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<mods:title id="45E42BCD683E3EF179F8F91CC0DBFE96">A taxonomic revision of the genus Paradynomene Sakai, 1963 (Crustacea: Decapoda: Brachyura: Dynomenidae)</mods:title>
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<subSubSection id="C34C46E5FF840C13FED2FD9B0A79B258" box="[264,596,593,619]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8BE9156EFF840C13FED2FD9B0A79B258" blockId="1.[264,596,593,619]" box="[264,596,593,619]" pageId="1" pageNumber="2">
<heading id="D0A1A202FF840C13FED2FD9B0A79B258" bold="true" box="[264,596,593,619]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<taxonomicName id="4C566EEDFF840C13FED2FD9B0A79B258" ID-CoL="7PRSS" authority="Sakai, 1963" authorityName="Sakai" authorityYear="1963" box="[264,596,593,619]" class="Malacostraca" family="Dynomenidae" genus="Paradynomene" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B922C97CFF840C13FED2FD9B0A79B258" bold="true" box="[264,596,593,619]" pageId="1" pageNumber="2">
<emphasis id="B922C97CFF840C13FED2FD9B0993B258" bold="true" box="[264,446,593,619]" italics="true" pageId="1" pageNumber="2">Paradynomene</emphasis>
<bibRefCitation id="EFC7689FFF840C13FE1CFD9B0A79B258" author="Sakai" box="[454,596,593,619]" pageId="1" pageNumber="24" refString="Sakai, T. (1963) Description of two new genera and fourteen new species of Japanese crabs from the collection of his Majesty the Emperor of Japan. Crustaceana, 5 (3), 213 - 233." type="journal article" year="1963">Sakai, 1963</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C34C46E5FF840C13FED2FD560B7CB280" box="[264,849,668,691]" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="8BE9156EFF840C13FED2FD560B7CB280" blockId="1.[264,849,668,691]" box="[264,849,668,691]" pageId="1" pageNumber="2">
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<taxonomicName id="4C566EEDFF840C13FED2FD560A48B280" ID-CoL="7PRSS" authority="Sakai, 1963: 230" authorityName="Sakai" authorityPageNumber="230" authorityYear="1963" box="[264,613,668,691]" class="Malacostraca" family="Dynomenidae" genus="Paradynomene" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B922C97CFF840C13FED2FD560984B280" box="[264,425,668,691]" italics="true" pageId="1" pageNumber="2">Paradynomene</emphasis>
<treatmentCitation id="0AF7337FFF840C13FE6AFD560A48B280" author="Sakai" box="[432,613,668,691]" page="230" pageId="1" pageNumber="2" year="1963">
<bibRefCitation id="EFC7689FFF840C13FE6AFD560A1FB280" author="Sakai" box="[432,562,668,691]" pageId="1" pageNumber="24" refString="Sakai, T. (1963) Description of two new genera and fourteen new species of Japanese crabs from the collection of his Majesty the Emperor of Japan. Crustaceana, 5 (3), 213 - 233." type="journal article" year="1963">Sakai, 1963</bibRefCitation>
: 230
</treatmentCitation>
</taxonomicName>
.—
<treatmentCitation id="0AF7337FFF840C13FD5EFD560B61B280" author="McLay" box="[644,844,668,691]" page="542" pageId="1" pageNumber="2" year="1999">
<bibRefCitation id="EFC7689FFF840C13FD5EFD560B3AB280" author="McLay" box="[644,791,668,691]" pageId="1" pageNumber="24" refString="McLay C. L. (1999) Crustacea Decapoda: Revision of the family Dynomenidae, In: A. Crosnier (ed.), Resultats des Campagnes MUSORSTOM Vo l u m e 2 0. Memoires du Museum national d'Histoire naturelle (Paris), 180, 427 - 569, figs. 1 - 40." type="journal article" year="1999">McLay, 1999</bibRefCitation>
: 542
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C34C46E5FF840C13FED2FD2C0A3DB5CA" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8BE9156EFF840C13FED2FD2C0A3DB5CA" blockId="1.[264,1324,742,1889]" pageId="1" pageNumber="2">
<emphasis id="B922C97CFF840C13FED2FD2C09ADB333" bold="true" box="[264,384,742,768]" pageId="1" pageNumber="2">Diagnosis</emphasis>
. Carapace sub­rectangular or sub­oval, surface markedly convex, areolate, regions distinct. Lateral carapace margins not well defined, sub­parallel or slightly convex, armed with irregular teeth. Frontal margin truncate, rostrum bi­ or tridentate. Eyes protected by well defined orbits, exposed dorsally. Sternal sutures 7 and 8 of female end well apart on low tubercles behind bases of third pereopods. Body sparsely setose except along margins of abdomen, which can be fringed with setae. Antennule may be concealed inside orbit at base of eyestalk. First article large, triangular, filling large part of ventral orbital region. Antennal flagella shorter than half of carapace width. All articles of antenna moveable, first segment (urinal) beaked medially. Third maxillipeds opercular, sharply angled, completely covering buccal cavern, separated at bases by sternite, basis, and ischium of endopod fused but always marked by shallow suture. Crista dentata with 1214 small teeth increasing in size distally, first five or six may be obsolete. Chelipeds robust, equal, stouter than ambulatory legs, last pair of legs very reduced, dactyl rudimentary, forming obsolete chelate mechanism with extension of propodus only in female. Branchial formula 19 gills + 7 epipods (for detailed branchial formula see
<bibRefCitation id="EFC7689FFF840C13FCE4FADD0BCDB502" author="McLay" box="[830,992,1303,1329]" pageId="1" pageNumber="24" refString="McLay C. L. (1999) Crustacea Decapoda: Revision of the family Dynomenidae, In: A. Crosnier (ed.), Resultats des Campagnes MUSORSTOM Vo l u m e 2 0. Memoires du Museum national d'Histoire naturelle (Paris), 180, 427 - 569, figs. 1 - 40." type="journal article" year="1999">McLay, 1999</bibRefCitation>
: 546). Abdomen of six segments and telson, folded loosely under thorax, uropods large, abdominal locking mechanism not functional. Both sexes with five pairs of pleopods, first pair rudimentary in female, last three pairs vestigial in male. First pair of male pleopods stout, setose semirolled tube with apical plate, second pair needle­like with linear row of tiny inset spines along anterior surface.
</paragraph>
</subSubSection>
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<emphasis id="B922C97CFF840C13FEE2F9CC09FAB613" bold="true" box="[312,471,1542,1568]" pageId="1" pageNumber="2">
<typeStatus id="54EDABCCFF840C13FEE2F9CC0958B613" box="[312,373,1542,1568]" pageId="1" pageNumber="2">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="4C566EEDFF840C13FE30F9CD0BE7B612" authority="Sakai, 1963" authorityName="Sakai" authorityYear="1963" box="[490,970,1543,1569]" class="Malacostraca" family="Dynomenidae" genus="Paradynomene" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B922C97CFF840C13FE30F9CD0B00B613" box="[490,813,1543,1568]" italics="true" pageId="1" pageNumber="2">Paradynomene tuberculata</emphasis>
<bibRefCitation id="EFC7689FFF840C13FCE3F9CD0BE7B612" author="Sakai" box="[825,970,1543,1569]" pageId="1" pageNumber="24" refString="Sakai, T. (1963) Description of two new genera and fourteen new species of Japanese crabs from the collection of his Majesty the Emperor of Japan. Crustaceana, 5 (3), 213 - 233." type="journal article" year="1963">Sakai, 1963</bibRefCitation>
</taxonomicName>
, by monotypy (gender feminine).
</paragraph>
<paragraph id="8BE9156EFF840C13FEE2F99C0C27B6AA" blockId="1.[264,1324,742,1889]" pageId="1" pageNumber="2">
<emphasis id="B922C97CFF840C13FEE2F99C0A24B643" bold="true" box="[312,521,1622,1648]" pageId="1" pageNumber="2">Included species</emphasis>
.
<taxonomicName id="4C566EEDFF840C13FDC6F99D0BD3B642" authority="Sakai, 1963" authorityName="Sakai" authorityYear="1963" box="[540,1022,1623,1649]" class="Malacostraca" family="Dynomenidae" genus="Paradynomene" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B922C97CFF840C13FDC6F99D0B4DB643" box="[540,864,1623,1648]" italics="true" pageId="1" pageNumber="2">Paradynomene tuberculata</emphasis>
<bibRefCitation id="EFC7689FFF840C13FCB6F99D0BD3B642" author="Sakai" box="[876,1022,1623,1649]" pageId="1" pageNumber="24" refString="Sakai, T. (1963) Description of two new genera and fourteen new species of Japanese crabs from the collection of his Majesty the Emperor of Japan. Crustaceana, 5 (3), 213 - 233." type="journal article" year="1963">Sakai, 1963</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C566EEDFF840C13FBD5F99D0C9CB643" box="[1039,1201,1623,1648]" class="Malacostraca" family="Dynomenidae" genus="Paradynomene" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="quasimodo" status="sp. nov.">
<emphasis id="B922C97CFF840C13FBD5F99D0C9CB643" box="[1039,1201,1623,1648]" italics="true" pageId="1" pageNumber="2">P. quasimodo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2117407FF840C13FB64F99D0D2EB642" box="[1214,1283,1623,1649]" pageId="1" pageNumber="2" rank="species">n. sp.</taxonomicNameLabel>
,
<taxonomicName id="4C566EEDFF840C13FACFF99D097BB6AB" class="Malacostraca" family="Dynomenidae" genus="Paradynomene" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="demon" status="sp. nov.">
<emphasis id="B922C97CFF840C13FACFF99D097BB6AB" italics="true" pageId="1" pageNumber="2">P. demon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2117407FF840C13FE87F9B509B3B6AA" box="[349,414,1663,1689]" pageId="1" pageNumber="2" rank="species">n. sp.</taxonomicNameLabel>
,
<taxonomicName id="4C566EEDFF840C13FE71F9B50A3EB6AB" box="[427,531,1663,1688]" class="Malacostraca" family="Dynomenidae" genus="Paradynomene" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="diablo" status="sp. nov.">
<emphasis id="B922C97CFF840C13FE71F9B50A3EB6AB" box="[427,531,1663,1688]" italics="true" pageId="1" pageNumber="2">P. diablo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2117407FF840C13FDC0F9B50A77B6AA" box="[538,602,1663,1689]" pageId="1" pageNumber="2" rank="species">n. sp.</taxonomicNameLabel>
,
<emphasis id="B922C97CFF840C13FDBDF9B50AECB6AB" box="[615,705,1663,1688]" italics="true" pageId="1" pageNumber="2">P. t e u f e l</emphasis>
n. sp., and
<emphasis id="B922C97CFF840C13FC91F9B50BEDB6AB" box="[843,960,1663,1688]" italics="true" pageId="1" pageNumber="2">P. ro t u n d a</emphasis>
n. sp.
</paragraph>
</subSubSection>
<subSubSection id="C34C46E5FF840C10FEE2F96C0CDDB325" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="description">
<paragraph id="8BE9156EFF840C10FEE2F96C0CDDB325" blockId="1.[264,1324,742,1889]" lastBlockId="2.[264,1324,284,790]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<emphasis id="B922C97CFF840C13FEE2F96C09F9B6F3" bold="true" box="[312,468,1702,1728]" pageId="1" pageNumber="2">Terminology</emphasis>
. A feature of the species of
<taxonomicName id="4C566EEDFF840C13FCFBF96D0BFFB6F3" box="[801,978,1703,1728]" class="Malacostraca" family="Dynomenidae" genus="Paradynomene" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B922C97CFF840C13FCFBF96D0BFFB6F3" box="[801,978,1703,1728]" italics="true" pageId="1" pageNumber="2">Paradynomene</emphasis>
</taxonomicName>
is the granular surface of the carapace and the presence of areolae that may be tipped by a distinct group of granules. These areolae range from slightly swollen areas not greatly elevated, to steep­sided, volcano­like mounds, tipped with a distinct group of granules. We have used the term &quot;tubercle&quot; to refer to those areolae that are clearly peaked with the highest elevation being marked by a group of granules. On the figures, these tubercles are marked by small circles. Thus for some species the number of swollen areas on the carapace may be greater than the given number of tubercles. We have used the terms &quot;gastric&quot;, &quot;cardiac&quot;, &quot;branchial,&quot; and 'intestinal&quot; as used by Rathbun (1930) in describing the location of these tubercles (
<figureCitation id="136D09EBFF870C10FECBFE760974B1E5" box="[273,345,444,470]" captionStart="FIGURE 1" captionStartId="2.[264,367,1560,1584]" captionTargetBox="[340,1195,931,1542]" captionTargetId="figure@2.[305,1282,887,1567]" captionTargetPageId="2" captionText="FIGURE 1. Carapace of Paradynomene showing major regions and the terminology used in the text." httpUri="https://zenodo.org/record/169349/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
). Another character that requires careful attention is the number of anterolateral teeth. The problem arises because the anterolateral margins are not well­defined and projections on, or placed higher, but overhanging the margin, are variable in size and sometimes the left side can differ from the right. A distinction needs to be made between teeth, which are granulated projections on the anterolateral margin, and granules, which are smaller and form part of the background carapace ornamentation. The number of teeth that are given in the text include only the maximum number of distinct projections and do not include the smaller granules. Posterolateral teeth are those on the margin posterior to the maximum carapace width. The terminology otherwise follows that of
<bibRefCitation id="EFC7689FFF870C10FBE4FD360CC1B325" author="McLay" box="[1086,1260,764,790]" pageId="2" pageNumber="24" refString="McLay C. L. (1999) Crustacea Decapoda: Revision of the family Dynomenidae, In: A. Crosnier (ed.), Resultats des Campagnes MUSORSTOM Vo l u m e 2 0. Memoires du Museum national d'Histoire naturelle (Paris), 180, 427 - 569, figs. 1 - 40." type="journal article" year="1999">McLay (1999)</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="DF2945E6FF870C10FED2F9D2091AB661" httpUri="https://zenodo.org/record/169349/files/figure.png" pageId="2" pageNumber="3" targetBox="[340,1195,931,1542]" targetPageId="2">
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<emphasis id="B922C97CFF870C10FED2F9D209A1B603" bold="true" box="[264,396,1560,1584]" pageId="2" pageNumber="3">FIGURE 1.</emphasis>
Carapace of
<taxonomicName id="4C566EEDFF870C10FDFBF9D20AEFB61C" box="[545,706,1560,1583]" class="Malacostraca" family="Dynomenidae" genus="Paradynomene" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B922C97CFF870C10FDFBF9D20AEFB61C" box="[545,706,1560,1583]" italics="true" pageId="2" pageNumber="3">Paradynomene</emphasis>
</taxonomicName>
showing major regions and the terminology used in the text.
</paragraph>
</caption>
<subSubSection id="C34C46E5FF870C11FEE2F96209DCB2DD" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="discussion">
<paragraph id="8BE9156EFF870C11FEE2F96209DCB2DD" blockId="2.[264,1323,1704,1890]" lastBlockId="3.[264,1324,284,750]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<emphasis id="B922C97CFF870C10FEE2F9620987B6F1" bold="true" box="[312,426,1704,1730]" pageId="2" pageNumber="3">Remarks</emphasis>
. The genus
<taxonomicName id="4C566EEDFF870C10FDE2F9630AC4B6F1" box="[568,745,1705,1730]" class="Malacostraca" family="Dynomenidae" genus="Paradynomene" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B922C97CFF870C10FDE2F9630AC4B6F1" box="[568,745,1705,1730]" italics="true" pageId="2" pageNumber="3">Paradynomene</emphasis>
</taxonomicName>
was erected by
<bibRefCitation id="EFC7689FFF870C10FC7FF9620C11B6F1" author="Sakai" box="[933,1084,1704,1730]" pageId="2" pageNumber="24" refString="Sakai, T. (1963) Description of two new genera and fourteen new species of Japanese crabs from the collection of his Majesty the Emperor of Japan. Crustaceana, 5 (3), 213 - 233." type="journal article" year="1963">Sakai (1963)</bibRefCitation>
for an unusual male dynomenid crab collected from Sagami Bay at
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.
</specimenCount>
Instead of being transversely oval or polygonal and slightly convex, like species of
<taxonomicName id="4C566EEDFF870C10FCE8F9330CACB721" authority="Desmarest, 1823" authorityName="Desmarest" authorityYear="1823" box="[818,1153,1784,1810]" class="Malacostraca" family="Dynomenidae" genus="Dynomene" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B922C97CFF870C10FCE8F9330B82B721" box="[818,943,1785,1810]" italics="true" pageId="2" pageNumber="3">Dynomene</emphasis>
<bibRefCitation id="EFC7689FFF870C10FC62F9320CACB721" author="Desmarest" box="[952,1153,1784,1810]" pageId="2" pageNumber="24" refString="Desmarest A. G. (1823) Crustaces Malacostraces, In: Dictionnaire des Sciences naturelles, 28, 138 - 425; atlas, vol. 4, pl. 1 - 58. Strasbourg &amp; Paris, F. G. Levrault et Le Normant." type="journal article" year="1823">Desmarest, 1823</bibRefCitation>
</taxonomicName>
, this crab was subquadrangular and markedly convex with tuberculate areolae, and the &quot;face&quot; more closely resembled dromiids belonging to
<taxonomicName id="4C566EEDFF870C10FD2BF8830C19B751" authority="McLay, 1993" authorityName="McLay" authorityYear="1993" box="[753,1076,1864,1890]" class="Malacostraca" family="Dromiidae" genus="Epigodromia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B922C97CFF870C10FD2BF8830BA6B751" box="[753,907,1865,1890]" italics="true" pageId="2" pageNumber="3">Epigodromia</emphasis>
McLay, 1993
</taxonomicName>
.
<bibRefCitation id="EFC7689FFF870C10FB9AF8820CFAB751" author="Sakai" box="[1088,1239,1864,1890]" pageId="2" pageNumber="24" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha Ltd, Tokyo. [In 3 volumes: (1) English text, xxxix + 773 pp., figs., 1 - 379, (2) plates, 16 p, pls. 1 - 251, (3) Japanese text, 461 p., figs 1 - 2]." type="book chapter" year="1976">Sakai (1976)</bibRefCitation>
subsequently reported on a second specimen (a female) from
<collectingCountry id="F34155FEFF860C11FC79FED60BC4B105" box="[931,1001,284,310]" name="Japan" pageId="3" pageNumber="4">Japan</collectingCountry>
. For many years they were the only known specimens. In his revision of the
<taxonomicName id="4C566EEDFF860C11FC91FE8E0C84B16D" authority="McLay (1999)" authorityName="McLay" authorityYear="1999" box="[843,1193,324,350]" class="Malacostraca" family="Dynomenidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">
Dynomenidae,
<bibRefCitation id="EFC7689FFF860C11FC25FE8E0C84B16D" author="McLay" box="[1023,1193,324,350]" pageId="3" pageNumber="24" refString="McLay C. L. (1999) Crustacea Decapoda: Revision of the family Dynomenidae, In: A. Crosnier (ed.), Resultats des Campagnes MUSORSTOM Vo l u m e 2 0. Memoires du Museum national d'Histoire naturelle (Paris), 180, 427 - 569, figs. 1 - 40." type="journal article" year="1999">McLay (1999)</bibRefCitation>
</taxonomicName>
assembled a collection of
<specimenCount id="9D50DEE7FF860C11FE6DFEA60A78B1B5" box="[439,597,364,390]" pageId="3" pageNumber="4" type="generic">19 specimens</specimenCount>
from
<collectingCountry id="F34155FEFF860C11FD47FEA60B7EB1B5" box="[669,851,364,390]" name="New Caledonia" pageId="3" pageNumber="4">New Caledonia</collectingCountry>
(including the
<collectingCountry id="F34155FEFF860C11FBDFFEA60C4FB1B5" box="[1029,1122,364,390]" name="Norfolk Island" pageId="3" pageNumber="4">Norfolk</collectingCountry>
Ridge), Chesterfield and Loyalty islands,
<collectingCountry id="F34155FEFF860C11FD97FE5E0AEFB19D" box="[589,706,404,430]" name="Indonesia" pageId="3" pageNumber="4">Indonesia</collectingCountry>
,
<collectingCountry id="F34155FEFF860C11FD09FE5E0B33B19D" box="[723,798,404,430]" name="Guam" pageId="3" pageNumber="4">Guam</collectingCountry>
, and the Gulf of Aden, all of which were referred to
<taxonomicName id="4C566EEDFF860C11FE4AFE760A1BB1E6" box="[400,566,444,469]" class="Malacostraca" family="Dynomenidae" genus="Paradynomene" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B922C97CFF860C11FE4AFE760A1BB1E6" box="[400,566,444,469]" italics="true" pageId="3" pageNumber="4">P. tuberculata</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFC7689FFF860C11FD92FE760AD8B1E5" author="McLay" box="[584,757,444,470]" pageId="3" pageNumber="24" refString="McLay, C. L. (2001) Dynomenidae and Dromiidae (Decapoda, Brachyura) from Guam, Philippine Islands, Tonga and Samoa. Zoosystema, 23 (4), 807 - 856." type="journal article" year="2001">McLay (2001)</bibRefCitation>
subsequently recorded further specimens from
<collectingCountry id="F34155FEFF860C11FED2FE2E0979B1CD" box="[264,340,484,510]" name="Guam" pageId="3" pageNumber="4">Guam</collectingCountry>
. Many of the locations were represented by only a few specimens, often small, making precise species determinations difficult. The recent specimens from the
<collectingCountry id="F34155FEFF860C11FB7DFDC60D06B215" box="[1191,1323,524,550]" name="Philippines" pageId="3" pageNumber="4">Philippines</collectingCountry>
facilitated a reassessment of the species definitions and allowed us to establish a narrower concept of
<taxonomicName id="4C566EEDFF860C11FE43FD960A6EB246" box="[409,579,604,629]" class="Malacostraca" family="Dynomenidae" genus="Paradynomene" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B922C97CFF860C11FE43FD960A6EB246" box="[409,579,604,629]" italics="true" pageId="3" pageNumber="4">P. tuberculata</emphasis>
</taxonomicName>
and to argue that the genus
<taxonomicName id="4C566EEDFF860C11FC64FD960C5DB246" box="[958,1136,604,629]" class="Malacostraca" family="Dynomenidae" genus="Paradynomene" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B922C97CFF860C11FC64FD960C5DB246" box="[958,1136,604,629]" italics="true" pageId="3" pageNumber="4">Paradynomene</emphasis>
</taxonomicName>
should not be regarded as monotypic. This revision means that six species are now recognized, five of which are new. Of the specimens studied by
<bibRefCitation id="EFC7689FFF860C11FCC0FD660BE5B2F5" author="McLay" box="[794,968,684,710]" pageId="3" pageNumber="24" refString="McLay C. L. (1999) Crustacea Decapoda: Revision of the family Dynomenidae, In: A. Crosnier (ed.), Resultats des Campagnes MUSORSTOM Vo l u m e 2 0. Memoires du Museum national d'Histoire naturelle (Paris), 180, 427 - 569, figs. 1 - 40." type="journal article" year="1999">McLay (1999)</bibRefCitation>
, only a few can be referred to
<emphasis id="B922C97CFF860C11FED2FD1E09C4B2DE" box="[264,489,724,749]" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="4C566EEDFF860C11FED2FD1E0986B2DE" box="[264,427,724,749]" class="Malacostraca" family="Dynomenidae" genus="Paradynomene" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" sensu="stricto" species="tuberculata">P. tuberculata</taxonomicName>
<taxonomicNameLabel id="A2117407FF860C11FE68FD1E09C4B2DE" box="[434,489,724,749]" pageId="3" pageNumber="4" sensu="stricto">s. str</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>