treatments-xml/data/03/BF/87/03BF87FB6372E840FF209E7743AD37F0.xml
2024-06-21 12:22:17 +02:00

160 lines
22 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="F7473365785978EDD35F134EC7E8E9AE" ID-DOI="10.11646/zootaxa.3616.5.1" ID-GBIF-Dataset="61a7f15d-e430-465e-b732-92e7486fec0b" ID-ISSN="1175-5326" ID-Zenodo-Dep="220125" ID-ZooBank="D7C95348-727D-499C-8E67-23C6ADFCAFC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460576386170" checkinUser="plazi" docAuthor="Arango, Claudia P. &amp; Brenneis, Georg" docDate="2013" docId="03BF87FB6372E840FF209E7743AD37F0" docLanguage="en" docName="zt03616p436.pdf" docOrigin="Zootaxa 3616 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudopallene brevicephala Staples 2008" docType="treatment" docVersion="8" lastPageNumber="418" masterDocId="FF86FF836363E852FFB7984241683574" masterDocTitle="New species of Australian Pseudopallene (Pycnogonida: Callipallenidae) based on live colouration, morphology and DNA" masterLastPageNumber="436" masterPageNumber="401" pageNumber="418" updateTime="1698323344127" updateUser="plazi">
<mods:mods id="1D13C99D92D7855B3FFCAE9AC2908DC0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="28A509EADDB0ED0DB154C4156F121E91">
<mods:title id="BCC59DD5C40D2D930FC33EB4559871A4">New species of Australian Pseudopallene (Pycnogonida: Callipallenidae) based on live colouration, morphology and DNA</mods:title>
</mods:titleInfo>
<mods:name id="045D34385D39C6BD68CD8787234972C6" type="personal">
<mods:role id="41EFE8E3835E9C1B92BEE36DECFAEB68">
<mods:roleTerm id="A116E7CBF51F994BB104D69000510B6F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AC92A368F72937E64B578C6EA1481E69">Arango, Claudia P.</mods:namePart>
</mods:name>
<mods:name id="C4D49608A636C02FF740A75D40F97678" type="personal">
<mods:role id="D412EE2B0D285FFC48B486BA921EBC70">
<mods:roleTerm id="463FB6431E1D7D58DBB707385EAA82E7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FC8D005DE0B6FCDDCE46FCF8296E303A">Brenneis, Georg</mods:namePart>
</mods:name>
<mods:typeOfResource id="8D3E5E8D82E58DE1EF062B6D75AB11D2">text</mods:typeOfResource>
<mods:relatedItem id="A1F9792B7D0770C63A684B2F05FEB11A" type="host">
<mods:titleInfo id="AFE2E43DA51D962FFD41C232831A50FB">
<mods:title id="96216D8CE05867DC19160F19C21ED0CB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E8AEFD491D620669738C8C91519A100C">
<mods:date id="CE72CB52650D493E8FB91DE90B85545C">2013</mods:date>
<mods:detail id="632C1B561A53585DAD558A5929AF217A" type="volume">
<mods:number id="7E99EA89F5C949F38F00C8A3C1766945">3616</mods:number>
</mods:detail>
<mods:detail id="9BD6F998E402C7C0361841109AD7F80B" type="issue">
<mods:number id="5CF96610B17462C677D817EA0E0987A1">5</mods:number>
</mods:detail>
<mods:extent id="D057966D59DCE5DC4CBDE6EDF24FF201" unit="page">
<mods:start id="2DF69479E43DAA01206BE862AFAB9286">401</mods:start>
<mods:end id="3D1D11C0EFF8A99D35877AD1349102E6">436</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F30D3A307A87D20766411086E7DA078B">journal article</mods:classification>
<mods:identifier id="25402933F0EADCFC27961248AE84ACFF" type="DOI">10.11646/zootaxa.3616.5.1</mods:identifier>
<mods:identifier id="5ACB4DE152D0CDDCBA67F90FC40C4E67" type="GBIF-Dataset">61a7f15d-e430-465e-b732-92e7486fec0b</mods:identifier>
<mods:identifier id="E1BD4B0608FE08DCE302E68D73607B62" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4E8EE826138CDC8954B550D8B5CC79BA" type="Zenodo-Dep">220125</mods:identifier>
<mods:identifier id="750478C45FED403D3C85E9905C476DB2" type="ZooBank">D7C95348-727D-499C-8E67-23C6ADFCAFC4</mods:identifier>
</mods:mods>
<treatment id="03BF87FB6372E840FF209E7743AD37F0" ID-DOI="http://doi.org/10.5281/zenodo.5678924" ID-GBIF-Taxon="119560085" ID-Zenodo-Dep="5678924" LSID="urn:lsid:plazi:treatment:03BF87FB6372E840FF209E7743AD37F0" httpUri="http://treatment.plazi.org/id/03BF87FB6372E840FF209E7743AD37F0" lastPageId="18" lastPageNumber="418" pageId="17" pageNumber="418">
<subSubSection id="C30C65666372E843FF209E77401B3304" pageId="17" pageNumber="418" type="nomenclature">
<paragraph id="8BA936ED6372E843FF209E7743F9333B" blockId="17.[151,657,1589,1648]" box="[151,657,1589,1615]" pageId="17" pageNumber="418">
<heading id="D0E181816372E843FF209E7743F9333B" bold="true" box="[151,657,1589,1615]" fontSize="11" level="1" pageId="17" pageNumber="418" reason="1">
<taxonomicName id="4C164D6E6372E843FF209E7743F9333B" ID-CoL="61cac803-77a9-4697-98cf-1eb39f97f3ab" authority="Staples, 2008" authorityName="Staples" authorityYear="2008" box="[151,657,1589,1615]" class="Pycnogonida" family="Callipallenidae" genus="Pseudopallene" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="brevicephala">
<emphasis id="B962EAFF6372E843FF209E7743F9333B" bold="true" box="[151,657,1589,1615]" pageId="17" pageNumber="418">
<emphasis id="B962EAFF6372E843FF209E774081333B" bold="true" box="[151,489,1589,1615]" italics="true" pageId="17" pageNumber="418">Pseudopallene brevicephala</emphasis>
Staples, 2008
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA936ED6372E843FF209E1A401B3304" blockId="17.[151,657,1589,1648]" box="[151,371,1624,1648]" pageId="17" pageNumber="418">
<figureCitation id="132D2A686372E843FF209E1A418C3304" box="[151,228,1624,1648]" captionStart="FIGURE 2" captionStartId="3.[151,250,1742,1765]" captionTargetBox="[196,1403,187,1717]" captionTargetId="figure@3.[186,1410,181,1725]" captionTargetPageId="3" captionText="FIGURE 2. Live colouration of some of the species of Pseudopallene and Stylopallene included in the study. A, fresh live bryozoans Orthoscuticella spp. with many Pseudopallene specimens; B, P. constricta sp. nov., male in ventral view with egg masses attached to the ovigers; C, P. variabilis - complex, frontal-ventral view, note black tips of proboscis and chela fingers; D, P. flava sp. nov. in its natural habitat; E, Close-up of P. brevicephala male characterized by uniform orange-red body and legs, bearing orange eggs and transparent post-embryonic stages; F, P. pachycheira attached to bryozoan, showing banding pattern and constrictions of long leg articles; G, P. h a r r i s i sp. nov. in its natural habitat showing red-and-yellow colouration; H, specimen from the P. variabilis - complex with yellow-and-red colouration attached to live bryozoan; I, Stylopallene cheilorhynchus showing characteristic live black-and-creamy-white banding, photograph taken in captivity. All specimens from Eaglehawk Neck, Tasmania except 2 G and 2 H collected at Bass point, New South Wales." httpUri="https://zenodo.org/record/220127/files/figure.png" pageId="17" pageNumber="418">Figs. 2</figureCitation>
E, 3G, 9AD
</paragraph>
</subSubSection>
<subSubSection id="C30C65666372E843FF209EE2454B3250" pageId="17" pageNumber="418" type="materials_examined">
<paragraph id="8BA936ED6372E843FF209EE2454B3250" blockId="17.[151,1436,1696,2009]" pageId="17" pageNumber="418">
<emphasis id="B962EAFF6372E843FF209EE240ED33CD" bold="true" box="[151,389,1696,1721]" pageId="17" pageNumber="418">Material examined:</emphasis>
(S922219) 1 sub-adult (TAS28),
<date id="FFA8102D6372E843FCBE9EE242F033CC" box="[777,920,1696,1720]" pageId="17" pageNumber="418" value="2007-01-26">26 Jan 2007</date>
, Fortescue Bay,
<quantity id="4CEE9B086372E843FBEF9EE245FB33CC" box="[1112,1171,1696,1720]" metricMagnitude="1" metricUnit="m" metricValue="2.1" pageId="17" pageNumber="418" unit="m" value="21.0">21 m</quantity>
depth in kelp forest;
<specimenCount id="9D10FD646372E843FA399EE2418B33A8" pageId="17" pageNumber="418" type="female">1 female</specimenCount>
(PSE7),
<specimenCount id="9D10FD646372E843FEFD9E8640FE33A8" box="[330,406,1732,1756]" pageId="17" pageNumber="418" type="male">1 male</specimenCount>
(PSE7a), 2 sub-adults (PSE7b, PSE12), (
<accessionNumber id="9445AB0E6372E843FCDF9E8642A833A8" box="[872,960,1732,1756]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S92221" pageId="17" pageNumber="418" type="EnaNcbi">S92221</accessionNumber>
) 2 post-embryonic stages (PSE12a, b) (no DNA available),
<date id="FFA8102D6372E843FEEE9EAA40993274" box="[345,497,1768,1792]" pageId="17" pageNumber="418" value="2009-11-26">Nov-26 2009</date>
, Patterson Arch, Eaglehawk Neck,
<quantity id="4CEE9B086372E843FC339EAA42803274" box="[900,1000,1768,1792]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="17" pageNumber="418" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">1015 m</quantity>
depth, on
<taxonomicName id="4C164D6E6372E843FBD69EAA44673274" box="[1121,1295,1768,1792]" class="Gymnolaemata" family="Catenicellidae" genus="Orthoscuticella" kingdom="Animalia" order="Cheilostomatida" pageId="17" pageNumber="418" phylum="Bryozoa" rank="genus">
<emphasis id="B962EAFF6372E843FBD69EAA44673274" box="[1121,1295,1768,1792]" italics="true" pageId="17" pageNumber="418">Orthoscuticella</emphasis>
</taxonomicName>
spp; 3 postembryonic stages (PSE9, PSE9a, b),
<date id="FFA8102D6372E843FD839F4E43A43250" box="[564,716,1804,1828]" pageId="17" pageNumber="418" value="2009-11-22">Nov-22 2009</date>
, Boulder Point,
<quantity id="4CEE9B086372E843FC369F4E42BE3250" box="[897,982,1804,1828]" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="17" pageNumber="418" unit="m" value="7.5" valueMax="10.0" valueMin="5.0">510 m</quantity>
depth.
</paragraph>
</subSubSection>
<subSubSection id="C30C65666372E840FF709F7343AD37F0" lastPageId="18" lastPageNumber="419" pageId="17" pageNumber="418" type="discussion">
<paragraph id="8BA936ED6372E840FF709F7343AD37F0" blockId="17.[151,1436,1696,2009]" lastBlockId="18.[151,1436,151,644]" lastPageId="18" lastPageNumber="419" pageId="17" pageNumber="418">
<emphasis id="B962EAFF6372E843FF709F734055323D" bold="true" box="[199,317,1841,1865]" pageId="17" pageNumber="418">Remarks:</emphasis>
The adult morphology of the Tasmanian specimens corresponds to the species description based on material from South
<collectingCountry id="F301767D6372E843FE2E9F1643683218" box="[409,512,1876,1900]" name="Australia" pageId="17" pageNumber="418">Australia</collectingCountry>
(Staples 2008), especially the diagnostic short neck (
<figureCitation id="132D2A686372E843FB2E9F16458B3218" box="[1177,1251,1876,1900]" captionStart="FIGURE 9" captionStartId="18.[151,250,1711,1734]" captionTargetBox="[155,1424,707,1680]" captionTargetId="figure@18.[151,1436,700,1691]" captionTargetPageId="18" captionText="FIGURE 9. Pseudopallene brevicephala Staples, 2008. A, B, female (S 92219); A, body in dorsal-lateral view showing the diagnostic short neck; B, frontal-ventral view of cheliphores, proboscis and cephalic margin; C, D, male (S 92219). C, distal articles of the oviger, strigilis, terminal claw and distal apophysis of fifth article; D, propodus and terminal claw of third walking leg. Scale bars on stereomicroscopic images = 1 mm." httpUri="https://zenodo.org/record/220134/files/figure.png" pageId="17" pageNumber="418">Fig. 9</figureCitation>
A), the distinct longitudinal cuticular division line on the cephalon (
<figureCitation id="132D2A686372E843FCBB9F3A423E32E4" box="[780,854,1912,1936]" captionStart="FIGURE 9" captionStartId="18.[151,250,1711,1734]" captionTargetBox="[155,1424,707,1680]" captionTargetId="figure@18.[151,1436,700,1691]" captionTargetPageId="18" captionText="FIGURE 9. Pseudopallene brevicephala Staples, 2008. A, B, female (S 92219); A, body in dorsal-lateral view showing the diagnostic short neck; B, frontal-ventral view of cheliphores, proboscis and cephalic margin; C, D, male (S 92219). C, distal articles of the oviger, strigilis, terminal claw and distal apophysis of fifth article; D, propodus and terminal claw of third walking leg. Scale bars on stereomicroscopic images = 1 mm." httpUri="https://zenodo.org/record/220134/files/figure.png" pageId="17" pageNumber="418">Fig. 9</figureCitation>
B), the oviger compound spine count (8:7:6:
<quantity id="4CEE9B086372E843FADF9F3A44F432E4" box="[1384,1436,1912,1936]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="17" pageNumber="418" unit="in" value="6.0">6 in</quantity>
specimen PSE7) (
<figureCitation id="132D2A686372E843FEE89FDE40C932C0" box="[351,417,1948,1972]" captionStart="FIGURE 9" captionStartId="18.[151,250,1711,1734]" captionTargetBox="[155,1424,707,1680]" captionTargetId="figure@18.[151,1436,700,1691]" captionTargetPageId="18" captionText="FIGURE 9. Pseudopallene brevicephala Staples, 2008. A, B, female (S 92219); A, body in dorsal-lateral view showing the diagnostic short neck; B, frontal-ventral view of cheliphores, proboscis and cephalic margin; C, D, male (S 92219). C, distal articles of the oviger, strigilis, terminal claw and distal apophysis of fifth article; D, propodus and terminal claw of third walking leg. Scale bars on stereomicroscopic images = 1 mm." httpUri="https://zenodo.org/record/220134/files/figure.png" pageId="17" pageNumber="418">Fig. 9</figureCitation>
C) and the propodus configuration (
<figureCitation id="132D2A686372E843FC849FDE421F32C0" box="[819,887,1948,1972]" captionStart="FIGURE 9" captionStartId="18.[151,250,1711,1734]" captionTargetBox="[155,1424,707,1680]" captionTargetId="figure@18.[151,1436,700,1691]" captionTargetPageId="18" captionText="FIGURE 9. Pseudopallene brevicephala Staples, 2008. A, B, female (S 92219); A, body in dorsal-lateral view showing the diagnostic short neck; B, frontal-ventral view of cheliphores, proboscis and cephalic margin; C, D, male (S 92219). C, distal articles of the oviger, strigilis, terminal claw and distal apophysis of fifth article; D, propodus and terminal claw of third walking leg. Scale bars on stereomicroscopic images = 1 mm." httpUri="https://zenodo.org/record/220134/files/figure.png" pageId="17" pageNumber="418">Fig. 9</figureCitation>
D). One character to add to the description of the species is the uniform red-orange colour on body and legs in live adults, sub-adults and post-embryonic stages at least for the Tasmanian specimens (
<figureCitation id="132D2A686371E840FD9C98D5430735DB" box="[555,623,151,175]" captionStart="FIGURE 2" captionStartId="3.[151,250,1742,1765]" captionTargetBox="[196,1403,187,1717]" captionTargetId="figure@3.[186,1410,181,1725]" captionTargetPageId="3" captionText="FIGURE 2. Live colouration of some of the species of Pseudopallene and Stylopallene included in the study. A, fresh live bryozoans Orthoscuticella spp. with many Pseudopallene specimens; B, P. constricta sp. nov., male in ventral view with egg masses attached to the ovigers; C, P. variabilis - complex, frontal-ventral view, note black tips of proboscis and chela fingers; D, P. flava sp. nov. in its natural habitat; E, Close-up of P. brevicephala male characterized by uniform orange-red body and legs, bearing orange eggs and transparent post-embryonic stages; F, P. pachycheira attached to bryozoan, showing banding pattern and constrictions of long leg articles; G, P. h a r r i s i sp. nov. in its natural habitat showing red-and-yellow colouration; H, specimen from the P. variabilis - complex with yellow-and-red colouration attached to live bryozoan; I, Stylopallene cheilorhynchus showing characteristic live black-and-creamy-white banding, photograph taken in captivity. All specimens from Eaglehawk Neck, Tasmania except 2 G and 2 H collected at Bass point, New South Wales." httpUri="https://zenodo.org/record/220127/files/figure.png" pageId="18" pageNumber="419">Fig. 2</figureCitation>
E). Clark (1963) described
<taxonomicName id="4C164D6E6371E840FC1498DB429735C4" box="[931,1023,152,176]" class="Pycnogonida" family="Callipallenidae" genus="Pseudopallene" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="dubia">
<emphasis id="B962EAFF6371E840FC1498DB429735C4" box="[931,1023,152,176]" italics="true" pageId="18" pageNumber="419">P. dubia</emphasis>
</taxonomicName>
based on material from Port Arthur,
<taxonomicName id="4C164D6E6371E840FF2098FE406D35A0" box="[151,261,188,212]" class="Pycnogonida" family="Callipallenidae" genus="Pseudopallene" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="18" pageNumber="432" phylum="Arthropoda" rank="species" species="tasmania">Tasmania</taxonomicName>
, with different
<typeStatus id="54AD884F6371E840FE0098FF409A35A1" box="[439,498,189,213]" pageId="18" pageNumber="419">types</typeStatus>
of chela shapes. Some specimens had a characteristic delicately-curved tube-like elongation of the movable finger, whereas others were lacking this feature. Staples (2005) already proved the
<taxonomicName id="4C164D6E6371E840FA3098A341BF3468" class="Pycnogonida" family="Callipallenidae" genus="Pseudopallene" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="dubia">
<emphasis id="B962EAFF6371E840FA3098A341BF3468" italics="true" pageId="18" pageNumber="419">P. dubia</emphasis>
</taxonomicName>
specimens to represent late developmental stages of
<taxonomicName id="4C164D6E6371E840FC81994642B33468" box="[822,987,260,284]" class="Pycnogonida" family="Callipallenidae" genus="Pseudopallene" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="B962EAFF6371E840FC81994642B33468" box="[822,987,260,284]" italics="true" pageId="18" pageNumber="419">Pseudopallene</emphasis>
</taxonomicName>
rather than fully grown adults, without assigning them to a species. So far, post-embryonic stages of different
<taxonomicName id="4C164D6E6371E840FC01996A45333434" box="[950,1115,296,320]" class="Pycnogonida" family="Callipallenidae" genus="Pseudopallene" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="B962EAFF6371E840FC01996A45333434" box="[950,1115,296,320]" italics="true" pageId="18" pageNumber="419">Pseudopallene</emphasis>
</taxonomicName>
species have been shown to exhibit the tube-like projection of the movable chela finger, e.g., in
<taxonomicName id="4C164D6E6371E840FC3F990F45C13410" authority="Staples, 2005" authorityName="Staples" authorityYear="2005" box="[904,1193,332,356]" class="Pycnogonida" family="Callipallenidae" genus="Pseudopallene" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="watsonae">
<emphasis id="B962EAFF6371E840FC3F990F45653410" box="[904,1037,333,356]" italics="true" pageId="18" pageNumber="419">P. watsonae</emphasis>
Staples, 2005
</taxonomicName>
or in
<taxonomicName id="4C164D6E6371E840FB5D990F441D3410" box="[1258,1397,333,356]" class="Pycnogonida" family="Callipallenidae" genus="Pseudopallene" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="constricta" status="sp. nov.">
<emphasis id="B962EAFF6371E840FB5D990F441D3410" box="[1258,1397,333,356]" italics="true" pageId="18" pageNumber="419">P. constricta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25157846371E840FACB990F41AF34F3" pageId="18" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
(see below, see also Brenneis
<emphasis id="B962EAFF6371E840FDAA9933433D34FC" box="[541,597,368,392]" italics="true" pageId="18" pageNumber="419">et al.</emphasis>
2011b). A detailed morphological description of our genetically identified orange-red post-embryonic stages of
<taxonomicName id="4C164D6E6371E840FD8299D7438934D8" box="[565,737,404,428]" class="Pycnogonida" family="Callipallenidae" genus="Pseudopallene" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="brevicephala">
<emphasis id="B962EAFF6371E840FD8299D7438934D8" box="[565,737,404,428]" italics="true" pageId="18" pageNumber="419">P. brevicephala</emphasis>
</taxonomicName>
from Tasmanian is still underway. However, first observations have shown that the movable chela finger of these individuals lacks the distal tube-like extension, similar to some of the specimens investigated by Clark (1963). This strongly indicates that (1) Clark confounded developmental stages of different species, his material probably including
<taxonomicName id="4C164D6E6371E840FCF59A434299376C" box="[834,1009,512,536]" class="Pycnogonida" family="Callipallenidae" genus="Pseudopallene" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="brevicephala">
<emphasis id="B962EAFF6371E840FCF59A434299376C" box="[834,1009,512,536]" italics="true" pageId="18" pageNumber="419">P. brevicephala</emphasis>
</taxonomicName>
besides other congeners and that (2) those post-embryonic stages having been cautiously assigned to
<taxonomicName id="4C164D6E6371E840FCCB9A6745433748" box="[892,1067,548,572]" class="Pycnogonida" family="Callipallenidae" genus="Pseudopallene" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="brevicephala">
<emphasis id="B962EAFF6371E840FCCB9A6745433748" box="[892,1067,548,572]" italics="true" pageId="18" pageNumber="419">P. brevicephala</emphasis>
</taxonomicName>
by Staples (2008) might in fact belong to another species, since they exhibit the distinctive tube-like elongation of the movable chela finger. This is the first record of
<taxonomicName id="4C164D6E6371E840FED69A2F436537F0" box="[353,525,620,644]" class="Pycnogonida" family="Callipallenidae" genus="Pseudopallene" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="brevicephala">
<emphasis id="B962EAFF6371E840FED69A2F436537F0" box="[353,525,620,644]" italics="true" pageId="18" pageNumber="419">P. brevicephala</emphasis>
</taxonomicName>
from
<taxonomicName id="4C164D6E6371E840FDE59A2E43A937F0" box="[594,705,620,644]" class="Pycnogonida" family="Callipallenidae" genus="Pseudopallene" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="18" pageNumber="432" phylum="Arthropoda" rank="species" species="tasmania">Tasmania</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>