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<document id="2A652BE2ACF05E9C566F5E58CBC78C52" ID-CLB-Dataset="51663" ID-DOI="10.5281/zenodo.34001" ID-GBIF-Dataset="18bd5418-ed8c-45a5-a662-5d2ebde01c75" checkinTime="1456484105736" checkinUser="donat" docAuthor="Ivan Löbl" docDate="2015" docId="03BF87FBFF91FFFF99E5942AFD09FBC3" docLanguage="en" docName="367-370_Löbl.pdf.imf" docOrigin="Mitteilungen Der Schweizerischen Entomologischen Gesellschaft 88" docTitle="Scaphisoma poussereaui Löbl, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="370" masterDocId="FF86FF83FF90FFFC9879957AFF96FF9F" masterDocTitle="Scaphisoma poussereaui sp. nov. from La Réunion (Coleoptera: Staphylinidae: Scaphidiinae), arange extension of the S. tricolor species group" masterLastPageNumber="370" masterPageNumber="367" pageNumber="368" updateTime="1732881642526" updateUser="ExternalLinkService">
<mods:mods id="7F9926B380F087E86EE441886DE17B0F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="824C5A7AB3DDF71185A502EC396218AD">
<mods:title id="E99389E04D48CE23DC7B8ADAB0F78894">Scaphisoma poussereaui sp. nov. from La Réunion (Coleoptera: Staphylinidae: Scaphidiinae), arange extension of the S. tricolor species group</mods:title>
</mods:titleInfo>
<mods:name id="6025768A67BB63E396690F5F34DB2D5A" type="personal">
<mods:role id="48542213217449B0D32D5168E6CA3386">
<mods:roleTerm id="6D5D0ACB2302DB850D8AB9A1125897EE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0F60072B7EEB0A8916E37EEF62DEB58F">Ivan Löbl</mods:namePart>
</mods:name>
<mods:typeOfResource id="3C2ACA4894932B2878CF02D6D88D6322">text</mods:typeOfResource>
<mods:relatedItem id="DDF961B309B1E019B0A3FA722BC1BCD4" type="host">
<mods:titleInfo id="08E4CBA63EE653D18BFD6CE972C112D2">
<mods:title id="D6951A66DD60D24CE8D80A736B6519A8">Mitteilungen Der Schweizerischen Entomologischen Gesellschaft</mods:title>
</mods:titleInfo>
<mods:part id="8F7378243C957D273EBEE15CBF5C658F">
<mods:date id="F77FC4E7687A4469A20D009DF7166E39">2015</mods:date>
<mods:detail id="546A763781C28690657529FC947ED4AE" type="volume">
<mods:number id="29B6D6900B16BAAA191B4FC1FC056945">88</mods:number>
</mods:detail>
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<mods:start id="FAFB7A6AF5DF7B994FCD6D431D5E54CF">367</mods:start>
<mods:end id="60005C660CBB105D0998D9D585CD5142">370</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="380E934E06638639A6FA1EE790FD01DE">journal article</mods:classification>
<mods:identifier id="E52C5A3B9E5D5BD83C3D42DC078DD70E" type="DOI">10.5281/zenodo.34001</mods:identifier>
</mods:mods>
<treatment id="03BF87FBFF91FFFF99E5942AFD09FBC3" ID-DOI="http://doi.org/10.5281/zenodo.6094551" ID-GBIF-Taxon="117135279" ID-Zenodo-Dep="6094551" LSID="urn:lsid:plazi:treatment:03BF87FBFF91FFFF99E5942AFD09FBC3" httpUri="http://treatment.plazi.org/id/03BF87FBFF91FFFF99E5942AFD09FBC3" lastPageId="3" lastPageNumber="370" pageId="1" pageNumber="368" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Scaphidiinae">
<subSubSection id="C30C6566FF91FFFD99E5942AFC9DFEF7" box="[412,779,336,360]" pageId="1" pageNumber="368" type="nomenclature">
<paragraph id="8BA936EDFF91FFFD99E5942AFC9DFEF7" box="[412,779,336,360]" pageId="1" pageNumber="368">
<heading id="D0E18181FF91FFFD99E5942AFC9DFEF7" bold="true" box="[412,779,336,360]" centered="true" fontSize="10" level="1" pageId="1" pageNumber="368" reason="4">
<emphasis id="B962EAFFFF91FFFD99E5942AFC9DFEF7" bold="true" box="[412,779,336,360]" pageId="1" pageNumber="368">
<taxonomicName id="4C164D6EFF91FFFD99E5942AFD3AFEF7" ID-CoL="6XSVJ" box="[412,684,336,360]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="368" phylum="Arthropoda" rank="species" species="poussereaui" status="sp. nov.">
<emphasis id="B962EAFFFF91FFFD99E5942AFD3AFEF7" bold="true" box="[412,684,336,360]" italics="true" pageId="1" pageNumber="368">Scaphisoma poussereaui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2515784FF91FFFD9ACD942AFC9DFEF7" box="[692,779,336,360]" pageId="1" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30C6566FF91FFFD9A5F9408FD17FE1A" box="[550,641,370,389]" pageId="1" pageNumber="368" type="description">
<paragraph id="8BA936EDFF91FFFD9A5F9408FD17FE1A" box="[550,641,370,389]" pageId="1" pageNumber="368">
(
<figureCitation id="132D2A68FF91FFFD9A559408FDEFFE1A" box="[556,633,370,389]" captionStart="Figs 1 – 3" captionStartId="2.[138,175,1120,1139]" captionTargetBox="[245,915,123,1102]" captionTargetId="figure@2.[245,915,122,1102]" captionTargetPageId="2" captionText="Figs 1 – 3. Aedeagus of Scaphisoma pousserreaui sp. nov., in ventral view (1), in lateral view (2), median lobe, without parameres in dorsal view (3). Scale bar = 0.2 mm." pageId="1" pageNumber="368">
Figs
<number id="3DDE24CAFF91FFFD9A209408FDEFFE1A" box="[601,633,370,389]" pageId="1" pageNumber="368" value="2.0" valueMax="3.0" valueMin="1.0">1–3</number>
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30C6566FF91FFFD98E094E3FCCCFD96" pageId="1" pageNumber="368" type="materials_examined">
<paragraph id="8BA936EDFF91FFFD98E094E3FD24FE51" pageId="1" pageNumber="368">
<materialsCitation id="3B7E3CB0FF91FFFD98E094E3FD38FE51" ID-GBIF-Occurrence="1249305262" collectingDate="2015-02-19" collectionCode="MNHN" collectorName="Coll. J. Poussereau" country="Réunion" location="/ Chemin de Centure Maison Boyer J. P. Souche Jamblon" pageId="1" pageNumber="368" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B962EAFFFF91FFFD98E094E3FF6DFE2E" bold="true" box="[153,251,409,433]" italics="true" pageId="1" pageNumber="368">
<typeStatus id="54AD884FFF91FFFD98E094E3FF6DFE2E" box="[153,251,409,433]" pageId="1" pageNumber="368">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="9D10FD64FF91FFFD997594E2FEB5FE2F" box="[268,291,408,432]" pageId="1" pageNumber="368" type="male">♂</specimenCount>
,
<collectingCountry id="F301767DFF91FFFD995194E3FE38FE2E" box="[296,430,407,433]" name="Réunion" pageId="1" pageNumber="368" value="Réunion">La Réunion</collectingCountry>
<number id="3DDE24CAFF91FFFD99C194E3FE76FE2E" box="[440,480,409,433]" pageId="1" pageNumber="368" value="974.0">974</number>
<collectorName id="26E3533BFF91FFFD999394E3FD2DFE2E" box="[490,699,409,433]" pageId="1" pageNumber="368">Coll. J. Poussereau</collectorName>
<location id="8EC96036FF91FFFD9ABC94E3FE56FE51" LSID="urn:lsid:plazi:treatment:03BF87FBFF91FFFF99E5942AFD09FBC3:8EC96036FF91FFFD9ABC94E3FE56FE51" country="Réunion" name="/ Chemin de Centure Maison Boyer J. P. Souche Jamblon" pageId="1" pageNumber="368">/Chemin de Centure Maison Boyer J.P. Souche Jamblon</location>
<collectingDate id="EFECE9C5FF91FFFD99BE94CCFDD6FE51" box="[455,576,438,462]" pageId="1" pageNumber="368">
<number id="3DDE24CAFF91FFFD99BE94CCFE74FE51" box="[455,482,438,462]" pageId="1" pageNumber="368" value="19.0">19</number>
0
<number id="3DDE24CAFF91FFFD998F94CCFD92FE51" box="[502,516,438,462]" pageId="1" pageNumber="368" value="2.0">2</number>
<number id="3DDE24CAFF91FFFD9A7294CCFDD6FE51" box="[523,576,438,462]" pageId="1" pageNumber="368" value="2015.0">2015</number>
</collectingDate>
(
<collectionCode id="ED07AE28FF91FFFD9A2994CCFD31FE51" box="[592,679,438,462]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="1" pageNumber="368">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA936EDFF91FFFD98E094AEFCCCFD96" pageId="1" pageNumber="368">
<materialsCitation id="3B7E3CB0FF91FFFD98E094AEFCC0FD96" ID-GBIF-Occurrence="1249305263" collectingDate="2015-02-24" collectionCode="CPJP, MHNG" collectorName="Coll. J. Poussereau" country="Réunion" location="Le Tampon 800 m maison Lebeau M. tronc grand natte" pageId="1" pageNumber="368" specimenCount="7" specimenCount-female="4" specimenCount-male="3" typeStatus="paratype">
<emphasis id="B962EAFFFF91FFFD98E094AEFE93FE74" bold="true" box="[153,261,468,491]" italics="true" pageId="1" pageNumber="368">
<typeStatus id="54AD884FFF91FFFD98E094AEFE93FE74" box="[153,261,468,491]" pageId="1" pageNumber="368">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D10FD64FF91FFFD997394AEFEC3FE74" box="[266,341,467,491]" pageId="1" pageNumber="368" type="male">
<number id="3DDE24CAFF91FFFD997394AEFE8CFE74" box="[266,282,468,491]" pageId="1" pageNumber="368" value="3.0">3</number>
♂♂
</specimenCount>
,
<specimenCount id="9D10FD64FF91FFFD992394AEFE32FE74" box="[346,420,465,491]" pageId="1" pageNumber="368" type="female">
<number id="3DDE24CAFF91FFFD992394AEFEFCFE74" box="[346,362,468,491]" pageId="1" pageNumber="368" value="4.0">4</number>
♀♀
</specimenCount>
,
<collectingCountry id="F301767DFF91FFFD99D094AEFDBFFE74" box="[425,553,465,491]" name="Réunion" pageId="1" pageNumber="368" value="Réunion">La Réunion</collectingCountry>
<number id="3DDE24CAFF91FFFD9A4994AEFDCEFE74" box="[560,600,468,491]" pageId="1" pageNumber="368" value="974.0">974</number>
<collectorName id="26E3533BFF91FFFD9A2694A9FCBDFE74" box="[607,811,467,491]" pageId="1" pageNumber="368">Coll. J. Poussereau</collectorName>
/
<location id="8EC96036FF91FFFD9B4394A9FD89FD96" LSID="urn:lsid:plazi:treatment:03BF87FBFF91FFFF99E5942AFD09FBC3:8EC96036FF91FFFD9B4394A9FD89FD96" country="Réunion" name="Le Tampon 800 m maison Lebeau M. tronc grand natte" pageId="1" pageNumber="368">
Le Tampon
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maison Lebeau M. tronc grand natte
</location>
<collectingDate id="EFECE9C5FF91FFFD9A5F948BFD36FD96" box="[550,672,497,521]" pageId="1" pageNumber="368">
<number id="3DDE24CAFF91FFFD9A5F948BFDD7FD96" box="[550,577,497,521]" pageId="1" pageNumber="368" value="24.0">24</number>
0
<number id="3DDE24CAFF91FFFD9A2F948BFDF5FD96" box="[598,611,497,521]" pageId="1" pageNumber="368" value="2.0">2</number>
<number id="3DDE24CAFF91FFFD9A13948BFD36FD96" box="[618,672,497,521]" pageId="1" pageNumber="368" value="2015.0">2015</number>
</collectingDate>
(
<collectionCode id="ED07AE28FF91FFFD9AD6948BFD7AFD96" box="[687,748,497,521]" pageId="1" pageNumber="368">CPJP</collectionCode>
,
<collectionCode id="ED07AE28FF91FFFD9A81948BFCD8FD96" box="[760,846,497,521]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="1" pageNumber="368">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30C6566FF91FFFF98AF9766FF4FFEDB" lastPageId="3" lastPageNumber="370" pageId="1" pageNumber="368" type="description">
<paragraph id="8BA936EDFF91FFFF98AF9766FD79FF2E" lastPageId="3" lastPageNumber="370" pageId="1" pageNumber="368">
<emphasis id="B962EAFFFF91FFFD98AF9766FEC2FDAC" bold="true" box="[214,340,540,563]" italics="true" pageId="1" pageNumber="368">Description</emphasis>
.Male. Length
<quantity id="4CEE9B08FF91FFFD99929766FD11FDAC" box="[491,647,540,563]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.05" metricValueMin="1.85" pageId="1" pageNumber="368" unit="mm" value="1.95" valueMax="2.05" valueMin="1.85">1.85–2.05 mm</quantity>
, width
<quantity id="4CEE9B08FF91FFFD9AA89766FCF8FDAC" box="[721,878,540,563]" metricMagnitude="-3" metricUnit="m" metricValue="1.365" metricValueMax="1.46" metricValueMin="1.27" pageId="1" pageNumber="368" unit="mm" value="1.365" valueMax="1.46" valueMin="1.27">1.27–1.46 mm</quantity>
. Head, mouthparts, antennae, pronotum, hypomera, and tarsi ochreous. Anterior two thirds of elytra dark brown to blackish, apical third of elytra yellowish. Mesoventrite and metaventrite with epimera and anepisterna reddish-brown. Coxae, femora, tibiae and abdomen light brown, slightly lighter than metaventrite. Antennae with relative length/width of antennomeres as:
<date id="FFA8102DFF91FFFD9A7397D5FDF6FD59" box="[522,608,686,710]" pageId="1" pageNumber="368" value="2013-03-08">
III
<number id="3DDE24CAFF91FFFD9A5497D4FDDFFD59" box="[557,585,686,710]" pageId="1" pageNumber="368" value="13.0">13</number>
/
<number id="3DDE24CAFF91FFFD9A2B97D4FDF6FD59" box="[594,608,686,710]" pageId="1" pageNumber="368" value="8.0">8</number>
</date>
:
<date id="FFA8102DFF91FFFD9A1597D5FD52FD59" box="[620,708,686,710]" pageId="1" pageNumber="368" value="1950-04-09">
IV
<number id="3DDE24CAFF91FFFD9AE897D4FD38FD59" box="[657,686,686,710]" pageId="1" pageNumber="368" value="50.0">50</number>
/
<number id="3DDE24CAFF91FFFD9ACE97D4FD52FD59" box="[695,708,686,710]" pageId="1" pageNumber="368" value="9.0">9</number>
</date>
:
<date id="FFA8102DFF91FFFD9AA997D4FC8EFD59" box="[720,792,686,710]" pageId="1" pageNumber="368" value="1973-05-09">
V
<number id="3DDE24CAFF91FFFD9A9F97D4FC94FD59" box="[742,770,686,710]" pageId="1" pageNumber="368" value="73.0">73</number>
/
<number id="3DDE24CAFF91FFFD9B7297D4FC8EFD59" box="[779,792,686,710]" pageId="1" pageNumber="368" value="9.0">9</number>
</date>
:
<date id="FFA8102DFF91FFFD9B5D97D5FC1FFD59" box="[804,905,686,710]" pageId="1" pageNumber="368" value="1960-06-10">
VI
<number id="3DDE24CAFF91FFFD9B3097D4FCF0FD59" box="[841,870,686,710]" pageId="1" pageNumber="368" value="60.0">60</number>
/
<number id="3DDE24CAFF91FFFD9B1697D4FC1FFD59" box="[879,905,686,710]" pageId="1" pageNumber="368" value="10.0">10</number>
</date>
:
<date id="FFA8102DFF91FFFD9BEC97D5FB95FD59" box="[917,1027,686,710]" pageId="1" pageNumber="368" value="1965-07-12">
VII
<number id="3DDE24CAFF91FFFD9BBA97D4FC49FD59" box="[963,991,686,710]" pageId="1" pageNumber="368" value="65.0">65</number>
/
<number id="3DDE24CAFF91FFFD9B9197D4FB95FD59" box="[1000,1027,686,710]" pageId="1" pageNumber="368" value="12.0">12</number>
</date>
:
<date id="FFA8102DFF91FFFD98E097B6FE94FD7C" box="[153,258,715,739]" pageId="1" pageNumber="368" value="1956-08-09">
VIII
<number id="3DDE24CAFF91FFFD98B797B1FF7DFD7C" box="[206,235,715,739]" pageId="1" pageNumber="368" value="56.0">56</number>
/
<number id="3DDE24CAFF91FFFD988D97B1FE94FD7C" box="[244,258,715,739]" pageId="1" pageNumber="368" value="9.0">9</number>
</date>
:
<date id="FFA8102DFF91FFFD997597B6FEE6FD7C" box="[268,368,715,739]" pageId="1" pageNumber="368" value="1968-09-12">
IX
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/
<number id="3DDE24CAFF91FFFD992C97B1FEE6FD7C" box="[341,368,715,739]" pageId="1" pageNumber="368" value="12.0">12</number>
</date>
:
<date id="FFA8102DFF91FFFD990297B1FE59FD7C" box="[379,463,715,739]" pageId="1" pageNumber="368" value="1965-10-15">
X
<number id="3DDE24CAFF91FFFD99E897B1FE3BFD7C" box="[401,429,715,739]" pageId="1" pageNumber="368" value="65.0">65</number>
/
<number id="3DDE24CAFF91FFFD99CF97B1FE59FD7C" box="[438,463,715,739]" pageId="1" pageNumber="368" value="15.0">15</number>
</date>
:
<date id="FFA8102DFF91FFFD99A397B6FDD7FD7C" box="[474,577,715,739]" pageId="1" pageNumber="368" value="1968-11-14">
XI
<number id="3DDE24CAFF91FFFD998497B1FD8CFD7C" box="[509,538,715,739]" pageId="1" pageNumber="368" value="68.0">68</number>
/
<number id="3DDE24CAFF91FFFD9A5B97B1FDABFD7C" box="[546,573,715,739]" pageId="1" pageNumber="368" value="14.0">14</number>
.
</date>
Pronotum and elytra not microsculptured. Pronotum with lateral margins evenly rounded, lateral margin carinae not visible in dorsal view, lateral striae very densely and distinctly punctate, disc with very fine and fairly dense punctation. Hypomera not microsculptured, with punctation very fine, hardly visible at
<number id="3DDE24CAFF91FFFD9905963BFE01FCC6" box="[380,407,833,857]" pageId="1" pageNumber="368" value="25.0">25</number>
times magnification. Exposed part of scutellum fairly large, about
<quantity id="4CEE9B08FF91FFFD98A59624FEADFCE9" box="[220,315,862,886]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="368" unit="mm" value="0.05">0.05 mm</quantity>
long. Elytra with lateral margins evenly rounded, lateral carinae in dorsal view concealed near bases, exposed posterior basal fourth to third; lateral striae very densely and fairly coarsely punctate; apical margins rounded, indistinctly crenulated near inner angles, inner apical angles situated posterior level of outer apical angles; sutural margins not raised, sutural striae fairly shallow, starting posterior to level of scutellar tip, subparallel; adsutural areas flat, with single puncture row; discal punctation of elytra dense and coarse except on yellowish apical area, fairly irregular, with punctures distinctly delimited, puncture intervals mostly distinctly larger than puncture diameters; punctation on light apical area very fine. Hind wings appearing fully developed. Mesanepisterna not microsculptured. Mesepimera as interval to mesocoxae and about
<number id="3DDE24CAFF91FFFD9A0391F9FD13FB04" box="[634,645,1155,1179]" pageId="1" pageNumber="368" value="3.0">3</number>
times as long as wide. Metaventrite with strigulate microsculpture; punctation very fine, distinct punctures in impressed antecoxal rows and punctures bordering submesocoxal lines excepted; punctures also fairly coarse on small areas near inner margin of metacoxae. Middle part of metaventrite convex, shallowly impressed apically. Submesocoxal lines parallel, submesocoxal areas about
<quantity id="4CEE9B08FF91FFFD99C2906CFD8DFAB1" box="[443,539,1302,1326]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="368" unit="mm" value="0.03">0.03 mm</quantity>
long, about as sixth of interval to metacoxae. Metanepisterna convex, impressed toward suture, below level of metaventrite along suture, narrowed anteriad, with margin mostly oblique, rounded near anterior angles. Protibiae and metatibiae straight, mesotibiae distinctly curved posterior between basal third and apex, about as thick as protibiae, thicker than metatibiae. Protarsomeres
<number id="3DDE24CAFF91FFFD994390D3FED4FA5E" box="[314,322,1449,1473]" pageId="1" pageNumber="368" value="1.0">1</number>
strongly widened, about as wide as apices of protibiae. Protarsomeres
<number id="3DDE24CAFF91FFFD989A90BCFF66FA41" box="[227,240,1478,1502]" pageId="1" pageNumber="368" value="2.0">2</number>
and
<number id="3DDE24CAFF91FFFD995A90BCFEB8FA41" box="[291,302,1478,1502]" pageId="1" pageNumber="368" value="3.0">3</number>
much narrower than protarsomere
<number id="3DDE24CAFF91FFFD9ACF90BCFD52FA41" box="[694,708,1478,1502]" pageId="1" pageNumber="368" value="1.0">1</number>
. Mesotarsomeres
<number id="3DDE24CAFF91FFFD9BF690BCFC01FA41" box="[911,919,1478,1502]" pageId="1" pageNumber="368" value="1.0">1</number>
strongly widened, wider than apices of tibiae. Mesotarsomere
<number id="3DDE24CAFF92FFFE9AA091C4FD73FB49" box="[729,741,1214,1238]" pageId="2" pageNumber="369" value="2.0">2</number>
slightly narrower than
<number id="3DDE24CAFF92FFFE9B9D91C4FC67FB49" box="[996,1009,1214,1238]" pageId="2" pageNumber="369" value="1.0">1</number>
, mesotarsomeres
<number id="3DDE24CAFF92FFFE994491A1FEDEFB6B" box="[317,328,1243,1268]" pageId="2" pageNumber="369" value="3.0">3</number>
weakly widened. Abdomen with strigulate microsculpture, very finely punctate except on anarrow area posterior to the intercoxal process and at margins of submetaxocal line. Submetacoxal lines convex, submetacoxal areas
<quantity id="4CEE9B08FF92FFFE9BA9906CFF20FAD3" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="369" unit="mm" value="0.06">0.06 mm</quantity>
, almost as third of interval to apical margin of sternite
<number id="3DDE24CAFF92FFFE9A87904EFC9AFAD3" box="[766,780,1332,1356]" pageId="2" pageNumber="369" value="1.0">1</number>
. Sternite
<number id="3DDE24CAFF92FFFE9B0B9049FCE8FAD3" box="[882,894,1331,1356]" pageId="2" pageNumber="369" value="6.0">6</number>
flattened in middle, extended apically to form large, broadly rounded lobe. Aedeagus (
<figureCitation id="132D2A68FF92FFFE9BE0902BFC63FAF6" box="[921,1013,1361,1385]" captionStart="Figs 1 – 3" captionStartId="2.[138,175,1120,1139]" captionTargetBox="[245,915,123,1102]" captionTargetId="figure@2.[245,915,122,1102]" captionTargetPageId="2" captionText="Figs 1 – 3. Aedeagus of Scaphisoma pousserreaui sp. nov., in ventral view (1), in lateral view (2), median lobe, without parameres in dorsal view (3). Scale bar = 0.2 mm." pageId="2" pageNumber="369">
Figs
<number id="3DDE24CAFF92FFFE9BB7902BFC63FAF6" box="[974,1013,1361,1385]" pageId="2" pageNumber="369" value="2.0" valueMax="3.0" valueMin="1.0">1–3</number>
</figureCitation>
)
<quantity id="4CEE9B08FF92FFFE98F39014FF7CFA19" box="[138,234,1390,1414]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" pageId="2" pageNumber="369" unit="mm" value="1.28">1.28 mm</quantity>
long. Basal bulb weakly sclerotized. Ventral branch of apical process of median lobe strongly inflexed lateroventrally, somewhat flattened, weakly narrowed toward tip in dorsal view. Parameres each with large and strongly sclerotized subbasal apophyses bearing membranous lobe. Internal sac with proximal, strongly bent, large tooth joined to moderately sclerotized area bearing three rods; apical part of internal sac very densely and very finely denticulate.
</paragraph>
<caption id="DF696665FF92FFFE98F3911AFC93FB15" pageId="2" pageNumber="369" targetBox="[245,915,123,1102]" targetPageId="2">
<paragraph id="8BA936EDFF92FFFE98F3911AFC93FB15" pageId="2" pageNumber="369">
Figs 1–3. Aedeagus of
<taxonomicName id="4C164D6EFF92FFFE9927911AFDD7FBEC" box="[350,577,1120,1139]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="369" phylum="Arthropoda" rank="species" species="poussereaui" status="sp. nov.">
<emphasis id="B962EAFFFF92FFFE9927911AFDD7FBEC" bold="true" box="[350,577,1120,1139]" italics="true" pageId="2" pageNumber="369">Scaphisoma pousserreaui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2515784FF92FFFE9A30911AFD05FBEC" box="[585,659,1120,1139]" pageId="2" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
, in ventral view (1), in lateral view (2), median lobe, without parameres in dorsal view (3). Scale bar = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BA936EDFF93FFFF98AF95CCFF4FFEDB" pageId="3" pageNumber="370">
Female. With most characters as in male, but: Pronotum slightly lighter than or as dark as anterior two thirds of elytra. Apical elytral margins almost oblique, with inner angles slightly prominent, tooth-like and very finely crenulated. Mesotibiae weakly bent, about as thick as metatibiae. All tarsomeres narrow. Sternite
<number id="3DDE24CAFF93FFFF9C799475FB98FEB9" box="[1024,1038,271,294]" pageId="3" pageNumber="370" value="6.0">6</number>
not lobed.
</paragraph>
</subSubSection>
<subSubSection id="C30C6566FF93FFFF98AF9433FEBCFEE1" pageId="3" pageNumber="370" type="etymology">
<paragraph id="8BA936EDFF93FFFF98AF9433FEBCFEE1" pageId="3" pageNumber="370">
<emphasis id="B962EAFFFF93FFFF98AF9433FEDEFEFE" bold="true" box="[214,328,329,353]" italics="true" pageId="3" pageNumber="370">Etymology</emphasis>
.The new species is named in honour of its collector, Jacques Poussereau., Dax, France
</paragraph>
</subSubSection>
<subSubSection id="C30C6566FF93FFFF98AF94FEFD09FBC3" pageId="3" pageNumber="370" type="discussion">
<paragraph id="8BA936EDFF93FFFF98AF94FEFD09FBC3" pageId="3" pageNumber="370">
<emphasis id="B962EAFFFF93FFFF98AF94FEFED0FE03" bold="true" box="[214,326,388,412]" italics="true" pageId="3" pageNumber="370">Comments</emphasis>
.The species is amember of the
<emphasis id="B962EAFFFF93FFFF9AD594FEFC8CFE03" bold="true" box="[684,794,388,412]" italics="true" pageId="3" pageNumber="370">S. tricolor</emphasis>
group defined in Löbl (
<number id="3DDE24CAFF93FFFF98DB94DBFF4DFE26" box="[162,219,417,441]" pageId="3" pageNumber="370" value="1975.0">1975</number>
:
<number id="3DDE24CAFF93FFFF989F94DBFE99FE26" box="[230,271,417,441]" pageId="3" pageNumber="370" value="272.0">272</number>
) containing the following species:
<emphasis id="B962EAFFFF93FFFF9AF394DBFC94FE26" bold="true" box="[650,770,417,441]" italics="true" pageId="3" pageNumber="370">S. affectum</emphasis>
Löbl,
<number id="3DDE24CAFF93FFFF9B3394DBFC17FE26" box="[842,897,417,441]" pageId="3" pageNumber="370" value="2015.0">2015</number>
,
<emphasis id="B962EAFFFF93FFFF9BF494DBFB98FE26" bold="true" box="[909,1038,417,441]" italics="true" pageId="3" pageNumber="370">S. aspectum</emphasis>
Löbl,
<number id="3DDE24CAFF93FFFF98A394C5FE84FE49" box="[218,274,447,470]" pageId="3" pageNumber="370" value="2015.0">2015</number>
,
<emphasis id="B962EAFFFF93FFFF996794C5FE3EFE49" bold="true" box="[286,424,447,470]" italics="true" pageId="3" pageNumber="370">S. borneense</emphasis>
Pic,
<number id="3DDE24CAFF93FFFF999994C5FD8EFE49" box="[480,536,447,470]" pageId="3" pageNumber="370" value="1916.0">1916</number>
,
<emphasis id="B962EAFFFF93FFFF9A5D94C5FD15FE49" bold="true" box="[548,643,447,470]" italics="true" pageId="3" pageNumber="370">S. chujoi</emphasis>
Löbl,
<number id="3DDE24CAFF93FFFF9AB594C5FC95FE49" box="[716,771,447,470]" pageId="3" pageNumber="370" value="1982.0">1982</number>
,
<emphasis id="B962EAFFFF93FFFF9B6994C5FCFBFE49" bold="true" box="[784,877,447,470]" italics="true" pageId="3" pageNumber="370">S. hajeki</emphasis>
Löbl,
<number id="3DDE24CAFF93FFFF9BCF94C5FC7BFE49" box="[950,1005,447,470]" pageId="3" pageNumber="370" value="2012.0">2012</number>
,
<emphasis id="B962EAFFFF93FFFF9B8394C5FF70FE6B" bold="true" italics="true" pageId="3" pageNumber="370">S. luctans</emphasis>
Löbl,
<number id="3DDE24CAFF93FFFF995294A6FEF5FE6B" box="[299,355,476,500]" pageId="3" pageNumber="370" value="1986.0">1986</number>
,
<emphasis id="B962EAFFFF93FFFF991494A6FE65FE6B" bold="true" box="[365,499,476,500]" italics="true" pageId="3" pageNumber="370">S. maculiger</emphasis>
Löbl,
<number id="3DDE24CAFF93FFFF9A4194A6FDE6FE6B" box="[568,624,476,500]" pageId="3" pageNumber="370" value="1875.0">1875</number>
,
<emphasis id="B962EAFFFF93FFFF9A0394A6FCB4FE6B" bold="true" box="[634,802,476,500]" italics="true" pageId="3" pageNumber="370">S. renominatum</emphasis>
Löbl,
<number id="3DDE24CAFF93FFFF9B1194A6FC09FE6B" box="[872,927,476,500]" pageId="3" pageNumber="370" value="1975.0">1975</number>
,
<emphasis id="B962EAFFFF93FFFF9BD094A6FB98FE6B" bold="true" box="[937,1038,476,500]" italics="true" pageId="3" pageNumber="370">S. tarsale</emphasis>
Löbl,
<number id="3DDE24CAFF93FFFF98A59483FE82FD8E" box="[220,276,505,529]" pageId="3" pageNumber="370" value="2015.0">2015</number>
,
<emphasis id="B962EAFFFF93FFFF995B9483FD9BFD8E" bold="true" box="[290,525,505,529]" italics="true" pageId="3" pageNumber="370">S. testaceomaculatum</emphasis>
(Pic,
<number id="3DDE24CAFF93FFFF9A2A9483FD1FFD8E" box="[595,649,505,529]" pageId="3" pageNumber="370" value="1915.0">1915</number>
) and
<emphasis id="B962EAFFFF93FFFF9AB29483FCACFD8E" bold="true" box="[715,826,505,529]" italics="true" pageId="3" pageNumber="370">S. tricolor</emphasis>
Heller,
<number id="3DDE24CAFF93FFFF9BEE9483FC46FD8E" box="[919,976,505,529]" pageId="3" pageNumber="370" value="1917.0">1917</number>
. Two additional species,
<emphasis id="B962EAFFFF93FFFF9911976DFE6FFDB1" bold="true" box="[360,505,535,558]" italics="true" pageId="3" pageNumber="370">S. dentipenne</emphasis>
Löbl,
<number id="3DDE24CAFF93FFFF9A3B976DFDEEFDB1" box="[578,632,535,558]" pageId="3" pageNumber="370" value="1971.0">1971</number>
and
<emphasis id="B962EAFFFF93FFFF9AD7976DFCCCFDB1" bold="true" box="[686,858,535,558]" italics="true" pageId="3" pageNumber="370">S. philippinense</emphasis>
Oberthür,
<number id="3DDE24CAFF93FFFF9BAB976DFB9FFDB1" box="[978,1033,535,558]" pageId="3" pageNumber="370" value="1883.0">1883</number>
, known only from females, may also be members of the group. The group was defined mainly by having conspicuously large aedeagi with moderately sclerotized basal bulb, usually bearing aventral ridge supporting robust external muscles, the apical process of median lobe is bifid, strongly sclerotized and asymmetrical, the parameres are complex and the expanded membranous apex is supported by strongly sclerotized apophyses, and the internal sac is complex, often containing a curved proximal tooth. The species of the
<emphasis id="B962EAFFFF93FFFF9A29979EFD2DFD63" bold="true" box="[592,699,740,764]" italics="true" pageId="3" pageNumber="370">S. tricolor</emphasis>
group have comparatively large bodies, long antennae, the metaventrite with antecoxal puncture rows, the abdominal microsculpture strigulate, the male sternite
<number id="3DDE24CAFF93FFFF9A0D9664FD16FCA9" box="[628,640,798,822]" pageId="3" pageNumber="370" value="6.0">6</number>
with alarge, broadly rounded lobe. Having shortened sutural striae of the elytra is another diagnostic feature for some species and in anumber of examined undescribed Philippine species (the striae are elongate in
<emphasis id="B962EAFFFF93FFFF996F960DFE1BFC11" bold="true" box="[278,397,887,910]" italics="true" pageId="3" pageNumber="370">S. affectum</emphasis>
,
<emphasis id="B962EAFFFF93FFFF99E3960DFD8CFC11" bold="true" box="[410,538,887,910]" italics="true" pageId="3" pageNumber="370">S. aspectum</emphasis>
,
<emphasis id="B962EAFFFF93FFFF9A5E960DFD13FC11" bold="true" box="[551,645,887,910]" italics="true" pageId="3" pageNumber="370">S. chujoi</emphasis>
,
<emphasis id="B962EAFFFF93FFFF9AEA960DFD79FC11" bold="true" box="[659,751,887,910]" italics="true" pageId="3" pageNumber="370">S. hajeki</emphasis>
,
<emphasis id="B962EAFFFF93FFFF9A85960DFC15FC11" bold="true" box="[764,899,887,910]" italics="true" pageId="3" pageNumber="370">S. maculiger</emphasis>
and
<emphasis id="B962EAFFFF93FFFF9BCE960DFEA3FC33" bold="true" italics="true" pageId="3" pageNumber="370">S. testaceomaculatum</emphasis>
). The apical margins of the elytra are sexually dimorphic for some species, and males may also differ from females by their pronotal coloration.
<taxonomicName id="4C164D6EFF93FFFF9BA796CBFEEAFC79" class="Insecta" family="Staphylinidae" genus="Scaphisoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="370" phylum="Arthropoda" rank="species" species="poussereaui">
<emphasis id="B962EAFFFF93FFFF9BA796CBFEEAFC79" bold="true" italics="true" pageId="3" pageNumber="370">Scaphisoma poussereaui</emphasis>
</taxonomicName>
is readily distinguished from all members of the group by its elytral coloration and the aedeagal characters, in particular by the shape of the ventral branch of the apical process inflexed strongly ventrolaterally. In addition to these characters,
<taxonomicName id="4C164D6EFF93FFFF992F915DFE63FBA1" box="[342,501,1063,1086]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="370" phylum="Arthropoda" rank="species" species="poussereaui">
<emphasis id="B962EAFFFF93FFFF992F915DFE63FBA1" bold="true" box="[342,501,1063,1086]" italics="true" pageId="3" pageNumber="370">S. poussereaui</emphasis>
</taxonomicName>
may be easily distinguished from its Mascarene congeners by the elytra with short sutural striae.
</paragraph>
</subSubSection>
</treatment>
</document>
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