120 lines
13 KiB
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120 lines
13 KiB
XML
<document id="504E621670019113505B365CA600F748" ID-DOI="10.11646/zootaxa.5296.3.6" ID-ISSN="1175-5326" ID-ZooBank="59AD1B4F-15B2-4DC0-A57E-2F6B57539D1A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685427549966" checkinUser="plazi" docAuthor="Bartel, Christian, Dunlop, Jason A. & Giribet, Gonzalo" docDate="2023" docId="03D878433941FD258BB5F8B5FB95FDF2" docLanguage="en" docName="zootaxa.5296.3.6.pdf" docOrigin="Zootaxa 5296 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5296.3.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptopsalis Thorell 1882" docType="treatment" docVersion="1" lastPageNumber="426" masterDocId="FFE1003B3945FD208B22FFC5FFC5FFCF" masterDocTitle="An unexpected diversity of Cyphophthalmi (Arachnida: Opiliones) in Upper Cretaceous Burmese amber" masterLastPageNumber="445" masterPageNumber="421" pageNumber="425" updateTime="1685427628447" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:titleInfo id="CB8A07BAB73307662EAE48503092A4C0">
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<mods:title id="46969074000A02E2C46D56C6122ED04A">An unexpected diversity of Cyphophthalmi (Arachnida: Opiliones) in Upper Cretaceous Burmese amber</mods:title>
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<mods:name id="7432F08A4ACD3D0B380F884A50674E3C" type="personal">
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<mods:namePart id="2C20B7973D678A9989BA6B0711BCE19A">Bartel, Christian</mods:namePart>
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<mods:roleTerm id="5131207E5EDB00066D59E36385853EFE">Author</mods:roleTerm>
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<mods:namePart id="3CD2C521EDECEF466D0F3E4C40161486">Dunlop, Jason A.</mods:namePart>
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<mods:namePart id="880E739B4F09BDAA837A4F9957261AB2">Giribet, Gonzalo</mods:namePart>
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<mods:title id="86661C1F0EE0EF67678430EA21B0BA48">Zootaxa</mods:title>
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<mods:date id="EFB0848E6A060D7E9EC796435B94E8F7">2023</mods:date>
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<mods:number id="CE32F7F5636883E95D6C0FEB9B22D2F4">2023-05-29</mods:number>
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<mods:number id="C01219F0FF30304D5EA30BA1411BA25E">5296</mods:number>
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<mods:url id="90CB4A9808A30C8ED7B521D62DB5BF19">http://dx.doi.org/10.11646/zootaxa.5296.3.6</mods:url>
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<mods:classification id="664140EDBEF0B83622E59A7E74C5BAFF">journal article</mods:classification>
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<mods:identifier id="BB9FD972DF062CD7A9827FCF05D5EE14" type="DOI">10.11646/zootaxa.5296.3.6</mods:identifier>
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<mods:identifier id="191BDB6D00F9295A4B36D96D548B1428" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03D878433941FD258BB5F8B5FB95FDF2" LSID="urn:lsid:plazi:treatment:03D878433941FD258BB5F8B5FB95FDF2" httpUri="http://treatment.plazi.org/id/03D878433941FD258BB5F8B5FB95FDF2" lastPageId="5" lastPageNumber="426" pageId="4" pageNumber="425">
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<subSubSection id="C36B9ADE3941FD248BB5F8B5FDE0F845" box="[151,549,1904,1931]" pageId="4" pageNumber="425" type="nomenclature">
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<paragraph id="8BCEC9553941FD248BB5F8B5FDE0F845" blockId="4.[151,549,1904,1931]" box="[151,549,1904,1931]" pageId="4" pageNumber="425">
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<heading id="D0867E393941FD248BB5F8B5FDE0F845" bold="true" box="[151,549,1904,1931]" fontSize="11" level="1" pageId="4" pageNumber="425" reason="1">
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<emphasis id="B90515473941FD248BB5F8B5FDE0F845" bold="true" box="[151,549,1904,1931]" pageId="4" pageNumber="425">
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Genus
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<taxonomicName id="4C71B2D63941FD248BCCF8B5FDE0F845" ID-CoL="7PMPD" ID-ENA="1276008" authority="Thorell, 1882" authorityName="Thorell" authorityYear="1882" box="[238,549,1904,1931]" class="Arachnida" family="Stylocellidae" genus="Leptopsalis" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="425" phylum="Arthropoda" rank="genus">
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<emphasis id="B90515473941FD248BCCF8B5FEB2F844" bold="true" box="[238,375,1904,1931]" italics="true" pageId="4" pageNumber="425">Leptopsalis</emphasis>
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<bibRefCitation id="EFE0B4A43941FD248A5DF8B4FDE0F845" author="Thorell, T." box="[383,549,1904,1931]" pageId="4" pageNumber="425" pagination="21 - 69" refId="ref14579" refString="Thorell, T. (1882) Descrizione di alcuni Aracnidi inferiori dell' Arcipelago Malese. Annali del Museo Civico di Storia Naturale di Genova, 18, 21 - 69." type="journal article" year="1882">Thorell, 1882</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B9ADE3941FD248BB5F87CFDC9F838" pageId="4" pageNumber="425" type="type_taxon">
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<paragraph id="8BCEC9553941FD248BB5F87CFDC9F838" blockId="4.[151,708,1976,2039]" pageId="4" pageNumber="425">
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<emphasis id="B90515473941FD248BB5F87CFEF7F81D" bold="true" box="[151,306,1976,2003]" pageId="4" pageNumber="425">
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<typeStatus id="54CA77F73941FD248BB5F87CFF17F81C" box="[151,210,1977,2003]" pageId="4" pageNumber="425">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C71B2D63941FD248A1BF87DFD05F81C" authority="Thorell, 1882" authorityName="Thorell" authorityYear="1882" box="[313,704,1976,2003]" class="Arachnida" family="Stylocellidae" genus="Leptopsalis" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="beccarii">
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<emphasis id="B90515473941FD248A1BF87DFDD8F81D" box="[313,541,1976,2002]" italics="true" pageId="4" pageNumber="425">Leptopsalis beccarii</emphasis>
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<bibRefCitation id="EFE0B4A43941FD248906F87DFD05F81C" author="Thorell, T." box="[548,704,1976,2003]" pageId="4" pageNumber="425" pagination="21 - 69" refId="ref14579" refString="Thorell, T. (1882) Descrizione di alcuni Aracnidi inferiori dell' Arcipelago Malese. Annali del Museo Civico di Storia Naturale di Genova, 18, 21 - 69." type="journal article" year="1882">Thorell, 1882</bibRefCitation>
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</taxonomicName>
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.
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<emphasis id="B90515473941FD248BE5F819FE87F839" bold="true" box="[199,322,2012,2038]" pageId="4" pageNumber="425">Diagnosis.</emphasis>
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See Clouse 2012.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B9ADE3940FD258BE5FF52FB95FDF2" pageId="5" pageNumber="426" type="discussion">
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<paragraph id="8BCEC9553940FD258BE5FF52FB95FDF2" blockId="5.[151,1437,150,573]" pageId="5" pageNumber="426">
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<emphasis id="B90515473940FD258BE5FF52FEFEFF7E" bold="true" box="[199,315,151,177]" pageId="5" pageNumber="426">Remarks.</emphasis>
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The
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<typeStatus id="54CA77F73940FD258A55FF52FE1CFF7E" box="[375,473,151,177]" pageId="5" pageNumber="426">holotype</typeStatus>
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of
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<taxonomicName id="4C71B2D63940FD258920FF52FD1EFF7E" authority="Bartel & Dunlop & Giribet, 2023" authorityName="Bartel & Dunlop & Giribet" authorityYear="2023" box="[514,731,151,177]" class="Arachnida" family="Stylocellidae" genus="Leptopsalis" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="426" phylum="Arthropoda" rank="species" species="breyeri" status="sp. nov.">
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<emphasis id="B90515473940FD258920FF52FD1EFF7E" box="[514,731,151,177]" italics="true" pageId="5" pageNumber="426">Leptopsalis breyeri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A236A83C3940FD2589C6FF52FCF9FF7E" box="[740,828,151,177]" pageId="5" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
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(MB.A.4457;
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<figureCitation id="134AD5D03940FD2588C1FF52FB8AFF7E" box="[995,1103,151,177]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="6.[151,250,1900,1924]" captionStartId-1="7.[151,250,794,818]" captionTargetBox-0="[151,1435,628,1865]" captionTargetBox-1="[151,1435,181,770]" captionTargetId-0="figure-130@6.[151,1435,628,1877]" captionTargetId-1="figure-20@7.[151,1435,181,770]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 3. Leptopsalis breyeri sp. nov., male holotype (MB.A.4457). A. Whole specimen, dorsolateral view. B. Same, ventral view. C. Camera lucida drawing of A. D. Same of B. Abbreviations: ad, adenostyle; ap, anal plate; ch, chelicerae; fe, femur; mt, metatarsus; oz, ozophore; pa, patella; pp, pedipalp; ta, tarsus; ti, tibia; tr, trochanter; legs numbered from I–IV. Scale bars equal 1 mm." captionText-1="FIGURE 4. Details of Leptopsalis breyeri sp. nov., male holotype (MB.A.4457).A. Close-up of tarsus IV. B. Close-up of ventral side of opisthosoma. Abbreviations: ad, adenostyle; ap, anal plate; sp, spiracle; leg IV numbered. Scale bars equal 0.5 mm." pageId="5" pageNumber="426">Figs. 3–4</figureCitation>
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) is an adult male, because of its large size and the presence of an adenostyle on tarsus IV. This specimen can be reliably identified as a member of the family
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<taxonomicName id="4C71B2D63940FD258A12FF1AFE03FF36" authorityName="Hansen & Sorensen" authorityYear="1904" box="[304,454,223,249]" class="Arachnida" family="Stylocellidae" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="426" phylum="Arthropoda" rank="family">Stylocellidae</taxonomicName>
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. The ornamented second cheliceral segment, type 2 ozophores, free sternites, C-shaped spiracles and the position of the fringed adenostyle are typical characters for representatives of this family. The possible presence of eyes and indistinctly curved opisthosomal sternal sulci combined with the absence of anal gland pores and extensive cheliceral sculpturing suggest affinities to the extant genus
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<taxonomicName id="4C71B2D63940FD258F55FE8EFB3DFEAA" authorityName="Thorell" authorityYear="1882" box="[1143,1272,331,357]" class="Arachnida" family="Stylocellidae" genus="Leptopsalis" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="426" phylum="Arthropoda" rank="genus">
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<emphasis id="B90515473940FD258F55FE8EFB3DFEAA" box="[1143,1272,331,357]" italics="true" pageId="5" pageNumber="426">Leptopsalis</emphasis>
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</taxonomicName>
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. Furthermore, the habitus of this fossil is somewhat similar to that of the
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<typeStatus id="54CA77F73940FD258860FEAAFC61FE46" box="[834,932,367,393]" pageId="5" pageNumber="426">holotype</typeStatus>
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of
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<taxonomicName id="4C71B2D63940FD2588ECFEAAFB52FE47" authority="Bartel & Dunlop & Giribet, 2023" authorityName="Bartel & Dunlop & Giribet" authorityYear="2023" box="[974,1175,367,393]" class="Arachnida" family="Stylocellidae" genus="Sirocellus" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="426" phylum="Arthropoda" rank="species" species="iunctus" status="sp. nov.">
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<emphasis id="B90515473940FD2588ECFEAAFB52FE47" box="[974,1175,367,393]" italics="true" pageId="5" pageNumber="426">Sirocellus iunctus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A236A83C3940FD258F82FEAAFB3CFE46" box="[1184,1273,367,393]" pageId="5" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
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(GPIH05128) described above. However, specimen MB.A.4457 is much larger and features a completely different adenostyle on tarsus IV. The latter cannot be observed in any of the extant species of
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<taxonomicName id="4C71B2D63940FD2588CBFE72FBAFFE1E" authorityName="Thorell" authorityYear="1882" box="[1001,1130,439,465]" class="Arachnida" family="Stylocellidae" genus="Leptopsalis" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="426" phylum="Arthropoda" rank="genus">
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<emphasis id="B90515473940FD2588CBFE72FBAFFE1E" box="[1001,1130,439,465]" italics="true" pageId="5" pageNumber="426">Leptopsalis</emphasis>
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</taxonomicName>
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, which often have a rather small adenostyle. Based on these characters, this fossil is provisionally placed as an extinct species of
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<taxonomicName id="4C71B2D63940FD258E36FE1EFA50FE3A" authorityName="Thorell" authorityYear="1882" box="[1300,1429,475,501]" class="Arachnida" family="Stylocellidae" genus="Leptopsalis" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="426" phylum="Arthropoda" rank="genus">
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<emphasis id="B90515473940FD258E36FE1EFA50FE3A" box="[1300,1429,475,501]" italics="true" pageId="5" pageNumber="426">Leptopsalis</emphasis>
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</taxonomicName>
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. However, we should keep in mind that some of the characters are possibly plesiomorphic, like the mostly straight opisthosomal sternal sulci, which is the predominant form observed in all our fossils.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |