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<document id="C2F1126C88BF0FA9334AA115B2F752D1" ID-DOI="10.1080/00222930902968809" ID-ISSN="1464-5262" ID-Zenodo-Dep="5216589" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629290479036" checkinUser="felipe" docAuthor="Ahrens, Dirk &amp; Fabrizi, Silvia" docDate="2009" docId="0399C459DB326F43FDA4FAB0585A0A1E" docLanguage="en" docName="JNatHist.43.25-26.1505-1584.pdf" docOrigin="Journal of Natural History 43 (25 - 26)" docSource="http://dx.doi.org/10.1080/00222930902968809" docStyle="DocumentStyle:FCA295E821436EF9307F50C7AC261B53.5:JNatHist.2008-2012.journal_article.1cover" docStyleId="FCA295E821436EF9307F50C7AC261B53" docStyleName="JNatHist.2008-2012.journal_article.1cover" docStyleVersion="5" docTitle="Gynaecoserica bocaki Ahrens &amp; Fabrizi 2009, sp. nov." docType="treatment" docVersion="3" lastPageNumber="1531" masterDocId="FFA0BC21DB2B6F58FFA8FF8A5B280E11" masterDocTitle="A review of the genus Gynaecoserica Brenske, 1896 (Coleoptera, Scarabaeidae, Sericini)" masterLastPageNumber="1584" masterPageNumber="1505" pageNumber="1529" updateTime="1699066727896" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1E9952FEE0B1E6B7863B9921B8A6E62E">A review of the genus Gynaecoserica Brenske, 1896 (Coleoptera, Scarabaeidae, Sericini)</mods:title>
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<mods:name id="F50C2C6160E173C130316E8BE87B0A31" type="personal">
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<mods:namePart id="37EB3107AEEBCC705E83601B704FC21B">Ahrens, Dirk</mods:namePart>
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<mods:namePart id="2853CAA769370923F13B098D873B2B82">Fabrizi, Silvia</mods:namePart>
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<mods:typeOfResource id="F43B33EC0384FFA059E13DB31B432696">text</mods:typeOfResource>
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<mods:title id="85C35F9226DB174832F76F9F2E708AAD">Journal of Natural History</mods:title>
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<mods:date id="1B5571D872899065B486113C2B333842">2009</mods:date>
<mods:detail id="35F719E9C2E8BE5896E3C5CB0424692C" type="pubDate">
<mods:number id="B49D40C07AF27E3EBAAE47C9820EECC7">2009-06-16</mods:number>
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<mods:detail id="C90328E0EFF8378150E19249C33F1A6F" type="volume">
<mods:number id="F46CB136D6EF5041E14F9A35A4C27E06">43</mods:number>
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<mods:number id="23F9F7E812C61AA138A30078734B3F54">25 - 26</mods:number>
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<mods:start id="61BBA3415A03728B674A191DB86E6639">1505</mods:start>
<mods:end id="DD4B99AB3DAB6ACE6E153CD7986D35F8">1584</mods:end>
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<mods:classification id="68ED31974430638018116D0FB7B11985">journal article</mods:classification>
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<mods:identifier id="1AF4F6D9EED63924CB446AA8BBB91930" type="ISSN">1464-5262</mods:identifier>
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<treatment id="0399C459DB326F43FDA4FAB0585A0A1E" LSID="urn:lsid:plazi:treatment:0399C459DB326F43FDA4FAB0585A0A1E" httpUri="http://treatment.plazi.org/id/0399C459DB326F43FDA4FAB0585A0A1E" lastPageId="27" lastPageNumber="1531" pageId="25" pageNumber="1529">
<subSubSection id="C32A26C4DB326F41FDA4FAB058740B43" box="[524,860,1338,1362]" pageId="25" pageNumber="1529" type="nomenclature">
<paragraph id="8B8F754FDB326F41FDA4FAB058740B43" blockId="25.[524,860,1338,1394]" box="[524,860,1338,1362]" pageId="25" pageNumber="1529">
<heading id="D0C7C223DB326F41FDA4FAB058740B43" box="[524,860,1338,1362]" centered="true" fontSize="36" level="2" pageId="25" pageNumber="1529" reason="7">
<taxonomicName id="4C300ECCDB326F41FDA4FAB059D50B43" authority="Ahrens &amp; Fabrizi, 2009" authorityName="Ahrens &amp; Fabrizi" authorityYear="2009" box="[524,765,1338,1362]" class="Insecta" family="Melolonthidae" genus="Gynaecoserica" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1529" phylum="Arthropoda" rank="species" species="bocaki" status="sp. nov.">
<emphasis id="B944A95DDB326F41FDA4FAB059D50B43" bold="true" box="[524,765,1338,1362]" italics="true" pageId="25" pageNumber="1529">Gynaecoserica bocaki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2771426DB326F41FCABFAB158740B43" box="[771,860,1339,1362]" pageId="25" pageNumber="1529" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A26C4DB326F43FD86FAD0585A0A1E" lastPageId="27" lastPageNumber="1531" pageId="25" pageNumber="1529" type="description">
<paragraph id="8B8F754FDB326F41FD86FAD058110B63" blockId="25.[524,860,1338,1394]" box="[558,825,1370,1394]" pageId="25" pageNumber="1529">
(
<figureCitation id="130B69CADB326F41FD9EFAD059F10B63" box="[566,729,1370,1394]" captionStart="Figure 1" captionStartId="8.[142,209,1258,1280]" captionTargetBox="[143,1101,178,1219]" captionTargetId="figure-182@8.[142,1102,177,1227]" captionTargetPageId="8" captionText="Figure 1. (AC) Gynaecoserica alma sp. nov. (holotype), (DF) G. amara sp. nov. (holotype), (G, H) G. arunachalensis sp. nov. (holotype), (J, K) G. barclayi sp. nov. (holotype), (LN) G. bicolorata sp. nov. (holotype), (OQ) G. bocaki sp. nov. (holotype). (A, D, G, J, L, O) Aedeagus, left side lateral view; (C, F, I, N, Q) aedeagus, left side lateral view; (B, E, H, K, M, P) parameres, dorsal view. Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5216591" httpUri="https://zenodo.org/record/5216591/files/figure.png" pageId="25" pageNumber="1529">Figures 1OQ</figureCitation>
,
<figureCitation id="130B69CADB326F41FD4CFAD0582E0B63" box="[740,774,1370,1394]" captionStart="Figure 6" captionStartId="13.[212,277,1536,1558]" captionTargetBox="[230,1154,172,1504]" captionTargetId="figure-88@13.[229,1155,171,1504]" captionTargetPageId="13" captionText="Figure 6. Habitus of the species (not to scale). (A) Gynaecoserica alma sp. nov. (5.8 mm; holotype), (B) G. amara sp. nov. (4.8 mm; holotype), (C) G. arunachalensis sp. nov. (5.1 mm; holotype), (D) G. bicolorata sp. nov. (4.7 mm; holotype), (E) G. bocaki sp. nov. (4.7 mm; holotype), (F) G. bomdilana sp. nov. (6.8 mm; holotype), (G) G. compacta sp. nov. (6.3 mm; holotype), (H) G. densipunctata sp. nov. (4.0 mm; holotype), (I) G. dirangensis sp. nov. (5.2 mm; holotype)." figureDoi="http://doi.org/10.5281/zenodo.5216601" httpUri="https://zenodo.org/record/5216601/files/figure.png" pageId="25" pageNumber="1529">6E</figureCitation>
,
<figureCitation id="130B69CADB326F41FCB9FAD058190B63" box="[785,817,1370,1394]" captionStart="Figure 9" captionStartId="16.[145,212,1609,1631]" captionTargetBox="[165,1082,174,1579]" captionTargetId="figure-40@16.[165,1084,173,1579]" captionTargetPageId="16" captionText="Figure 9. Distribution of Gynaecoserica species (A, upper panel; B, lower panel). “?” in (B) indicates the not precisely localized collecting locality of G. gisionensis sp. nov. in Vientiane province in Laos." figureDoi="http://doi.org/10.5281/zenodo.5216607" httpUri="https://zenodo.org/record/5216607/files/figure.png" pageId="25" pageNumber="1529">9B</figureCitation>
)
</paragraph>
<paragraph id="8B8F754FDB326F41FF7CFA285AF70BAB" blockId="25.[212,479,1442,1466]" box="[212,479,1442,1466]" pageId="25" pageNumber="1529">
<emphasis id="B944A95DDB326F41FF7CFA285AF70BAB" box="[212,479,1442,1466]" italics="true" pageId="25" pageNumber="1529">Type material examined</emphasis>
</paragraph>
<paragraph id="8B8F754FDB326F41FF7CFA40597A0813" blockId="25.[212,1171,1481,1538]" pageId="25" pageNumber="1529">
<materialsCitation id="3B587F12DB326F41FF7CFA4058650BF3" box="[212,845,1481,1506]" collectingDate="1990-06" collectionCode="TICB" country="China" location="China" pageId="25" pageNumber="1529" specimenCount="1" typeStatus="holotype">
<typeStatus id="548BCBEDDB326F41FF7CFA405A6B0BF3" box="[212,323,1482,1506]" pageId="25" pageNumber="1529" type="holotype">Holotype</typeStatus>
. Ƌ “
<collectingCountry id="F32735DFDB326F41FEDEFA405A960BF3" box="[374,446,1482,1506]" name="China" pageId="25" pageNumber="1529">China</collectingCountry>
<date id="FF8E538FDB326F41FE6EFA4059370BF3" box="[454,543,1482,1506]" pageId="25" pageNumber="1529" value="1990-06">
<collectingDate id="EFCAAA67DB326F41FE6EFA4059370BF3" box="[454,543,1482,1506]" pageId="25" pageNumber="1529" value="1990-06">VI.1990</collectingDate>
</date>
Dali Leg. Bocák” (
<collectionCode id="ED21ED8ADB326F41FD57FA40586D0BF3" box="[767,837,1482,1506]" pageId="25" pageNumber="1529">TICB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B587F12DB326F41FCF0FA4059660813" collectingDate="1990-06" collectionCode="TICB, BMNH, CA" country="China" location="China" pageId="25" pageNumber="1529" specimenCount="9" specimenCount-female="9" typeStatus="paratype">
<typeStatus id="548BCBEDDB326F41FCF0FA4058E40BF3" box="[856,972,1482,1506]" pageId="25" pageNumber="1529" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="9D36BEC6DB326F41FC7FFA405F300BF3" box="[983,1048,1481,1506]" pageId="25" pageNumber="1529" type="female">9 ♀♀</specimenCount>
same data as holotype (
<collectionCode id="ED21ED8ADB326F41FECCFA605A810813" box="[356,425,1514,1538]" pageId="25" pageNumber="1529">TICB</collectionCode>
,
<collectionCode id="ED21ED8ADB326F41FE1CFA6059270810" box="[436,527,1514,1537]" pageId="25" pageNumber="1529">BMNH</collectionCode>
,
<collectionCode id="ED21ED8ADB326F41FDB2FA60596E0813" box="[538,582,1514,1538]" country="USA" name="Chicago Academy of Sciences" pageId="25" pageNumber="1529" type="Museum">CA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8F754FDB326F41FF7CF9B85A7C085B" blockId="25.[212,340,1586,1610]" box="[212,340,1586,1610]" pageId="25" pageNumber="1529">
<emphasis id="B944A95DDB326F41FF7CF9B85A7C085B" box="[212,340,1586,1610]" italics="true" pageId="25" pageNumber="1529">Description</emphasis>
</paragraph>
<paragraph id="8B8F754FDB326F42FF7CF9D05AF60ECE" blockId="25.[212,1171,1626,1682]" lastBlockId="26.[144,1104,167,1696]" lastPageId="26" lastPageNumber="1530" pageId="25" pageNumber="1529">
Length
<quantity id="4CC8D8AADB326F41FE9AF9D05AA20860" box="[306,394,1626,1650]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="25" pageNumber="1529" unit="mm" value="4.7">4.7 mm</quantity>
, length of elytra
<quantity id="4CC8D8AADB326F41FDCBF9D059930860" box="[611,699,1626,1650]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="25" pageNumber="1529" unit="mm" value="3.1">3.1 mm</quantity>
, width
<quantity id="4CC8D8AADB326F41FCB3F9D0585B0860" box="[795,883,1626,1650]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="25" pageNumber="1529" unit="mm" value="2.4">2.4 mm</quantity>
. Body moderately oval, surface light reddish brown with disc of pronotum and lateral elytral margins widely darker, legs and labroclypeus yellowish brown, dorsal surface except head moderately dull and sparsely setose.
</paragraph>
<paragraph id="8B8F754FDB316F42FF12FF6D59ED0C8E" blockId="26.[144,1104,167,1696]" pageId="26" pageNumber="1530">
Labroclypeus almost semicircular, distinctly wider than long, widest at base, lateral margins strongly curved and strongly convergent, anterior angles obsolete, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anterior margin narrowly sinuate at the middle; surface weakly convex and shiny, finely and densely punctate, distance between punctures equal to their diameter, with a few coarse punctures behind anterior margin each bearing a long, erect seta; frontoclypeal suture feebly incised and weakly curved medially; smooth area in front of eye approximately 1.2 times as wide as long; ocular canthus short and slender, densely minutely punctate, without a fine terminal seta. Frons completely moderately shiny, with fine, moderately dense punctures, glabrous except a few short setae beside eyes and behind lateral frontoclypeal suture. Eyes small, ratio of diameter: interocular width 0.54. Antenna yellow, composed of 10 antennomeres; club with four antennomeres, as long as the remaining antennomeres combined.
<collectingRegion id="49F4BBADDB316F42FF38FD025BDA0C8E" box="[144,242,648,671]" country="Myanmar" name="Chin" pageId="26" pageNumber="1530">Mentum</collectingRegion>
weakly elevated and flattened anteriorly.
</paragraph>
<paragraph id="8B8F754FDB316F42FF12FD22599A0DAE" blockId="26.[144,1104,167,1696]" pageId="26" pageNumber="1530">Pronotum moderately wide, widest at base, lateral margins weakly curved and weakly convergent anteriorly, in anterior half more strongly curved and strongly convergent towards the strongly produced and sharp anterior angles, posterior angles blunt, anterior margin convexly produced medially, with a broad marginal line, basal margin without marginal line; surface with dense and fine punctures, with minute setae in punctures; anterior border sparsely setaceous, lateral border almost glabrous; hypomeron distinctly margined at base but not ventrally produced. Scutellum shorttriangular, with fine and very dense punctures, basally with a large smooth triangular area, microscopic setae present in the punctures.</paragraph>
<paragraph id="8B8F754FDB316F42FF12FC4D59AE0A6E" blockId="26.[144,1104,167,1696]" pageId="26" pageNumber="1530">Elytra short and oval, widest at middle, striae moderately impressed, finely and densely punctate, intervals weakly convex, with fine and moderately dense punctures along the striae, punctures with fine microscopic setae, odd intervals with numerous coarse punctures bearing a robust white seta; epipleural edge robust and sharp ending at the strongly curved external apical angle of elytra, epipleura densely setaceous, apical border without short microtrichomes.</paragraph>
<paragraph id="8B8F754FDB316F42FF12FB0D5ACA0BAE" blockId="26.[144,1104,167,1696]" pageId="26" pageNumber="1530">Ventral surface dull, with fine and moderately dense punctures, sparsely setose, metacoxa only laterally with a few strong setae; each abdominal sternite with indistinct transverse row of coarse punctures bearing short setae between fine, dense punctation, penultimate sternite apically with a shiny smooth sclerotized border, which is one-sixth as long as sternite, last sternite medially a little shorter than the penultimate one. Mesosternum between mesocoxae as wide as mesofemur, with irregularly scattered very strong setae. Ratio of length of metepisternum: metacoxa: 1: 1.6. Pygidium convex, densely mixed, finely and coarsely punctate, without smooth midline, with sparse robust adjacent yellowish setae in the coarse punctures, otherwise only with minute setae in the punctures.</paragraph>
<paragraph id="8B8F754FDB316F43FF12FA425A840FAE" blockId="26.[144,1104,167,1696]" lastBlockId="27.[212,1172,167,479]" lastPageId="27" lastPageNumber="1531" pageId="26" pageNumber="1530">Legs robust and short; femora dull, with two longitudinal rows of setae, finely and sparsely punctate; metafemur shiny, sharply margined anteriorly and without a submarginal serrate line, posterior margin weakly convex and glabrous, ventrally only weakly widened in apical half and not serrate, dorsally finely serrate, with moderately short setae. Metatibia broad and moderately short, ventromedially convexly widened, widest at middle, ratio width: length: 1: 3.2, dorsally distinctly carinate, with two groups of spines, basal one at one-third, apical one at two-thirds of metatibial length, basally with a few single, fine spines in the punctures; external face longitudinally convex, with very sparse, fine punctures, glabrous; ventrally edged and serrate, with three strong, equally distant spines, internal face not punctate and smooth, apex interiorly near tarsal articulation sharply and deeply truncate. Tarsomeres dorsally glabrous and impunctate, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrate ridge, beside which is a fine longitudinal carina, first metatarsomere as long as the following two tarsomeres combined and slightly less than twice as long as the upper tibial spur. Protibia very short, bidentate, protarsal claws symmetrical.</paragraph>
<paragraph id="8B8F754FDB306F43FF56FE4D59A70FCE" blockId="27.[212,1172,167,479]" box="[254,655,455,479]" pageId="27" pageNumber="1531">
Aedeagus shown in
<figureCitation id="130B69CADB306F43FE48FE4D59140FCE" box="[480,572,455,479]" captionStart="Figure 1" captionStartId="8.[142,209,1258,1280]" captionTargetBox="[143,1101,178,1219]" captionTargetId="figure-182@8.[142,1102,177,1227]" captionTargetPageId="8" captionText="Figure 1. (AC) Gynaecoserica alma sp. nov. (holotype), (DF) G. amara sp. nov. (holotype), (G, H) G. arunachalensis sp. nov. (holotype), (J, K) G. barclayi sp. nov. (holotype), (LN) G. bicolorata sp. nov. (holotype), (OQ) G. bocaki sp. nov. (holotype). (A, D, G, J, L, O) Aedeagus, left side lateral view; (C, F, I, N, Q) aedeagus, left side lateral view; (B, E, H, K, M, P) parameres, dorsal view. Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5216591" httpUri="https://zenodo.org/record/5216591/files/figure.png" pageId="27" pageNumber="1531">Figure 1</figureCitation>
(OQ).
</paragraph>
<paragraph id="8B8F754FDB306F43FF7CFDAF5A690C2C" blockId="27.[212,321,549,573]" box="[212,321,549,573]" pageId="27" pageNumber="1531">
<emphasis id="B944A95DDB306F43FF7CFDAF5A690C2C" box="[212,321,549,573]" italics="true" pageId="27" pageNumber="1531">Diagnosis</emphasis>
</paragraph>
<paragraph id="8B8F754FDB306F43FF7CFDC759E30CD4" blockId="27.[212,1172,589,709]" pageId="27" pageNumber="1531">
<taxonomicName id="4C300ECCDB306F43FF7CFDC75AED0C74" authority="Ahrens &amp; Fabrizi, 2009" authorityName="Ahrens &amp; Fabrizi" authorityYear="2009" box="[212,453,589,613]" class="Insecta" family="Melolonthidae" genus="Gynaecoserica" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="1531" phylum="Arthropoda" rank="species" species="bocaki" status="sp. nov.">
<emphasis id="B944A95DDB306F43FF7CFDC75AED0C74" box="[212,453,589,613]" italics="true" pageId="27" pageNumber="1531">Gynaecoserica bocaki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2771426DB306F43FE65FDC759000C75" box="[461,552,589,612]" pageId="27" pageNumber="1531" rank="species">sp. nov.</taxonomicNameLabel>
is externally rather similar to
<taxonomicName id="4C300ECCDB306F43FC2DFDC75FBC0C74" authority="(Brenske)" authorityName="Ahrens &amp; Fabrizi" authorityYear="2009" baseAuthorityName="Brenske" baseAuthorityYear="1896" box="[901,1172,589,613]" class="Insecta" family="Melolonthidae" genus="Gynaecoserica" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="1531" phylum="Arthropoda" rank="species" species="marginipes">
<emphasis id="B944A95DDB306F43FC2DFDC75F370C74" box="[901,1055,589,613]" italics="true" pageId="27" pageNumber="1531">G. marginipes</emphasis>
(Brenske)
</taxonomicName>
but antennal club in male shorter and with four antennomeres only. From all other species with four-segmented club it differs by the small size (&lt;
<quantity id="4CC8D8AADB306F43FC68FD075F290CB4" box="[960,1025,653,677]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="1531" unit="mm" value="5.0">5 mm</quantity>
) and by the absence of a lateral apophysis of phallobase.
</paragraph>
<paragraph id="8B8F754FDB306F43FF7CFC805A160D33" blockId="27.[212,318,778,802]" box="[212,318,778,802]" pageId="27" pageNumber="1531">
<emphasis id="B944A95DDB306F43FF7CFC805A160D33" box="[212,318,778,802]" italics="true" pageId="27" pageNumber="1531">Variation</emphasis>
</paragraph>
<paragraph id="8B8F754FDB306F43FF7CFCB85ABB0D9B" blockId="27.[212,1172,818,906]" pageId="27" pageNumber="1531">
Length 4.75.0 mm, length of elytra
<quantity id="4CC8D8AADB306F43FDC6FCB859DD0D5B" box="[622,757,818,842]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="27" pageNumber="1531" unit="mm" value="3.0" valueMax="3.1" valueMin="2.9">2.93.1 mm</quantity>
, width
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. Antennal club in female with three antennomeres and distinctly shorter than the remaining antennomeres combined.
</paragraph>
<paragraph id="8B8F754FDB306F43FF7CFC455A650DF6" blockId="27.[212,333,975,999]" box="[212,333,975,999]" pageId="27" pageNumber="1531">
<emphasis id="B944A95DDB306F43FF7CFC455A650DF6" box="[212,333,975,999]" italics="true" pageId="27" pageNumber="1531">Etymology</emphasis>
</paragraph>
<paragraph id="8B8F754FDB306F43FF7CFC7D585A0A1E" blockId="27.[212,882,1015,1039]" box="[212,882,1015,1039]" pageId="27" pageNumber="1531">Named for L. Bocák, the successful collector of the species.</paragraph>
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</treatment>
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