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<document id="1C58E51E52E0914CC62CD3C954F0FCB1" ID-DOI="10.11646/zootaxa.5323.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8204101" ID-ZooBank="0ABC4CFF-9D2A-449A-B7C2-CFE16DF16004" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1690887012857" checkinUser="plazi" docAuthor="Soares, Matheus M. M., Runyon, Justin B., Capellari, Renato S. &amp; Ale-Rocha, Rosaly" docDate="2023" docId="03E62055FF900014FF7BF968FEF534FA" docLanguage="en" docName="zootaxa.5323.2.1.pdf" docOrigin="Zootaxa 5323 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5323.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pelastoneurus arcuatus Loew 1861" docType="treatment" docVersion="4" lastPageNumber="153" masterDocId="FFDF582DFF910016FFECFFBDFFDB3247" masterDocTitle="Review of Paraclius arcuatus species-group (Diptera, Dolichopodidae) with description of two new species" masterLastPageNumber="182" masterPageNumber="151" pageNumber="152" updateTime="1691081561564" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C1AA561ADAEAE1C56D116C4C943E3152">Review of Paraclius arcuatus species-group (Diptera, Dolichopodidae) with description of two new species</mods:title>
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<mods:namePart id="8908E40DA4C01E313C59BDE81A60495E">Soares, Matheus M. M.</mods:namePart>
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<mods:namePart id="45C8DFF7C45D736D782502303A42FE71">Runyon, Justin B.</mods:namePart>
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<mods:affiliation id="252BE69A4DA4E1D56ACB00603B1E2E71">Rocky Mountain Research Station, USDA Forest Service, 1648 S. 7 th Avenue, Bozeman, Montana 59717, USA. &amp; justin. runyon @ usda. gov; https: // orcid. org / 0000 - 0002 - 0271 - 0511</mods:affiliation>
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<mods:namePart id="E59050FFE756515E29FA2131B836BF74">Capellari, Renato S.</mods:namePart>
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<mods:affiliation id="F915D44CD11564352F049D4823207612">Instituto Federal do Triângulo Mineiro, Campus Uberaba, Rua João Batista Ribeiro 4000, Distrito Industrial II, Uberaba, Minas Gerais 38064 - 790, Brazil. &amp; rscapellari @ gmail. com; https: // orcid. org / 0000 - 0002 - 8410 - 9235</mods:affiliation>
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<mods:namePart id="73C4FB644CCA0C70AA93BC1683EC88B8">Ale-Rocha, Rosaly</mods:namePart>
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<mods:affiliation id="F82EE4A9924BC9587B992F53E9818572">Coordenação de Biodiversidade (COBIO, INPA), Av. André Araújo, 2936, Petrópolis, CEP 69067 - 375, Manaus, Amazonas, Brazil. &amp; alerocha @ inpa. gov. br; https: // orcid. org / 0000 - 0001 - 9874 - 9770</mods:affiliation>
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<paragraph id="8BF09143FF900017FF7BF968FDF234B7" blockId="1.[151,553,1749,1776]" box="[151,553,1749,1776]" pageId="1" pageNumber="152">
<heading id="D0B8262FFF900017FF7BF968FDF234B7" bold="true" box="[151,553,1749,1776]" fontSize="11" level="1" pageId="1" pageNumber="152" reason="1">
<emphasis id="B93B4D51FF900017FF7BF968FDF234B7" bold="true" box="[151,553,1749,1776]" pageId="1" pageNumber="152">
The
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<emphasis id="B93B4D51FF900017FF23F96BFEE234B7" bold="true" box="[207,313,1750,1776]" italics="true" pageId="1" pageNumber="152">arcuatus</emphasis>
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-group of
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<emphasis id="B93B4D51FF900017FE59F968FDF234B7" bold="true" box="[437,553,1749,1776]" italics="true" pageId="1" pageNumber="152">Paraclius</emphasis>
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<paragraph id="8BF09143FF900014FF7BF8A0FA4733CE" blockId="1.[151,1437,1821,2028]" lastBlockId="2.[151,1437,150,1725]" lastPageId="2" lastPageNumber="153" pageId="1" pageNumber="152">
<emphasis id="B93B4D51FF900017FF7BF8A0FE1E3570" bold="true" box="[151,453,1821,1847]" pageId="1" pageNumber="152">Diagnosis (males). Head.</emphasis>
Face gradually narrowing below (funnel-shaped), the narrowest point slightly wider than the mid ocellus (
<emphasis id="B93B4D51FF900017FE62F8FFFE6D351C" box="[398,438,1858,1883]" italics="true" pageId="1" pageNumber="152">e.g.</emphasis>
<figureCitation id="13748DC6FF900017FE52F8FCFDC1351C" box="[446,538,1857,1883]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). AB. Habitus, lateral and dorsal views respectively; C. Labels; DE. Head, anterior and lateral views, respectively; FH. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="1" pageNumber="152">Figs 1D</figureCitation>
,
<figureCitation id="13748DC6FF900017FDC4F8FCFD90351C" box="[552,587,1857,1883]" captionStart="FIGURE 2" captionStartId="5.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-18@5.[151,1436,181,1730]" captionTargetPageId="5" captionText="FIGURE 2. Paraclius arcuatus (Loew, 1861), A, D, E, F male specimen from Honduras (CNC), B, C male specimen from Mexico (NMNH). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D.Abdomen and hypopyium, lateral view; E. Left antenna, inner view; F. Left wing.Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R2+3 = second branch of radius; R = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.8204107" httpUri="https://zenodo.org/record/8204107/files/figure.png" pageId="1" pageNumber="152">2C</figureCitation>
,
<figureCitation id="13748DC6FF900017FDB4F8FCFDA0351C" box="[600,635,1857,1883]" captionStart="FIGURE 7" captionStartId="11.[151,250,1939,1963]" captionTargetBox="[158,1430,771,1915]" captionTargetId="figure-209@11.[158,1430,771,1915]" captionTargetPageId="11" captionText="FIGURE 7. Paraclius angustipennis Van Duzee, 1929, male holotype (NMNH). AB. Habitus, lateral and dorsal views, respectively; C. Abdominal segments 5 and 6; D. Labels; EF. Head, anterior and anterolateral views, respectively. Images B, C, E originally lacked scale bars. Photographs by Alyssa Seemann, NMNH." figureDoi="http://doi.org/10.5281/zenodo.8204125" httpUri="https://zenodo.org/record/8204125/files/figure.png" pageId="1" pageNumber="152">7E</figureCitation>
). Clypeus short, about 1/4 as long as face (
<emphasis id="B93B4D51FF900017FB89F8FFFB56351C" box="[1125,1165,1858,1883]" italics="true" pageId="1" pageNumber="152">e.g.</emphasis>
<figureCitation id="13748DC6FF900017FB79F8FCFB2A351C" box="[1173,1265,1857,1883]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). AB. Habitus, lateral and dorsal views respectively; C. Labels; DE. Head, anterior and lateral views, respectively; FH. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="1" pageNumber="152">Figs 1D</figureCitation>
,
<figureCitation id="13748DC6FF900017FB13F8FCFAF9351C" box="[1279,1314,1857,1883]" captionStart="FIGURE 2" captionStartId="5.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-18@5.[151,1436,181,1730]" captionTargetPageId="5" captionText="FIGURE 2. Paraclius arcuatus (Loew, 1861), A, D, E, F male specimen from Honduras (CNC), B, C male specimen from Mexico (NMNH). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D.Abdomen and hypopyium, lateral view; E. Left antenna, inner view; F. Left wing.Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R2+3 = second branch of radius; R = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.8204107" httpUri="https://zenodo.org/record/8204107/files/figure.png" pageId="1" pageNumber="152">2C</figureCitation>
,
<figureCitation id="13748DC6FF900017FAC3F8FCFA8A351C" box="[1327,1361,1857,1883]" captionStart="FIGURE 8" captionStartId="12.[151,250,1973,1997]" captionTargetBox="[151,1436,489,1950]" captionTargetId="figure-189@12.[151,1436,489,1950]" captionTargetPageId="12" captionText="FIGURE 8. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D. Abdomen, lateral view; E. Left antenna, outer view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204127" httpUri="https://zenodo.org/record/8204127/files/figure.png" pageId="1" pageNumber="152">8C</figureCitation>
,
<figureCitation id="13748DC6FF900017FAB2F8FCFA54351C" box="[1374,1423,1857,1883]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., AE male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; CD. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="1" pageNumber="152">11C</figureCitation>
). Palpus oval, short, with 1 strong apical seta, shorter than palpus (absent in
<taxonomicName id="4C4FEAC0FF900017FC0CF8DBFBE53538" authority="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha, 2023" authorityName="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha" authorityYear="2023" box="[992,1086,1893,1920]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="152" phylum="Arthropoda" rank="species" species="xibun" status="sp. nov.">
<emphasis id="B93B4D51FF900017FC0CF8DBFC2A3538" box="[992,1009,1894,1919]" italics="true" pageId="1" pageNumber="152">P</emphasis>
.
<emphasis id="B93B4D51FF900017FBECF8D8FBE53538" box="[1024,1086,1893,1919]" italics="true" pageId="1" pageNumber="152">xibun</emphasis>
</taxonomicName>
<emphasis id="B93B4D51FF900017FBA9F8D8FB443538" bold="true" box="[1093,1183,1893,1919]" pageId="1" pageNumber="152">
<taxonomicNameLabel id="A208F02AFF900017FBA9F8D8FB443538" box="[1093,1183,1893,1919]" pageId="1" pageNumber="152" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). Antenna dark brown to black; postpedicel suboval, subtriangular or subrectangular; arista-like stylus two-segmented, shorter than eye height, first segment usually arched, second segment long, about 23X longer than first segment and with dense and short microtrichia (
<emphasis id="B93B4D51FF900017FE81F86FFE4E35AC" box="[365,405,2002,2027]" italics="true" pageId="1" pageNumber="152">e.g.</emphasis>
<figureCitation id="13748DC6FF900017FE70F86CFE2835AC" box="[412,499,2001,2027]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). AB. Habitus, lateral and dorsal views respectively; C. Labels; DE. Head, anterior and lateral views, respectively; FH. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="1" pageNumber="152">Figs 1E</figureCitation>
,
<figureCitation id="13748DC6FF900017FE12F86CFDC435AC" box="[510,543,2001,2027]" captionStart="FIGURE 2" captionStartId="5.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-18@5.[151,1436,181,1730]" captionTargetPageId="5" captionText="FIGURE 2. Paraclius arcuatus (Loew, 1861), A, D, E, F male specimen from Honduras (CNC), B, C male specimen from Mexico (NMNH). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D.Abdomen and hypopyium, lateral view; E. Left antenna, inner view; F. Left wing.Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R2+3 = second branch of radius; R = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.8204107" httpUri="https://zenodo.org/record/8204107/files/figure.png" pageId="1" pageNumber="152">2E</figureCitation>
,
<figureCitation id="13748DC6FF900017FDC6F86CFD9635AC" box="[554,589,2001,2027]" captionStart="FIGURE 8" captionStartId="12.[151,250,1973,1997]" captionTargetBox="[151,1436,489,1950]" captionTargetId="figure-189@12.[151,1436,489,1950]" captionTargetPageId="12" captionText="FIGURE 8. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D. Abdomen, lateral view; E. Left antenna, outer view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204127" httpUri="https://zenodo.org/record/8204127/files/figure.png" pageId="1" pageNumber="152">8E</figureCitation>
).
<emphasis id="B93B4D51FF900017FDB3F86CFD6435AC" bold="true" box="[607,703,2001,2027]" pageId="1" pageNumber="152">Thorax.</emphasis>
Mesonotum with 5 pairs of strong dorsocentral setae; acrostichals short, biseriate, ending at level of the fourth posterior dorsocentral seta (
<emphasis id="B93B4D51FF930014FC5FFF2AFC0032F7" box="[947,987,151,176]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FC0DFF2AFBE132F6" box="[993,1082,151,177]" captionStart="FIGURE 8" captionStartId="12.[151,250,1973,1997]" captionTargetBox="[151,1436,489,1950]" captionTargetId="figure-189@12.[151,1436,489,1950]" captionTargetPageId="12" captionText="FIGURE 8. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D. Abdomen, lateral view; E. Left antenna, outer view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204127" httpUri="https://zenodo.org/record/8204127/files/figure.png" pageId="2" pageNumber="153">Figs 8B</figureCitation>
,
<figureCitation id="13748DC6FF930014FBA9FF2AFBAD32F6" box="[1093,1142,151,177]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., AE male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; CD. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="2" pageNumber="153">11B</figureCitation>
).
<emphasis id="B93B4D51FF930014FB64FF2BFB0B32F7" bold="true" box="[1160,1232,150,176]" pageId="2" pageNumber="153">Wing.</emphasis>
Long and narrow; anal lobe barely developed; vein M
<subScript id="17CB9306FF930014FDCFFF70FDF0329C" attach="left" box="[547,555,205,219]" fontSize="6" pageId="2" pageNumber="153">1</subScript>
strongly bowed towards R
<subScript id="17CB9306FF930014FCB6FF70FCAF329C" attach="left" box="[858,884,205,219]" fontSize="6" pageId="2" pageNumber="153">4+5</subScript>
, weakly to strongly concave posteriorly; vein M
<subScript id="17CB9306FF930014FA78FF70FA47329C" attach="left" box="[1428,1436,205,219]" fontSize="6" pageId="2" pageNumber="153">1</subScript>
ending before distal portion of wing apex; vein M
<subScript id="17CB9306FF930014FD2EFF4CFD1132B8" attach="left" box="[706,714,241,255]" fontSize="6" pageId="2" pageNumber="153">4</subScript>
longer than dm-m (
<emphasis id="B93B4D51FF930014FC45FF62FC0A32BF" box="[937,977,223,248]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FC34FF62FBF332BE" box="[984,1064,223,249]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). AB. Habitus, lateral and dorsal views respectively; C. Labels; DE. Head, anterior and lateral views, respectively; FH. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="2" pageNumber="153">Figs 1I</figureCitation>
,
<figureCitation id="13748DC6FF930014FBD8FF62FB8932BE" box="[1076,1106,223,249]" captionStart="FIGURE 2" captionStartId="5.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-18@5.[151,1436,181,1730]" captionTargetPageId="5" captionText="FIGURE 2. Paraclius arcuatus (Loew, 1861), A, D, E, F male specimen from Honduras (CNC), B, C male specimen from Mexico (NMNH). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D.Abdomen and hypopyium, lateral view; E. Left antenna, inner view; F. Left wing.Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R2+3 = second branch of radius; R = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.8204107" httpUri="https://zenodo.org/record/8204107/files/figure.png" pageId="2" pageNumber="153">2F</figureCitation>
,
<figureCitation id="13748DC6FF930014FBB1FF62FBA532BE" box="[1117,1150,223,249]" captionStart="FIGURE 8" captionStartId="12.[151,250,1973,1997]" captionTargetBox="[151,1436,489,1950]" captionTargetId="figure-189@12.[151,1436,489,1950]" captionTargetPageId="12" captionText="FIGURE 8. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D. Abdomen, lateral view; E. Left antenna, outer view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204127" httpUri="https://zenodo.org/record/8204127/files/figure.png" pageId="2" pageNumber="153">8F</figureCitation>
).
<emphasis id="B93B4D51FF930014FB7DFF62FB0B32BE" bold="true" box="[1169,1232,223,249]" pageId="2" pageNumber="153">Legs.</emphasis>
Legs unmodified, only ventral surface of tarsus I covered with weak whitish pile (
<emphasis id="B93B4D51FF930014FCB4FEBEFC5B335B" box="[856,896,259,284]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
,
<taxonomicName id="4C4FEAC0FF930014FC61FEBEFBD6335B" authorityName="Loew" authorityYear="1861" box="[909,1037,259,285]" class="Insecta" family="Dolichopodidae" genus="Pelastoneurus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="B93B4D51FF930014FC61FEBEFC45335B" box="[909,926,259,284]" italics="true" pageId="2" pageNumber="153">P</emphasis>
.
<emphasis id="B93B4D51FF930014FC40FEBEFBD6335B" box="[940,1037,259,284]" italics="true" pageId="2" pageNumber="153">arcuatus</emphasis>
</taxonomicName>
). Femur I with anteroventral row of short setae increasing in length at apex; femur II with anteroventral row of setae, ending in 23 more conspicuous setae and with 1 strong anterior preapical seta at 5/6; femur III laterally compressed, with anteroventral and dorsal rows of setae, anteroventral row increasing in length towards apex, and 1 strong anterior preapical seta at 5/6.
</paragraph>
<paragraph id="8BF09143FF930014FF7BFE2FFCD73682" blockId="2.[151,1437,150,1725]" pageId="2" pageNumber="153">
<emphasis id="B93B4D51FF930014FF7BFE2FFEC833EB" bold="true" box="[151,275,402,428]" pageId="2" pageNumber="153">Abdomen.</emphasis>
Tergites 15 setose, tergite 6 either with short or long and strong lateral setae (
<emphasis id="B93B4D51FF930014FB97FE2EFB7833EB" box="[1147,1187,403,428]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FB45FE2EFAD833EA" box="[1193,1283,403,429]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). AB. Habitus, lateral and dorsal views respectively; C. Labels; DE. Head, anterior and lateral views, respectively; FH. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="2" pageNumber="153">Figs 1H</figureCitation>
,
<figureCitation id="13748DC6FF930014FAE3FE2EFAE833EA" box="[1295,1331,403,429]" captionStart="FIGURE 8" captionStartId="12.[151,250,1973,1997]" captionTargetBox="[151,1436,489,1950]" captionTargetId="figure-189@12.[151,1436,489,1950]" captionTargetPageId="12" captionText="FIGURE 8. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D. Abdomen, lateral view; E. Left antenna, outer view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204127" httpUri="https://zenodo.org/record/8204127/files/figure.png" pageId="2" pageNumber="153">8D</figureCitation>
,
<figureCitation id="13748DC6FF930014FAD2FE2EFA8633EA" box="[1342,1373,403,429]" captionStart="FIGURE 9" captionStartId="14.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1615]" captionTargetId="figure-20@14.[151,1436,181,1615]" captionTargetPageId="14" captionText="FIGURE 9. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Abdomen, ventral view; B. Hypopygium, left lateral view; C. Internal appendages, left lateral view; D. Apical part of phallus; E. Apical edge of epandrium; F. Tergite 6, lateral view; G. Right cercus, outer view; H. Left ventral surstylus, outer view; I. Hypandrium, ventral view; JK. Surstyli, left and right lateral views, respectively. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204131" httpUri="https://zenodo.org/record/8204131/files/figure.png" pageId="2" pageNumber="153">9F</figureCitation>
,
<figureCitation id="13748DC6FF930014FA8BFE2EFA4F33EA" box="[1383,1428,403,429]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., AE male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; CD. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="2" pageNumber="153">11E</figureCitation>
) (bare in
<taxonomicName id="4C4FEAC0FF930014FF15FE0AFEA03397" authorityName="Loew" authorityYear="1861" box="[249,379,439,465]" class="Insecta" family="Dolichopodidae" genus="Pelastoneurus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="B93B4D51FF930014FF15FE0AFED13397" box="[249,266,439,464]" italics="true" pageId="2" pageNumber="153">P</emphasis>
.
<emphasis id="B93B4D51FF930014FEF6FE0AFEA03397" box="[282,379,439,464]" italics="true" pageId="2" pageNumber="153">arcuatus</emphasis>
</taxonomicName>
(
<figureCitation id="13748DC6FF930014FE60FE0AFE3C3396" box="[396,487,439,465]" captionStart="FIGURE 2" captionStartId="5.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-18@5.[151,1436,181,1730]" captionTargetPageId="5" captionText="FIGURE 2. Paraclius arcuatus (Loew, 1861), A, D, E, F male specimen from Honduras (CNC), B, C male specimen from Mexico (NMNH). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D.Abdomen and hypopyium, lateral view; E. Left antenna, inner view; F. Left wing.Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R2+3 = second branch of radius; R = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.8204107" httpUri="https://zenodo.org/record/8204107/files/figure.png" pageId="2" pageNumber="153">Fig. 2D</figureCitation>
)), segment 7 forming a long (usually as long as hypopygium) and well sclerotized peduncle; sternite 6 plate-like, weakly connected posteriorly to tergite 6 (
<emphasis id="B93B4D51FF930014FC09FE66FBD633B3" box="[997,1037,475,500]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FBFBFE66FBAE33B2" box="[1047,1141,475,501]" captionStart="FIGURE 3" captionStartId="6.[151,250,1458,1482]" captionTargetBox="[151,1436,181,1435]" captionTargetId="figure-20@6.[151,1436,181,1435]" captionTargetPageId="6" captionText="FIGURE 3. Paraclius arcuatus (Loew, 1861), identified male specimens from Honduras (CNC). A. Abdomen, ventral view; B. Tergite 7 and Sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Internal appendages, left lateral view; E. Hypandrium, ventral view; F. Dorsal lobe of surstylus, outer view; G. Ventral lobe of surstylus, outer view. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct br = proctiger brush; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204111" httpUri="https://zenodo.org/record/8204111/files/figure.png" pageId="2" pageNumber="153">Figs 3A</figureCitation>
,
<figureCitation id="13748DC6FF930014FB68FE66FB7233B2" box="[1156,1193,475,501]" captionStart="FIGURE 9" captionStartId="14.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1615]" captionTargetId="figure-20@14.[151,1436,181,1615]" captionTargetPageId="14" captionText="FIGURE 9. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Abdomen, ventral view; B. Hypopygium, left lateral view; C. Internal appendages, left lateral view; D. Apical part of phallus; E. Apical edge of epandrium; F. Tergite 6, lateral view; G. Right cercus, outer view; H. Left ventral surstylus, outer view; I. Hypandrium, ventral view; JK. Surstyli, left and right lateral views, respectively. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204131" httpUri="https://zenodo.org/record/8204131/files/figure.png" pageId="2" pageNumber="153">9A</figureCitation>
,
<figureCitation id="13748DC6FF930014FB5AFE66FB3033B2" box="[1206,1259,475,501]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="2" pageNumber="153">12A</figureCitation>
). Hypopygium: Epandrium narrow, about 2.53X longer than high, with lateral ridge forming an acute apicoventral projection. Apicoventral epandrial lobe not developed, but recognizable by two apicoventral epandrial setae (
<emphasis id="B93B4D51FF930014FAEAFD9EFAF5307B" box="[1286,1326,547,572]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FAD5FD9EFA4C307A" box="[1337,1431,547,573]" captionStart="FIGURE 3" captionStartId="6.[151,250,1458,1482]" captionTargetBox="[151,1436,181,1435]" captionTargetId="figure-20@6.[151,1436,181,1435]" captionTargetPageId="6" captionText="FIGURE 3. Paraclius arcuatus (Loew, 1861), identified male specimens from Honduras (CNC). A. Abdomen, ventral view; B. Tergite 7 and Sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Internal appendages, left lateral view; E. Hypandrium, ventral view; F. Dorsal lobe of surstylus, outer view; G. Ventral lobe of surstylus, outer view. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct br = proctiger brush; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204111" httpUri="https://zenodo.org/record/8204111/files/figure.png" pageId="2" pageNumber="153">Figs 3G</figureCitation>
,
<figureCitation id="13748DC6FF930014FF7BFDFAFF633026" box="[151,184,583,609]" captionStart="FIGURE 9" captionStartId="14.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1615]" captionTargetId="figure-20@14.[151,1436,181,1615]" captionTargetPageId="14" captionText="FIGURE 9. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Abdomen, ventral view; B. Hypopygium, left lateral view; C. Internal appendages, left lateral view; D. Apical part of phallus; E. Apical edge of epandrium; F. Tergite 6, lateral view; G. Right cercus, outer view; H. Left ventral surstylus, outer view; I. Hypandrium, ventral view; JK. Surstyli, left and right lateral views, respectively. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204131" httpUri="https://zenodo.org/record/8204131/files/figure.png" pageId="2" pageNumber="153">9E</figureCitation>
,
<figureCitation id="13748DC6FF930014FF29FDFAFF213026" box="[197,250,583,609]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="2" pageNumber="153">12D</figureCitation>
); basal apicoventral epandrial seta usually longer than ventral lobe of surstylus. Ventral lobe of surstylus somewhat digitiform or L-shaped, weakly to strongly curved ventrally, with 1 stout modified apical seta (
<emphasis id="B93B4D51FF930014FAD0FDD6FABF30C3" box="[1340,1380,619,644]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FA80FDD6FF6130EE" captionStart="FIGURE 9" captionStartId="14.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1615]" captionTargetId="figure-20@14.[151,1436,181,1615]" captionTargetPageId="14" captionText="FIGURE 9. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Abdomen, ventral view; B. Hypopygium, left lateral view; C. Internal appendages, left lateral view; D. Apical part of phallus; E. Apical edge of epandrium; F. Tergite 6, lateral view; G. Right cercus, outer view; H. Left ventral surstylus, outer view; I. Hypandrium, ventral view; JK. Surstyli, left and right lateral views, respectively. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204131" httpUri="https://zenodo.org/record/8204131/files/figure.png" pageId="2" pageNumber="153">Figs 9C</figureCitation>
,
<figureCitation id="13748DC6FF930014FF2AFD32FF2230EE" box="[198,249,655,681]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="2" pageNumber="153">12D</figureCitation>
,
<figureCitation id="13748DC6FF930014FEE8FD32FEE230EE" box="[260,313,655,681]" captionStart="FIGURE 16" captionStartId="23.[151,250,1794,1818]" captionTargetBox="[272,1315,381,1770]" captionTargetId="figure-119@23.[272,1315,381,1770]" captionTargetPageId="23" captionText="FIGURE 16. Paraclius pumilio Loew, 1872, identified male specimen from Texas (CNC). A. Abdomen, ventral view; B. Tergite 7 and sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium, outer view; E. Internal appendages, left lateral view; F. Dorsal lobe of surstylus, outer view; G. Hypandrium, ventral view; H. Cercus, dorsal view; I. Tergite 6, lateral view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204149" httpUri="https://zenodo.org/record/8204149/files/figure.png" pageId="2" pageNumber="153">16D</figureCitation>
). Dorsal lobe of surstylus shorter than ventral lobe and weakly sclerotized, usually with strong setae at ventral and/or apical edges (
<emphasis id="B93B4D51FF930014FE17FD0EFDF8308B" box="[507,547,691,716]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FDC0FD0EFD58308A" box="[556,643,691,717]" captionStart="FIGURE 3" captionStartId="6.[151,250,1458,1482]" captionTargetBox="[151,1436,181,1435]" captionTargetId="figure-20@6.[151,1436,181,1435]" captionTargetPageId="6" captionText="FIGURE 3. Paraclius arcuatus (Loew, 1861), identified male specimens from Honduras (CNC). A. Abdomen, ventral view; B. Tergite 7 and Sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Internal appendages, left lateral view; E. Hypandrium, ventral view; F. Dorsal lobe of surstylus, outer view; G. Ventral lobe of surstylus, outer view. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct br = proctiger brush; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204111" httpUri="https://zenodo.org/record/8204111/files/figure.png" pageId="2" pageNumber="153">Figs 3F</figureCitation>
,
<figureCitation id="13748DC6FF930014FD7DFD0EFD0A308A" box="[657,721,691,717]" captionStart="FIGURE 9" captionStartId="14.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1615]" captionTargetId="figure-20@14.[151,1436,181,1615]" captionTargetPageId="14" captionText="FIGURE 9. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Abdomen, ventral view; B. Hypopygium, left lateral view; C. Internal appendages, left lateral view; D. Apical part of phallus; E. Apical edge of epandrium; F. Tergite 6, lateral view; G. Right cercus, outer view; H. Left ventral surstylus, outer view; I. Hypandrium, ventral view; JK. Surstyli, left and right lateral views, respectively. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204131" httpUri="https://zenodo.org/record/8204131/files/figure.png" pageId="2" pageNumber="153">9J, K</figureCitation>
,
<figureCitation id="13748DC6FF930014FD33FD0EFCC8308A" box="[735,787,691,717]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="2" pageNumber="153">12D</figureCitation>
). Phallus divided into two narrow and strong sclerotized arms, coated with a thin membrane, with small bumps or spinules at base of exposed part, apical part with dentiform or rounded processes (
<emphasis id="B93B4D51FF930014FE79FD46FE663153" box="[405,445,763,788]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FE29FD46FDFA3152" box="[453,545,763,789]" captionStart="FIGURE 3" captionStartId="6.[151,250,1458,1482]" captionTargetBox="[151,1436,181,1435]" captionTargetId="figure-20@6.[151,1436,181,1435]" captionTargetPageId="6" captionText="FIGURE 3. Paraclius arcuatus (Loew, 1861), identified male specimens from Honduras (CNC). A. Abdomen, ventral view; B. Tergite 7 and Sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Internal appendages, left lateral view; E. Hypandrium, ventral view; F. Dorsal lobe of surstylus, outer view; G. Ventral lobe of surstylus, outer view. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct br = proctiger brush; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204111" httpUri="https://zenodo.org/record/8204111/files/figure.png" pageId="2" pageNumber="153">Figs 3D</figureCitation>
,
<figureCitation id="13748DC6FF930014FDC1FD46FD823152" box="[557,601,763,789]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="2" pageNumber="153">12F</figureCitation>
,
<figureCitation id="13748DC6FF930014FD88FD46FD4F3152" box="[612,660,763,789]" captionStart="FIGURE 16" captionStartId="23.[151,250,1794,1818]" captionTargetBox="[272,1315,381,1770]" captionTargetId="figure-119@23.[272,1315,381,1770]" captionTargetPageId="23" captionText="FIGURE 16. Paraclius pumilio Loew, 1872, identified male specimen from Texas (CNC). A. Abdomen, ventral view; B. Tergite 7 and sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium, outer view; E. Internal appendages, left lateral view; F. Dorsal lobe of surstylus, outer view; G. Hypandrium, ventral view; H. Cercus, dorsal view; I. Tergite 6, lateral view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204149" httpUri="https://zenodo.org/record/8204149/files/figure.png" pageId="2" pageNumber="153">16E</figureCitation>
,
<figureCitation id="13748DC6FF930014FD4CFD46FD143152" box="[672,719,763,789]" captionStart="FIGURE 20" captionStartId="26.[151,250,1827,1851]" captionTargetBox="[222,1366,525,1804]" captionTargetId="figure-203@26.[222,1366,525,1804]" captionTargetPageId="26" captionText="FIGURE 20. Paraclius xibun sp. nov., male paratype (CNC). A. Abdomen, ventral view; B. Tergite 7 and sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium, outer view; E. Internal appendages, left lateral view; F. Hypandrium, ventral view; G. Cercus, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; pgt = postgonite; ph = phallus; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204167" httpUri="https://zenodo.org/record/8204167/files/figure.png" pageId="2" pageNumber="153">20E</figureCitation>
) (with fringe of spines in
<taxonomicName id="4C4FEAC0FF930014FC1FFD46FAEA3152" authority="Van Duzee" authorityName="Van Duzee" authorityYear="1929" box="[1011,1329,763,789]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="angustipennis">
<emphasis id="B93B4D51FF930014FC1FFD46FBDF3153" box="[1011,1028,763,788]" italics="true" pageId="2" pageNumber="153">P</emphasis>
.
<emphasis id="B93B4D51FF930014FBFFFD46FB743153" box="[1043,1199,763,788]" italics="true" pageId="2" pageNumber="153">angustipennis</emphasis>
Van Duzee
</taxonomicName>
(
<figureCitation id="13748DC6FF930014FAACFD46FF76317E" captionStart="FIGURE 9" captionStartId="14.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1615]" captionTargetId="figure-20@14.[151,1436,181,1615]" captionTargetPageId="14" captionText="FIGURE 9. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Abdomen, ventral view; B. Hypopygium, left lateral view; C. Internal appendages, left lateral view; D. Apical part of phallus; E. Apical edge of epandrium; F. Tergite 6, lateral view; G. Right cercus, outer view; H. Left ventral surstylus, outer view; I. Hypandrium, ventral view; JK. Surstyli, left and right lateral views, respectively. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204131" httpUri="https://zenodo.org/record/8204131/files/figure.png" pageId="2" pageNumber="153">Fig. 9C, D</figureCitation>
)). Hypandrium as long as exposed part of phallus, strongly sclerotized, trough-shaped, with weak connection to epandrium basally, free laterally, apex bifurcate and weakly sclerotized (
<emphasis id="B93B4D51FF930014FC19FCFEFBC6311B" box="[1013,1053,835,860]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FBCAFCFEFB5A311A" box="[1062,1153,835,861]" captionStart="FIGURE 3" captionStartId="6.[151,250,1458,1482]" captionTargetBox="[151,1436,181,1435]" captionTargetId="figure-20@6.[151,1436,181,1435]" captionTargetPageId="6" captionText="FIGURE 3. Paraclius arcuatus (Loew, 1861), identified male specimens from Honduras (CNC). A. Abdomen, ventral view; B. Tergite 7 and Sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Internal appendages, left lateral view; E. Hypandrium, ventral view; F. Dorsal lobe of surstylus, outer view; G. Ventral lobe of surstylus, outer view. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct br = proctiger brush; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204111" httpUri="https://zenodo.org/record/8204111/files/figure.png" pageId="2" pageNumber="153">Figs 3E</figureCitation>
,
<figureCitation id="13748DC6FF930014FB63FCFEFB73311A" box="[1167,1192,835,861]" captionStart="FIGURE 9" captionStartId="14.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1615]" captionTargetId="figure-20@14.[151,1436,181,1615]" captionTargetPageId="14" captionText="FIGURE 9. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Abdomen, ventral view; B. Hypopygium, left lateral view; C. Internal appendages, left lateral view; D. Apical part of phallus; E. Apical edge of epandrium; F. Tergite 6, lateral view; G. Right cercus, outer view; H. Left ventral surstylus, outer view; I. Hypandrium, ventral view; JK. Surstyli, left and right lateral views, respectively. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204131" httpUri="https://zenodo.org/record/8204131/files/figure.png" pageId="2" pageNumber="153">9I</figureCitation>
,
<figureCitation id="13748DC6FF930014FB5AFCFEFB33311A" box="[1206,1256,835,861]" captionStart="FIGURE 16" captionStartId="23.[151,250,1794,1818]" captionTargetBox="[272,1315,381,1770]" captionTargetId="figure-119@23.[272,1315,381,1770]" captionTargetPageId="23" captionText="FIGURE 16. Paraclius pumilio Loew, 1872, identified male specimen from Texas (CNC). A. Abdomen, ventral view; B. Tergite 7 and sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium, outer view; E. Internal appendages, left lateral view; F. Dorsal lobe of surstylus, outer view; G. Hypandrium, ventral view; H. Cercus, dorsal view; I. Tergite 6, lateral view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204149" httpUri="https://zenodo.org/record/8204149/files/figure.png" pageId="2" pageNumber="153">16G</figureCitation>
) (apex strongly sclerotized in
<taxonomicName id="4C4FEAC0FF930014FEDEFCDAFE5631C6" authority="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha, 2023" authorityName="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha" authorityYear="2023" box="[306,397,871,897]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="xibun" status="sp. nov.">
<emphasis id="B93B4D51FF930014FEDEFCDAFE9831C7" box="[306,323,871,896]" italics="true" pageId="2" pageNumber="153">P</emphasis>
.
<emphasis id="B93B4D51FF930014FEA3FCDAFE5631C6" box="[335,397,871,897]" italics="true" pageId="2" pageNumber="153">xibun</emphasis>
</taxonomicName>
<emphasis id="B93B4D51FF930014FE7FFCDAFE3031C6" bold="true" box="[403,491,871,897]" pageId="2" pageNumber="153">
<taxonomicNameLabel id="A208F02AFF930014FE7FFCDAFE3031C6" box="[403,491,871,897]" pageId="2" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13748DC6FF930014FE15FCDAFD8031C6" box="[505,603,871,897]" captionStart="FIGURE 20" captionStartId="26.[151,250,1827,1851]" captionTargetBox="[222,1366,525,1804]" captionTargetId="figure-203@26.[222,1366,525,1804]" captionTargetPageId="26" captionText="FIGURE 20. Paraclius xibun sp. nov., male paratype (CNC). A. Abdomen, ventral view; B. Tergite 7 and sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium, outer view; E. Internal appendages, left lateral view; F. Hypandrium, ventral view; G. Cercus, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; pgt = postgonite; ph = phallus; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204167" httpUri="https://zenodo.org/record/8204167/files/figure.png" pageId="2" pageNumber="153">Fig. 20F</figureCitation>
)), somewhat arrow-shaped, lateral margin usually with dentiform preapical processes, base of hypandrium projecting up inside epandrial capsule, cradling the phallus. Sperm pump short, well sclerotized, somewhat kidney-shaped, pipe-shaped or C-shaped, about 1/2 to 1/3 as long as ejaculatory apodeme (
<emphasis id="B93B4D51FF930014FF4DFC6EFF1231AB" box="[161,201,979,1004]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FF23FC6EFEF131AA" box="[207,298,979,1005]" captionStart="FIGURE 3" captionStartId="6.[151,250,1458,1482]" captionTargetBox="[151,1436,181,1435]" captionTargetId="figure-20@6.[151,1436,181,1435]" captionTargetPageId="6" captionText="FIGURE 3. Paraclius arcuatus (Loew, 1861), identified male specimens from Honduras (CNC). A. Abdomen, ventral view; B. Tergite 7 and Sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Internal appendages, left lateral view; E. Hypandrium, ventral view; F. Dorsal lobe of surstylus, outer view; G. Ventral lobe of surstylus, outer view. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct br = proctiger brush; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204111" httpUri="https://zenodo.org/record/8204111/files/figure.png" pageId="2" pageNumber="153">Figs 3D</figureCitation>
,
<figureCitation id="13748DC6FF930014FED9FC6EFE8C31AA" box="[309,343,979,1005]" captionStart="FIGURE 9" captionStartId="14.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1615]" captionTargetId="figure-20@14.[151,1436,181,1615]" captionTargetPageId="14" captionText="FIGURE 9. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Abdomen, ventral view; B. Hypopygium, left lateral view; C. Internal appendages, left lateral view; D. Apical part of phallus; E. Apical edge of epandrium; F. Tergite 6, lateral view; G. Right cercus, outer view; H. Left ventral surstylus, outer view; I. Hypandrium, ventral view; JK. Surstyli, left and right lateral views, respectively. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204131" httpUri="https://zenodo.org/record/8204131/files/figure.png" pageId="2" pageNumber="153">9C</figureCitation>
,
<figureCitation id="13748DC6FF930014FE8FFC6EFE4F31AA" box="[355,404,979,1005]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="2" pageNumber="153">12E</figureCitation>
). Ejaculatory apodeme usually strongly sclerotized (weakly sclerotized in
<taxonomicName id="4C4FEAC0FF930014FB20FC6EFAE531AA" authority="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha, 2023" authorityName="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha" authorityYear="2023" box="[1228,1342,979,1005]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="brooksi" status="sp. nov.">
<emphasis id="B93B4D51FF930014FB20FC6EFB0631AB" box="[1228,1245,979,1004]" italics="true" pageId="2" pageNumber="153">P</emphasis>
.
<emphasis id="B93B4D51FF930014FB07FC6EFAE531AA" box="[1259,1342,979,1005]" italics="true" pageId="2" pageNumber="153">brooksi</emphasis>
</taxonomicName>
<emphasis id="B93B4D51FF930014FAA8FC6EFA4731AA" bold="true" box="[1348,1436,979,1005]" pageId="2" pageNumber="153">
<taxonomicNameLabel id="A208F02AFF930014FAA8FC6EFA4731AA" box="[1348,1436,979,1005]" pageId="2" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13748DC6FF930014FF73FC4AFED93656" box="[159,258,1015,1041]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="2" pageNumber="153">Fig. 12E</figureCitation>
)), long and narrow, about 1/2 as long as epandrium and slightly widening apically (
<emphasis id="B93B4D51FF930014FB71FC4AFB1E3657" box="[1181,1221,1015,1040]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FB20FC4AFAFD3656" box="[1228,1318,1015,1041]" captionStart="FIGURE 3" captionStartId="6.[151,250,1458,1482]" captionTargetBox="[151,1436,181,1435]" captionTargetId="figure-20@6.[151,1436,181,1435]" captionTargetPageId="6" captionText="FIGURE 3. Paraclius arcuatus (Loew, 1861), identified male specimens from Honduras (CNC). A. Abdomen, ventral view; B. Tergite 7 and Sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Internal appendages, left lateral view; E. Hypandrium, ventral view; F. Dorsal lobe of surstylus, outer view; G. Ventral lobe of surstylus, outer view. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct br = proctiger brush; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204111" httpUri="https://zenodo.org/record/8204111/files/figure.png" pageId="2" pageNumber="153">Figs 3D</figureCitation>
,
<figureCitation id="13748DC6FF930014FADDFC4AFA8F3656" box="[1329,1364,1015,1041]" captionStart="FIGURE 9" captionStartId="14.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1615]" captionTargetId="figure-20@14.[151,1436,181,1615]" captionTargetPageId="14" captionText="FIGURE 9. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Abdomen, ventral view; B. Hypopygium, left lateral view; C. Internal appendages, left lateral view; D. Apical part of phallus; E. Apical edge of epandrium; F. Tergite 6, lateral view; G. Right cercus, outer view; H. Left ventral surstylus, outer view; I. Hypandrium, ventral view; JK. Surstyli, left and right lateral views, respectively. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204131" httpUri="https://zenodo.org/record/8204131/files/figure.png" pageId="2" pageNumber="153">9C</figureCitation>
,
<figureCitation id="13748DC6FF930014FAB3FC4AFA4B3656" box="[1375,1424,1015,1041]" captionStart="FIGURE 16" captionStartId="23.[151,250,1794,1818]" captionTargetBox="[272,1315,381,1770]" captionTargetId="figure-119@23.[272,1315,381,1770]" captionTargetPageId="23" captionText="FIGURE 16. Paraclius pumilio Loew, 1872, identified male specimen from Texas (CNC). A. Abdomen, ventral view; B. Tergite 7 and sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium, outer view; E. Internal appendages, left lateral view; F. Dorsal lobe of surstylus, outer view; G. Hypandrium, ventral view; H. Cercus, dorsal view; I. Tergite 6, lateral view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204149" httpUri="https://zenodo.org/record/8204149/files/figure.png" pageId="2" pageNumber="153">16E</figureCitation>
). Proctiger weakly sclerotized, dorsoventrally flattened (
<emphasis id="B93B4D51FF930014FCFDFBA6FCE23673" box="[785,825,1051,1076]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FCA8FBA6FC753672" box="[836,942,1051,1077]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="2" pageNumber="153">Figs 12E</figureCitation>
,
<figureCitation id="13748DC6FF930014FC51FBA6FC373672" box="[957,1004,1051,1077]" captionStart="FIGURE 16" captionStartId="23.[151,250,1794,1818]" captionTargetBox="[272,1315,381,1770]" captionTargetId="figure-119@23.[272,1315,381,1770]" captionTargetPageId="23" captionText="FIGURE 16. Paraclius pumilio Loew, 1872, identified male specimen from Texas (CNC). A. Abdomen, ventral view; B. Tergite 7 and sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium, outer view; E. Internal appendages, left lateral view; F. Dorsal lobe of surstylus, outer view; G. Hypandrium, ventral view; H. Cercus, dorsal view; I. Tergite 6, lateral view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204149" httpUri="https://zenodo.org/record/8204149/files/figure.png" pageId="2" pageNumber="153">16E</figureCitation>
) (modified in an undivided proctiger brush in
<taxonomicName id="4C4FEAC0FF930014FF1BFB82FEAE361F" authorityName="Loew" authorityYear="1861" box="[247,373,1087,1113]" class="Insecta" family="Dolichopodidae" genus="Pelastoneurus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="B93B4D51FF930014FF1BFB82FED3361F" box="[247,264,1087,1112]" italics="true" pageId="2" pageNumber="153">P</emphasis>
.
<emphasis id="B93B4D51FF930014FEF8FB82FEAE361F" box="[276,373,1087,1112]" italics="true" pageId="2" pageNumber="153">arcuatus</emphasis>
</taxonomicName>
(
<figureCitation id="13748DC6FF930014FE6FFB82FE00361E" box="[387,475,1087,1113]" captionStart="FIGURE 3" captionStartId="6.[151,250,1458,1482]" captionTargetBox="[151,1436,181,1435]" captionTargetId="figure-20@6.[151,1436,181,1435]" captionTargetPageId="6" captionText="FIGURE 3. Paraclius arcuatus (Loew, 1861), identified male specimens from Honduras (CNC). A. Abdomen, ventral view; B. Tergite 7 and Sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Internal appendages, left lateral view; E. Hypandrium, ventral view; F. Dorsal lobe of surstylus, outer view; G. Ventral lobe of surstylus, outer view. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct br = proctiger brush; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204111" httpUri="https://zenodo.org/record/8204111/files/figure.png" pageId="2" pageNumber="153">Fig. 3D</figureCitation>
)). Cercus short, suboval or drop-shaped, about 1/3 to 1/2 as long as epandrium, lacking basolateral tail (
<emphasis id="B93B4D51FF930014FEA6FBDEFEA9363B" box="[330,370,1123,1148]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FE95FBDEFE0F363A" box="[377,468,1123,1149]" captionStart="FIGURE 2" captionStartId="5.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-18@5.[151,1436,181,1730]" captionTargetPageId="5" captionText="FIGURE 2. Paraclius arcuatus (Loew, 1861), A, D, E, F male specimen from Honduras (CNC), B, C male specimen from Mexico (NMNH). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D.Abdomen and hypopyium, lateral view; E. Left antenna, inner view; F. Left wing.Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R2+3 = second branch of radius; R = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.8204107" httpUri="https://zenodo.org/record/8204107/files/figure.png" pageId="2" pageNumber="153">Figs 2D</figureCitation>
,
<figureCitation id="13748DC6FF930014FE33FBDEFDD8363A" box="[479,515,1123,1149]" captionStart="FIGURE 9" captionStartId="14.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1615]" captionTargetId="figure-20@14.[151,1436,181,1615]" captionTargetPageId="14" captionText="FIGURE 9. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Abdomen, ventral view; B. Hypopygium, left lateral view; C. Internal appendages, left lateral view; D. Apical part of phallus; E. Apical edge of epandrium; F. Tergite 6, lateral view; G. Right cercus, outer view; H. Left ventral surstylus, outer view; I. Hypandrium, ventral view; JK. Surstyli, left and right lateral views, respectively. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204131" httpUri="https://zenodo.org/record/8204131/files/figure.png" pageId="2" pageNumber="153">9G</figureCitation>
,
<figureCitation id="13748DC6FF930014FDE3FBDEFD9A363A" box="[527,577,1123,1149]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="2" pageNumber="153">12B</figureCitation>
).
<emphasis id="B93B4D51FF930014FDB8FBDEFD66363A" bold="true" box="[596,701,1123,1149]" pageId="2" pageNumber="153">Females.</emphasis>
Similar to male in habitus and chaetotaxy, differing mainly by the face and clypeus wider (both wider than ocellar tubercle), with eyes almost parallel sided and the short postpedicel, about as long as wide (
<emphasis id="B93B4D51FF930014FE75FB16FE1A3683" box="[409,449,1195,1220]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FE24FB16FD983682" box="[456,579,1195,1221]" captionStart="FIGURE 4" captionStartId="9.[151,250,582,606]" captionTargetBox="[151,1436,181,558]" captionTargetId="figure-18@9.[151,1436,181,558]" captionTargetPageId="9" captionText="FIGURE 4. Paraclius arcuatus (Loew, 1861), identified female specimen from Mexico (CNC). A. Habitus, lateral view; B. Head, anterior view; C. Left antenna, inner view." figureDoi="http://doi.org/10.5281/zenodo.8204113" httpUri="https://zenodo.org/record/8204113/files/figure.png" pageId="2" pageNumber="153">Figs 4B, C</figureCitation>
,
<figureCitation id="13748DC6FF930014FDA3FB16FD7B3682" box="[591,672,1195,1221]" captionStart="FIGURE 10" captionStartId="15.[151,250,717,741]" captionTargetBox="[151,1436,237,694]" captionTargetId="figure-42@15.[151,1436,237,694]" captionTargetPageId="15" captionText="FIGURE 10. Paraclius angustipennis Van Duzee, 1929, identified female specimen from Mexico (CNC). A. Habitus, lateral view; B. Head, anterior view; C. Left antenna, outer view." figureDoi="http://doi.org/10.5281/zenodo.8204133" httpUri="https://zenodo.org/record/8204133/files/figure.png" pageId="2" pageNumber="153">10B, C</figureCitation>
,
<figureCitation id="13748DC6FF930014FD40FB16FD243682" box="[684,767,1195,1221]" captionStart="FIGURE 13" captionStartId="19.[151,250,655,679]" captionTargetBox="[151,1436,181,632]" captionTargetId="figure-18@19.[151,1436,181,632]" captionTargetPageId="19" captionText="FIGURE 13. Paraclius brooksi sp. nov, identified female from Mexico (CNC). A. Habitus, lateral view; B. Head, anterior view; C. Right antenna, outer view." figureDoi="http://doi.org/10.5281/zenodo.8204141" httpUri="https://zenodo.org/record/8204141/files/figure.png" pageId="2" pageNumber="153">13B, C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C355C2C8FF930014FF2BFB72FDFB3776" pageId="2" pageNumber="153" type="description">
<paragraph id="8BF09143FF930014FF2BFB72FDFB3776" blockId="2.[151,1437,150,1725]" pageId="2" pageNumber="153">
<emphasis id="B93B4D51FF930014FF2BFB72FE4A36AF" bold="true" box="[199,401,1230,1257]" pageId="2" pageNumber="153">Included species.</emphasis>
<taxonomicName id="4C4FEAC0FF930014FE74FB72FD6C36AE" authority="(Loew, 1861)" baseAuthorityName="Loew" baseAuthorityYear="1861" box="[408,695,1231,1257]" class="Insecta" family="Dolichopodidae" genus="Pelastoneurus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="B93B4D51FF930014FE74FB72FE7236AF" box="[408,425,1231,1256]" italics="true" pageId="2" pageNumber="153">P</emphasis>
.
<emphasis id="B93B4D51FF930014FE5BFB72FDC336AF" box="[439,536,1231,1256]" italics="true" pageId="2" pageNumber="153">arcuatus</emphasis>
(
<bibRefCitation id="EFDEECB2FF930014FDCAFB72FD7436AE" author="Loew, H." box="[550,687,1231,1257]" pageId="2" pageNumber="153" refId="ref21901" refString="Loew, H. (1861) Neue Beitrage zur Kenntniss der Dipteren. Achter Beitrag. Mittler &amp; Sohn, Berlin, 100 pp." type="book" year="1861">Loew, 1861</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C4FEAC0FF930014FD2FFB72FC1936AE" authority="Loew, 1872" authorityName="Loew" authorityYear="1872" box="[707,962,1231,1257]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="pumilio">
<emphasis id="B93B4D51FF930014FD2FFB72FD0F36AF" box="[707,724,1231,1256]" italics="true" pageId="2" pageNumber="153">P</emphasis>
.
<emphasis id="B93B4D51FF930014FD0EFB72FCE336AE" box="[738,824,1231,1257]" italics="true" pageId="2" pageNumber="153">pumilio</emphasis>
<bibRefCitation id="EFDEECB2FF930014FCD2FB72FC1936AE" author="Loew, H." box="[830,962,1231,1257]" pageId="2" pageNumber="153" pagination="49 - 124" refId="ref21967" refString="Loew, H. (1872) Diptera Americae septentrionalis indigena. Centuria decima. Berliner Entomologische Zeitschrift, 16 (1), 49 - 124. https: // doi. org / 10.1002 / mmnd. 18720160110" type="journal article" year="1872">Loew, 1872</bibRefCitation>
</taxonomicName>
and the following newly ascribed species:
<taxonomicName id="4C4FEAC0FF930014FF7BFB4EFDB1374A" authority="Van Duzee, 1933" authorityName="Van Duzee" authorityYear="1933" box="[151,618,1267,1293]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="angusticauda">
<emphasis id="B93B4D51FF930014FF7BFB4EFE7A374A" box="[151,417,1267,1293]" italics="true" pageId="2" pageNumber="153">Paraclius angusticauda</emphasis>
<bibRefCitation id="EFDEECB2FF930014FE44FB4EFDB1374A" author="Van Duzee, M. C." box="[424,618,1267,1293]" pageId="2" pageNumber="153" pagination="1 - 27" refId="ref22526" refString="Van Duzee, M. C. (1933) New Dolichopidae from North America with notes on several described species. American Museum Novitates, 599, 1 - 27, 52 figs." type="journal article" year="1933">Van Duzee, 1933</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C4FEAC0FF930014FD98FB4EFC2D374A" authority="Van Duzee, 1929" authorityName="Van Duzee" authorityYear="1929" box="[628,1014,1267,1293]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="angustipennis">
<emphasis id="B93B4D51FF930014FD98FB4EFD5E374B" box="[628,645,1267,1292]" italics="true" pageId="2" pageNumber="153">P</emphasis>
.
<emphasis id="B93B4D51FF930014FD7FFB4EFCF4374B" box="[659,815,1267,1292]" italics="true" pageId="2" pageNumber="153">angustipennis</emphasis>
<bibRefCitation id="EFDEECB2FF930014FCD8FB4EFC2D374A" author="Van Duzee, M. C." box="[820,1014,1267,1293]" pageId="2" pageNumber="153" pagination="1 - 64" refId="ref22434" refString="Van Duzee, M. C. (1929) Tropical American Diptera or two-winged flies of the family Dolichopodidae from Central and South America. Proceedings of the United States of National Museum, 74 (10), 1 - 64. https: // doi. org / 10.5479 / si. 00963801.74 - 2755.1" type="journal article" year="1929">Van Duzee, 1929</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C4FEAC0FF930014FBEDFB4EFBA9374A" authority="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha, 2023" authorityName="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha" authorityYear="2023" box="[1025,1138,1267,1293]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="brooksi" status="sp. nov.">
<emphasis id="B93B4D51FF930014FBEDFB4EFBC9374B" box="[1025,1042,1267,1292]" italics="true" pageId="2" pageNumber="153">P</emphasis>
.
<emphasis id="B93B4D51FF930014FBF3FB4EFBA9374A" box="[1055,1138,1267,1293]" italics="true" pageId="2" pageNumber="153">brooksi</emphasis>
</taxonomicName>
<emphasis id="B93B4D51FF930014FB94FB4EFB0A374A" bold="true" box="[1144,1233,1267,1293]" pageId="2" pageNumber="153">
<taxonomicNameLabel id="A208F02AFF930014FB94FB4EFB0A374A" box="[1144,1233,1267,1293]" pageId="2" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C4FEAC0FF930014FB33FB4EFEF33776" authority="Van Duzee, 1930" authorityName="Van Duzee" authorityYear="1930" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B93B4D51FF930014FB33FB4EFB2B374B" box="[1247,1264,1267,1292]" italics="true" pageId="2" pageNumber="153">P</emphasis>
.
<emphasis id="B93B4D51FF930014FB11FB4EFAB1374A" box="[1277,1386,1267,1293]" italics="true" pageId="2" pageNumber="153">elongatus</emphasis>
<bibRefCitation id="EFDEECB2FF930014FA9DFB4EFEF33776" author="Van Duzee, M. C." pageId="2" pageNumber="153" pagination="53 - 55" refId="ref22492" refString="Van Duzee, M. C. (1930) New species of Dolichopodidae from North America (Diptera). Entomological News, 41, 53 - 55 + 70 - 73." type="journal article" year="1930">Van Duzee, 1930</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C4FEAC0FF930014FE8EFAAAFE643776" authority="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha, 2023" authorityName="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha" authorityYear="2023" box="[354,447,1303,1329]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="xibun" status="sp. nov.">
<emphasis id="B93B4D51FF930014FE8EFAAAFEA83777" box="[354,371,1303,1328]" italics="true" pageId="2" pageNumber="153">P</emphasis>
.
<emphasis id="B93B4D51FF930014FE6DFAAAFE643776" box="[385,447,1303,1329]" italics="true" pageId="2" pageNumber="153">xibun</emphasis>
</taxonomicName>
<emphasis id="B93B4D51FF930014FE2AFAAAFDFB3776" bold="true" box="[454,544,1303,1329]" pageId="2" pageNumber="153">
<taxonomicNameLabel id="A208F02AFF930014FE2AFAAAFDFB3776" box="[454,544,1303,1329]" pageId="2" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C355C2C8FF930014FF2BFA86FCDF346A" pageId="2" pageNumber="153" type="discussion">
<paragraph id="8BF09143FF930014FF2BFA86FCDF346A" blockId="2.[151,1437,150,1725]" pageId="2" pageNumber="153">
<emphasis id="B93B4D51FF930014FF2BFA86FEE03712" bold="true" box="[199,315,1339,1365]" pageId="2" pageNumber="153">Remarks.</emphasis>
<bibRefCitation id="EFDEECB2FF930014FEACFA86FE3A3712" author="Brooks, S. E." box="[320,481,1338,1365]" pageId="2" pageNumber="153" pagination="1 - 158" refId="ref21534" refString="Brooks, S. E. (2005) Systematics and phylogeny of Dolichopodinae (Diptera: Dolichopodidae). Zootaxa, 857 (1), 1 - 158. https: // doi. org / 10.11646 / zootaxa. 857.1.1" type="journal article" year="2005">Brooks (2005)</bibRefCitation>
was the first to mention the
<taxonomicName id="4C4FEAC0FF930014FCF8FA86FC3C3713" baseAuthorityName="Loew" baseAuthorityYear="1861" box="[788,999,1339,1365]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="B93B4D51FF930014FCF8FA86FC3C3713" box="[788,999,1339,1365]" italics="true" pageId="2" pageNumber="153">Paraclius arcuatus</emphasis>
</taxonomicName>
species-group (
<emphasis id="B93B4D51FF930014FB7AFA86FA483713" box="[1174,1427,1339,1365]" italics="true" pageId="2" pageNumber="153">
<taxonomicName id="4C4FEAC0FF930014FB7AFA86FAD83712" authorityName="Loew" authorityYear="1864" box="[1174,1283,1339,1365]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus" sensu="stricto">Paraclius</taxonomicName>
<taxonomicNameLabel id="A208F02AFF930014FAE4FA86FA483713" box="[1288,1427,1339,1364]" pageId="2" pageNumber="153" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
) composed of the following species:
<taxonomicName id="4C4FEAC0FF930014FDC6FAE2FD25373F" baseAuthorityName="Loew" baseAuthorityYear="1861" box="[554,766,1375,1401]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="B93B4D51FF930014FDC6FAE2FD25373F" box="[554,766,1375,1401]" italics="true" pageId="2" pageNumber="153">Paraclius arcuatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4FEAC0FF930014FCE0FAE2FC5A373E" authorityName="Loew" authorityYear="1872" box="[780,897,1375,1401]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="pumilio">
<emphasis id="B93B4D51FF930014FCE0FAE2FCC6373F" box="[780,797,1375,1400]" italics="true" pageId="2" pageNumber="153">P</emphasis>
.
<emphasis id="B93B4D51FF930014FCC7FAE2FC5A373E" box="[811,897,1375,1401]" italics="true" pageId="2" pageNumber="153">pumilio</emphasis>
</taxonomicName>
and “
<taxonomicName id="4C4FEAC0FF930014FC2FFAE2FBB1373E" box="[963,1130,1375,1401]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="B93B4D51FF930014FC2FFAE2FBEB373E" box="[963,1072,1375,1401]" italics="true" pageId="2" pageNumber="153">Paraclius</emphasis>
sp. 1
</taxonomicName>
”. However, after gathering information from descriptions, photos of
<typeStatus id="54F42FE1FF930014FD88FA3EFD4437DA" box="[612,671,1411,1437]" pageId="2" pageNumber="153">types</typeStatus>
and identified material of nearly all New World
<taxonomicName id="4C4FEAC0FF930014FB2CFA3EFAF637DA" authorityName="Loew" authorityYear="1864" box="[1216,1325,1411,1437]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="B93B4D51FF930014FB2CFA3EFAF637DA" box="[1216,1325,1411,1437]" italics="true" pageId="2" pageNumber="153">Paraclius</emphasis>
</taxonomicName>
we found additional species that fit this group. The
<taxonomicName id="4C4FEAC0FF930014FD86FA1AFCE43787" baseAuthorityName="Loew" baseAuthorityYear="1861" box="[618,831,1447,1473]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="B93B4D51FF930014FD86FA1AFCE43787" box="[618,831,1447,1473]" italics="true" pageId="2" pageNumber="153">Paraclius arcuatus</emphasis>
</taxonomicName>
species-group is a small lineage whose monophyly is supported by the characters discussed below (see Discussion section), mainly by the male face narrowing below, clypeus short, about 1/4 as long as face, the absence of a developed apicoventral epandrial lobe, the acute apical ridge of epandrium and the trough-shaped hypandrium.
</paragraph>
</subSubSection>
<subSubSection id="C355C2C8FF930014FF2BF98BFEF534FA" pageId="2" pageNumber="153" type="distribution">
<paragraph id="8BF09143FF930014FF2BF98BFEF534FA" blockId="2.[151,1437,150,1725]" pageId="2" pageNumber="153">
<emphasis id="B93B4D51FF930014FF2BF98BFEBA3417" bold="true" box="[199,353,1590,1616]" pageId="2" pageNumber="153">Distribution.</emphasis>
The group is mostly restricted to Central and North America, with some species reaching the Caribbean Islands. The distribution range is from Northern
<collectingCountry id="F358D1D3FF930014FCCAF9E6FC873432" box="[806,860,1627,1653]" name="United States of America" pageId="2" pageNumber="153">USA</collectingCountry>
to
<collectingCountry id="F358D1D3FF930014FC91F9E6FC2D3432" box="[893,1014,1627,1653]" name="Costa Rica" pageId="2" pageNumber="153">Costa Rica</collectingCountry>
in Central America and the Caribbean islands of
<collectingCountry id="F358D1D3FF930014FEE1F9C2FE9034DE" box="[269,331,1663,1689]" name="Cuba" pageId="2" pageNumber="153">Cuba</collectingCountry>
,
<collectingCountry id="F358D1D3FF930014FEB4F9C2FE6E34DE" box="[344,437,1663,1689]" name="Jamaica" pageId="2" pageNumber="153">Jamaica</collectingCountry>
,
<collectingCountry id="F358D1D3FF930014FE2EF9C2FDFF34DE" box="[450,548,1663,1689]" name="Grenada" pageId="2" pageNumber="153">Grenada</collectingCountry>
, and
<collectingCountry id="F358D1D3FF930014FD8DF9C2FC3B34DE" box="[609,992,1663,1689]" name="Saint Vincent and the Grenadines" pageId="2" pageNumber="153">Saint Vincent and the Grenadines</collectingCountry>
, not reaching mainland South America (
<figureCitation id="13748DC6FF930014FF73F91EFF2C34FA" box="[159,247,1699,1725]" captionStart="FIGURE 22" captionStartId="28.[151,250,1998,2022]" captionTargetBox="[151,1436,1021,1974]" captionTargetId="figure-154@28.[151,1436,1021,1974]" captionTargetPageId="28" captionText="FIGURE 22. Known geographic distribution of Paraclius arcuatus (Loew, 1861) and P. pumilio Loew, 1872." figureDoi="http://doi.org/10.5281/zenodo.8204173" httpUri="https://zenodo.org/record/8204173/files/figure.png" pageId="2" pageNumber="153">Figs 22</figureCitation>
,
<figureCitation id="13748DC6FF930014FEEEF91EFEF934FA" box="[258,290,1699,1725]" captionStart="FIGURE 23" captionStartId="29.[151,250,1033,1057]" captionTargetBox="[151,1436,237,1009]" captionTargetId="figure-36@29.[151,1436,237,1009]" captionTargetPageId="29" captionText="FIGURE 23. Known geographic distribution of Paraclius angusticauda Van Duzee, 1933, P. angustipennis Van Duzee, 1929, P. brooksi sp. nov., P. elongatus Van Duzee, 1930 and P. xibun sp. nov." figureDoi="http://doi.org/10.5281/zenodo.8204175" httpUri="https://zenodo.org/record/8204175/files/figure.png" pageId="2" pageNumber="153">23</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>