoda" rank="species" species="angustialata" status="sp. nov.">
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<emphasis id="B9713575FFD52544FF72FB724FB559FA" bold="true" box="[151,463,1182,1208]" italics="true" pageId="1" pageNumber="595">Xenoberotha angustialata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A242880EFFD52544FE32FB724C4F59FA" box="[471,565,1182,1208]" pageId="1" pageNumber="595" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBAE967FFD52544FF72FB2D4E835998" blockId="1.[151,565,1182,1242]" box="[151,249,1217,1242]" pageId="1" pageNumber="595">
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<figureCitation id="133EF5E2FFD52544FF72FB2D4EA65998" box="[151,220,1217,1242]" captionStart="FIGURE 1" captionStartId="2.[151,249,1933,1955]" captionTargetBox="[151,1436,447,1884]" captionTargetId="figure@2.[151,1436,447,1908]" captionTargetPageId="2" captionText="FIGURE 1. Xenoberotha angustialata gen. et sp. nov., holotype UCM 80385. A, the specimen as preserved (part). B, same (counterpart); both wetted with ethanol. C, the venation of the left forewing (converted to standard view, with apex to the right). Grey shadings show the color pattern. h, hypocauda; ph, pseudohypocauda. Scale bars = 1 mm." httpUri="https://zenodo.org/record/265183/files/figure.png" pageId="1" pageNumber="595">Figs 1</figureCitation>
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,
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<figureCitation id="133EF5E2FFD52544FF0FFB2E4E835998" box="[234,249,1218,1242]" captionStart="FIGURE 2" captionStartId="3.[151,249,1125,1147]" captionTargetBox="[151,1436,193,1103]" captionTargetId="figure@3.[151,1436,193,1103]" captionTargetPageId="3" captionText="FIGURE 2. Xenoberotha angustialata gen. et sp. nov., holotype UCM 80385. Head and forelegs, ventral view (wetted with ethanol). cx, coxa; fm, femur; sc, scapus; ti, tibia. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/265184/files/figure.png" pageId="1" pageNumber="595">2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31FBAECFFD52544FF72FAE54C255894" pageId="1" pageNumber="595" type="materials_examined">
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<paragraph id="8BBAE967FFD52544FF72FAE54FD15828" blockId="1.[151,1436,1289,2034]" pageId="1" pageNumber="595">
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<emphasis id="B9713575FFD52544FF72FAE54F715860" bold="true" box="[151,267,1289,1314]" pageId="1" pageNumber="595">Holotype.</emphasis>
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UCM 80385a,b (part, counterpart), collected on
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<date id="FFBBCFA7FFD52544FCA9FAE64A7C5860" box="[844,1030,1290,1314]" pageId="1" pageNumber="595" value="2009-08-11">11 August 2009</date>
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by David Kohls, and deposited in UCM. An incomplete female specimen with the left forewing relatively well preserved, and the right hind wing fragmentarily preserved.
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</paragraph>
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<paragraph id="8BBAE967FFD52544FF22FA994C255894" blockId="1.[151,1436,1289,2034]" pageId="1" pageNumber="595">
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<materialsCitation id="3B6DE33AFFD52544FF22FA994C255894" ID-GBIF-Occurrence="1426520868" collectionCode="UCM" country="United States of America" county="Garfield County" location="Parachute Creek" municipality="Eocene" pageId="1" pageNumber="595" specimenCount="1" stateProvince="Colorado" typeStatus="holotype">
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<emphasis id="B9713575FFD52544FF22FA994F8F58CC" bold="true" box="[199,501,1397,1422]" pageId="1" pageNumber="595">
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<typeStatus id="54BE57C5FFD52544FF22FA994F7858CC" box="[199,258,1397,1422]" pageId="1" pageNumber="595">Type</typeStatus>
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locality and horizon
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</emphasis>
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.
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<collectionCode id="ED1471A2FFD52544FDE7FA9A4C3858CC" box="[514,578,1398,1422]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="1" pageNumber="595">UCM</collectionCode>
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locality 2009063 (‘Claudia’s Place’), situated
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<locationDeviation id="9A08F045FFD52544FBA2FA994EB658F0" pageId="1" pageNumber="595">between East Fork and Middle Fork</locationDeviation>
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of
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<location id="8EDABFBCFFD52544FF1AFA754FBB58F0" LSID="urn:lsid:plazi:treatment:03AC5871FFD52546FF72FB724AB358F3:8EDABFBCFFD52544FF1AFA754FBB58F0" box="[255,449,1433,1458]" country="United States of America" county="Garfield County" municipality="Eocene" name="Parachute Creek" pageId="1" pageNumber="595" stateProvince="Colorado">Parachute Creek</location>
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;
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<collectingCounty id="62DB91EBFFD52544FE31FA754CE958F0" box="[468,659,1433,1458]" pageId="1" pageNumber="595">Garfield County</collectingCounty>
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,
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<collectingRegion id="49C12785FFD52544FD41FA754D7558F0" box="[676,783,1433,1458]" country="United States of America" name="Colorado" pageId="1" pageNumber="595">Colorado</collectingRegion>
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,
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<collectingCountry id="F312A9F7FFD52544FCC4FA764D1558F0" box="[801,879,1434,1458]" name="United States of America" pageId="1" pageNumber="595">U.S.A.</collectingCountry>
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<location id="8EDABFBCFFD52544FC99FA754A4658F0" LSID="urn:lsid:plazi:treatment:03AC5871FFD52546FF72FB724AB358F3:8EDABFBCFFD52544FC99FA754A4658F0" box="[892,1084,1433,1458]" country="United States of America" county="Garfield County" municipality="Eocene" name="Parachute Creek" pageId="1" pageNumber="595" stateProvince="Colorado">Parachute Creek</location>
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Member of the
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<location id="8EDABFBCFFD52544FAE9FA764F755894" LSID="urn:lsid:plazi:treatment:03AC5871FFD52546FF72FB724AB358F3:8EDABFBCFFD52544FAE9FA764F755894" country="United States of America" county="Garfield County" municipality="Eocene" name="Green River Formation" pageId="1" pageNumber="595" stateProvince="Colorado">
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<location id="8EDABFBCFFD52544FAE9FA764BE658F0" LSID="urn:lsid:plazi:treatment:03AC5871FFD52546FF72FB724AB358F3:8EDABFBCFFD52544FAE9FA764BE658F0" box="[1292,1436,1433,1458]" country="United States of America" county="Garfield County" municipality="Eocene" name="Green River" pageId="1" pageNumber="595" stateProvince="Colorado">
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Green
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<location id="8EDABFBCFFD52544FABBFA754BE658F0" LSID="urn:lsid:plazi:treatment:03AC5871FFD52546FF72FB724AB358F3:8EDABFBCFFD52544FABBFA754BE658F0" box="[1374,1436,1433,1458]" country="United States of America" county="Garfield County" municipality="Eocene" name="River Formation" pageId="1" pageNumber="595" stateProvince="Colorado">River</location>
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</location>
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Formation
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</location>
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; early
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<collectingMunicipality id="6BDE731DFFD52544FEBCFA524FD15897" box="[345,427,1470,1493]" pageId="1" pageNumber="595">Eocene</collectingMunicipality>
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(late Ypresian).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31FBAECFFD52544FF22FA0D4E955B5C" pageId="1" pageNumber="595" type="etymology">
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<paragraph id="8BBAE967FFD52544FF22FA0D4E955B5C" blockId="1.[151,1436,1289,2034]" pageId="1" pageNumber="595">
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<emphasis id="B9713575FFD52544FF22FA0D4F3458B8" bold="true" box="[199,334,1505,1530]" pageId="1" pageNumber="595">Etymology.</emphasis>
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From the Latin
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<emphasis id="B9713575FFD52544FDEAFA0E4C0958BB" box="[527,627,1506,1529]" italics="true" pageId="1" pageNumber="595">angustus</emphasis>
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, narrow, and
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<emphasis id="B9713575FFD52544FCF1FA0D4D2258BB" box="[788,856,1505,1529]" italics="true" pageId="1" pageNumber="595">alatus</emphasis>
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, winged, referring to the narrow forewings of the species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31FBAECFFD52546FF22F9C54AB358F3" lastPageId="3" lastPageNumber="597" pageId="1" pageNumber="595" type="description">
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<paragraph id="8BBAE967FFD52544FF22F9C54A665BCB" blockId="1.[151,1436,1289,2034]" pageId="1" pageNumber="595">
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<emphasis id="B9713575FFD52544FF22F9C54F235B00" bold="true" box="[199,345,1577,1602]" pageId="1" pageNumber="595">Description.</emphasis>
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Female. Head poorly preserved except for well-preserved eyes. Antennae filiform, rather stout; scapus very short, transverse (length shorter than diameter); pedicellus not visible (or does not differ from basal flagellomeres); proximal flagellomeres transverse, covered with fine short setae.
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</paragraph>
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<paragraph id="8BBAE967FFD52544FF22F9794B6B5B90" blockId="1.[151,1436,1289,2034]" pageId="1" pageNumber="595">Thorax very poorly preserved; details not visible. Abdomen apically with poorly preserved, long anteroventrally directed processes (probably hypocaudae of gonocoxite 9 or pseudohypocaudae of tergite 9).</paragraph>
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<paragraph id="8BBAE967FFD52544FF22F9314D9D5A7C" blockId="1.[151,1436,1289,2034]" pageId="1" pageNumber="595">
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Foreleg: coxae relatively short, stout, densely covered with short, dilated setae resembling scales (
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<figureCitation id="133EF5E2FFD52544FAA0F9314BF75BB4" box="[1349,1421,1757,1782]" captionStart="FIGURE 2" captionStartId="3.[151,249,1125,1147]" captionTargetBox="[151,1436,193,1103]" captionTargetId="figure@3.[151,1436,193,1103]" captionTargetPageId="3" captionText="FIGURE 2. Xenoberotha angustialata gen. et sp. nov., holotype UCM 80385. Head and forelegs, ventral view (wetted with ethanol). cx, coxa; fm, femur; sc, scapus; ti, tibia. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/265184/files/figure.png" pageId="1" pageNumber="595">Fig. 2</figureCitation>
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); femora moderately stout, with few preserved, rather short setae; tibiae nearly as long as femur, densely covered with fine long setae, mainly in distal part. Mid-, hind legs poorly preserved.
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</paragraph>
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<paragraph id="8BBAE967FFD52546FF22F8A54F1C58CF" blockId="1.[151,1436,1289,2034]" lastBlockId="3.[151,1436,1216,1457]" lastPageId="3" lastPageNumber="597" pageId="1" pageNumber="595">
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Forewing very narrow, with hind margin distally excised; ca.
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<quantity id="4CFD4482FFD52544FC7EF8A54A795A23" box="[923,1027,1865,1890]" metricMagnitude="-3" metricUnit="m" metricValue="6.35" pageId="1" pageNumber="595" unit="mm" value="6.35">6.35 mm</quantity>
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long,
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<quantity id="4CFD4482FFD52544FBB4F8A54AC35A23" box="[1105,1209,1865,1890]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="1" pageNumber="595" unit="mm" value="1.55">1.55 mm</quantity>
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wide. Costal space narrow; subcostal veinlets not preserved except one, once forked. ScP, RA probably distally fused; ScP+RA long, entering margin slightly proximad wing apex; some distal veinlets of ScP+RA with terminal fork. Three crossveins detected in RA space: 2ra-rp poorly preserved, located far proximad origin of RP1; 3ra-rp stout, dark, located between origins of RP2, RP3; 4ra-rp poorly preserved, located far distad presumable fusion of ScP, RA. RP originated far from wing base, with four branches. RP1 originating far distad origin RP; forked at outer gradate series, both branches shallowly forked; RP2 to RP4 distally with only shallow branching. 2r-m poorly preserved, connecting RP, MA rather far for fork of M. M appears not fused basally with R; forked slightly distad origin of RP; intramedial space rather broad basally. MA, MP similarly branched distally: both relatively deeply forked at gradate series, then all four branches shallowly forked one to four times. Cu dividing into CuA, CuP near wing base. Anterior trace of CuA distally shallowly forked, pectinate with four rather long branches, one short shallowly forked branch; two proximal-most branches originate from one point. CuP deeply forked, somewhat proximad fork of M; distal part not preserved. AA1 with three proximal-most branches originating from one point; distal part not preserved. AA2 probably pectinate, but poorly preserved (only one branch preserved). AA3 not detected. Outer (fourth) gradate series of crossveins nearly parallel to hind margin; four crossveins detected connecting RP2 to CuA. Trichosors fragmentarily preserved along hind margin. Color pattern: five crossveins of gradate series broadly margined with brown (i.e., 4ra-rp, 4rp1-rp2, 4r-m, 4im, 4m-cu); some distal forks of CuA, MA and MP inside pale brown.
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</paragraph>
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<caption id="DF7AB9EFFFD62547FF72F8614A275AA3" httpUri="https://zenodo.org/record/265183/files/figure.png" pageId="2" pageNumber="596" targetBox="[151,1436,447,1884]" targetPageId="2">
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<paragraph id="8BBAE967FFD62547FF72F8614A275AA3" blockId="2.[151,1437,1933,2017]" pageId="2" pageNumber="596">
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<emphasis id="B9713575FFD62547FF72F8614F745AE0" bold="true" box="[151,270,1933,1955]" pageId="2" pageNumber="596">FIGURE 1</emphasis>
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.
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<taxonomicName id="4C0592E4FFD62547FEF9F8614C5E5AE0" box="[284,548,1933,1954]" class="Insecta" family="Berothidae" genus="Xenoberotha" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="596" phylum="Arthropoda" rank="species" species="angustialata">
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<emphasis id="B9713575FFD62547FEF9F8614C5E5AE0" box="[284,548,1933,1954]" italics="true" pageId="2" pageNumber="596">Xenoberotha angustialata</emphasis>
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</taxonomicName>
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<emphasis id="B9713575FFD62547FDC9F8624CB45AE1" bold="true" box="[556,718,1934,1955]" pageId="2" pageNumber="596">gen. et sp. nov.</emphasis>
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, holotype UCM 80385. A, the specimen as preserved (part). B, same (counterpart); both wetted with ethanol. C, the venation of the left forewing (converted to standard view, with apex to the right). Grey shadings show the color pattern. h, hypocauda; ph, pseudohypocauda. Scale bars = 1 mm.
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</paragraph>
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</caption>
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<caption id="DF7AB9EFFFD72546FF72FB894D2459DB" httpUri="https://zenodo.org/record/265184/files/figure.png" pageId="3" pageNumber="597" targetBox="[151,1436,193,1103]" targetPageId="3">
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<paragraph id="8BBAE967FFD72546FF72FB894D2459DB" blockId="3.[151,1436,1124,1177]" pageId="3" pageNumber="597">
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<emphasis id="B9713575FFD72546FF72FB894F745938" bold="true" box="[151,270,1125,1147]" pageId="3" pageNumber="597">FIGURE 2</emphasis>
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.
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<taxonomicName id="4C0592E4FFD72546FEFEFB894C5E5938" box="[283,548,1125,1146]" class="Insecta" family="Berothidae" genus="Xenoberotha" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="597" phylum="Arthropoda" rank="species" species="angustialata">
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<emphasis id="B9713575FFD72546FEFEFB894C5E5938" box="[283,548,1125,1146]" italics="true" pageId="3" pageNumber="597">Xenoberotha angustialata</emphasis>
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</taxonomicName>
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<emphasis id="B9713575FFD72546FDCEFB894CB75938" bold="true" box="[555,717,1125,1146]" pageId="3" pageNumber="597">gen. et sp. nov.</emphasis>
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, holotype UCM 80385. Head and forelegs, ventral view (wetted with ethanol). cx, coxa; fm, femur; sc, scapus; ti, tibia. Scale bar = 0.5 mm.
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</paragraph>
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</caption>
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<paragraph id="8BBAE967FFD72546FF22FA744AB358F3" blockId="3.[151,1436,1216,1457]" box="[199,1225,1432,1457]" pageId="3" pageNumber="597">Basal portion of right hind wing fragmentarily, poorly preserved, venation hard to interpret.</paragraph>
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</subSubSection>
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</treatment>
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</document> |