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<document id="A1C2AF38BAA6A7B4680718C8EE4C329E" ID-DOI="10.5281/zenodo.171314" ID-GBIF-Dataset="9fcc3e54-c294-42b8-8f12-75919ae4f461" ID-ISSN="1175­5326" ID-Zenodo-Dep="171314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459720424464" checkinUser="plazi" docAuthor="Saether, Ole A." docDate="2005" docId="03BF87F4FF96257DFE9EFCB1017AFC0E" docLanguage="en" docName="zt00974.pdf" docOrigin="Zootaxa 974" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Orthocladius (Mesorthocladius) nimidens Saether, 2005, sp. n." docType="treatment" docVersion="8" lastPageNumber="42" masterDocId="FF86FF8CFFB52554FF96FF950554FF80" masterDocTitle="A new subgenus and new species of Orthocladius van der Wulp, with a phylogenetic evaluation of the validity of the subgenera of the genus (Diptera: Chironomidae)" masterLastPageNumber="56" masterPageNumber="1" pageNumber="36" updateTime="1698209064823" updateUser="plazi">
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<mods:title id="94566328F56BD778AF3752DE6FE9955D">A new subgenus and new species of Orthocladius van der Wulp, with a phylogenetic evaluation of the validity of the subgenera of the genus (Diptera: Chironomidae)</mods:title>
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<mods:namePart id="1F8DB3E3672613F0A99F7D20F7C8C070">Saether, Ole A.</mods:namePart>
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<mods:date id="7389941DF46D08CB1C4A1B3C448F5669">2005</mods:date>
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<treatment id="03BF87F4FF96257DFE9EFCB1017AFC0E" ID-DOI="http://doi.org/10.5281/zenodo.6266204" ID-GBIF-Taxon="119335508" ID-Zenodo-Dep="6266204" LSID="urn:lsid:plazi:treatment:03BF87F4FF96257DFE9EFCB1017AFC0E" httpUri="http://treatment.plazi.org/id/03BF87F4FF96257DFE9EFCB1017AFC0E" lastPageId="41" lastPageNumber="42" pageId="35" pageNumber="36">
<subSubSection id="C30C6569FF962577FE9EFCB1065CFCE1" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="8BA936E2FF962577FE9EFCB1061EFCBE" blockId="35.[264,842,804,905]" box="[264,842,804,830]" pageId="35" pageNumber="36">
<heading id="D0E1818EFF962577FE9EFCB1061EFCBE" bold="true" box="[264,842,804,830]" fontSize="11" level="1" pageId="35" pageNumber="36" reason="1">
<emphasis id="B962EAF0FF962577FE9EFCB1061EFCBE" bold="true" box="[264,842,804,830]" pageId="35" pageNumber="36">
<taxonomicName id="4C164D61FF962577FE9EFCB10655FCBE" ID-CoL="8D49P" box="[264,769,804,830]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="nimidens" status="sp. nov." subGenus="Mesorthocladius">
<emphasis id="B962EAF0FF962577FE9EFCB104F2FCBE" bold="true" box="[264,422,804,830]" italics="true" pageId="35" pageNumber="36">Orthocladius</emphasis>
(
<emphasis id="B962EAF0FF962577FE21FCB107D5FCBE" bold="true" box="[439,641,804,830]" italics="true" pageId="35" pageNumber="36">Mesorthocladius</emphasis>
)
<emphasis id="B962EAF0FF962577FD04FCB10655FCBE" bold="true" box="[658,769,804,830]" italics="true" pageId="35" pageNumber="36">nimidens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A251578BFF962577FC9EFCB1061EFCBE" box="[776,842,804,830]" pageId="35" pageNumber="36" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="8BA936E2FF962577FE9EFCD2065CFCE1" blockId="35.[264,842,804,905]" box="[264,776,839,865]" pageId="35" pageNumber="36">
<emphasis id="B962EAF0FF962577FE9EFCD20419FCE0" box="[264,333,839,864]" italics="true" pageId="35" pageNumber="36">O. sp.</emphasis>
“Jacobsen” (
<bibRefCitation id="EF874B13FF962577FE71FCD20654FCE1" author="Epler" box="[487,768,839,865]" pageId="35" pageNumber="55" refString="Epler, J. H. (2001) Identification manual for the larval Chironomidae (Diptera) of North and South Carolina. A guide to the taxonomy of the midges of the southeastern United States, including Florida. Special Publication SJ 2001 - SP 13. North Carolina Department of Environment and Natural Resources, Raleigh, NC, and St. Johns River Water Management District, Palatka, FL. 526 pp." type="book" year="2001">Epler 2001: 7.99, 7.109</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30C6569FF962577FE9EFCFA0656FC09" box="[264,770,879,905]" pageId="35" pageNumber="36" type="description">
<paragraph id="8BA936E2FF962577FE9EFCFA0656FC09" blockId="35.[264,842,804,905]" box="[264,770,879,905]" pageId="35" pageNumber="36">
(
<figureCitation id="132D2A67FF962577FE87FCFA04CAFC09" box="[273,414,879,905]" captionStart="FIGURES 13 22" captionStartId="31.[264,383,1768,1792]" captionTargetBox="[285,1303,643,1752]" captionTargetId="figure@31.[269,1318,618,1765]" captionTargetPageId="31" captionText="FIGURES 13 22. Orthocladius (Mesorthocladius) spp., female imagines, genitalia, 13 17: O. (M.) lamellatus sp. n., 13: ventral view, 14: ventrolateral lobe, 15: dorsomesal lobe, 16: apodeme lobe, 17: dorsal view, 18 22: O. (M.) nimidens sp. n., 18: ventral view, 19: ventrolateral lobe, 20: dorsomesal lobe, 21: apodeme lobe, 22 dorsal view." httpUri="https://zenodo.org/record/171318/files/figure.png" pageId="35" pageNumber="36">Figs. 1822</figureCitation>
,
<figureCitation id="132D2A67FF962577FE3CFCFA04A0FC09" box="[426,500,879,905]" captionStart="FIGURES 23 27" captionStartId="32.[264,382,1814,1838]" captionTargetBox="[362,1271,556,1802]" captionTargetId="figure@32.[312,1277,554,1804]" captionTargetPageId="32" captionText="FIGURES 23 27. Orthocladius (Mesorthocladius) spp., pupae, 23: O. (M.) lamellatus sp. n., tergites, 24: thoracic horn, 25 27: O. (M.) nimidens sp. n., 25: tergites, 26: thoracic horn, 27: frontal apotome." httpUri="https://zenodo.org/record/171319/files/figure.png" pageId="35" pageNumber="36">2527</figureCitation>
,
<figureCitation id="132D2A67FF962577FD96FCFA0718FC09" box="[512,588,879,905]" captionStart="FIGURES 28 34" captionStartId="34.[264,383,1688,1712]" captionTargetBox="[308,1302,296,1678]" captionTargetId="figure@34.[269,1318,280,1682]" captionTargetPageId="34" captionText="FIGURES 28 34. Orthocladius spp., larvae, 28: Orthocladius (Mesorthocladius) frigidus (Zetterstedt) labrum, epipharyngeal area and premandible, 29, 30: Orthocladius (Orthocladius) nitidoscutellatus Lundström, 29: antenna, 30: labrum, epipharyngeal area and premandible, 31, 32: Orthocladius (Mesorthocladius) lamellatus sp. n., 31: antenna, 32: labrum, epipharyngeal area and premandible, 33 34: Orthocladius (Mesorthocladius) nimidens sp. n., 31: antenna, 32: labrum, epipharyngeal area and premandible." httpUri="https://zenodo.org/record/171320/files/figure.png" pageId="35" pageNumber="36">33, 34</figureCitation>
,
<figureCitation id="132D2A67FF962577FDCEFCFA07F0FC09" box="[600,676,879,905]" captionStart="FIGURES 35 40" captionStartId="36.[264,383,1691,1715]" captionTargetBox="[281,1294,319,1672]" captionTargetId="figure@36.[269,1318,299,1687]" captionTargetPageId="36" captionText="FIGURES 35 40. Orthocladius spp., larvae, 35, 36: Orthocladius (Mesorthocladius) lamellatus sp. n., 35: mandible, 36: mentum, 37 38: Orthocladius (Mesorthocladius) nimidens sp. n., 37: mandible, 38: mentum, 39 40: Orthocladius (Orthocladius) nitidoscutellatus Lundström, 39: mandible, 40: mentum" httpUri="https://zenodo.org/record/171321/files/figure.png" pageId="35" pageNumber="36">37, 38</figureCitation>
,
<figureCitation id="132D2A67FF962577FD26FCFA07AEFC09" box="[688,762,879,905]" captionStart="FIGURES 41 46" captionStartId="38.[264,383,1655,1679]" captionTargetBox="[282,1308,321,1626]" captionTargetId="figure@38.[269,1318,307,1652]" captionTargetPageId="38" captionText="FIGURES 41 46. Orthocladius (Mesorthocladius) nimidens sp. n., male imago, 41: head, 42: thorax, 43: wing, 44: hypopygium, 45: inner margin of gonocoxite, 46: gonostylus, different view." httpUri="https://zenodo.org/record/171322/files/figure.png" pageId="35" pageNumber="36">4146</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30C6569FF962577FE9EFC2A0657F951" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="8BA936E2FF962577FE9EFC2A04FDFC58" blockId="35.[264,1323,959,1145]" box="[264,425,959,984]" pageId="35" pageNumber="36">
<emphasis id="B962EAF0FF962577FE9EFC2A04FDFC58" box="[264,425,959,984]" italics="true" pageId="35" pageNumber="36">
<typeStatus id="54AD8840FF962577FE9EFC2A046BFC58" box="[264,319,959,984]" pageId="35" pageNumber="36">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="8BA936E2FF962577FEAEFC7206ACFBF9" blockId="35.[264,1323,959,1145]" pageId="35" pageNumber="36">
<typeStatus id="54AD8840FF962577FEAEFC7204F1FB81" box="[312,421,999,1025]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
ɗ reared from larva,
<collectingCountry id="F3017672FF962577FD3EFC7207B3FB81" box="[680,743,999,1025]" name="United States of America" pageId="35" pageNumber="36">USA</collectingCountry>
: Ohio, Franklin Co., Sharon Woods Park, woodland trickle,
<date id="FFA81022FF962577FE36FB9A048BFBA9" box="[416,479,1039,1065]" pageId="35" pageNumber="36" value="1928-03">28 iii</date>
emerged
<date id="FFA81022FF962577FDC0FB9A0798FBA9" box="[598,716,1039,1065]" pageId="35" pageNumber="36" value="1987-04-02">2 iv 1987</date>
, M. J. Bolton (
<collectionCode id="ED07AE27FF962577FC15FB9A068EFBA9" box="[899,986,1039,1065]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="35" pageNumber="36">ZMBN</collectionCode>
<typeStatus id="54AD8840FF962577FC75FB9A0149FBA9" box="[995,1053,1039,1065]" pageId="35" pageNumber="36">Type</typeStatus>
No.407).
<typeStatus id="54AD8840FF962577FB0FFB9A005BFBA9" box="[1177,1295,1039,1065]" pageId="35" pageNumber="36" type="paratype">Paratypes</typeStatus>
: 3 mature ɗ pupae reared from larva, 3 mature Ψ pupae reared from larva, 3 pupal exuviae,
<specimenCount id="9D10FD6BFF962577FA8AFBA20406FBF9" pageId="35" pageNumber="36" type="larva">3 larvae</specimenCount>
, as
<typeStatus id="54AD8840FF962577FEE8FBCA04B1FBF9" box="[382,485,1119,1145]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
except
<date id="FFA81022FF962577FDA9FBCA07BEFBF9" box="[575,746,1119,1145]" pageId="35" pageNumber="36" value="1987-04-22" valueMax="1987-04-26" valueMin="1987-04-22">2226 iv 1987</date>
, M. J. Bolton (
<collectionCode id="ED07AE27FF962577FC0EFBCA06A4FBF9" box="[920,1008,1119,1145]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="35" pageNumber="36">ZMBN</collectionCode>
)
</paragraph>
<paragraph id="8BA936E2FF962577FE9EFB3A04CFFA99" pageId="35" pageNumber="36">
<table id="F916C442FF96DAABFE9EFB3A007EFA99" box="[264,1322,1199,1305]" gridcols="14" gridrows="3" pageId="35" pageNumber="36">
<tr id="352634A0FF96DAABFE9EFB3A007EFB48" box="[264,1322,1199,1224]" gridrow="0" pageId="35" pageNumber="36" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76F75DDCFF96DAABFE9EFB3A0490FB48" box="[264,452,1199,1224]" gridcol="0" gridrow="0" pageId="35" pageNumber="36">
<emphasis id="B962EAF0FF962577FE9EFB3A04D1FB48" box="[264,389,1199,1224]" italics="true" pageId="35" pageNumber="36">Etymology</emphasis>
</th>
</tr>
<tr id="352634A0FF96DAABFE9EFB42007EFB71" box="[264,1322,1239,1265]" gridrow="1" pageId="35" pageNumber="36">
<th id="76F75DDCFF96DAABFE9EFB420490FB71" box="[264,452,1239,1265]" gridcol="0" gridrow="1" pageId="35" pageNumber="36">From Latin,</th>
<td id="76F75DDCFF96DAABFE5AFB420775FB71" box="[460,545,1239,1265]" gridcol="1" gridrow="1" pageId="35" pageNumber="36">
<emphasis id="B962EAF0FF962577FE5AFB42074EFB70" box="[460,538,1239,1264]" italics="true" pageId="35" pageNumber="36">nimius</emphasis>
,
</td>
<td id="76F75DDCFF96DAABFDBCFB4207F5FB71" box="[554,673,1239,1265]" gridcol="2" gridrow="1" pageId="35" pageNumber="36">excessive,</td>
<td id="76F75DDCFF96DAABFD3FFB420787FB71" box="[681,723,1239,1265]" gridcol="3" gridrow="1" pageId="35" pageNumber="36">and</td>
<td id="76F75DDCFF96DAABFD4AFB42064CFB71" box="[732,792,1239,1265]" gridcol="4" gridrow="1" pageId="35" pageNumber="36">
<emphasis id="B962EAF0FF962577FD4AFB420645FB70" box="[732,785,1239,1264]" italics="true" pageId="35" pageNumber="36">dens</emphasis>
,
</td>
<td id="76F75DDCFF96DAABFCB6FB420637FB71" box="[800,867,1239,1265]" gridcol="5" gridrow="1" pageId="35" pageNumber="36">tooth,</td>
<td id="76F75DDCFF96DAABFCFDFB420685FB71" box="[875,977,1239,1265]" gridcol="6" gridrow="1" pageId="35" pageNumber="36">referring</td>
<td id="76F75DDCFF96DAABFC4FFB4206A4FB71" box="[985,1008,1239,1265]" gridcol="7" gridrow="1" pageId="35" pageNumber="36">to</td>
<td id="76F75DDCFF96DAABFC6EFB42014FFB71" box="[1016,1051,1239,1265]" gridcol="8" gridrow="1" pageId="35" pageNumber="36">the</td>
<td id="76F75DDCFF96DAABFBB2FB420138FB71" box="[1060,1132,1239,1265]" gridcol="9" gridrow="1" pageId="35" pageNumber="36">1517</td>
<td id="76F75DDCFF96DAABFBE3FB4201FAFB71" box="[1141,1198,1239,1265]" gridcol="10" gridrow="1" pageId="35" pageNumber="36">teeth</td>
<td id="76F75DDCFF96DAABFB20FB42019AFB71" box="[1206,1230,1239,1265]" gridcol="11" gridrow="1" pageId="35" pageNumber="36">of</td>
<td id="76F75DDCFF96DAABFB40FB4201AEFB71" box="[1238,1274,1239,1265]" gridcol="12" gridrow="1" pageId="35" pageNumber="36">the</td>
<tr id="352634A0FF96DAABFE9EFB6A007EFA99" box="[264,1322,1279,1305]" gridrow="2" pageId="35" pageNumber="36" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76F75DDCFF96DAABFE9EFB6A0490FA99" box="[264,452,1279,1305]" gridcol="0" gridrow="2" pageId="35" pageNumber="36">
<td id="76F75DDCFF96DAABFA94FB42007EFB71" box="[1282,1322,1239,1265]" gridcol="13" gridrow="1" pageId="35" pageNumber="36">larval</td>
mentum.
</th>
</tr>
</tr>
</table>
</paragraph>
<paragraph id="8BA936E2FF962577FE9EFADA0759FAE8" blockId="35.[264,1323,1359,1745]" box="[264,525,1359,1384]" pageId="35" pageNumber="36">
<emphasis id="B962EAF0FF962577FE9EFADA0759FAE8" box="[264,525,1359,1384]" italics="true" pageId="35" pageNumber="36">Diagnostic characters</emphasis>
</paragraph>
<paragraph id="8BA936E2FF962577FEAEFAE20657F951" blockId="35.[264,1323,1359,1745]" pageId="35" pageNumber="36">The male imago differs from all other species of the genus by means of the triangular anal point. The apical wart of the seminal capsule, the evenly wide spermathecal ducts with several loops, and the lack of an anterior projection of the dorsomesal lobe of gonapophysis VIII all are unique within the genus. The combination of prominent frontal warts, granulose pedes spurii B on segment II, thoracic horn covered with spinules, and minute lateral threads but no apical spines on the anal lobe separates the pupa from other members of the genus. The mentum with 1517 teeth combined with a bifid premandible, very broad median tooth of the mentum, and presence of seta interna of the mandible separate the larva from other members of the genus.</paragraph>
</subSubSection>
<caption id="DF69666AFF912570FE9EF90E04DCF89B" httpUri="https://zenodo.org/record/171321/files/figure.png" pageId="36" pageNumber="37" targetBox="[281,1294,319,1672]" targetPageId="36">
<paragraph id="8BA936E2FF912570FE9EF90E04DCF89B" blockId="36.[264,1323,1691,1819]" pageId="36" pageNumber="37">
<emphasis id="B962EAF0FF912570FE9EF90E0486F933" bold="true" box="[264,466,1691,1715]" pageId="36" pageNumber="37">FIGURES 3540.</emphasis>
<taxonomicName id="4C164D61FF912570FE4AF909073CF933" box="[476,616,1692,1715]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B962EAF0FF912570FE4AF909073CF933" box="[476,616,1692,1715]" italics="true" pageId="36" pageNumber="37">Orthocladius</emphasis>
</taxonomicName>
spp., larvae, 35, 36:
<taxonomicName id="4C164D61FF912570FCC0F909007FF933" box="[854,1323,1692,1715]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lamellatus" status="sp. nov." subGenus="Mesorthocladius">
<emphasis id="B962EAF0FF912570FCC0F90906B7F933" box="[854,995,1692,1715]" italics="true" pageId="36" pageNumber="37">Orthocladius</emphasis>
(
<emphasis id="B962EAF0FF912570FC63F90901FCF933" box="[1013,1192,1692,1715]" italics="true" pageId="36" pageNumber="37">Mesorthocladius</emphasis>
)
<emphasis id="B962EAF0FF912570FB2AF909007FF933" box="[1212,1323,1692,1715]" italics="true" pageId="36" pageNumber="37">lamellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A251578BFF912570FE9EF92B0416F955" box="[264,322,1726,1749]" pageId="36" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
, 35: mandible, 36: mentum, 3738:
<taxonomicName id="4C164D61FF912570FD54F92A01D4F956" box="[706,1152,1726,1750]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="nimidens" status="sp. nov." subGenus="Mesorthocladius">
<emphasis id="B962EAF0FF912570FD54F92A061BF956" box="[706,847,1727,1750]" italics="true" pageId="36" pageNumber="37">Orthocladius</emphasis>
(
<emphasis id="B962EAF0FF912570FCC8F92A0145F956" box="[862,1041,1727,1750]" italics="true" pageId="36" pageNumber="37">Mesorthocladius</emphasis>
)
<emphasis id="B962EAF0FF912570FBB6F92A01D4F956" box="[1056,1152,1727,1750]" italics="true" pageId="36" pageNumber="37">nimidens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A251578BFF912570FB10F92B0194F955" box="[1158,1216,1726,1749]" pageId="36" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
, 37: mandible, 38: mentum, 3940:
<taxonomicName id="4C164D61FF912570FDBEF97701DAF978" authority="Lundstrom" authorityName="Lundstrom" box="[552,1166,1761,1785]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="nitidoscutellatus" subGenus="Orthocladius">
<emphasis id="B962EAF0FF912570FDBEF97707E1F979" box="[552,693,1762,1785]" italics="true" pageId="36" pageNumber="37">Orthocladius</emphasis>
(
<emphasis id="B962EAF0FF912570FD52F9770605F979" box="[708,849,1762,1785]" italics="true" pageId="36" pageNumber="37">Orthocladius</emphasis>
)
<emphasis id="B962EAF0FF912570FCF6F9770144F979" box="[864,1040,1762,1785]" italics="true" pageId="36" pageNumber="37">nitidoscutellatus</emphasis>
Lundström
</taxonomicName>
, 39: mandible, 40: mentum
</paragraph>
</caption>
<subSubSection id="C30C6569FF90257CFEAEFE8904ADF8DB" lastPageId="40" lastPageNumber="41" pageId="37" pageNumber="38" type="description">
<paragraph id="8BA936E2FF902571FEAEFE8906CCFEB6" blockId="37.[264,1324,284,1081]" box="[312,920,284,310]" pageId="37" pageNumber="38">
<emphasis id="B962EAF0FF902571FEAEFE890490FEB5" box="[312,452,284,309]" italics="true" pageId="37" pageNumber="38">Male imago</emphasis>
(
<emphasis id="B962EAF0FF902571FE43FE8904B0FEB5" box="[469,484,284,309]" italics="true" pageId="37" pageNumber="38">n</emphasis>
= 34 except when otherwise stated)
</paragraph>
<paragraph id="8BA936E2FF902571FEAEFED1074FFE2E" blockId="37.[264,1324,284,1081]" pageId="37" pageNumber="38">
Total length 3.354.30,
<quantity id="4CEE9B07FF902571FDC0FED10791FEDE" box="[598,709,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" pageId="37" pageNumber="38" unit="mm" value="3.95">3.95 mm</quantity>
. Wing length
<quantity id="4CEE9B07FF902571FCE6FED1014FFEDE" box="[880,1051,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="1.8399999999999999" metricValueMax="1.96" metricValueMin="1.72" pageId="37" pageNumber="38" unit="mm" value="1.8399999999999999" valueMax="1.96" valueMin="1.72">1.721.96 mm</quantity>
(2). Total length/wing length 1.952.07 (2). Wing length/length of profemur 2.442.49 (2). Coloration brown with dark brown vittae.
</paragraph>
<paragraph id="8BA936E2FF902571FEAEFE29011BFDF6" blockId="37.[264,1324,284,1081]" pageId="37" pageNumber="38">
Head (
<figureCitation id="132D2A67FF902571FE1FFE2904B1FE56" box="[393,485,444,470]" captionStart="FIGURES 41 46" captionStartId="38.[264,383,1655,1679]" captionTargetBox="[282,1308,321,1626]" captionTargetId="figure@38.[269,1318,307,1652]" captionTargetPageId="38" captionText="FIGURES 41 46. Orthocladius (Mesorthocladius) nimidens sp. n., male imago, 41: head, 42: thorax, 43: wing, 44: hypopygium, 45: inner margin of gonocoxite, 46: gonostylus, different view." httpUri="https://zenodo.org/record/171322/files/figure.png" pageId="37" pageNumber="38">Fig. 41</figureCitation>
). AR 1.761.92, 1.84. Ultimate flagellomere 567652, 614 µm long. Temporal
<taxonomicName id="4C164D61FF902571FEE8FE7104ECFE7E" box="[382,440,484,510]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
1114, including 45 inner verticals, 45 outer verticals, and 34 postorbitals. Clypeus with 58, 6
<taxonomicName id="4C164D61FF902571FDB6FD99070BFDA6" box="[544,607,524,550]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
. Cibarial pump, tentorium and stipes as in
<figureCitation id="132D2A67FF902571FBC6FD9901F1FDA6" box="[1104,1189,524,550]" captionStart="FIGURES 41 46" captionStartId="38.[264,383,1655,1679]" captionTargetBox="[282,1308,321,1626]" captionTargetId="figure@38.[269,1318,307,1652]" captionTargetPageId="38" captionText="FIGURES 41 46. Orthocladius (Mesorthocladius) nimidens sp. n., male imago, 41: head, 42: thorax, 43: wing, 44: hypopygium, 45: inner margin of gonocoxite, 46: gonostylus, different view." httpUri="https://zenodo.org/record/171322/files/figure.png" pageId="37" pageNumber="38">Fig. 41</figureCitation>
. Tentorium 195206 µm (2) long, 49 (2) µm wide. Stipes 146158 µm (2) long, 5660 µm (2) wide. Palpomere lengths (in µm): 3447, 5364, 75101, 6886, 113135 (2).
</paragraph>
<paragraph id="8BA936E2FF902571FEAEFD110432FD6E" blockId="37.[264,1324,284,1081]" pageId="37" pageNumber="38">
Thorax (
<figureCitation id="132D2A67FF902571FE0AFD1104A1FD1E" box="[412,501,644,670]" captionStart="FIGURES 41 46" captionStartId="38.[264,383,1655,1679]" captionTargetBox="[282,1308,321,1626]" captionTargetId="figure@38.[269,1318,307,1652]" captionTargetPageId="38" captionText="FIGURES 41 46. Orthocladius (Mesorthocladius) nimidens sp. n., male imago, 41: head, 42: thorax, 43: wing, 44: hypopygium, 45: inner margin of gonocoxite, 46: gonostylus, different view." httpUri="https://zenodo.org/record/171322/files/figure.png" pageId="37" pageNumber="38">Fig. 42</figureCitation>
). Antepronotum with 68, 7
<taxonomicName id="4C164D61FF902571FCD2FD1106D6FD1E" box="[836,898,644,670]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
. Dorsocentrals 1518, 16, uni­biserial; acrostichals 611, 8; prealars 78, 8; 1 supraalar. Scutellum with 1417, 16
<taxonomicName id="4C164D61FF902571FB27FD3901BBFD46" box="[1201,1263,684,710]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
; unibiserial.
</paragraph>
<paragraph id="8BA936E2FF902571FEAEFD6907FCFCE6" blockId="37.[264,1324,284,1081]" pageId="37" pageNumber="38">
Wing (
<figureCitation id="132D2A67FF902571FE1EFD6904B5FC96" box="[392,481,764,790]" captionStart="FIGURES 41 46" captionStartId="38.[264,383,1655,1679]" captionTargetBox="[282,1308,321,1626]" captionTargetId="figure@38.[269,1318,307,1652]" captionTargetPageId="38" captionText="FIGURES 41 46. Orthocladius (Mesorthocladius) nimidens sp. n., male imago, 41: head, 42: thorax, 43: wing, 44: hypopygium, 45: inner margin of gonocoxite, 46: gonostylus, different view." httpUri="https://zenodo.org/record/171322/files/figure.png" pageId="37" pageNumber="38">Fig. 43</figureCitation>
). Anal lobe moderately to strongly produced. VR 1.031.06 (2). Costal extension 3849 µm (2) long. Brachiolum with 1 seta, R with 79, 8
<taxonomicName id="4C164D61FF902571FBCFFCB101C3FCBE" box="[1113,1175,804,830]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
; other veins bare. Squama with 1724, 19
<taxonomicName id="4C164D61FF902571FDF3FCD907F7FCE6" box="[613,675,844,870]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
.
</paragraph>
<paragraph id="8BA936E2FF902571FEAEFCE107B2FBB9" blockId="37.[264,1324,284,1081]" pageId="37" pageNumber="38">
Legs (
<emphasis id="B962EAF0FF902571FE17FCE104C4FC0D" box="[385,400,884,909]" italics="true" pageId="37" pageNumber="38">n</emphasis>
= 2). Spur of front tibia 7194 µm long, spurs of middle tibia 3445 and 2441 µm long, of hind tibia 6883 and 2638 µm long. Width at apex of front and middle tibia each 4556 µm, of hind tibia 5668 µm. Pseudospurs present on ta1 of mid and hind leg, ta2 of mid leg and occasionally ta2 of hind leg, 2632 µm long. Sensilla chaetica absent. Lengths (in µm) and proportions of legs:
</paragraph>
<paragraph id="8BA936E2FF902571FEFBFBF90199FB6F" pageId="37" pageNumber="38">
<table id="F916C442FF90DAABFE83FBF90199FB75" box="[277,1229,1132,1269]" gridcols="5" gridrows="4" pageId="37" pageNumber="38">
<tr id="352634A0FF90DAABFE83FBF90199FB06" box="[277,1229,1132,1158]" gridrow="0" pageId="37" pageNumber="38">
<th id="76F75DDCFF90DAABFE83FBF90495FB06" box="[277,449,1132,1158]" gridcol="0" gridrow="0" pageId="37" pageNumber="38">Fe</th>
<th id="76F75DDCFF90DAABFDBAFBF907DFFB06" box="[556,651,1132,1158]" gridcol="1" gridrow="0" pageId="37" pageNumber="38">ti</th>
<th id="76F75DDCFF90DAABFD6AFBF90604FB06" box="[764,848,1132,1158]" gridcol="2" gridrow="0" pageId="37" pageNumber="38">ta1</th>
<th id="76F75DDCFF90DAABFC2DFBF9015BFB06" box="[955,1039,1132,1158]" gridcol="3" gridrow="0" pageId="37" pageNumber="38">ta2</th>
<th id="76F75DDCFF90DAABFBEFFBF90199FB06" box="[1145,1229,1132,1158]" gridcol="4" gridrow="0" pageId="37" pageNumber="38">ta3</th>
</tr>
<tr id="352634A0FF90DAABFE83FB040199FB2B" box="[277,1229,1169,1195]" gridrow="1" pageId="37" pageNumber="38">
<th id="76F75DDCFF90DAABFE83FB040495FB2B" box="[277,449,1169,1195]" gridcol="0" gridrow="1" pageId="37" pageNumber="38">p1 690803</th>
<td id="76F75DDCFF90DAABFDBAFB0407DFFB2B" box="[556,651,1169,1195]" gridcol="1" gridrow="1" pageId="37" pageNumber="38">775926</td>
<td id="76F75DDCFF90DAABFD6AFB040604FB2B" box="[764,848,1169,1195]" gridcol="2" gridrow="1" pageId="37" pageNumber="38">548624</td>
<td id="76F75DDCFF90DAABFC2DFB04015BFB2B" box="[955,1039,1169,1195]" gridcol="3" gridrow="1" pageId="37" pageNumber="38">340378</td>
<td id="76F75DDCFF90DAABFBEFFB040199FB2B" box="[1145,1229,1169,1195]" gridcol="4" gridrow="1" pageId="37" pageNumber="38">236255</td>
</tr>
<tr id="352634A0FF90DAABFE83FB230199FB50" box="[277,1229,1206,1232]" gridrow="2" pageId="37" pageNumber="38">
<th id="76F75DDCFF90DAABFE83FB230495FB50" box="[277,449,1206,1232]" gridcol="0" gridrow="2" pageId="37" pageNumber="38">p2 690813</th>
<td id="76F75DDCFF90DAABFDBAFB2307DFFB50" box="[556,651,1206,1232]" gridcol="1" gridrow="2" pageId="37" pageNumber="38">699851</td>
<td id="76F75DDCFF90DAABFD6AFB230604FB50" box="[764,848,1206,1232]" gridcol="2" gridrow="2" pageId="37" pageNumber="38">340435</td>
<td id="76F75DDCFF90DAABFC2DFB23015BFB50" box="[955,1039,1206,1232]" gridcol="3" gridrow="2" pageId="37" pageNumber="38">198208</td>
<td id="76F75DDCFF90DAABFBEFFB230199FB50" box="[1145,1229,1206,1232]" gridcol="4" gridrow="2" pageId="37" pageNumber="38">170189</td>
</tr>
<tr id="352634A0FF90DAABFE83FB4F0199FB75" box="[277,1229,1242,1269]" gridrow="3" pageId="37" pageNumber="38">
<th id="76F75DDCFF90DAABFE83FB4F0495FB75" box="[277,449,1242,1269]" gridcol="0" gridrow="3" pageId="37" pageNumber="38">p3 778888</th>
<td id="76F75DDCFF90DAABFDBAFB4F07DFFB75" box="[556,651,1242,1269]" gridcol="1" gridrow="3" pageId="37" pageNumber="38">8511021</td>
<td id="76F75DDCFF90DAABFD6AFB4F0604FB75" box="[764,848,1242,1269]" gridcol="2" gridrow="3" pageId="37" pageNumber="38">454517</td>
<td id="76F75DDCFF90DAABFC2DFB4F015BFB75" box="[955,1039,1242,1269]" gridcol="3" gridrow="3" pageId="37" pageNumber="38">265322</td>
<td id="76F75DDCFF90DAABFBEFFB4F0199FB75" box="[1145,1229,1242,1269]" gridcol="4" gridrow="3" pageId="37" pageNumber="38">198246</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA936E2FF902571FEFBFAB2019FFA2B" pageId="37" pageNumber="38">
<table id="F916C442FF90DAABFE83FAB201A1FA30" box="[277,1269,1319,1456]" gridcols="7" gridrows="4" pageId="37" pageNumber="38">
<tr id="352634A0FF90DAABFE83FAB201A1FAC1" box="[277,1269,1319,1345]" gridrow="0" pageId="37" pageNumber="38" rowspan-0="1">
<th id="76F75DDCFF90DAABFEFBFAB20495FAC1" box="[365,449,1319,1345]" gridcol="1" gridrow="0" pageId="37" pageNumber="38">ta4</th>
<th id="76F75DDCFF90DAABFDBAFAB20720FAC1" box="[556,628,1319,1345]" gridcol="2" gridrow="0" pageId="37" pageNumber="38">ta5</th>
<th id="76F75DDCFF90DAABFD5EFAB2067CFAC1" box="[712,808,1319,1345]" gridcol="3" gridrow="0" pageId="37" pageNumber="38">LR</th>
<th id="76F75DDCFF90DAABFCE3FAB20682FAC1" box="[885,982,1319,1345]" gridcol="4" gridrow="0" pageId="37" pageNumber="38">BV</th>
<th id="76F75DDCFF90DAABFB87FAB2012CFAC1" box="[1041,1144,1319,1345]" gridcol="5" gridrow="0" pageId="37" pageNumber="38">SV</th>
<th id="76F75DDCFF90DAABFB3BFAB201A1FAC1" box="[1197,1269,1319,1345]" gridcol="6" gridrow="0" pageId="37" pageNumber="38">BR</th>
</tr>
<tr id="352634A0FF90DAABFE83FAD901A1FAE6" box="[277,1269,1356,1382]" gridrow="1" pageId="37" pageNumber="38">
<th id="76F75DDCFF90DAABFE83FAD90478FAE6" box="[277,300,1356,1382]" gridcol="0" gridrow="1" pageId="37" pageNumber="38">p1</th>
<td id="76F75DDCFF90DAABFEFBFAD90495FAE6" box="[365,449,1356,1382]" gridcol="1" gridrow="1" pageId="37" pageNumber="38">151170</td>
<td id="76F75DDCFF90DAABFDBAFAD90720FAE6" box="[556,628,1356,1382]" gridcol="2" gridrow="1" pageId="37" pageNumber="38">99118</td>
<td id="76F75DDCFF90DAABFD5EFAD9067CFAE6" box="[712,808,1356,1382]" gridcol="3" gridrow="1" pageId="37" pageNumber="38">0.670.71</td>
<td id="76F75DDCFF90DAABFCE3FAD90682FAE6" box="[885,982,1356,1382]" gridcol="4" gridrow="1" pageId="37" pageNumber="38">2.452.55</td>
<td id="76F75DDCFF90DAABFB87FAD9012CFAE6" box="[1041,1144,1356,1382]" gridcol="5" gridrow="1" pageId="37" pageNumber="38">2.672. 77</td>
<td id="76F75DDCFF90DAABFB3BFAD901A1FAE6" box="[1197,1269,1356,1382]" gridcol="6" gridrow="1" pageId="37" pageNumber="38">2.42.5</td>
</tr>
<tr id="352634A0FF90DAABFE83FAE401A1FA0B" box="[277,1269,1393,1419]" gridrow="2" pageId="37" pageNumber="38">
<th id="76F75DDCFF90DAABFE83FAE40478FA0B" box="[277,300,1393,1419]" gridcol="0" gridrow="2" pageId="37" pageNumber="38">p2</th>
<td id="76F75DDCFF90DAABFEFBFAE40495FA0B" box="[365,449,1393,1419]" gridcol="1" gridrow="2" pageId="37" pageNumber="38">113132</td>
<td id="76F75DDCFF90DAABFDBAFAE40720FA0B" box="[556,628,1393,1419]" gridcol="2" gridrow="2" pageId="37" pageNumber="38">95109</td>
<td id="76F75DDCFF90DAABFD5EFAE4067CFA0B" box="[712,808,1393,1419]" gridcol="3" gridrow="2" pageId="37" pageNumber="38">0.490.51</td>
<td id="76F75DDCFF90DAABFCE3FAE40682FA0B" box="[885,982,1393,1419]" gridcol="4" gridrow="2" pageId="37" pageNumber="38">2.953.34</td>
<td id="76F75DDCFF90DAABFB87FAE4012CFA0B" box="[1041,1144,1393,1419]" gridcol="5" gridrow="2" pageId="37" pageNumber="38">3.834.08</td>
<td id="76F75DDCFF90DAABFB3BFAE401A1FA0B" box="[1197,1269,1393,1419]" gridcol="6" gridrow="2" pageId="37" pageNumber="38">2.9</td>
</tr>
<tr id="352634A0FF90DAABFE83FA0301A1FA30" box="[277,1269,1430,1456]" gridrow="3" pageId="37" pageNumber="38">
<th id="76F75DDCFF90DAABFE83FA030478FA30" box="[277,300,1430,1456]" gridcol="0" gridrow="3" pageId="37" pageNumber="38">p3</th>
<td id="76F75DDCFF90DAABFEFBFA030495FA30" box="[365,449,1430,1456]" gridcol="1" gridrow="3" pageId="37" pageNumber="38">123161</td>
<td id="76F75DDCFF90DAABFDBAFA030720FA30" box="[556,628,1430,1456]" gridcol="2" gridrow="3" pageId="37" pageNumber="38">99118</td>
<td id="76F75DDCFF90DAABFD5EFA03067CFA30" box="[712,808,1430,1456]" gridcol="3" gridrow="3" pageId="37" pageNumber="38">0.530.56</td>
<td id="76F75DDCFF90DAABFCE3FA030682FA30" box="[885,982,1430,1456]" gridcol="4" gridrow="3" pageId="37" pageNumber="38">2.952.96</td>
<td id="76F75DDCFF90DAABFB87FA03012CFA30" box="[1041,1144,1430,1456]" gridcol="5" gridrow="3" pageId="37" pageNumber="38">3.373.45</td>
<td id="76F75DDCFF90DAABFB3BFA0301A1FA30" box="[1197,1269,1430,1456]" gridcol="6" gridrow="3" pageId="37" pageNumber="38">3.1</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA936E2FF902571FEAEF99807E9F897" blockId="37.[264,1323,1549,1815]" pageId="37" pageNumber="38">
Hypopygium (
<figureCitation id="132D2A67FF902571FE75F9980727F9A7" box="[483,627,1549,1575]" captionStart="FIGURES 41 46" captionStartId="38.[264,383,1655,1679]" captionTargetBox="[282,1308,321,1626]" captionTargetId="figure@38.[269,1318,307,1652]" captionTargetPageId="38" captionText="FIGURES 41 46. Orthocladius (Mesorthocladius) nimidens sp. n., male imago, 41: head, 42: thorax, 43: wing, 44: hypopygium, 45: inner margin of gonocoxite, 46: gonostylus, different view." httpUri="https://zenodo.org/record/171322/files/figure.png" pageId="37" pageNumber="38">Figs. 4446</figureCitation>
). Anal point 86120, 109 µm long, triangular with pointed apex. Tergite IX including anal point with 1620, 19
<taxonomicName id="4C164D61FF902571FC14F9A00694F9CF" box="[898,960,1589,1615]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
; laterosternite IX with 710, 9
<taxonomicName id="4C164D61FF902571FE9EF9C80412F9F7" box="[264,326,1629,1655]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
. Phallapodeme 124135, 130 µm long; transverse sternapodeme 94131 µm long, oral projections moderately developed. Gonocoxite 266311, 288 µm long; inferior volsella with ventral part reduced, not extended beyond dorsal part. Gonostylus 113131, 122 µm long; crista dorsalis low or absent; megaseta 1115 µm long. Virga absent. HR 2.342.37, 2.36; HV 2.973.50, 3.24.
</paragraph>
<caption id="DF69666AFF932572FE9EF9E201A9F931" httpUri="https://zenodo.org/record/171322/files/figure.png" pageId="38" pageNumber="39" targetBox="[282,1308,321,1626]" targetPageId="38">
<paragraph id="8BA936E2FF932572FE9EF9E201A9F931" blockId="38.[264,1323,1655,1713]" pageId="38" pageNumber="39">
<emphasis id="B962EAF0FF932572FE9EF9E2049BF90F" bold="true" box="[264,463,1655,1679]" pageId="38" pageNumber="39">FIGURES 4146.</emphasis>
<taxonomicName id="4C164D61FF932572FE40F9E206C1F90E" box="[470,917,1655,1678]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="nimidens" status="sp. nov." subGenus="Mesorthocladius">
<emphasis id="B962EAF0FF932572FE40F9E20737F90E" box="[470,611,1655,1678]" italics="true" pageId="38" pageNumber="39">Orthocladius</emphasis>
(
<emphasis id="B962EAF0FF932572FDE4F9E20671F90E" box="[626,805,1655,1678]" italics="true" pageId="38" pageNumber="39">Mesorthocladius</emphasis>
)
<emphasis id="B962EAF0FF932572FCA3F9E206C1F90E" box="[821,917,1655,1678]" italics="true" pageId="38" pageNumber="39">nimidens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A251578BFF932572FC0AF9E20683F90E" box="[924,983,1655,1678]" pageId="38" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
, male imago, 41: head, 42: thorax, 43: wing, 44: hypopygium, 45: inner margin of gonocoxite, 46: gonostylus, different view.
</paragraph>
</caption>
<paragraph id="8BA936E2FF932572FEAEF975076CF979" blockId="38.[264,1323,1759,1905]" box="[312,568,1759,1785]" pageId="38" pageNumber="39">
<emphasis id="B962EAF0FF932572FEAEF97504B4F979" box="[312,480,1760,1785]" italics="true" pageId="38" pageNumber="39">Female imago</emphasis>
(
<emphasis id="B962EAF0FF932572FE67F9750754F979" box="[497,512,1760,1785]" italics="true" pageId="38" pageNumber="39">n</emphasis>
= 3)
</paragraph>
<paragraph id="8BA936E2FF932572FEAEF892070DF8F1" blockId="38.[264,1323,1759,1905]" pageId="38" pageNumber="39">
Total length
<quantity id="4CEE9B07FF932572FE41F89207DDF8A1" box="[471,649,1799,1825]" metricMagnitude="-3" metricUnit="m" metricValue="4.01" metricValueMax="4.19" metricValueMin="3.83" pageId="38" pageNumber="39" unit="mm" value="4.01" valueMax="4.19" valueMin="3.83">3.834.19 mm</quantity>
. Wing length
<quantity id="4CEE9B07FF932572FCABF89206BBF8A1" box="[829,1007,1799,1825]" metricMagnitude="-3" metricUnit="m" metricValue="2.1149999999999998" metricValueMax="2.28" metricValueMin="1.95" pageId="38" pageNumber="39" unit="mm" value="2.1149999999999998" valueMax="2.28" valueMin="1.95">1.952.28 mm</quantity>
. Total length/wing length 1.802.05. Wing length/length of profemur 2.612.84. Coloration not as dark as in male with more distinct markings.
</paragraph>
<paragraph id="8BA936E2FF922573FEAEFE890421FE56" blockId="39.[264,1323,284,886]" pageId="39" pageNumber="40">
Head. AR 0.460.49. Flagellomere lengths (in µm): 90101, 56, 4956, 6068, 124 135. Coronal suture
<quantity id="4CEE9B07FF922573FE68FED107C0FEDE" box="[510,660,324,350]" metricMagnitude="2" metricUnit="m" metricValue="1.205" metricValueMax="1.28" metricValueMin="1.13" pageId="39" pageNumber="40" unit="m" value="120.5" valueMax="128.0" valueMin="113.0">113128 m</quantity>
long. Temporal
<taxonomicName id="4C164D61FF922573FCCAFED106C3FEDE" box="[860,919,324,350]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
1416, including 46 inner verticals, 67 outer verticals, and 45 postorbitals. Clypeus with 812
<taxonomicName id="4C164D61FF922573FBB4FEF90134FE06" box="[1058,1120,364,390]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
. Tentorium 154 191 µm long, 3849 µm wide. Palpomere lengths (in µ m): 4145, 5364, 90124, 7994, 129143.
</paragraph>
<paragraph id="8BA936E2FF922573FEAEFE7104A7FDA6" blockId="39.[264,1323,284,886]" pageId="39" pageNumber="40">
Thorax. Dorsocentrals 1419, acrostichals 617, prealars 710. Scutellum with 1821 partly biserial
<taxonomicName id="4C164D61FF922573FE27FD9904BBFDA6" box="[433,495,524,550]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
.
</paragraph>
<paragraph id="8BA936E2FF922573FEAEFDA10733FD23" blockId="39.[264,1323,284,886]" pageId="39" pageNumber="40">
Wing. Anal lobe very slightly produced. VR 1.061.11. Costal extension 5660 µm long. Brachiolum with 13
<taxonomicName id="4C164D61FF922573FDCCFDC907C3FDF6" box="[602,663,604,630]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
, R with 7918, R1 with 611. R4+5 with 1118
<taxonomicName id="4C164D61FF922573FB7FFDC90073FDF6" box="[1257,1319,604,630]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
. Squama with 1724, 19
<taxonomicName id="4C164D61FF922573FDB3FD1C0737FD23" box="[549,611,649,675]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
.
</paragraph>
<paragraph id="8BA936E2FF922573FEAEFD24066AFCF6" blockId="39.[264,1323,284,886]" pageId="39" pageNumber="40">Legs. Spur of front tibia 4145 µm long, spurs of middle tibia 3038 and 2834 µm long, of hind tibia 6885 and 2638 µm long. Width at apex of front tibia 4556 µm, of middle tibia 5362 µm, of hind tibia 6071 µm. Pseudospurs present on ta1 of mid and hind leg, ta2 of mid leg and occasionally ta2 of hind leg, 3034 µm long. Sensilla chaetica absent. Lengths (in µm) and proportions of legs:</paragraph>
<paragraph id="8BA936E2FF922573FEFBFC3C0199FBAD" pageId="39" pageNumber="40">
<table id="F916C442FF92DAABFE83FC3C0199FBB2" box="[277,1229,937,1074]" gridcols="5" gridrows="4" pageId="39" pageNumber="40">
<tr id="352634A0FF92DAABFE83FC3C0199FC43" box="[277,1229,937,963]" gridrow="0" pageId="39" pageNumber="40">
<th id="76F75DDCFF92DAABFE83FC3C0495FC43" box="[277,449,937,963]" gridcol="0" gridrow="0" pageId="39" pageNumber="40">Fe</th>
<th id="76F75DDCFF92DAABFDBAFC3C07DFFC43" box="[556,651,937,963]" gridcol="1" gridrow="0" pageId="39" pageNumber="40">ti</th>
<th id="76F75DDCFF92DAABFD6AFC3C0604FC43" box="[764,848,937,963]" gridcol="2" gridrow="0" pageId="39" pageNumber="40">ta1</th>
<th id="76F75DDCFF92DAABFC2DFC3C015BFC43" box="[955,1039,937,963]" gridcol="3" gridrow="0" pageId="39" pageNumber="40">ta2</th>
<th id="76F75DDCFF92DAABFBEFFC3C0199FC43" box="[1145,1229,937,963]" gridcol="4" gridrow="0" pageId="39" pageNumber="40">ta3</th>
</tr>
<tr id="352634A0FF92DAABFE83FC5B0199FC68" box="[277,1229,974,1000]" gridrow="1" pageId="39" pageNumber="40">
<th id="76F75DDCFF92DAABFE83FC5B0495FC68" box="[277,449,974,1000]" gridcol="0" gridrow="1" pageId="39" pageNumber="40">p1 747803</th>
<td id="76F75DDCFF92DAABFDBAFC5B07DFFC68" box="[556,651,974,1000]" gridcol="1" gridrow="1" pageId="39" pageNumber="40">841983</td>
<td id="76F75DDCFF92DAABFD6AFC5B0604FC68" box="[764,848,974,1000]" gridcol="2" gridrow="1" pageId="39" pageNumber="40">510614</td>
<td id="76F75DDCFF92DAABFC2DFC5B015BFC68" box="[955,1039,974,1000]" gridcol="3" gridrow="1" pageId="39" pageNumber="40">331350</td>
<td id="76F75DDCFF92DAABFBEFFC5B0199FC68" box="[1145,1229,974,1000]" gridcol="4" gridrow="1" pageId="39" pageNumber="40">227246</td>
</tr>
<tr id="352634A0FF92DAABFE83FC660199FB8D" box="[277,1229,1011,1037]" gridrow="2" pageId="39" pageNumber="40">
<th id="76F75DDCFF92DAABFE83FC660495FB8D" box="[277,449,1011,1037]" gridcol="0" gridrow="2" pageId="39" pageNumber="40">p2 756851</th>
<td id="76F75DDCFF92DAABFDBAFC6607DFFB8D" box="[556,651,1011,1037]" gridcol="1" gridrow="2" pageId="39" pageNumber="40">803973</td>
<td id="76F75DDCFF92DAABFD6AFC660604FB8D" box="[764,848,1011,1037]" gridcol="2" gridrow="2" pageId="39" pageNumber="40">378435</td>
<td id="76F75DDCFF92DAABFC2DFC66015BFB8D" box="[955,1039,1011,1037]" gridcol="3" gridrow="2" pageId="39" pageNumber="40">217236</td>
<td id="76F75DDCFF92DAABFBEFFC660199FB8D" box="[1145,1229,1011,1037]" gridcol="4" gridrow="2" pageId="39" pageNumber="40">165180</td>
</tr>
<tr id="352634A0FF92DAABFE83FB8D0199FBB2" box="[277,1229,1048,1074]" gridrow="3" pageId="39" pageNumber="40">
<th id="76F75DDCFF92DAABFE83FB8D0495FBB2" box="[277,449,1048,1074]" gridcol="0" gridrow="3" pageId="39" pageNumber="40">p3 813936</th>
<td id="76F75DDCFF92DAABFDBAFB8D07DFFBB2" box="[556,651,1048,1074]" gridcol="1" gridrow="3" pageId="39" pageNumber="40">9541153</td>
<td id="76F75DDCFF92DAABFD6AFB8D0604FBB2" box="[764,848,1048,1074]" gridcol="2" gridrow="3" pageId="39" pageNumber="40">529643</td>
<td id="76F75DDCFF92DAABFC2DFB8D015BFBB2" box="[955,1039,1048,1074]" gridcol="3" gridrow="3" pageId="39" pageNumber="40">312369</td>
<td id="76F75DDCFF92DAABFBEFFB8D0199FBB2" box="[1145,1229,1048,1074]" gridcol="4" gridrow="3" pageId="39" pageNumber="40">227265</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA936E2FF922573FEFBFBF001B7FB68" pageId="39" pageNumber="40">
<table id="F916C442FF92DAABFE83FBF001B7FB6D" box="[277,1251,1125,1261]" gridcols="7" gridrows="4" pageId="39" pageNumber="40">
<tr id="352634A0FF92DAABFE83FBF001B7FBFF" box="[277,1251,1125,1151]" gridrow="0" pageId="39" pageNumber="40" rowspan-0="1">
<th id="76F75DDCFF92DAABFEFBFBF00495FBFF" box="[365,449,1125,1151]" gridcol="1" gridrow="0" pageId="39" pageNumber="40">ta4</th>
<th id="76F75DDCFF92DAABFE6EFBF0071EFBFF" box="[504,586,1125,1151]" gridcol="2" gridrow="0" pageId="39" pageNumber="40">ta5</th>
<th id="76F75DDCFF92DAABFD02FBF007A0FBFF" box="[660,756,1125,1151]" gridcol="3" gridrow="0" pageId="39" pageNumber="40">LR</th>
<th id="76F75DDCFF92DAABFCD7FBF006F5FBFF" box="[833,929,1125,1151]" gridcol="4" gridrow="0" pageId="39" pageNumber="40">BV</th>
<th id="76F75DDCFF92DAABFB96FBF00134FBFF" box="[1024,1120,1125,1151]" gridcol="5" gridrow="0" pageId="39" pageNumber="40">SV</th>
<th id="76F75DDCFF92DAABFB0AFBF001B7FBFF" box="[1180,1251,1125,1151]" gridcol="6" gridrow="0" pageId="39" pageNumber="40">BR</th>
</tr>
<tr id="352634A0FF92DAABFE83FB1C01B7FB24" box="[277,1251,1161,1188]" gridrow="1" pageId="39" pageNumber="40">
<th id="76F75DDCFF92DAABFE83FB1C0478FB24" box="[277,300,1161,1188]" gridcol="0" gridrow="1" pageId="39" pageNumber="40">p1</th>
<td id="76F75DDCFF92DAABFEFBFB1C0495FB24" box="[365,449,1161,1188]" gridcol="1" gridrow="1" pageId="39" pageNumber="40">142161</td>
<td id="76F75DDCFF92DAABFE6EFB1C071EFB24" box="[504,586,1161,1188]" gridcol="2" gridrow="1" pageId="39" pageNumber="40">104113</td>
<td id="76F75DDCFF92DAABFD02FB1C07A0FB24" box="[660,756,1161,1188]" gridcol="3" gridrow="1" pageId="39" pageNumber="40">0.610.63</td>
<td id="76F75DDCFF92DAABFCD7FB1C06F5FB24" box="[833,929,1161,1188]" gridcol="4" gridrow="1" pageId="39" pageNumber="40">2.582.76</td>
<td id="76F75DDCFF92DAABFB96FB1C0134FB24" box="[1024,1120,1161,1188]" gridcol="5" gridrow="1" pageId="39" pageNumber="40">2.913.13</td>
<td id="76F75DDCFF92DAABFB0AFB1C01B7FB24" box="[1180,1251,1161,1188]" gridcol="6" gridrow="1" pageId="39" pageNumber="40">1.61.9</td>
</tr>
<tr id="352634A0FF92DAABFE83FB3B01B7FB48" box="[277,1251,1198,1224]" gridrow="2" pageId="39" pageNumber="40">
<th id="76F75DDCFF92DAABFE83FB3B0478FB48" box="[277,300,1198,1224]" gridcol="0" gridrow="2" pageId="39" pageNumber="40">p2</th>
<td id="76F75DDCFF92DAABFEFBFB3B0495FB48" box="[365,449,1198,1224]" gridcol="1" gridrow="2" pageId="39" pageNumber="40">104123</td>
<td id="76F75DDCFF92DAABFE6EFB3B071EFB48" box="[504,586,1198,1224]" gridcol="2" gridrow="2" pageId="39" pageNumber="40">95109</td>
<td id="76F75DDCFF92DAABFD02FB3B07A0FB48" box="[660,756,1198,1224]" gridcol="3" gridrow="2" pageId="39" pageNumber="40">0.450.48</td>
<td id="76F75DDCFF92DAABFCD7FB3B06F5FB48" box="[833,929,1198,1224]" gridcol="4" gridrow="2" pageId="39" pageNumber="40">3.123.49</td>
<td id="76F75DDCFF92DAABFB96FB3B0134FB48" box="[1024,1120,1198,1224]" gridcol="5" gridrow="2" pageId="39" pageNumber="40">4.024.20</td>
<td id="76F75DDCFF92DAABFB0AFB3B01B7FB48" box="[1180,1251,1198,1224]" gridcol="6" gridrow="2" pageId="39" pageNumber="40">1.41.8</td>
</tr>
<tr id="352634A0FF92DAABFE83FB4601B7FB6D" box="[277,1251,1235,1261]" gridrow="3" pageId="39" pageNumber="40">
<th id="76F75DDCFF92DAABFE83FB460478FB6D" box="[277,300,1235,1261]" gridcol="0" gridrow="3" pageId="39" pageNumber="40">p3</th>
<td id="76F75DDCFF92DAABFEFBFB460495FB6D" box="[365,449,1235,1261]" gridcol="1" gridrow="3" pageId="39" pageNumber="40">132151</td>
<td id="76F75DDCFF92DAABFE6EFB46071EFB6D" box="[504,586,1235,1261]" gridcol="2" gridrow="3" pageId="39" pageNumber="40">104113</td>
<td id="76F75DDCFF92DAABFD02FB4607A0FB6D" box="[660,756,1235,1261]" gridcol="3" gridrow="3" pageId="39" pageNumber="40">0.540.56</td>
<td id="76F75DDCFF92DAABFCD7FB4606F5FB6D" box="[833,929,1235,1261]" gridcol="4" gridrow="3" pageId="39" pageNumber="40">2.893.04</td>
<td id="76F75DDCFF92DAABFB96FB460134FB6D" box="[1024,1120,1235,1261]" gridcol="5" gridrow="3" pageId="39" pageNumber="40">3.103.39</td>
<td id="76F75DDCFF92DAABFB0AFB4601B7FB6D" box="[1180,1251,1235,1261]" gridcol="6" gridrow="3" pageId="39" pageNumber="40">1.42.6</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA936E2FF922573FEAEFAB6079AF9D5" blockId="39.[264,1323,1315,1861]" pageId="39" pageNumber="40">
Abdomen. Tergite VIII with 3050
<taxonomicName id="4C164D61FF922573FD63FAB60666FABD" box="[757,818,1315,1341]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
, sternite VIII with 3539
<taxonomicName id="4C164D61FF922573FBEBFAB601ECFABD" box="[1149,1208,1315,1341]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
.Genitalia (
<figureCitation id="132D2A67FF922573FE87FADE04F5FAE5" box="[273,417,1355,1381]" captionStart="FIGURES 13 22" captionStartId="31.[264,383,1768,1792]" captionTargetBox="[285,1303,643,1752]" captionTargetId="figure@31.[269,1318,618,1765]" captionTargetPageId="31" captionText="FIGURES 13 22. Orthocladius (Mesorthocladius) spp., female imagines, genitalia, 13 17: O. (M.) lamellatus sp. n., 13: ventral view, 14: ventrolateral lobe, 15: dorsomesal lobe, 16: apodeme lobe, 17: dorsal view, 18 22: O. (M.) nimidens sp. n., 18: ventral view, 19: ventrolateral lobe, 20: dorsomesal lobe, 21: apodeme lobe, 22 dorsal view." httpUri="https://zenodo.org/record/171318/files/figure.png" pageId="39" pageNumber="40">Figs. 1822</figureCitation>
). Tergite IX strongly divided with margins well delimited, with 3340
<taxonomicName id="4C164D61FF922573FB7FFADE0073FAE5" box="[1257,1319,1355,1381]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
. Gonocoxite with 1419
<taxonomicName id="4C164D61FF922573FDB7FAE6070BFA0D" box="[545,607,1395,1421]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
. Cercus 169188 µm long. Seminal capsule pear­shaped with more or less well developed apical wart, 105130 µm long including about 45 µm long tapering neck, 6468 µm wide, sclerotised in anterior 7594 µm. Spermathecal ducts with several loops and apparently separate openings. Notum 98 µm long. Dorsomesal lobe of gonapophysis VIII without anterior projection. Labia with very fine microtrichiae medially, apodeme of apodeme lobe strong.
</paragraph>
<paragraph id="8BA936E2FF922573FEAEF9F6069EF925" blockId="39.[264,1323,1315,1861]" pageId="39" pageNumber="40">
<emphasis id="B962EAF0FF922573FEAEF9F60422F9FC" box="[312,374,1635,1660]" italics="true" pageId="39" pageNumber="40">Pupa</emphasis>
(
<emphasis id="B962EAF0FF922573FE1CF9F604CDF9FC" box="[394,409,1635,1660]" italics="true" pageId="39" pageNumber="40">n</emphasis>
= 910) Total length 4.295.78,
<quantity id="4CEE9B07FF922573FCBCF9F606CDF9FD" box="[810,921,1635,1661]" metricMagnitude="-3" metricUnit="m" metricValue="4.89" pageId="39" pageNumber="40" unit="mm" value="4.89">4.89 mm</quantity>
. Exuviae pale brown with darker thorax and apex of anal lobe and blackish brown apophyses.
</paragraph>
<paragraph id="8BA936E2FF92257CFEAEF92604D7FE11" blockId="39.[264,1323,1315,1861]" lastBlockId="40.[264,1323,284,1883]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
Cephalothorax. Frontal warts (
<figureCitation id="132D2A67FF922573FD31F9260654F94D" box="[679,768,1715,1741]" captionStart="FIGURES 23 27" captionStartId="32.[264,382,1814,1838]" captionTargetBox="[362,1271,556,1802]" captionTargetId="figure@32.[312,1277,554,1804]" captionTargetPageId="32" captionText="FIGURES 23 27. Orthocladius (Mesorthocladius) spp., pupae, 23: O. (M.) lamellatus sp. n., tergites, 24: thoracic horn, 25 27: O. (M.) nimidens sp. n., 25: tergites, 26: thoracic horn, 27: frontal apotome." httpUri="https://zenodo.org/record/171319/files/figure.png" pageId="39" pageNumber="40">Fig. 27</figureCitation>
) relatively pronounced. Frontal
<taxonomicName id="4C164D61FF922573FB10F9260194F94D" box="[1158,1216,1715,1741]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
64139, 103 µm long. Thoracic horn (
<figureCitation id="132D2A67FF922573FDFFF94E0794F975" box="[617,704,1755,1781]" captionStart="FIGURES 23 27" captionStartId="32.[264,382,1814,1838]" captionTargetBox="[362,1271,556,1802]" captionTargetId="figure@32.[312,1277,554,1804]" captionTargetPageId="32" captionText="FIGURES 23 27. Orthocladius (Mesorthocladius) spp., pupae, 23: O. (M.) lamellatus sp. n., tergites, 24: thoracic horn, 25 27: O. (M.) nimidens sp. n., 25: tergites, 26: thoracic horn, 27: frontal apotome." httpUri="https://zenodo.org/record/171319/files/figure.png" pageId="39" pageNumber="40">Fig. 26</figureCitation>
) 266428, 363 µm long; 4968, 56 µm wide; 5.46 7.85, 6.54 times as long as wide; 0.951.23, 1.11 times as long as anal macrosetae. Precorneal
<taxonomicName id="4C164D61FF922573FED6F8BE042FF8C5" box="[320,379,1835,1861]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
respectively 68113, 96 µm; 86169, 133 µm; and 5698, 84 µm long. Anterior dorsocentral (Dc1) 71120, 95 µm long; Dc2, 53105, 75 µm; Dc3 7194, 69 µm; and Dc4 79116, 94 µm long. Distances (in µm): Dc1Dc
<date id="FFA81022FF9D257CFCD4FEC306C8FEE3" box="[834,924,329,359]" pageId="40" pageNumber="41" value="1960-02-30">2 3060</date>
, 46; Dc2Dc
<date id="FFA81022FF9D257CFBA7FEC301D8FEE3" box="[1073,1164,329,359]" pageId="40" pageNumber="41" value="1964-03-26">3 2664</date>
, 49; Dc3Dc4 3471, 55.
</paragraph>
<paragraph id="8BA936E2FF9D257CFEAEFE0A0600FC1B" blockId="40.[264,1323,284,1883]" pageId="40" pageNumber="41">
Abdomen (
<figureCitation id="132D2A67FF9D257CFE2DFE0A0747FE39" box="[443,531,415,441]" captionStart="FIGURES 23 27" captionStartId="32.[264,382,1814,1838]" captionTargetBox="[362,1271,556,1802]" captionTargetId="figure@32.[312,1277,554,1804]" captionTargetPageId="32" captionText="FIGURES 23 27. Orthocladius (Mesorthocladius) spp., pupae, 23: O. (M.) lamellatus sp. n., tergites, 24: thoracic horn, 25 27: O. (M.) nimidens sp. n., 25: tergites, 26: thoracic horn, 27: frontal apotome." httpUri="https://zenodo.org/record/171319/files/figure.png" pageId="40" pageNumber="41">Fig. 25</figureCitation>
). Tergite I bare, tergite II with weak median and anterolateral group shagreen and stronger posterior shagreen; strong, extensive shagreen on tergites IIIVI, posterior spinules on tergite III stronger than anterior ones, of about the same size on IV VI, spinule bands confluent with anterior shagreen reaching about as far lateral as posterior shagreen; tergite VII with weak median shagreen, VIII and IX with anterior group shagreen. Sternite I bare; II with weak lateral shagreen, IIV with stronger lateral, VI with lateral, anterolateral and posteromedian shagreen, VII with fewer anterolateral and posteromedian spinules, VIII with anterolateral VVII with lateral plus anterolateral, VIII and IX with anterolateral shagreen. Pedes spurii A on sternites IVVII. Pedes spurii B well developed, granulose, on tergite II only. Caudal hooklets on tergite
<date id="FFA81022FF9D257CFBB3FC92010AFCA1" box="[1061,1118,775,801]" pageId="40" pageNumber="41" value="1961-02">II 61</date>
160, 111; in 45 rows. Segments IIVI each with 3
<taxonomicName id="4C164D61FF9D257CFD37FCBA07A3FCC9" box="[673,759,815,841]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="setae">L setae</taxonomicName>
, VII with 4, and VIII with 45
<taxonomicName id="4C164D61FF9D257CFBFFFCBA0195FCC9" box="[1129,1217,815,841]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="setae">L setae</taxonomicName>
. Lengths of L1L5
<taxonomicName id="4C164D61FF9D257CFEE3FCC204E4FCF1" box="[373,432,855,881]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
on segment VIII (in µm) as: 79165, 123; 86199, 138; 90184, 132; 139 240, 186; 105120 (present in 2 of
<specimenCount id="9D10FD6BFF9D257CFD32FC14061DFC1B" box="[676,841,897,923]" pageId="40" pageNumber="41" type="generic">10 specimens</specimenCount>
).
</paragraph>
<paragraph id="8BA936E2FF9D257CFEAEFC3C0111FB93" blockId="40.[264,1323,284,1883]" pageId="40" pageNumber="41">Anal lobe 338401, 374 µm long; each lobe with 914, 12 fine lateral spines; no posterior spines. Anal macrosetae subequal in length; 281375, 326 µm long; 0.790.94, 0.87 as long as anal lobe. Male genital sac reaching about to apex of anal lobe.</paragraph>
<paragraph id="8BA936E2FF9D257CFEAEFBB406B9FBBB" blockId="40.[264,1323,284,1883]" box="[312,1005,1057,1083]" pageId="40" pageNumber="41">
<emphasis id="B962EAF0FF9D257CFEAEFBB4074DFBBA" box="[312,537,1057,1082]" italics="true" pageId="40" pageNumber="41">Fourth instar larva</emphasis>
(
<emphasis id="B962EAF0FF9D257CFDBDFBB4076EFBBA" box="[555,570,1057,1082]" italics="true" pageId="40" pageNumber="41">n</emphasis>
= 89 except when otherwise stated)
</paragraph>
<paragraph id="8BA936E2FF9D257CFEAEFBDC06DEFB0B" blockId="40.[264,1323,284,1883]" pageId="40" pageNumber="41">
Total length about 4.88.1,
<quantity id="4CEE9B07FF9D257CFDECFBDC078CFBE3" box="[634,728,1097,1123]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="40" pageNumber="41" unit="mm" value="6.2">6.2 mm</quantity>
. Head capsule 0.480.53,
<quantity id="4CEE9B07FF9D257CFB9FFBDC0126FBE3" box="[1033,1138,1097,1123]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="40" pageNumber="41" unit="mm" value="0.51">0.51 mm</quantity>
long. Postmentum 225263, 240 µm long. Head capsule light brown.
</paragraph>
<paragraph id="8BA936E2FF9D257CFEAEFB0C076DF9EB" blockId="40.[264,1323,284,1883]" pageId="40" pageNumber="41">
Head. Antenna as in
<figureCitation id="132D2A67FF9D257CFDB8FB0C07D1FB33" box="[558,645,1177,1203]" captionStart="FIGURES 28 34" captionStartId="34.[264,383,1688,1712]" captionTargetBox="[308,1302,296,1678]" captionTargetId="figure@34.[269,1318,280,1682]" captionTargetPageId="34" captionText="FIGURES 28 34. Orthocladius spp., larvae, 28: Orthocladius (Mesorthocladius) frigidus (Zetterstedt) labrum, epipharyngeal area and premandible, 29, 30: Orthocladius (Orthocladius) nitidoscutellatus Lundström, 29: antenna, 30: labrum, epipharyngeal area and premandible, 31, 32: Orthocladius (Mesorthocladius) lamellatus sp. n., 31: antenna, 32: labrum, epipharyngeal area and premandible, 33 34: Orthocladius (Mesorthocladius) nimidens sp. n., 31: antenna, 32: labrum, epipharyngeal area and premandible." httpUri="https://zenodo.org/record/171320/files/figure.png" pageId="40" pageNumber="41">Fig. 31</figureCitation>
. Length of antennal segments (in µm): 7386, 79; 1517, 16; 47, 6; 34, 4; 56, 6. AR 2.382.81, 2.62. Ring organ 1319, 17 µm from base; basal setal mark 1625, 20 µm from base; apical setal mark 2035, 28 m from base. Basal antennal segment 2025, 22 µm wide; blade 2026, 23 µm (6) long; accessory blade10 17, 14 µm (8) long. Lauterborn organs 57, 6 µm (7) long; style 59, 7 µm long. Labrum (
<figureCitation id="132D2A67FF9D257CFE87FAF4043CFAFB" box="[273,360,1377,1403]" captionStart="FIGURES 28 34" captionStartId="34.[264,383,1688,1712]" captionTargetBox="[308,1302,296,1678]" captionTargetId="figure@34.[269,1318,280,1682]" captionTargetPageId="34" captionText="FIGURES 28 34. Orthocladius spp., larvae, 28: Orthocladius (Mesorthocladius) frigidus (Zetterstedt) labrum, epipharyngeal area and premandible, 29, 30: Orthocladius (Orthocladius) nitidoscutellatus Lundström, 29: antenna, 30: labrum, epipharyngeal area and premandible, 31, 32: Orthocladius (Mesorthocladius) lamellatus sp. n., 31: antenna, 32: labrum, epipharyngeal area and premandible, 33 34: Orthocladius (Mesorthocladius) nimidens sp. n., 31: antenna, 32: labrum, epipharyngeal area and premandible." httpUri="https://zenodo.org/record/171320/files/figure.png" pageId="40" pageNumber="41">Fig. 32</figureCitation>
) with one branch of S I split into 26 weak branches, S II double on one side in one specimen, 12 chaetae and 5 spinulae. Chaetulae laterales smooth. Premandible bifid (
<figureCitation id="132D2A67FF9D257CFE87FA24043EFA4B" box="[273,362,1457,1483]" captionStart="FIGURES 28 34" captionStartId="34.[264,383,1688,1712]" captionTargetBox="[308,1302,296,1678]" captionTargetId="figure@34.[269,1318,280,1682]" captionTargetPageId="34" captionText="FIGURES 28 34. Orthocladius spp., larvae, 28: Orthocladius (Mesorthocladius) frigidus (Zetterstedt) labrum, epipharyngeal area and premandible, 29, 30: Orthocladius (Orthocladius) nitidoscutellatus Lundström, 29: antenna, 30: labrum, epipharyngeal area and premandible, 31, 32: Orthocladius (Mesorthocladius) lamellatus sp. n., 31: antenna, 32: labrum, epipharyngeal area and premandible, 33 34: Orthocladius (Mesorthocladius) nimidens sp. n., 31: antenna, 32: labrum, epipharyngeal area and premandible." httpUri="https://zenodo.org/record/171320/files/figure.png" pageId="40" pageNumber="41">Fig. 32</figureCitation>
), 98109, 105 µm (8) long; vestigial brush indicated.
<taxonomicName id="4C164D61FF9D257CFC65FA2401F2FA4B" box="[1011,1190,1457,1483]" class="Bivalvia" family="Pectinidae" genus="Pecten" kingdom="Animalia" order="Pectinoida" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="galearis">Pecten galearis</taxonomicName>
apparently absent. Mandible (
<figureCitation id="132D2A67FF9D257CFE73FA4C076FFA73" box="[485,571,1497,1523]" captionStart="FIGURES 35 40" captionStartId="36.[264,383,1691,1715]" captionTargetBox="[281,1294,319,1672]" captionTargetId="figure@36.[269,1318,299,1687]" captionTargetPageId="36" captionText="FIGURES 35 40. Orthocladius spp., larvae, 35, 36: Orthocladius (Mesorthocladius) lamellatus sp. n., 35: mandible, 36: mentum, 37 38: Orthocladius (Mesorthocladius) nimidens sp. n., 37: mandible, 38: mentum, 39 40: Orthocladius (Orthocladius) nitidoscutellatus Lundström, 39: mandible, 40: mentum" httpUri="https://zenodo.org/record/171321/files/figure.png" pageId="40" pageNumber="41">Fig. 37</figureCitation>
) 176210, 189 µm long; seta interna of 78 branches. Mentum (
<figureCitation id="132D2A67FF9D257CFE87F9940433F99B" box="[273,359,1537,1563]" captionStart="FIGURES 35 40" captionStartId="36.[264,383,1691,1715]" captionTargetBox="[281,1294,319,1672]" captionTargetId="figure@36.[269,1318,299,1687]" captionTargetPageId="36" captionText="FIGURES 35 40. Orthocladius spp., larvae, 35, 36: Orthocladius (Mesorthocladius) lamellatus sp. n., 35: mandible, 36: mentum, 37 38: Orthocladius (Mesorthocladius) nimidens sp. n., 37: mandible, 38: mentum, 39 40: Orthocladius (Orthocladius) nitidoscutellatus Lundström, 39: mandible, 40: mentum" httpUri="https://zenodo.org/record/171321/files/figure.png" pageId="40" pageNumber="41">Fig. 38</figureCitation>
) 161180, 169 µm wide; with 15 or 17 teeth, median tooth 3547, 40 µm wide; ventromental plate 1016, 14 µm wide; about 4 very weak
<taxonomicName id="4C164D61FF9D257CFC5DF9BC0152F9C3" box="[971,1030,1577,1603]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
present underneath ventromental plates on cardo.
</paragraph>
<paragraph id="8BA936E2FF9D257CFEAEF9EC04ADF8DB" blockId="40.[264,1323,284,1883]" pageId="40" pageNumber="41">
Abdomen. Claws of anterior parapods strongly serrated. Longest body
<taxonomicName id="4C164D61FF9D257CFBEAF9EC01E3F913" box="[1148,1207,1657,1683]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
131278, 186 µm long. Posterior parapods 225323, 280 µm (5) long. Procercus 4156, 48 µm high; 2645, 39 µm wide. Anal
<taxonomicName id="4C164D61FF9D257CFDD5F95C0729F963" box="[579,637,1737,1763]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
544675, 618 µm long; supraanal seta 143206, 167 µm (8) long, 0.240.31, 0.27 (8) as long as anal
<taxonomicName id="4C164D61FF9D257CFC8DF964060DF88B" box="[795,857,1777,1803]" class="Insecta" family="Chironomidae" genus="Orthocladius" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="setae">setae</taxonomicName>
. Longer anal tubules 151188, 162 µm (6) long, 75113, 94 µm (6) wide at base; shorter anal tubules 105143 µm (3) long, 5668 µm (3) wide at base.
</paragraph>
</subSubSection>
<subSubSection id="C30C6569FF9C257DFE9EFE89017AFC0E" pageId="41" pageNumber="42" type="discussion">
<paragraph id="8BA936E2FF9C257DFE9EFE8904D7FEB5" blockId="41.[264,1323,284,670]" box="[264,387,284,309]" pageId="41" pageNumber="42">
<emphasis id="B962EAF0FF9C257DFE9EFE8904D7FEB5" box="[264,387,284,309]" italics="true" pageId="41" pageNumber="42">Comments</emphasis>
</paragraph>
<paragraph id="8BA936E2FF9C257DFEAEFED10772FD1E" blockId="41.[264,1323,284,670]" pageId="41" pageNumber="42">
The female imago is unique within the genus, with the apical wart on the seminal capsule, spermathecal ducts with several loops but no widened portions, dorsomesal lobe without the anterior projection characteristic for the
<emphasis id="B962EAF0FF9C257DFC19FE0101E2FE2D" box="[911,1206,404,429]" italics="true" pageId="41" pageNumber="42">Cricotopus­Orthocladius</emphasis>
group of genera, and no discernible rami. The genitalia are surprisingly similar to those of some
<taxonomicName id="4C164D61FF9C257DFE9EFE71075BFE7E" authority="Waltl. Other" authorityName="Waltl. Other" box="[264,527,484,510]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B962EAF0FF9C257DFE9EFE710424FE7D" box="[264,368,484,509]" italics="true" pageId="41" pageNumber="42">Diamesa</emphasis>
Waltl. Other
</taxonomicName>
apparent autapomorphies within the genus are the triangular anal point of the male imago and the rounded, instead of concave, posterior part of the larval ventromental plates. However, the pupa shares several synapomorphies with other members of the subgenus such as prominent frontal warts, granulose pedes spurii B, and lateral threads on the anal lobe.
</paragraph>
<paragraph id="8BA936E2FF9C257DFE9EFD410778FD6D" blockId="41.[264,1323,724,910]" box="[264,556,724,749]" pageId="41" pageNumber="42">
<emphasis id="B962EAF0FF9C257DFE9EFD410778FD6D" box="[264,556,724,749]" italics="true" pageId="41" pageNumber="42">Ecology and distribution</emphasis>
</paragraph>
<paragraph id="8BA936E2FF9C257DFEAEFD69017AFC0E" blockId="41.[264,1323,724,910]" pageId="41" pageNumber="42">The species inhabits vernal woodland shallow pools and springs. This habitat is only present in the spring when the groundwater level is high enough to supply them, or possibly they are created where an impermeable clay lens is pooling snow­melt and spring rains. The species is known from Ohio, Indiana, and North Carolina.</paragraph>
</subSubSection>
</treatment>
</document>