1015 lines
158 KiB
XML
1015 lines
158 KiB
XML
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<mods:title id="2829E92CADD2AE3D7FE8A16061F6A032">Description of two new species of goby-associated snapping shrimps from the tropical western Pacific (Decapoda: Alpheidae: Alpheus)</mods:title>
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<treatment id="03BF87EB05130510B4B18CFFFCF31956" ID-DOI="http://doi.org/10.5281/zenodo.5888954" ID-GBIF-Taxon="192501151" ID-Zenodo-Dep="5888954" LSID="urn:lsid:plazi:treatment:03BF87EB05130510B4B18CFFFCF31956" httpUri="http://treatment.plazi.org/id/03BF87EB05130510B4B18CFFFCF31956" lastPageId="15" lastPageNumber="288" pageId="8" pageNumber="281">
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<paragraph id="8BA936FD05130517B4B18CFFFE791DF6" blockId="8.[151,431,151,213]" box="[151,431,151,178]" pageId="8" pageNumber="281">
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<heading id="D0E1819105130517B4B18CFFFE791DF6" bold="true" box="[151,431,151,178]" fontSize="11" level="1" pageId="8" pageNumber="281" reason="1">
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<emphasis id="B962EAEF05130517B4B18CFFFE791DF6" bold="true" box="[151,431,151,178]" pageId="8" pageNumber="281">
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<taxonomicName id="4C164D7E05130517B4B18CFFFE9C1DF5" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[151,330,151,178]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="281" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
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<emphasis id="B962EAEF05130517B4B18CFFFE9C1DF5" bold="true" box="[151,330,151,178]" italics="true" pageId="8" pageNumber="281">Alpheus sciolii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A251579405130517B5778CFFFE791DF6" box="[337,431,151,177]" pageId="8" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C30C657605130517B4B18CD3FE971D92" box="[151,321,187,213]" pageId="8" pageNumber="281" type="description">
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<paragraph id="8BA936FD05130517B4B18CD3FE971D92" blockId="8.[151,431,151,213]" box="[151,321,187,213]" pageId="8" pageNumber="281">
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(
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<figureCitation id="132D2A7805130517B4B98CD3FEDA1D92" box="[159,268,187,213]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStartId-0="9.[151,250,1839,1863]" captionStartId-1="10.[151,250,1709,1733]" captionStartId-2="11.[151,250,1665,1689]" captionStartId-3="13.[151,250,1880,1904]" captionTargetBox-0="[169,1413,191,1801]" captionTargetBox-1="[165,1420,203,1676]" captionTargetBox-2="[190,1400,792,1619]" captionTargetBox-3="[151,1435,181,1856]" captionTargetId-0="figure-15@9.[151,1436,181,1816]" captionTargetId-1="figure-17@10.[151,1436,181,1686]" captionTargetId-2="figure-271@11.[151,1436,777,1642]" captionTargetId-3="figure-15@13.[151,1435,181,1856]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="13" captionText-0="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." captionText-1="FIGURE 5. Alpheus sciolii sp. nov., paratype, male (cl 9.3 mm) from New Georgia, Solomon Islands (MNHN-IU-2018-5669): A, major (right) cheliped, lateral; B, same, chela, mesial; C, same, detail of ventromesial margin of merus, mesial, slender setae omitted; D, same, dactylus and pollex, lateral, setae omitted; E, minor (left) cheliped, carpus and chela, mesial; F, same, chela, lateral." captionText-2="FIGURE 6. Alpheus sciolii sp. nov., non-type male (cl 14.5 mm) from Nha Trang Bay, Vietnam (MNHN-IU-2018-5671):A, left (major) cheliped, carpus and chela, lateral; B, right (minor) cheliped, chela, lateral; C, telson, dorsal, plumose setae omitted." captionText-3="FIGURE 7. Alpheus sciolii sp. nov., holotype male (cl 8.6 mm) from New Georgia, Solomon Islands (MNHN-IU-2018-5667) [A, B]; paratype female (cl 10.7 mm) from Heron Island, Queensland, Australia (MNHN-IU-2018-5668) [C]: A, C, living shrimps, dorsal; B, same, lateral. Photographs by the author." figureDoi-0="http://doi.org/10.5281/zenodo.5881105" figureDoi-1="http://doi.org/10.5281/zenodo.5881107" figureDoi-2="http://doi.org/10.5281/zenodo.5881113" figureDoi-3="http://doi.org/10.5281/zenodo.5881119" httpUri-0="https://zenodo.org/record/5881105/files/figure.png" httpUri-1="https://zenodo.org/record/5881107/files/figure.png" httpUri-2="https://zenodo.org/record/5881113/files/figure.png" httpUri-3="https://zenodo.org/record/5881119/files/figure.png" pageId="8" pageNumber="281">Figs. 4–7</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A7805130517B5318CD3FEEE1D92" box="[279,312,187,213]" captionStart="FIGURE 8" captionStartId="14.[151,250,1545,1569]" captionTargetBox="[158,1430,181,1521]" captionTargetId="figure-17@14.[158,1430,181,1521]" captionTargetPageId="14" captionText="FIGURE 8. Some goby-associated species of Alpheus mentioned in the text, photographed in situ: A, Alpheus thompsoni sp. nov. associated with Amblyeleotris wheeleri, Anilao, Philippines; B, same, associated with Amblyeleotris guttata, Sulawesi, Indonesia; C, Alpheus sciolii sp. nov. associated with Cryptocentrus inexplicatus, Papua New Guinea; D, Alpheus sp. aff. A. sciolii sp. nov. associated with Amblyeleotris steinitzi, Bali, Indonesia; E, Alpheus sp. cf. mannarensis Purushothaman, Abhilash, Kumar & Lal, 2021, Papua New Guinea; F, Alpheus bellulus Miya & Miyake, 1969, Sulawesi, Indonesia. Photographs courtesy of Andrey Ryanskiy (A–C, E, F) and Yury Ivanov (D)." figureDoi="http://doi.org/10.5281/zenodo.5881121" httpUri="https://zenodo.org/record/5881121/files/figure.png" pageId="8" pageNumber="281">8C</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30C657605130517B4B18D68FC8E1CF6" pageId="8" pageNumber="281" type="reference_group">
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<paragraph id="8BA936FD05130517B4B18D68FCE11C71" blockId="8.[151,1412,255,433]" pageId="8" pageNumber="281">
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<taxonomicName id="4C164D7E05130517B4B18D68FE891C5F" ID-CoL="85LFJ" authorityName="Miya & Miyake" authorityYear="1969" box="[151,351,256,280]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="281" phylum="Arthropoda" rank="species" species="bellulus">
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||
<emphasis id="B962EAEF05130517B4B18D68FF3F1C5F" box="[151,233,256,280]" italics="true" pageId="8" pageNumber="281">Alpheus</emphasis>
|
||
cf.
|
||
<emphasis id="B962EAEF05130517B5298D68FE891C5F" box="[271,351,256,280]" italics="true" pageId="8" pageNumber="281">bellulus</emphasis>
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</taxonomicName>
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||
—
|
||
<bibRefCitation id="EF874B0C05130517B5A08D68FDE41C5F" author="Marin, I." box="[390,562,256,280]" pageId="8" pageNumber="281" pagination="375 - 390" refId="ref11334" refString="Marin, I. (2008) Description of two new species from the genera Palaemonella Dana, 1852 and Vir Holthuis, 1952 (Crustacea: Caridea: Palaemonidae: Pontoniinae). Zoologische Mededelingen, 82, 375 - 390." type="journal article" year="2008">Marin 2008: 382</bibRefCitation>
|
||
, fig. 9b (as
|
||
<taxonomicName id="4C164D7E05130517B68F8D68FCC31C5F" ID-CoL="85LFJ" authorityName="Miya & Miyake" authorityYear="1969" box="[681,789,256,280]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="281" phylum="Arthropoda" rank="species" species="bellulus">
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||
<emphasis id="B962EAEF05130517B68F8D68FCC31C5F" box="[681,789,256,280]" italics="true" pageId="8" pageNumber="281">A. bellulus</emphasis>
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||
</taxonomicName>
|
||
in the figure legend) [not
|
||
<taxonomicName id="4C164D7E05130517B0398D68FAAC1C50" ID-CoL="85LFJ" authority="Miya & Miyake, 1969" authorityName="Miya & Miyake" authorityYear="1969" box="[1055,1402,255,280]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="281" phylum="Arthropoda" rank="species" species="bellulus">
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<emphasis id="B962EAEF05130517B0398D68FB5D1C5F" box="[1055,1163,256,280]" italics="true" pageId="8" pageNumber="281">A. bellulus</emphasis>
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||
<bibRefCitation id="EF874B0C05130517B0B48D68FAAC1C50" author="Miya, Y. & Miyake, S." box="[1170,1402,255,280]" pageId="8" pageNumber="281" pagination="307 - 314" refId="ref11409" refString="Miya, Y. & Miyake, S. (1969) Description of Alpheus bellulus sp. nov. associated with gobies from Japan (Crustacea, Decapoda, Alpheidae). Publications of the Seto Marine Biological Laboratory, 16, 307 - 314. https: // doi. org / 10.5134 / 175556" type="journal article" year="1969">Miya & Miyake, 1969</bibRefCitation>
|
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</taxonomicName>
|
||
].
|
||
<taxonomicName id="4C164D7E05130517B4B18D76FE5D1C71" authorityName="Miya & Miyake" authorityYear="1969" box="[151,395,286,310]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="281" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B962EAEF05130517B4B18D76FF3F1C71" box="[151,233,286,310]" italics="true" pageId="8" pageNumber="281">Alpheus</emphasis>
|
||
sp. aff.
|
||
<emphasis id="B962EAEF05130517B51D8D76FE5D1C71" box="[315,395,286,310]" italics="true" pageId="8" pageNumber="281">bellulus</emphasis>
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</taxonomicName>
|
||
—
|
||
<bibRefCitation id="EF874B0C05130517B5898D76FD091C71" author="Anker, A. & De Grave, S." box="[431,735,286,310]" pageId="8" pageNumber="281" pagination="145 - 150" refId="ref10292" refString="Anker, A. & De Grave, S. (2019) Further records of burrow-associated palaemonid shrimps (Decapoda: Palaemonidae). Zootaxa, 4612 (1), 145 - 150. https: // doi. org / 10.11646 / zootaxa. 4612.1.13" type="journal article" year="2019">Anker & De Grave 2019: 148</bibRefCitation>
|
||
, fig. 4C.
|
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</paragraph>
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<paragraph id="8BA936FD05130517B4B18D55FD1B1C12" blockId="8.[151,1412,255,433]" box="[151,717,317,341]" pageId="8" pageNumber="281">
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<taxonomicName id="4C164D7E05130517B4B18D55FE471C12" ID-CoL="85LH6" authorityName="De Man" authorityYear="1909" box="[151,401,317,341]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="281" phylum="Arthropoda" rank="species" species="djiboutensis">
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<emphasis id="B962EAEF05130517B4B18D55FF3F1C12" box="[151,233,317,341]" italics="true" pageId="8" pageNumber="281">Alpheus</emphasis>
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aff.
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<emphasis id="B962EAEF05130517B5318D55FE471C12" box="[279,401,317,341]" italics="true" pageId="8" pageNumber="281">djiboutensis</emphasis>
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</taxonomicName>
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—
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<treatmentCitation id="0AB710EC05130517B59E8D55FDA21C12" author="Hurt, C. & Hultgren, K. M. & Anker, A. & Lemmon, A. R. & Moriarty Lemmon, E. & Bracken-Grissom, H." box="[440,628,317,341]" page="2" pageId="8" pageNumber="281" year="2021">
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<bibRefCitation id="EF874B0C05130517B59E8D55FDA21C12" author="Hurt, C. & Hultgren, K. M. & Anker, A. & Lemmon, A. R. & Moriarty Lemmon, E. & Bracken-Grissom, H." box="[440,628,317,341]" pageId="8" pageNumber="281" pagination="1 - 11" refId="ref11003" refString="Hurt, C., Hultgren, K. M., Anker, A., Lemmon, A. R., Moriarty Lemmon, E. & Bracken-Grissom, H. (2021) First worldwide molecular phylogeny of the morphologically and ecologically hyperdiversified snapping shrimp genus Alpheus (Malacostraca: Decapoda). Molecular Phylogenetics and Evolution, 158 (197080), 1 - 11. https: // doi. org / 10.1016 / j. ympev. 2021.107080" type="journal article" year="2021">
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Hurt
|
||
<emphasis id="B962EAEF05130517B5CB8D56FDF61C12" box="[493,544,317,341]" italics="true" pageId="8" pageNumber="281">et al.</emphasis>
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2021: 2
|
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</bibRefCitation>
|
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</treatmentCitation>
|
||
, fig. 1D.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BA936FD05130517B4B18D34FD111C33" blockId="8.[151,1412,255,433]" box="[151,711,347,372]" pageId="8" pageNumber="281">
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<treatmentCitationGroup id="AB0611D305130517B4B18D34FD111C33" box="[151,711,347,372]" pageId="8" pageNumber="281">
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<taxonomicName id="4C164D7E05130517B4B18D34FEDA1C34" box="[151,268,348,372]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="281" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B962EAEF05130517B4B18D34FF3F1C33" box="[151,233,348,372]" italics="true" pageId="8" pageNumber="281">Alpheus</emphasis>
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||
sp.
|
||
</taxonomicName>
|
||
—
|
||
<treatmentCitation id="0AB710EC05130517B5148D34FE2C1C34" author="Debelius, H." box="[306,506,347,372]" page="153" pageId="8" pageNumber="281" year="2001">
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<bibRefCitation id="EF874B0C05130517B5148D34FE2C1C34" author="Debelius, H." box="[306,506,347,372]" pageId="8" pageNumber="281" refId="ref10810" refString="Debelius, H. (2001) Crustacea guide of the world. IKAN, Frankfurt, 321 pp." type="book" year="2001">Debelius 2001: 153</bibRefCitation>
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, colour photograph.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BA936FD05130517B4B18D12FD3E1CD5" blockId="8.[151,1412,255,433]" box="[151,744,378,402]" pageId="8" pageNumber="281">
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<treatmentCitationGroup id="AB0611D305130517B4B18D12FD3E1CD5" box="[151,744,378,402]" pageId="8" pageNumber="281">
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<taxonomicName id="4C164D7E05130517B4B18D12FEC91CD5" box="[151,287,378,402]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="281" phylum="Arthropoda" rank="species" species="undefined-2">
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<emphasis id="B962EAEF05130517B4B18D12FF3F1CD5" box="[151,233,378,402]" italics="true" pageId="8" pageNumber="281">Alpheus</emphasis>
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||
sp. 2
|
||
</taxonomicName>
|
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—
|
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<treatmentCitation id="0AB710EC05130517B5638D12FDCA1CD5" author="Minemizu, R." box="[325,540,378,402]" page="101" pageId="8" pageNumber="281" year="2013">
|
||
<bibRefCitation id="EF874B0C05130517B5638D12FDCA1CD5" author="Minemizu, R." box="[325,540,378,402]" pageId="8" pageNumber="281" refId="ref11377" refString="Minemizu, R. (2013) Coral reef shrimps of Indo-West Pacific. Bun-ichi Sogo Shuppan, Co. Ltd., Tokyo, 144 pp. [in Japanese]" type="book" year="2013">Minemizu 2013: 101</bibRefCitation>
|
||
</treatmentCitation>
|
||
, colour photograph.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BA936FD05130517B4B18DF1FC8E1CF6" blockId="8.[151,1412,255,433]" box="[151,856,409,433]" pageId="8" pageNumber="281">
|
||
<treatmentCitationGroup id="AB0611D305130517B4B18DF1FC8E1CF6" box="[151,856,409,433]" pageId="8" pageNumber="281">
|
||
<taxonomicName id="4C164D7E05130517B4B18DF1FEC91CF6" box="[151,287,409,433]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="281" phylum="Arthropoda" rank="species" species="undefined-6">
|
||
<emphasis id="B962EAEF05130517B4B18DF1FF3F1CF6" box="[151,233,409,433]" italics="true" pageId="8" pageNumber="281">Alpheus</emphasis>
|
||
sp. 6
|
||
</taxonomicName>
|
||
—
|
||
<treatmentCitation id="0AB710EC05130517B5638DF1FDB91CF6" author="Kuiter, R. H. & Debelius, H." box="[325,623,409,433]" page="152" pageId="8" pageNumber="281" year="2009">
|
||
<bibRefCitation id="EF874B0C05130517B5638DF1FDB91CF6" author="Kuiter, R. H. & Debelius, H." box="[325,623,409,433]" pageId="8" pageNumber="281" refId="ref11307" refString="Kuiter, R. H. & Debelius, H. (2009) World atlas of marine fauna. IKAN, Frankfurt, 725 pp." type="book" year="2009">Kuiter & Debelius 2009: 152</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 4 colour photographs.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C657605130517B4B18DB7FB351EC2" pageId="8" pageNumber="281" type="materials_examined">
|
||
<paragraph id="8BA936FD05130517B4B18DB7FB5B1E7A" blockId="8.[151,1437,479,1910]" pageId="8" pageNumber="281">
|
||
<emphasis id="B962EAEF05130517B4B18DB7FEEA1CBE" bold="true" box="[151,316,479,505]" pageId="8" pageNumber="281">Type material</emphasis>
|
||
.
|
||
<materialsCitation id="3B7E3CA005130517B56F8DB7FAC91F5A" ID-GBIF-Occurrence="3450196301" collectingDate="2016-09-14" collectorName="A. Anker" country="Solomon Islands" county="Hopei Island" elevation="8" location="New" municipality="Munda" pageId="8" pageNumber="281" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="54AD885F05130517B56F8DB7FE601CBE" box="[329,438,479,505]" pageId="8" pageNumber="281" type="holotype">Holotype</typeStatus>
|
||
: male (cl
|
||
<quantity id="4CEE9B1805130517B6388DB7FDA11CBD" box="[542,631,479,506]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="8" pageNumber="281" unit="mm" value="8.6">8.6 mm</quantity>
|
||
), MNHN-IU-2018-5667,
|
||
<collectingCountry id="F301766D05130517B7B28DB7FB871CBE" box="[916,1105,479,505]" name="Solomon Islands" pageId="8" pageNumber="281">Solomon Islands</collectingCountry>
|
||
,
|
||
<location id="8EC9602605130517B07D8D88FB461CBD" LSID="urn:lsid:plazi:treatment:03BF87EB05130510B4B18CFFFCF31956:8EC9602605130517B07D8D88FB461CBD" box="[1115,1168,480,506]" country="Solomon Islands" county="Hopei Island" municipality="Munda" name="New" pageId="8" pageNumber="281">New</location>
|
||
Georgia,
|
||
<collectingMunicipality id="6BCDAC8705130517B0DA8DB7FA981CBE" box="[1276,1358,479,505]" pageId="8" pageNumber="281">Munda</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62C84E7105130517B17E8DB7FF081F5A" pageId="8" pageNumber="281">Hopei Island</collectingCounty>
|
||
, shallow sand flat, suction (yabby) pump, depth
|
||
<quantity id="4CEE9B1805130517B6D78E6BFC9B1F59" box="[753,845,515,542]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="8" pageNumber="281" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">
|
||
<elevation id="003BD1CE05130517B6D78E6BFC9B1F59" box="[753,845,515,542]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="8" pageNumber="281" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 m</elevation>
|
||
</quantity>
|
||
, leg.
|
||
<collectorName id="26E3532B05130517B7A28E6BFC3D1F5A" box="[900,1003,515,541]" pageId="8" pageNumber="281">A. Anker</collectorName>
|
||
,
|
||
<date id="FFA8103D05130517B7D28E6BFBA41F5A" box="[1012,1138,515,541]" pageId="8" pageNumber="281" value="2016-09-14">
|
||
<collectingDate id="EFECE9D505130517B7D28E6BFBA41F5A" box="[1012,1138,515,541]" pageId="8" pageNumber="281" value="2016-09-14">14.09.2016</collectingDate>
|
||
</date>
|
||
[fcn SOL-106]
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B7E3CA005130517B10E8E6CFAF61FEA" ID-GBIF-Occurrence="3450196305" collectingDate="2016-09-19" collectingDateMax="2016-09-28" collectingDateMin="2016-09-19" collectorName="A. Anker" country="Solomon Islands" elevation="1" location="Heron Island" pageId="8" pageNumber="281" specimenCount="1" specimenCount-male="1" stateProvince="Queensland" typeStatus="paratype">
|
||
<typeStatus id="54AD885F05130517B10E8E6CFA4F1F59" box="[1320,1433,516,542]" pageId="8" pageNumber="281" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D10FD7405130517B4B18E4FFF341F06" box="[151,226,551,577]" pageId="8" pageNumber="281" type="male">1 male</specimenCount>
|
||
(cl
|
||
<quantity id="4CEE9B1805130517B5288E4FFEBE1F05" box="[270,360,551,578]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="8" pageNumber="281" unit="mm" value="9.3">9.3 mm</quantity>
|
||
), MNHN-IU-2018-5669,
|
||
<collectingCountry id="F301766D05130517B6A18E4FFC931F06" box="[647,837,551,577]" name="Solomon Islands" pageId="8" pageNumber="281">Solomon Islands</collectingCountry>
|
||
,
|
||
<collectorName id="26E3532B05130517B7778E40FC501F05" box="[849,902,552,578]" pageId="8" pageNumber="281">New</collectorName>
|
||
Georgia,
|
||
<collectorName id="26E3532B05130517B7D28E4FFB901F06" box="[1012,1094,551,577]" pageId="8" pageNumber="281">Munda</collectorName>
|
||
,
|
||
<collectorName id="26E3532B05130517B0778E4FFAD61F06" box="[1105,1280,551,577]" hasSafeAffix="true" pageId="8" pageNumber="281">Sosuhite Island</collectorName>
|
||
, shallow sand flat with coral rubble, depth
|
||
<quantity id="4CEE9B1805130517B5FB8E23FDEA1F21" box="[477,572,587,614]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="8" pageNumber="281" unit="m" value="1.25" valueMax="2.0" valueMin="0.5">
|
||
<elevation id="003BD1CE05130517B5FB8E23FDEA1F21" box="[477,572,587,614]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="8" pageNumber="281" unit="m" value="1.25" valueMax="2.0" valueMin="0.5">0.5–2 m</elevation>
|
||
</quantity>
|
||
, in burrow under large piece of coral rubble, leg.
|
||
<collectorName id="26E3532B05130517B0558E23FB0B1F22" box="[1139,1245,587,613]" pageId="8" pageNumber="281">A. Anker</collectorName>
|
||
,
|
||
<date id="FFA8103D05130517B0CF8E23FAB11F22" box="[1257,1383,587,613]" pageId="8" pageNumber="281" value="2016-09-19">
|
||
<collectingDate id="EFECE9D505130517B0CF8E23FAB11F22" box="[1257,1383,587,613]" pageId="8" pageNumber="281" value="2016-09-19">19.09.2016</collectingDate>
|
||
</date>
|
||
[fcn SOL-047]; 1 ov. female (cl
|
||
<quantity id="4CEE9B1805130517B5E98E07FDEE1FCD" box="[463,568,623,650]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="8" pageNumber="281" unit="mm" value="10.7">10.7 mm</quantity>
|
||
), MNHN-IU-2018-5668,
|
||
<collectingCountry id="F301766D05130517B77E8E07FC141FCE" box="[856,962,623,649]" name="Australia" pageId="8" pageNumber="281">Australia</collectingCountry>
|
||
,
|
||
<collectingRegion id="49D2F81F05130517B7EB8E07FB821FCE" box="[973,1108,623,649]" country="Australia" name="Queensland" pageId="8" pageNumber="281">Queensland</collectingRegion>
|
||
,
|
||
<location id="8EC9602605130517B0468E18FB221FCE" LSID="urn:lsid:plazi:treatment:03BF87EB05130510B4B18CFFFCF31956:8EC9602605130517B0468E18FB221FCE" box="[1120,1268,623,650]" country="Solomon Islands" name="Heron Island" pageId="8" pageNumber="281" stateProvince="Queensland">Heron Island</location>
|
||
, southern side, shallow reef flat, coarse sand, suction (yabby) pump,
|
||
<quantity id="4CEE9B1805130517B6D58EFBFC871FE9" box="[755,849,659,686]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="8" pageNumber="281" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">
|
||
<elevation id="003BD1CE05130517B6D58EFBFC871FE9" box="[755,849,659,686]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="8" pageNumber="281" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 m</elevation>
|
||
</quantity>
|
||
, leg.
|
||
<collectorName id="26E3532B05130517B7AA8EFBFC201FEA" box="[908,1014,659,685]" pageId="8" pageNumber="281">A. Anker</collectorName>
|
||
,
|
||
<date id="FFA8103D05130517B0278EFBFBA91FEA" box="[1025,1151,659,685]" pageId="8" pageNumber="281" value="2016-09-28">
|
||
<collectingDate id="EFECE9D505130517B0278EFBFBA91FEA" box="[1025,1151,659,685]" pageId="8" pageNumber="281" value="2016-09-28">28.09.2016</collectingDate>
|
||
</date>
|
||
[fcn HE-011]
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="9D10FD7405130517B10D8EFBFAA11FEA" box="[1323,1399,659,685]" pageId="8" pageNumber="281" type="male">1 male</specimenCount>
|
||
(cl
|
||
<quantity id="4CEE9B1805130517B4B18EDFFF261F95" box="[151,240,695,722]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="8" pageNumber="281" unit="mm" value="6.6">6.6 mm</quantity>
|
||
),
|
||
<materialsCitation id="3B7E3CA005130517B5248EDFFB651FB2" ID-GBIF-Occurrence="3450196306" collectingDate="2016-09-24" collectorName="A. Anker" country="Australia" elevation="3" location="Heron Island" pageId="8" pageNumber="281" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="paratype">
|
||
<specimenCount id="9D10FD7405130517B5248EDFFEB41F96" box="[258,354,695,721]" pageId="8" pageNumber="281" type="female">1 female</specimenCount>
|
||
(cl
|
||
<quantity id="4CEE9B1805130517B5AA8EDFFE331F95" box="[396,485,695,722]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="8" pageNumber="281" unit="mm" value="8.4">8.4 mm</quantity>
|
||
), MNHN-IU-2018-5670,
|
||
<collectingCountry id="F301766D05130517B7248EDFFCBA1F96" box="[770,876,695,721]" name="Australia" pageId="8" pageNumber="281">Australia</collectingCountry>
|
||
,
|
||
<collectingRegion id="49D2F81F05130517B7508EDFFC2B1F96" box="[886,1021,695,721]" country="Australia" name="Queensland" pageId="8" pageNumber="281">Queensland</collectingRegion>
|
||
,
|
||
<location id="8EC9602605130517B0218ED0FB4C1F96" LSID="urn:lsid:plazi:treatment:03BF87EB05130510B4B18CFFFCF31956:8EC9602605130517B0218ED0FB4C1F96" box="[1031,1178,695,722]" country="Australia" name="Heron Island" pageId="8" pageNumber="281" stateProvince="Queensland">Heron Island</location>
|
||
, southern side, shallow reef flat, coarse sand, suction (yabby) pump,
|
||
<quantity id="4CEE9B1805130517B6AA8EB3FD3C1FB1" box="[652,746,731,758]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="8" pageNumber="281" unit="m" value="0.25" valueMax="0.5" valueMin="0.0">
|
||
<elevation id="003BD1CE05130517B6AA8EB3FD3C1FB1" box="[652,746,731,758]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="8" pageNumber="281" unit="m" value="0.25" valueMax="0.5" valueMin="0.0">0–0.5 m</elevation>
|
||
</quantity>
|
||
, leg.
|
||
<collectorName id="26E3532B05130517B7058EB3FC5D1FB2" box="[803,907,731,757]" pageId="8" pageNumber="281">A. Anker</collectorName>
|
||
,
|
||
<date id="FFA8103D05130517B7B38EB3FBC51FB2" box="[917,1043,731,757]" pageId="8" pageNumber="281" value="2016-09-24">
|
||
<collectingDate id="EFECE9D505130517B7B38EB3FBC51FB2" box="[917,1043,731,757]" pageId="8" pageNumber="281" value="2016-09-24">24.09.2016</collectingDate>
|
||
</date>
|
||
[fcn HE-120]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B7E3CA005130517B09B8EB3FB5F1E7A" ID-GBIF-Occurrence="3450196303" collectingDate="2016-09-25" collectorName="A. Anker" country="Australia" elevation="3" location="Heron Island" pageId="8" pageNumber="281" specimenCount="1" specimenCount-male="1" stateProvince="Queensland" typeStatus="paratype">
|
||
<specimenCount id="9D10FD7405130517B09B8EB3FADE1FB2" box="[1213,1288,731,757]" pageId="8" pageNumber="281" type="male">1 male</specimenCount>
|
||
(cl 16.0 mm, missing both chelipeds), MNHN-IU-2018-5525,
|
||
<collectingCountry id="F301766D05130517B6E68E97FCFC1E5E" box="[704,810,767,793]" name="Australia" pageId="8" pageNumber="281">Australia</collectingCountry>
|
||
,
|
||
<collectingRegion id="49D2F81F05130517B7108E97FC6B1E5E" box="[822,957,767,793]" country="Australia" name="Queensland" pageId="8" pageNumber="281">Queensland</collectingRegion>
|
||
,
|
||
<location id="8EC9602605130517B7EC8F68FB891E5E" LSID="urn:lsid:plazi:treatment:03BF87EB05130510B4B18CFFFCF31956:8EC9602605130517B7EC8F68FB891E5E" box="[970,1119,767,794]" country="Australia" name="Heron Island" pageId="8" pageNumber="281" stateProvince="Queensland">Heron Island</location>
|
||
, southern side, shallow reef flat, coarse sand, suction (yabby) pump,
|
||
<quantity id="4CEE9B1805130517B6788F4BFD6A1E79" box="[606,700,803,830]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="8" pageNumber="281" unit="m" value="0.25" valueMax="0.5" valueMin="0.0">
|
||
<elevation id="003BD1CE05130517B6788F4BFD6A1E79" box="[606,700,803,830]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="8" pageNumber="281" unit="m" value="0.25" valueMax="0.5" valueMin="0.0">0–0.5 m</elevation>
|
||
</quantity>
|
||
, leg.
|
||
<collectorName id="26E3532B05130517B6D18F4BFC891E7A" box="[759,863,803,829]" pageId="8" pageNumber="281">A. Anker</collectorName>
|
||
,
|
||
<date id="FFA8103D05130517B74C8F4BFC3E1E79" box="[874,1000,803,830]" pageId="8" pageNumber="281" value="2016-09-25">
|
||
<collectingDate id="EFECE9D505130517B74C8F4BFC3E1E79" box="[874,1000,803,830]" pageId="8" pageNumber="281" value="2016-09-25">25.09.2016</collectingDate>
|
||
</date>
|
||
[fcn HE-026]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA936FD05130517B4E18F2FFB351EC2" blockId="8.[151,1437,479,1910]" pageId="8" pageNumber="281">
|
||
<emphasis id="B962EAEF05130517B4E18F2FFE621E26" bold="true" box="[199,436,839,865]" pageId="8" pageNumber="281">Additional material</emphasis>
|
||
.
|
||
<materialsCitation id="3B7E3CA005130517B5E28F2FFB091EC2" ID-GBIF-Occurrence="3450196304" collectingDate="2006-12-07" collectorName="I. N. Marin" country="Vietnam" county="Tre Bay" location="Nha Trang Bay" municipality="Tre Island" pageId="8" pageNumber="281" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="9D10FD7405130517B5E28F2FFDC61E26" box="[452,528,839,865]" pageId="8" pageNumber="281" type="male">1 male</specimenCount>
|
||
(cl
|
||
<quantity id="4CEE9B1805130517B6198F2FFD7F1E25" box="[575,681,839,866]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="8" pageNumber="281" unit="mm" value="14.5">14.5 mm</quantity>
|
||
), MNHN-IU-2018-5671,
|
||
<collectingCountry id="F301766D05130517B7ED8F2FFBFB1E26" box="[971,1069,839,865]" name="Vietnam" pageId="8" pageNumber="281">Vietnam</collectingCountry>
|
||
,
|
||
<location id="8EC9602605130517B01F8F2FFB3E1E25" LSID="urn:lsid:plazi:treatment:03BF87EB05130510B4B18CFFFCF31956:8EC9602605130517B01F8F2FFB3E1E25" box="[1081,1256,839,866]" country="Vietnam" county="Tre Bay" municipality="Tre Island" name="Nha Trang Bay" pageId="8" pageNumber="281">Nha Trang Bay</location>
|
||
,
|
||
<collectingMunicipality id="6BCDAC8705130517B0D38F20FABC1E26" box="[1269,1386,839,866]" pageId="8" pageNumber="281">Tre Island</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62C84E7105130517B1508F20FF131EC1" pageId="8" pageNumber="281">Tre Bay</collectingCounty>
|
||
, intertidal area near mangrove, suction (yabby) pump, with goby, leg.
|
||
<collectorName id="26E3532B05130517B7FE8F04FB841EC2" box="[984,1106,875,902]" pageId="8" pageNumber="281">I.N. Marin</collectorName>
|
||
,
|
||
<date id="FFA8103D05130517B07B8F03FB091EC2" box="[1117,1247,875,901]" pageId="8" pageNumber="281" value="2006-12-07">
|
||
<collectingDate id="EFECE9D505130517B07B8F03FB091EC2" box="[1117,1247,875,901]" pageId="8" pageNumber="281" value="2006-12-07">12.07.2006</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C657605130513B4E18FE7FBE51EC6" lastPageId="12" lastPageNumber="285" pageId="8" pageNumber="281" type="description">
|
||
<paragraph id="8BA936FD05130517B4E18FE7FDAE19C5" blockId="8.[151,1437,479,1910]" pageId="8" pageNumber="281">
|
||
<emphasis id="B962EAEF05130517B4E18FE7FE871EEE" bold="true" box="[199,337,911,937]" pageId="8" pageNumber="281">Description</emphasis>
|
||
. Carapace not setose, not pubescent, pitted (
|
||
<figureCitation id="132D2A7805130517B77E8FE7FC011EED" box="[856,983,911,938]" captionStart="FIGURE 4" captionStartId="9.[151,250,1839,1863]" captionTargetBox="[169,1413,191,1801]" captionTargetId="figure-15@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="8" pageNumber="281">Fig. 4A, B</figureCitation>
|
||
). Rostrum well developed, wide, about 1.7 times as long as wide at base, subacute distally, just reaching distal third of first article of antennular peduncle, pointing straight-forward in lateral view; rostral carina well marked, rounded dorsally, gently sloping into welldemarcated adrostral furrows, continuing well beyond base of orbital hoods, gradually flattening, not reaching midlength of carapace (
|
||
<figureCitation id="132D2A7805130517B5578877FE3F197D" box="[369,489,1055,1082]" captionStart="FIGURE 4" captionStartId="9.[151,250,1839,1863]" captionTargetBox="[169,1413,191,1801]" captionTargetId="figure-15@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="8" pageNumber="281">Fig. 4A, B</figureCitation>
|
||
). Orbital hoods swollen, feebly projecting anteriorly in lateral view, unarmed; frontal margin between rostrum and orbital hood shallowly concave (
|
||
<figureCitation id="132D2A7805130517B771882BFC051919" box="[855,979,1091,1118]" captionStart="FIGURE 4" captionStartId="9.[151,250,1839,1863]" captionTargetBox="[169,1413,191,1801]" captionTargetId="figure-15@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="8" pageNumber="281">Fig. 4A, B</figureCitation>
|
||
). Pterygostomial angle rounded; cardiac notch well developed, deep (
|
||
<figureCitation id="132D2A7805130517B5FF880FFDED19C6" box="[473,571,1127,1153]" captionStart="FIGURE 4" captionStartId="9.[151,250,1839,1863]" captionTargetBox="[169,1413,191,1801]" captionTargetId="figure-15@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="8" pageNumber="281">Figs. 4B</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A7805130517B6618800FDBD19C5" box="[583,619,1128,1154]" captionStart="FIGURE 7" captionStartId="13.[151,250,1880,1904]" captionTargetBox="[151,1435,181,1856]" captionTargetId="figure-15@13.[151,1435,181,1856]" captionTargetPageId="13" captionText="FIGURE 7. Alpheus sciolii sp. nov., holotype male (cl 8.6 mm) from New Georgia, Solomon Islands (MNHN-IU-2018-5667) [A, B]; paratype female (cl 10.7 mm) from Heron Island, Queensland, Australia (MNHN-IU-2018-5668) [C]: A, C, living shrimps, dorsal; B, same, lateral. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.5881119" httpUri="https://zenodo.org/record/5881119/files/figure.png" pageId="8" pageNumber="281">7B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA936FD05130517B4E188E3FAD0183A" blockId="8.[151,1437,479,1910]" pageId="8" pageNumber="281">
|
||
Telson very broad, subrectangular, gently tapering distally, about 1.8 times as long as maximal width, with lateral margins not noticeably constricted; dorsal surface with two pairs of stout cuspidate setae both inserted far from lateral margin, first pair at about 0.5 telson length, second pair at about 0.7 of telson length; posterior margin broadly rounded, with row of short spiniform setae above plumose setae (less conspicuous or with slender spiniform setae in some specimens); posterolateral angles each with one pair of spiniform setae (typically, sometimes with only one spiniform seta), mesial ones stouter and two to three times as long as lateral ones [posterolateral spiniform setae somewhat aberrant in the illustrated
|
||
<typeStatus id="54AD885F05130517B648890CFD181839" box="[622,718,1380,1406]" pageId="8" pageNumber="281" type="paratype">paratype</typeStatus>
|
||
(
|
||
<figureCitation id="132D2A7805130517B6F8890BFCE0183A" box="[734,822,1379,1405]" captionStart="FIGURE 4" captionStartId="9.[151,250,1839,1863]" captionTargetBox="[169,1413,191,1801]" captionTargetId="figure-15@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="8" pageNumber="281">Fig. 4C</figureCitation>
|
||
), see
|
||
<figureCitation id="132D2A7805130517B753890BFC1F183A" box="[885,969,1379,1405]" captionStart="FIGURE 6" captionStartId="11.[151,250,1665,1689]" captionTargetBox="[190,1400,792,1619]" captionTargetId="figure-271@11.[151,1436,777,1642]" captionTargetPageId="11" captionText="FIGURE 6. Alpheus sciolii sp. nov., non-type male (cl 14.5 mm) from Nha Trang Bay, Vietnam (MNHN-IU-2018-5671):A, left (major) cheliped, carpus and chela, lateral; B, right (minor) cheliped, chela, lateral; C, telson, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881113" httpUri="https://zenodo.org/record/5881113/files/figure.png" pageId="8" pageNumber="281">Fig. 6C</figureCitation>
|
||
for more typical condition].
|
||
</paragraph>
|
||
<paragraph id="8BA936FD05130517B4E189E0FA5E1BD9" blockId="8.[151,1437,479,1910]" pageId="8" pageNumber="281">
|
||
Eyes well developed, with large, normally pigmented corneas (
|
||
<figureCitation id="132D2A7805130517B7A689EFFBD418E5" box="[896,1026,1415,1442]" captionStart="FIGURE 1" captionStartId="3.[151,250,1814,1838]" captionTargetBox="[153,1420,187,1767]" captionTargetId="figure-15@3.[151,1436,181,1790]" captionTargetPageId="3" captionText="FIGURE 1. Alpheus thompsoni sp. nov., holotype, female (cl 13.3 mm) from Madang, Papua New Guinea (MNHN-2013- 16813): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal portion of propodus and dactylus, ventrolateral, slender setae on propodus omitted; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventromesial; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881097" httpUri="https://zenodo.org/record/5881097/files/figure.png" pageId="8" pageNumber="281">Figs. 1A, B</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A7805130517B02B89E0FBCB18E5" box="[1037,1053,1416,1442]" captionStart="FIGURE 7" captionStartId="13.[151,250,1880,1904]" captionTargetBox="[151,1435,181,1856]" captionTargetId="figure-15@13.[151,1435,181,1856]" captionTargetPageId="13" captionText="FIGURE 7. Alpheus sciolii sp. nov., holotype male (cl 8.6 mm) from New Georgia, Solomon Islands (MNHN-IU-2018-5667) [A, B]; paratype female (cl 10.7 mm) from Heron Island, Queensland, Australia (MNHN-IU-2018-5668) [C]: A, C, living shrimps, dorsal; B, same, lateral. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.5881119" httpUri="https://zenodo.org/record/5881119/files/figure.png" pageId="8" pageNumber="281">7</figureCitation>
|
||
). Antennular peduncle moderately elongate and stout; stylocerite feebly swollen laterally, distal part slenderer, ending in sharp point, latter not reaching distal margin of first article; ventromesial carina with large subtriangular tooth; second article about 2.7 times as long as wide; lateral antennular flagellum with secondary ramus fused to main ramus over most of its length, with numerous groups of aesthetascs distally, starting from about 14th subdivision (
|
||
<figureCitation id="132D2A7805130517B0698A7FFB261B75" box="[1103,1264,1559,1586]" captionStart="FIGURE 4" captionStartId="9.[151,250,1839,1863]" captionTargetBox="[169,1413,191,1801]" captionTargetId="figure-15@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="8" pageNumber="281">Fig. 4A, B, D</figureCitation>
|
||
). Antenna with basicerite fairly stout, its distoventral margin armed with large, anteriorly projecting, sharp tooth; scaphocerite with moderately broad blade distinctly exceeded by sharp distolateral tooth, lateral margin shallowly concave; scaphocerite reaching distinctly beyond end of antennular peduncle and slightly beyond carpocerite (
|
||
<figureCitation id="132D2A7805130517B1278AEBFAAC1BD9" box="[1281,1402,1667,1694]" captionStart="FIGURE 4" captionStartId="9.[151,250,1839,1863]" captionTargetBox="[169,1413,191,1801]" captionTargetId="figure-15@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="8" pageNumber="281">Fig. 4A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA936FD05130517B4E18ACFFCE51A32" blockId="8.[151,1437,479,1910]" pageId="8" pageNumber="281">
|
||
Mouthparts not dissected, generally typical for genus in external observation. Third maxilliped relatively stout; coxa with bluntly projecting lateral plate; antepenultimate article flattened ventrolaterally, with distinct ridge running parallel to dorsal margin on lateral surface and rugose mesial margin, about 3.9 times as long as high; penultimate article very short, cup-shaped, bulging ventrally, its distoventral margin with few moderately long setae, its dorsal margin with row of elongate setae; ultimate article densely furnished with rows of very long, stiff setae, dorsal and apical setae longest; arthrobranch well developed (
|
||
<figureCitation id="132D2A7805130517B6E98B33FCF11A32" box="[719,807,1883,1909]" captionStart="FIGURE 4" captionStartId="9.[151,250,1839,1863]" captionTargetBox="[169,1413,191,1801]" captionTargetId="figure-15@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="8" pageNumber="281">Fig. 4E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF69667505120516B4B18B47FD4D1A9F" ID-DOI="http://doi.org/10.5281/zenodo.5881105" ID-Zenodo-Dep="5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="9" pageNumber="282" startId="9.[151,250,1839,1863]" targetBox="[169,1413,191,1801]" targetPageId="9">
|
||
<paragraph id="8BA936FD05120516B4B18B47FD4D1A9F" blockId="9.[151,1437,1839,2008]" pageId="9" pageNumber="282">
|
||
<emphasis id="B962EAEF05120516B4B18B47FED91A0F" bold="true" box="[151,271,1839,1864]" pageId="9" pageNumber="282">FIGURE 4</emphasis>
|
||
.
|
||
<taxonomicName id="4C164D7E05120516B53B8B58FE631A0F" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[285,437,1840,1864]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="282" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
|
||
<emphasis id="B962EAEF05120516B53B8B58FE631A0F" box="[285,437,1840,1864]" italics="true" pageId="9" pageNumber="282">Alpheus sciolii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05120516B59B8B58FDD91A0F" bold="true" box="[445,527,1840,1864]" pageId="9" pageNumber="282">
|
||
<taxonomicNameLabel id="A251579405120516B59B8B58FDD91A0F" box="[445,527,1840,1864]" pageId="9" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF69667505110515B4B18AC5FF081A75" ID-DOI="http://doi.org/10.5281/zenodo.5881107" ID-Zenodo-Dep="5881107" httpUri="https://zenodo.org/record/5881107/files/figure.png" pageId="10" pageNumber="283" startId="10.[151,250,1709,1733]" targetBox="[165,1420,203,1676]" targetPageId="10">
|
||
<paragraph id="8BA936FD05110515B4B18AC5FF081A75" blockId="10.[151,1437,1709,1842]" pageId="10" pageNumber="283">
|
||
<emphasis id="B962EAEF05110515B4B18AC5FEDA1B81" bold="true" box="[151,268,1709,1734]" pageId="10" pageNumber="283">FIGURE 5</emphasis>
|
||
.
|
||
<taxonomicName id="4C164D7E05110515B53E8AC6FE7B1B81" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[280,429,1710,1734]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="283" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
|
||
<emphasis id="B962EAEF05110515B53E8AC6FE7B1B81" box="[280,429,1710,1734]" italics="true" pageId="10" pageNumber="283">Alpheus sciolii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05110515B5958AC6FDD51B81" bold="true" box="[435,515,1710,1734]" pageId="10" pageNumber="283">
|
||
<taxonomicNameLabel id="A251579405110515B5958AC6FDD51B81" box="[435,515,1710,1734]" pageId="10" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, male (cl 9.3 mm) from New Georgia, Solomon Islands (MNHN-IU-2018-5669): A, major (right) cheliped, lateral; B, same, chela, mesial; C, same, detail of ventromesial margin of merus, mesial, slender setae omitted; D, same, dactylus and pollex, lateral, setae omitted; E, minor (left) cheliped, carpus and chela, mesial; F, same, chela, lateral.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA936FD05110514B4E18B08FA4A1CCE" blockId="10.[151,1436,1888,2022]" lastBlockId="11.[151,1437,151,753]" lastPageId="11" lastPageNumber="284" pageId="10" pageNumber="283">
|
||
Male major cheliped robust; ischium short, stout, smooth; merus very stout, trigonal in cross-section, about twice as long as maximal width, distodorsal margin ending bluntly, ventromesial margin somewhat rugose, with more or less pronounced, blunt, distal tooth and row of five or six spiniform setae; carpus short, distally widening, cup-shaped; chela not particularly elongate, fringed with long setae along dorsal and ventral surfaces; palm strongly compressed, stout, subrectangular in cross-section, length / maximal height ratio 1.8–2.1 (lower in larger males); surfaces relatively smooth, without granules; dorsal margin with well-marked transverse groove subdistally, without marked longitudinal ridges; ventral margin almost straight; lateral surface with shallow depression distally, near base of pollex; fingers subequal in length, about half-length of palm, not twisted or significantly deviating from chela axis; dactylus distally rounded, plunger somewhat reduced, poorly demarcated from anterior cutting edge, latter slightly bulging; adhesive disks very small (
|
||
<figureCitation id="132D2A7805100514B70E8D23FC7E1C22" box="[808,936,330,357]" captionStart="FIGURE 5" captionStartId="10.[151,250,1709,1733]" captionTargetBox="[165,1420,203,1676]" captionTargetId="figure-17@10.[151,1436,181,1686]" captionTargetPageId="10" captionText="FIGURE 5. Alpheus sciolii sp. nov., paratype, male (cl 9.3 mm) from New Georgia, Solomon Islands (MNHN-IU-2018-5669): A, major (right) cheliped, lateral; B, same, chela, mesial; C, same, detail of ventromesial margin of merus, mesial, slender setae omitted; D, same, dactylus and pollex, lateral, setae omitted; E, minor (left) cheliped, carpus and chela, mesial; F, same, chela, lateral." figureDoi="http://doi.org/10.5281/zenodo.5881107" httpUri="https://zenodo.org/record/5881107/files/figure.png" pageId="11" pageNumber="284">Fig. 5A–D</figureCitation>
|
||
; see also
|
||
<figureCitation id="132D2A7805100514B0398D23FBAA1C21" box="[1055,1148,331,358]" captionStart="FIGURE 6" captionStartId="11.[151,250,1665,1689]" captionTargetBox="[190,1400,792,1619]" captionTargetId="figure-271@11.[151,1436,777,1642]" captionTargetPageId="11" captionText="FIGURE 6. Alpheus sciolii sp. nov., non-type male (cl 14.5 mm) from Nha Trang Bay, Vietnam (MNHN-IU-2018-5671):A, left (major) cheliped, carpus and chela, lateral; B, right (minor) cheliped, chela, lateral; C, telson, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881113" httpUri="https://zenodo.org/record/5881113/files/figure.png" pageId="11" pageNumber="284">Fig. 6A</figureCitation>
|
||
). Female major cheliped generally similar to male major cheliped, smaller, weaker; chela with somewhat different proportions (see
|
||
<figureCitation id="132D2A7805100514B11E8D07FA591CCE" box="[1336,1423,367,393]" captionStart="FIGURE 7" captionStartId="13.[151,250,1880,1904]" captionTargetBox="[151,1435,181,1856]" captionTargetId="figure-15@13.[151,1435,181,1856]" captionTargetPageId="13" captionText="FIGURE 7. Alpheus sciolii sp. nov., holotype male (cl 8.6 mm) from New Georgia, Solomon Islands (MNHN-IU-2018-5667) [A, B]; paratype female (cl 10.7 mm) from Heron Island, Queensland, Australia (MNHN-IU-2018-5668) [C]: A, C, living shrimps, dorsal; B, same, lateral. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.5881119" httpUri="https://zenodo.org/record/5881119/files/figure.png" pageId="11" pageNumber="284">Fig. 7C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA936FD05100514B4E18DFBFE881FB6" blockId="11.[151,1437,151,753]" pageId="11" pageNumber="284">
|
||
Male minor cheliped with ischium short, smooth; merus similar to that of major cheliped in size and stoutness, trigonal in cross-section, distodorsal angle blunt, ventromesial margin slightly rugose distally, usually without distal tooth, armed with spiniform setae; carpus distinctly longer than that of major cheliped, distally widening, cupshaped; chela not elongate, not swollen, fringed with long setae along dorsal and ventral surfaces; palm compressed, subrectangular in cross-section, length / maximal height ratio 1.3–1.5 (lower in larger males); surfaces smooth, not granulated; dorsal margin without groove; ventral margin broadly convex; fingers subequal in length, about 1.3 times length of palm, not twisted, slightly gaping when closed, with balaeniceps crests lined with setae (better developed in larger males); adhesive disks greatly reduced (
|
||
<figureCitation id="132D2A7805100514B7648EE7FC6C1FEE" box="[834,954,654,681]" captionStart="FIGURE 5" captionStartId="10.[151,250,1709,1733]" captionTargetBox="[165,1420,203,1676]" captionTargetId="figure-17@10.[151,1436,181,1686]" captionTargetPageId="10" captionText="FIGURE 5. Alpheus sciolii sp. nov., paratype, male (cl 9.3 mm) from New Georgia, Solomon Islands (MNHN-IU-2018-5669): A, major (right) cheliped, lateral; B, same, chela, mesial; C, same, detail of ventromesial margin of merus, mesial, slender setae omitted; D, same, dactylus and pollex, lateral, setae omitted; E, minor (left) cheliped, carpus and chela, mesial; F, same, chela, lateral." figureDoi="http://doi.org/10.5281/zenodo.5881107" httpUri="https://zenodo.org/record/5881107/files/figure.png" pageId="11" pageNumber="284">Fig. 5E, F</figureCitation>
|
||
; see also
|
||
<figureCitation id="132D2A7805100514B00E8EE7FB541FED" box="[1064,1154,655,682]" captionStart="FIGURE 6" captionStartId="11.[151,250,1665,1689]" captionTargetBox="[190,1400,792,1619]" captionTargetId="figure-271@11.[151,1436,777,1642]" captionTargetPageId="11" captionText="FIGURE 6. Alpheus sciolii sp. nov., non-type male (cl 14.5 mm) from Nha Trang Bay, Vietnam (MNHN-IU-2018-5671):A, left (major) cheliped, carpus and chela, lateral; B, right (minor) cheliped, chela, lateral; C, telson, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881113" httpUri="https://zenodo.org/record/5881113/files/figure.png" pageId="11" pageNumber="284">Fig. 6B</figureCitation>
|
||
). Female minor cheliped generally similar, but with weaker chela, latter with balaeniceps ridges and setae on fingers poorly developed or lacking (
|
||
<figureCitation id="132D2A7805100514B4DF8EBFFE871FB6" box="[249,337,727,753]" captionStart="FIGURE 7" captionStartId="13.[151,250,1880,1904]" captionTargetBox="[151,1435,181,1856]" captionTargetId="figure-15@13.[151,1435,181,1856]" captionTargetPageId="13" captionText="FIGURE 7. Alpheus sciolii sp. nov., holotype male (cl 8.6 mm) from New Georgia, Solomon Islands (MNHN-IU-2018-5667) [A, B]; paratype female (cl 10.7 mm) from Heron Island, Queensland, Australia (MNHN-IU-2018-5668) [C]: A, C, living shrimps, dorsal; B, same, lateral. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.5881119" httpUri="https://zenodo.org/record/5881119/files/figure.png" pageId="11" pageNumber="284">Fig. 7C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF69667505100514B4B18AE9FA541BFA" ID-DOI="http://doi.org/10.5281/zenodo.5881113" ID-Zenodo-Dep="5881113" httpUri="https://zenodo.org/record/5881113/files/figure.png" pageId="11" pageNumber="284" startId="11.[151,250,1665,1689]" targetBox="[190,1400,792,1619]" targetPageId="11">
|
||
<paragraph id="8BA936FD05100514B4B18AE9FA541BFA" blockId="11.[151,1436,1665,1725]" pageId="11" pageNumber="284">
|
||
<emphasis id="B962EAEF05100514B4B18AE9FEDA1BDE" bold="true" box="[151,268,1665,1689]" pageId="11" pageNumber="284">FIGURE 6</emphasis>
|
||
.
|
||
<taxonomicName id="4C164D7E05100514B5318AE9FE7A1BDE" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[279,428,1665,1689]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="284" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
|
||
<emphasis id="B962EAEF05100514B5318AE9FE7A1BDE" box="[279,428,1665,1689]" italics="true" pageId="11" pageNumber="284">Alpheus sciolii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05100514B5978AE9FDD61BDE" bold="true" box="[433,512,1665,1689]" pageId="11" pageNumber="284">
|
||
<taxonomicNameLabel id="A251579405100514B5978AE9FDD61BDE" box="[433,512,1665,1689]" pageId="11" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, non-type male (cl 14.5 mm) from Nha Trang Bay, Vietnam (MNHN-IU-2018-5671): A, left (major) cheliped, carpus and chela, lateral; B, right (minor) cheliped, chela, lateral; C, telson, dorsal, plumose setae omitted.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA936FD05100513B4E18A8DFCAA1C06" blockId="11.[151,1437,1765,2043]" lastBlockId="12.[151,1437,151,1870]" lastPageId="12" lastPageNumber="285" pageId="11" pageNumber="284">
|
||
Second pereiopod slender; ischium and merus subequal in length; carpus with five subarticles, first noticeably longer than second, ratio of carpal subarticles approximately equal to 3.0/2.5/1.0/1.0/1.7; chela longer than distalmost carpal subarticle (
|
||
<figureCitation id="132D2A7805100514B5878B45FE2E1A00" box="[417,504,1837,1863]" captionStart="FIGURE 4" captionStartId="9.[151,250,1839,1863]" captionTargetBox="[169,1413,191,1801]" captionTargetId="figure-15@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="11" pageNumber="284">Fig. 4F</figureCitation>
|
||
). Third pereiopod fairly robust; ischium with stout spiniform seta on ventrolateral surface; merus about 4.1 times as long as maximal width, ventromesial margin unarmed distally; carpus about 0.6 length of merus, much more slender than merus, unarmed; propodus subequal to carpus, setose, with at least 10 spiniform setae along ventral margin, including distal pair adjacent to dactylar base; dactylus slightly more than halflength of propodus, gradually curving distally, subspatulate, moderately broadened, flattened ventrally, with distinct longitudinal keel separating ventral and dorsal surfaces, with scarce setae (
|
||
<figureCitation id="132D2A7805100514B7D28B89FB871ABC" box="[1012,1105,2017,2043]" captionStart="FIGURE 4" captionStartId="9.[151,250,1839,1863]" captionTargetBox="[169,1413,191,1801]" captionTargetId="figure-15@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="11" pageNumber="284">Fig. 4G</figureCitation>
|
||
). Fourth pereiopod generally similar to third pereiopod, more slender. Fifth pereiopod much more slender than third and fourth pereiopods; ischium with small spiniform seta on ventrolateral surface; merus about 5.7 times as long as wide; carpus slightly more slender than merus, about 0.9 length of merus; propodus subequal to carpus, with rows of serrulate setae forming cleaning brush on distal third of ventrolateral surface, and at least eight spiniform setae along ventromesial margin, including distal pair adjacent to dactylar base (
|
||
<figureCitation id="132D2A7805170513B6D88D4FFCB81C06" box="[766,878,295,321]" captionStart="FIGURE 4" captionStartId="9.[151,250,1839,1863]" captionTargetBox="[169,1413,191,1801]" captionTargetId="figure-15@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="12" pageNumber="285">Fig. 4H, I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA936FD05170513B4E18D23FBD51CB2" blockId="12.[151,1437,151,1870]" pageId="12" pageNumber="285">
|
||
Second pleopod with appendices masculina and interna equal in length, former with long stiff setae (
|
||
<figureCitation id="132D2A7805170513B1188D23FA591C22" box="[1342,1423,331,357]" captionStart="FIGURE 4" captionStartId="9.[151,250,1839,1863]" captionTargetBox="[169,1413,191,1801]" captionTargetId="figure-15@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="12" pageNumber="285">Fig. 4J</figureCitation>
|
||
). Uropod with mesial and lateral lobes of protopod each ending in sharp (lateral) or blunt (mesial) distal tooth; exopod broad, broadly rounded distally, with short triangular distolateral tooth; diaeresis straight for most part, except for rounded lobe adjacent to slender spiniform seta, latter not reaching level of distal margin of exopod; endopod much narrower than exopod, with row of spiniform setae on distal margin (
|
||
<figureCitation id="132D2A7805170513B7BA8DB3FC211CB2" box="[924,1015,475,501]" captionStart="FIGURE 4" captionStartId="9.[151,250,1839,1863]" captionTargetBox="[169,1413,191,1801]" captionTargetId="figure-15@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="12" pageNumber="285">Fig. 4K</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA936FD05170513B4E18D96FBE51EC6" blockId="12.[151,1437,151,1870]" pageId="12" pageNumber="285">
|
||
<emphasis id="B962EAEF05170513B4E18D96FEA91F5E" bold="true" box="[199,383,510,537]" pageId="12" pageNumber="285">Colour pattern</emphasis>
|
||
. General background colour white with pale yellowish tinge; carapace with several (three to four) broad bands, running transversely dorsally and more obliquely laterally, some broadly croissant-shaped; carapace flanks with very large brown to almost black spot (quickly fading in captured individuals); pleon with narrower, transversely-obliquely running, brown bands, usually two per pleonite; posterior end of carapace and first pleonite with conspicuous, white, dorsal “saddle”; fourth and fifth pleonites with transverse, whitish, dorsal patches; chelipeds whitish or greyish, with irregular, brown, green-brown or green-bluish patches; distal half of fingers hyaline-white; antennules and antennae whitish with brownish spots and streaks; antennular flagella pale yellow proximally, more intense yellow distally; antennal flagella pale yellow proximally, uncoloured distally; second pereiopods and walking legs greyish-blue with yellow markings around articulations; pleopods brownish yellow; uropods whitish with some brown-red bands or spots, mainly on endopod; telson yellow whitish proximally, with brown-red pattern distally (
|
||
<figureCitation id="132D2A7805170513B5EF8F0FFDCF1EC6" box="[457,537,871,897]" captionStart="FIGURE 7" captionStartId="13.[151,250,1880,1904]" captionTargetBox="[151,1435,181,1856]" captionTargetId="figure-15@13.[151,1435,181,1856]" captionTargetPageId="13" captionText="FIGURE 7. Alpheus sciolii sp. nov., holotype male (cl 8.6 mm) from New Georgia, Solomon Islands (MNHN-IU-2018-5667) [A, B]; paratype female (cl 10.7 mm) from Heron Island, Queensland, Australia (MNHN-IU-2018-5668) [C]: A, C, living shrimps, dorsal; B, same, lateral. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.5881119" httpUri="https://zenodo.org/record/5881119/files/figure.png" pageId="12" pageNumber="285">Figs. 7</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A7805170513B6038F0FFD9E1EC6" box="[549,584,871,897]" captionStart="FIGURE 8" captionStartId="14.[151,250,1545,1569]" captionTargetBox="[158,1430,181,1521]" captionTargetId="figure-17@14.[158,1430,181,1521]" captionTargetPageId="14" captionText="FIGURE 8. Some goby-associated species of Alpheus mentioned in the text, photographed in situ: A, Alpheus thompsoni sp. nov. associated with Amblyeleotris wheeleri, Anilao, Philippines; B, same, associated with Amblyeleotris guttata, Sulawesi, Indonesia; C, Alpheus sciolii sp. nov. associated with Cryptocentrus inexplicatus, Papua New Guinea; D, Alpheus sp. aff. A. sciolii sp. nov. associated with Amblyeleotris steinitzi, Bali, Indonesia; E, Alpheus sp. cf. mannarensis Purushothaman, Abhilash, Kumar & Lal, 2021, Papua New Guinea; F, Alpheus bellulus Miya & Miyake, 1969, Sulawesi, Indonesia. Photographs courtesy of Andrey Ryanskiy (A–C, E, F) and Yury Ivanov (D)." figureDoi="http://doi.org/10.5281/zenodo.5881121" httpUri="https://zenodo.org/record/5881121/files/figure.png" pageId="12" pageNumber="285">8C</figureCitation>
|
||
; see also
|
||
<bibRefCitation id="EF874B0C05170513B6978F0FFC1C1EC6" author="Anker, A. & De Grave, S." box="[689,970,871,897]" pageId="12" pageNumber="285" pagination="145 - 150" refId="ref10292" refString="Anker, A. & De Grave, S. (2019) Further records of burrow-associated palaemonid shrimps (Decapoda: Palaemonidae). Zootaxa, 4612 (1), 145 - 150. https: // doi. org / 10.11646 / zootaxa. 4612.1.13" type="journal article" year="2019">Anker & De Grave 2019</bibRefCitation>
|
||
: fig. 4C).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C657605170513B4E18FE3FC711E8E" pageId="12" pageNumber="285" type="etymology">
|
||
<paragraph id="8BA936FD05170513B4E18FE3FC711E8E" blockId="12.[151,1437,151,1870]" pageId="12" pageNumber="285">
|
||
<emphasis id="B962EAEF05170513B4E18FE3FE9E1EE2" bold="true" box="[199,328,907,933]" pageId="12" pageNumber="285">Etymology</emphasis>
|
||
. This new species is named after Justin Scioli, PhD cand., the author’s great friend and collaborator in projects involving taxonomy and phylogenetics of alpheid shrimps.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C657605170513B4E18FBAFACD1972" pageId="12" pageNumber="285" type="distribution">
|
||
<paragraph id="8BA936FD05170513B4E18FBAFACD1972" blockId="12.[151,1437,151,1870]" pageId="12" pageNumber="285">
|
||
<emphasis id="B962EAEF05170513B4E18FBAFE8F1EAB" bold="true" box="[199,345,978,1004]" pageId="12" pageNumber="285">Distribution</emphasis>
|
||
. Western Pacific: confirmed records from
|
||
<collectingCountry id="F301766D05170513B7638FBBFBD01EAA" box="[837,1030,979,1005]" name="Solomon Islands" pageId="12" pageNumber="285">Solomon Islands</collectingCountry>
|
||
,
|
||
<collectingCountry id="F301766D05170513B0348FBBFBAF1EAA" box="[1042,1145,979,1005]" name="Australia" pageId="12" pageNumber="285">Australia</collectingCountry>
|
||
(Great Barrier Reef) and
|
||
<collectingCountry id="F301766D05170513B4B18F9FFF201956" box="[151,246,1015,1041]" name="Vietnam" pageId="12" pageNumber="285">Vietnam</collectingCountry>
|
||
(present study); photographic records from the
|
||
<collectingCountry id="F301766D05170513B7228F9FFC541956" box="[772,898,1015,1041]" name="Philippines" pageId="12" pageNumber="285">Philippines</collectingCountry>
|
||
(
|
||
<collectingRegion id="49D2F81F05170513B7B68F9FFC231955" box="[912,1013,1015,1042]" country="Philippines" name="Palawan" pageId="12" pageNumber="285">Palawan</collectingRegion>
|
||
),
|
||
<collectingCountry id="F301766D05170513B0208F9FFBA51956" box="[1030,1139,1015,1041]" name="Indonesia" pageId="12" pageNumber="285">Indonesia</collectingCountry>
|
||
(
|
||
<collectingRegion id="49D2F81F05170513B0A78F9FFB611956" box="[1153,1207,1015,1041]" country="Indonesia" name="Jawa Timur" pageId="12" pageNumber="285">Java</collectingRegion>
|
||
,
|
||
<collectingCountry id="F301766D05170513B0E68F9FFADF1956" box="[1216,1289,1015,1041]" name="Indonesia" pageId="12" pageNumber="285">Flores</collectingCountry>
|
||
,
|
||
<collectingRegion id="49D2F81F05170513B1358F9FFA931955" box="[1299,1349,1015,1042]" country="Indonesia" name="Bali" pageId="12" pageNumber="285">Bali</collectingRegion>
|
||
),
|
||
<collectingCountry id="F301766D05170513B17E8F9FFEF21972" name="Papua New Guinea" pageId="12" pageNumber="285">Papua New Guinea</collectingCountry>
|
||
and
|
||
<collectingCountry id="F301766D05170513B57C8873FE4E1972" box="[346,408,1051,1077]" name="Palau" pageId="12" pageNumber="285">Palau</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF874B0C05170513B5818873FD981972" author="Debelius, H." box="[423,590,1051,1077]" pageId="12" pageNumber="285" refId="ref10810" refString="Debelius, H. (2001) Crustacea guide of the world. IKAN, Frankfurt, 321 pp." type="book" year="2001">Debelius 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B0C05170513B67C8873FCE71972" author="Randall, J. E. & Shao, K. T. & Chen, J. P." box="[602,817,1051,1077]" pageId="12" pageNumber="285" pagination="26 - 34" refId="ref11787" refString="Randall, J. E., Shao, K. T. & Chen, J. P. (2007) Two new shrimp gobies of the genus Ctenogobiops (Perciformes: Gobiidae), from the Western Pacific. Zoological Studies, 46, 26 - 34." type="journal article" year="2007">Randall et al. 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B0C05170513B71B8873FB9B1972" author="Kuiter, R. H. & Debelius, H." box="[829,1101,1051,1077]" pageId="12" pageNumber="285" refId="ref11307" refString="Kuiter, R. H. & Debelius, H. (2009) World atlas of marine fauna. IKAN, Frankfurt, 725 pp." type="book" year="2009">Kuiter & Debelius 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B0C05170513B07F8873FAD81972" author="Hurt, C. & Hultgren, K. M. & Anker, A. & Lemmon, A. R. & Moriarty Lemmon, E. & Bracken-Grissom, H." box="[1113,1294,1051,1077]" pageId="12" pageNumber="285" pagination="1 - 11" refId="ref11003" refString="Hurt, C., Hultgren, K. M., Anker, A., Lemmon, A. R., Moriarty Lemmon, E. & Bracken-Grissom, H. (2021) First worldwide molecular phylogeny of the morphologically and ecologically hyperdiversified snapping shrimp genus Alpheus (Malacostraca: Decapoda). Molecular Phylogenetics and Evolution, 158 (197080), 1 - 11. https: // doi. org / 10.1016 / j. ympev. 2021.107080" type="journal article" year="2021">
|
||
Hurt
|
||
<emphasis id="B962EAEF05170513B0B58873FB1A1972" box="[1171,1228,1051,1077]" italics="true" pageId="12" pageNumber="285">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C657605170513B4E18857FEFD19AE" pageId="12" pageNumber="285" type="biology_ecology">
|
||
<paragraph id="8BA936FD05170513B4E18857FEFD19AE" blockId="12.[151,1437,151,1870]" pageId="12" pageNumber="285">
|
||
<emphasis id="B962EAEF05170513B4E18857FEF0191E" bold="true" box="[199,294,1087,1113]" pageId="12" pageNumber="285">Ecology</emphasis>
|
||
. Shallow coral reefs and adjacent reef flats and lagoons; associated with gobies, typically
|
||
<taxonomicName id="4C164D7E05170513B0DA8857FE94193A" authority="(Herre)" baseAuthorityName="Herre" baseAuthorityYear="1936" class="Actinopterygii" family="Gobiidae" genus="Cryptocentrus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="285" phylum="Chordata" rank="species" species="cinctus">
|
||
<emphasis id="B962EAEF05170513B0DA8857FF30193B" italics="true" pageId="12" pageNumber="285">Cryptocentrus cinctus</emphasis>
|
||
(Herre)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C164D7E05170513B56D880BFD951939" authority="(Playfair)" baseAuthorityName="Playfair" baseAuthorityYear="1867" box="[331,579,1123,1150]" class="Actinopterygii" family="Gobiidae" genus="Cryptocentrus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="285" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="B962EAEF05170513B56D880BFE19193A" box="[331,463,1123,1149]" italics="true" pageId="12" pageNumber="285">C. fasciatus</emphasis>
|
||
(Playfair)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C164D7E05170513B66A880BFCB2193A" authority="Bleeker" authorityName="Bleeker" authorityYear="1876" box="[588,868,1123,1149]" class="Actinopterygii" family="Gobiidae" genus="Cryptocentrus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="285" phylum="Chordata" rank="species" species="leptocephalus">
|
||
<emphasis id="B962EAEF05170513B66A880BFCDE193A" box="[588,776,1123,1149]" italics="true" pageId="12" pageNumber="285">C. leptocephalus</emphasis>
|
||
Bleeker
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C164D7E05170513B7BE880BFB4C193A" authority="(Herre)" baseAuthorityName="Herre" baseAuthorityYear="1934" box="[920,1178,1123,1149]" class="Actinopterygii" family="Gobiidae" genus="Cryptocentrus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="285" phylum="Chordata" rank="species" species="inexplicatus">
|
||
<emphasis id="B962EAEF05170513B7BE880BFBE9193A" box="[920,1087,1123,1149]" italics="true" pageId="12" pageNumber="285">C. inexplicatus</emphasis>
|
||
(Herre)
|
||
</taxonomicName>
|
||
, but also
|
||
<taxonomicName id="4C164D7E05170513B127880BFECB19E7" class="Actinopterygii" family="Gobiidae" genus="Ctenogobiops" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="285" phylum="Chordata" rank="species" species="pomastictus">
|
||
<emphasis id="B962EAEF05170513B127880BFECB19E7" italics="true" pageId="12" pageNumber="285">Ctenogobiops pomastictus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="132D2A7805170513B50A88EFFE5319E6" box="[300,389,1159,1185]" captionStart="FIGURE 8" captionStartId="14.[151,250,1545,1569]" captionTargetBox="[158,1430,181,1521]" captionTargetId="figure-17@14.[158,1430,181,1521]" captionTargetPageId="14" captionText="FIGURE 8. Some goby-associated species of Alpheus mentioned in the text, photographed in situ: A, Alpheus thompsoni sp. nov. associated with Amblyeleotris wheeleri, Anilao, Philippines; B, same, associated with Amblyeleotris guttata, Sulawesi, Indonesia; C, Alpheus sciolii sp. nov. associated with Cryptocentrus inexplicatus, Papua New Guinea; D, Alpheus sp. aff. A. sciolii sp. nov. associated with Amblyeleotris steinitzi, Bali, Indonesia; E, Alpheus sp. cf. mannarensis Purushothaman, Abhilash, Kumar & Lal, 2021, Papua New Guinea; F, Alpheus bellulus Miya & Miyake, 1969, Sulawesi, Indonesia. Photographs courtesy of Andrey Ryanskiy (A–C, E, F) and Yury Ivanov (D)." figureDoi="http://doi.org/10.5281/zenodo.5881121" httpUri="https://zenodo.org/record/5881121/files/figure.png" pageId="12" pageNumber="285">Fig. 8C</figureCitation>
|
||
; see also
|
||
<bibRefCitation id="EF874B0C05170513B5C988EFFD4319E6" author="Debelius, H." box="[495,661,1159,1185]" pageId="12" pageNumber="285" refId="ref10810" refString="Debelius, H. (2001) Crustacea guide of the world. IKAN, Frankfurt, 321 pp." type="book" year="2001">Debelius 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B0C05170513B68788EFFCAF19E6" author="Randall, J. E. & Shao, K. T. & Chen, J. P." box="[673,889,1159,1185]" pageId="12" pageNumber="285" pagination="26 - 34" refId="ref11787" refString="Randall, J. E., Shao, K. T. & Chen, J. P. (2007) Two new shrimp gobies of the genus Ctenogobiops (Perciformes: Gobiidae), from the Western Pacific. Zoological Studies, 46, 26 - 34." type="journal article" year="2007">
|
||
Randall
|
||
<emphasis id="B962EAEF05170513B6D988EFFCEE19E6" box="[767,824,1159,1185]" italics="true" pageId="12" pageNumber="285">et al.</emphasis>
|
||
2007
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B0C05170513B7A388EFFB4019E6" author="Kuiter, R. H. & Debelius, H." box="[901,1174,1159,1185]" pageId="12" pageNumber="285" refId="ref11307" refString="Kuiter, R. H. & Debelius, H. (2009) World atlas of marine fauna. IKAN, Frankfurt, 725 pp." type="book" year="2009">Kuiter & Debelius 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B0C05170513B08488EFFA8E19E6" author="Hurt, C. & Hultgren, K. M. & Anker, A. & Lemmon, A. R. & Moriarty Lemmon, E. & Bracken-Grissom, H." box="[1186,1368,1159,1185]" pageId="12" pageNumber="285" pagination="1 - 11" refId="ref11003" refString="Hurt, C., Hultgren, K. M., Anker, A., Lemmon, A. R., Moriarty Lemmon, E. & Bracken-Grissom, H. (2021) First worldwide molecular phylogeny of the morphologically and ecologically hyperdiversified snapping shrimp genus Alpheus (Malacostraca: Decapoda). Molecular Phylogenetics and Evolution, 158 (197080), 1 - 11. https: // doi. org / 10.1016 / j. ympev. 2021.107080" type="journal article" year="2021">
|
||
Hurt
|
||
<emphasis id="B962EAEF05170513B0FB88EFFAD819E6" box="[1245,1294,1159,1185]" italics="true" pageId="12" pageNumber="285">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
); may also harbour the “commensal” palaemonid shrimp,
|
||
<taxonomicName id="4C164D7E05170513B6C088C3FBB21982" authority="Marin, 2008" authorityName="Marin" authorityYear="2008" box="[742,1124,1195,1221]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="285" phylum="Arthropoda" rank="species" species="aliska">
|
||
<emphasis id="B962EAEF05170513B6C088C3FC181982" box="[742,974,1195,1221]" italics="true" pageId="12" pageNumber="285">Palaemonella aliska</emphasis>
|
||
<bibRefCitation id="EF874B0C05170513B7FE88C3FBB21982" author="Marin, I." box="[984,1124,1195,1221]" pageId="12" pageNumber="285" pagination="375 - 390" refId="ref11334" refString="Marin, I. (2008) Description of two new species from the genera Palaemonella Dana, 1852 and Vir Holthuis, 1952 (Crustacea: Caridea: Palaemonidae: Pontoniinae). Zoologische Mededelingen, 82, 375 - 390." type="journal article" year="2008">Marin, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF874B0C05170513B05388C3FAD61982" author="Marin, I." box="[1141,1280,1195,1221]" pageId="12" pageNumber="285" pagination="375 - 390" refId="ref11334" refString="Marin, I. (2008) Description of two new species from the genera Palaemonella Dana, 1852 and Vir Holthuis, 1952 (Crustacea: Caridea: Palaemonidae: Pontoniinae). Zoologische Mededelingen, 82, 375 - 390." type="journal article" year="2008">Marin 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B0C05170513B12B88C3FEF619AE" author="Anker, A. & De Grave, S." pageId="12" pageNumber="285" pagination="145 - 150" refId="ref10292" refString="Anker, A. & De Grave, S. (2019) Further records of burrow-associated palaemonid shrimps (Decapoda: Palaemonidae). Zootaxa, 4612 (1), 145 - 150. https: // doi. org / 10.11646 / zootaxa. 4612.1.13" type="journal article" year="2019">Anker & De Grave 2019</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C657605170510B4E1889BFCF31956" lastPageId="15" lastPageNumber="288" pageId="12" pageNumber="285" type="discussion">
|
||
<paragraph id="8BA936FD05170513B4E1889BFAB718A2" blockId="12.[151,1437,151,1870]" pageId="12" pageNumber="285">
|
||
<emphasis id="B962EAEF05170513B4E1889BFEE2184A" bold="true" box="[199,308,1267,1293]" pageId="12" pageNumber="285">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C164D7E05170513B562889BFE3D184A" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[324,491,1267,1293]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="285" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
|
||
<emphasis id="B962EAEF05170513B562889BFE3D184A" box="[324,491,1267,1293]" italics="true" pageId="12" pageNumber="285">Alpheus sciolii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05170513B5D2889BFD86184A" bold="true" box="[500,592,1267,1293]" pageId="12" pageNumber="285">
|
||
<taxonomicNameLabel id="A251579405170513B5D2889BFD86184A" box="[500,592,1267,1293]" pageId="12" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has a unique and unmistakable colour pattern (
|
||
<figureCitation id="132D2A7805170513B056889BFB17184A" box="[1136,1217,1267,1293]" captionStart="FIGURE 7" captionStartId="13.[151,250,1880,1904]" captionTargetBox="[151,1435,181,1856]" captionTargetId="figure-15@13.[151,1435,181,1856]" captionTargetPageId="13" captionText="FIGURE 7. Alpheus sciolii sp. nov., holotype male (cl 8.6 mm) from New Georgia, Solomon Islands (MNHN-IU-2018-5667) [A, B]; paratype female (cl 10.7 mm) from Heron Island, Queensland, Australia (MNHN-IU-2018-5668) [C]: A, C, living shrimps, dorsal; B, same, lateral. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.5881119" httpUri="https://zenodo.org/record/5881119/files/figure.png" pageId="12" pageNumber="285">Figs. 7</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A7805170513B0E9889BFB27184A" box="[1231,1265,1267,1293]" captionStart="FIGURE 8" captionStartId="14.[151,250,1545,1569]" captionTargetBox="[158,1430,181,1521]" captionTargetId="figure-17@14.[158,1430,181,1521]" captionTargetPageId="14" captionText="FIGURE 8. Some goby-associated species of Alpheus mentioned in the text, photographed in situ: A, Alpheus thompsoni sp. nov. associated with Amblyeleotris wheeleri, Anilao, Philippines; B, same, associated with Amblyeleotris guttata, Sulawesi, Indonesia; C, Alpheus sciolii sp. nov. associated with Cryptocentrus inexplicatus, Papua New Guinea; D, Alpheus sp. aff. A. sciolii sp. nov. associated with Amblyeleotris steinitzi, Bali, Indonesia; E, Alpheus sp. cf. mannarensis Purushothaman, Abhilash, Kumar & Lal, 2021, Papua New Guinea; F, Alpheus bellulus Miya & Miyake, 1969, Sulawesi, Indonesia. Photographs courtesy of Andrey Ryanskiy (A–C, E, F) and Yury Ivanov (D)." figureDoi="http://doi.org/10.5281/zenodo.5881121" httpUri="https://zenodo.org/record/5881121/files/figure.png" pageId="12" pageNumber="285">8C</figureCitation>
|
||
), characterised by the dark-brown or olive-brown transverse banding and the presence of a very large dark spot on the flanks of the carapace. As in the previous species, the white “saddle” marking the limit between the carapace and the pleon is quite conspicuous. This colour pattern (
|
||
<figureCitation id="132D2A7805170513B64A8937FD66183E" box="[620,688,1375,1401]" captionStart="FIGURE 7" captionStartId="13.[151,250,1880,1904]" captionTargetBox="[151,1435,181,1856]" captionTargetId="figure-15@13.[151,1435,181,1856]" captionTargetPageId="13" captionText="FIGURE 7. Alpheus sciolii sp. nov., holotype male (cl 8.6 mm) from New Georgia, Solomon Islands (MNHN-IU-2018-5667) [A, B]; paratype female (cl 10.7 mm) from Heron Island, Queensland, Australia (MNHN-IU-2018-5668) [C]: A, C, living shrimps, dorsal; B, same, lateral. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.5881119" httpUri="https://zenodo.org/record/5881119/files/figure.png" pageId="12" pageNumber="285">Fig. 7</figureCitation>
|
||
; see also
|
||
<bibRefCitation id="EF874B0C05170513B73A8937FBFA183E" author="Kuiter, R. H. & Debelius, H." box="[796,1068,1375,1401]" pageId="12" pageNumber="285" refId="ref11307" refString="Kuiter, R. H. & Debelius, H. (2009) World atlas of marine fauna. IKAN, Frankfurt, 725 pp." type="book" year="2009">Kuiter & Debelius 2009</bibRefCitation>
|
||
, as
|
||
<taxonomicName id="4C164D7E05170513B0708937FB38183E" box="[1110,1262,1375,1401]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="285" phylum="Arthropoda" rank="species" species="undefined-6">
|
||
<emphasis id="B962EAEF05170513B0708937FB66183E" box="[1110,1200,1375,1401]" italics="true" pageId="12" pageNumber="285">Alpheus</emphasis>
|
||
sp. 6
|
||
</taxonomicName>
|
||
, and
|
||
<bibRefCitation id="EF874B0C05170513B10F8937FF0418DA" author="Minemizu, R." pageId="12" pageNumber="285" refId="ref11377" refString="Minemizu, R. (2013) Coral reef shrimps of Indo-West Pacific. Bun-ichi Sogo Shuppan, Co. Ltd., Tokyo, 144 pp. [in Japanese]" type="book" year="2013">Minemizu 2013</bibRefCitation>
|
||
, as
|
||
<taxonomicName id="4C164D7E05170513B52489EBFE7718DA" box="[258,417,1411,1437]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="285" phylum="Arthropoda" rank="species" species="undefined-2">
|
||
<emphasis id="B962EAEF05170513B52489EBFE8A18DA" box="[258,348,1411,1437]" italics="true" pageId="12" pageNumber="285">Alpheus</emphasis>
|
||
sp. 2
|
||
</taxonomicName>
|
||
) is very different from the colour patterns of
|
||
<taxonomicName id="4C164D7E05170513B79A89EBFBE118DA" authorityName="Miya & Miyake" authorityYear="1969" box="[956,1079,1411,1437]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="285" phylum="Arthropoda" rank="species" species="bellulus">
|
||
<emphasis id="B962EAEF05170513B79A89EBFBE118DA" box="[956,1079,1411,1437]" italics="true" pageId="12" pageNumber="285">A. bellulus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C164D7E05170513B06E89EBFB3918DA" authorityName="Chace" authorityYear="1988" box="[1096,1263,1411,1437]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="285" phylum="Arthropoda" rank="species" species="macellarius">
|
||
<emphasis id="B962EAEF05170513B06E89EBFB3918DA" box="[1096,1263,1411,1437]" italics="true" pageId="12" pageNumber="285">A. macellarius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C164D7E05170513B12789EBFAA418DA" authorityName="Bruce" authorityYear="1994" box="[1281,1394,1411,1437]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="285" phylum="Arthropoda" rank="species" species="fenneri">
|
||
<emphasis id="B962EAEF05170513B12789EBFAA418DA" box="[1281,1394,1411,1437]" italics="true" pageId="12" pageNumber="285">A. fenneri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C164D7E05170513B1A289EBFEF31887" authorityName="Purushothaman, Abhilash, Kumar & Lal" authorityYear="2021" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="285" phylum="Arthropoda" rank="species" species="mannarensis">
|
||
<emphasis id="B962EAEF05170513B1A289EBFEF31887" italics="true" pageId="12" pageNumber="285">A. mannarensis</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EF874B0C05170513B54389CFFDB51886" author="Miya, Y. & Miyake, S." box="[357,611,1447,1473]" pageId="12" pageNumber="285" pagination="307 - 314" refId="ref11409" refString="Miya, Y. & Miyake, S. (1969) Description of Alpheus bellulus sp. nov. associated with gobies from Japan (Crustacea, Decapoda, Alpheidae). Publications of the Seto Marine Biological Laboratory, 16, 307 - 314. https: // doi. org / 10.5134 / 175556" type="journal article" year="1969">Miya & Miyake 1969</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B0C05170513B65789CFFCCF1886" author="Debelius, H." box="[625,793,1447,1473]" pageId="12" pageNumber="285" refId="ref10810" refString="Debelius, H. (2001) Crustacea guide of the world. IKAN, Frankfurt, 321 pp." type="book" year="2001">Debelius 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B0C05170513B70E89CFFB961886" author="Kuiter, R. H. & Debelius, H." box="[808,1088,1447,1473]" pageId="12" pageNumber="285" refId="ref11307" refString="Kuiter, R. H. & Debelius, H. (2009) World atlas of marine fauna. IKAN, Frankfurt, 725 pp." type="book" year="2009">Kuiter & Debelius 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B0C05170513B06889CFFAD01886" author="Minemizu, R." box="[1102,1286,1447,1473]" pageId="12" pageNumber="285" refId="ref11377" refString="Minemizu, R. (2013) Coral reef shrimps of Indo-West Pacific. Bun-ichi Sogo Shuppan, Co. Ltd., Tokyo, 144 pp. [in Japanese]" type="book" year="2013">Minemizu 2013</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B0C05170513B13489CFFF0418A2" author="Anker, A. & Pratama, I. S. & Firdaus, M. & Rahayu, D. L." pageId="12" pageNumber="285" pagination="301 - 342" refId="ref10185" refString="Anker, A., Pratama, I. S., Firdaus, M. & Rahayu, D. L. (2015) On some interesting marine decapod crustaceans (Alpheidae, Laomediidae, Strahlaxiidae) from Lombok, Indonesia. Zootaxa, 3911, 301 - 342. https: // doi. org / 10.11646 / zootaxa. 3911.3.1" type="journal article" year="2015">
|
||
Anker
|
||
<emphasis id="B962EAEF05170513B14489CFFA431886" box="[1378,1429,1447,1473]" italics="true" pageId="12" pageNumber="285">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B0C05170513B4F889A3FDDB18A2" author="Purushothaman, P. & Abhilash, C. P. & Ajith Kumar, T. T. & Lal, K. K." box="[222,525,1483,1509]" pageId="12" pageNumber="285" pagination="127 - 135" refId="ref11627" refString="Purushothaman, P., Abhilash, C. P., Ajith Kumar, T. T. & Lal, K. K. (2021) A new alpheid shrimp, Alpheus mannarensis sp. nov. (Crustacea; Decapoda; Alpheidae) from Gulf of Mannar, southern India. Zootaxa, 5026 (1), 127 - 135. https: // doi. org / 10.11646 / zootaxa. 5026.1.5" type="journal article" year="2021">
|
||
Purushothaman
|
||
<emphasis id="B962EAEF05170513B5B589A3FE1218A2" box="[403,452,1483,1509]" italics="true" pageId="12" pageNumber="285">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
; for
|
||
<taxonomicName id="4C164D7E05170513B66789A3FD6F18A2" authorityName="Miya & Miyake" authorityYear="1969" box="[577,697,1483,1509]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="285" phylum="Arthropoda" rank="species" species="bellulus">
|
||
<emphasis id="B962EAEF05170513B66789A3FD6F18A2" box="[577,697,1483,1509]" italics="true" pageId="12" pageNumber="285">A. bellulus</emphasis>
|
||
</taxonomicName>
|
||
see also
|
||
<figureCitation id="132D2A7805170513B73889A3FCA618A2" box="[798,880,1483,1509]" captionStart="FIGURE 8" captionStartId="14.[151,250,1545,1569]" captionTargetBox="[158,1430,181,1521]" captionTargetId="figure-17@14.[158,1430,181,1521]" captionTargetPageId="14" captionText="FIGURE 8. Some goby-associated species of Alpheus mentioned in the text, photographed in situ: A, Alpheus thompsoni sp. nov. associated with Amblyeleotris wheeleri, Anilao, Philippines; B, same, associated with Amblyeleotris guttata, Sulawesi, Indonesia; C, Alpheus sciolii sp. nov. associated with Cryptocentrus inexplicatus, Papua New Guinea; D, Alpheus sp. aff. A. sciolii sp. nov. associated with Amblyeleotris steinitzi, Bali, Indonesia; E, Alpheus sp. cf. mannarensis Purushothaman, Abhilash, Kumar & Lal, 2021, Papua New Guinea; F, Alpheus bellulus Miya & Miyake, 1969, Sulawesi, Indonesia. Photographs courtesy of Andrey Ryanskiy (A–C, E, F) and Yury Ivanov (D)." figureDoi="http://doi.org/10.5281/zenodo.5881121" httpUri="https://zenodo.org/record/5881121/files/figure.png" pageId="12" pageNumber="285">Fig. 8F</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C164D7E05170513B78989A3FB9518A2" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[943,1091,1483,1509]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="285" phylum="Arthropoda" rank="species" species="thompsoni" status="sp. nov.">
|
||
<emphasis id="B962EAEF05170513B78989A3FB9518A2" box="[943,1091,1483,1509]" italics="true" pageId="12" pageNumber="285">A. thompsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05170513B06C89A3FB7518A2" bold="true" box="[1098,1187,1483,1509]" pageId="12" pageNumber="285">
|
||
<taxonomicNameLabel id="A251579405170513B06C89A3FB7518A2" box="[1098,1187,1483,1509]" pageId="12" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="132D2A7805170513B09489A3FAD718A2" box="[1202,1281,1483,1509]" captionStart="FIGURE 3" captionStartId="5.[151,250,1700,1724]" captionTargetBox="[151,1435,381,1677]" captionTargetId="figure-102@5.[151,1435,381,1677]" captionTargetPageId="5" captionText="FIGURE 3. Alpheus thompsoni sp. nov., holotype, female (cl 13.3 mm) from Madang, Papua New Guinea (MNHN-2013- 16813): A, shrimp in life, dorsal; B, same, lateral. Photographs by the author, taken during Our Planet Reviewed—Papua Niugini Expedition (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5881103" httpUri="https://zenodo.org/record/5881103/files/figure.png" pageId="12" pageNumber="285">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A7805170513B12889A3FA8518A2" box="[1294,1363,1483,1509]" captionStart="FIGURE 8" captionStartId="14.[151,250,1545,1569]" captionTargetBox="[158,1430,181,1521]" captionTargetId="figure-17@14.[158,1430,181,1521]" captionTargetPageId="14" captionText="FIGURE 8. Some goby-associated species of Alpheus mentioned in the text, photographed in situ: A, Alpheus thompsoni sp. nov. associated with Amblyeleotris wheeleri, Anilao, Philippines; B, same, associated with Amblyeleotris guttata, Sulawesi, Indonesia; C, Alpheus sciolii sp. nov. associated with Cryptocentrus inexplicatus, Papua New Guinea; D, Alpheus sp. aff. A. sciolii sp. nov. associated with Amblyeleotris steinitzi, Bali, Indonesia; E, Alpheus sp. cf. mannarensis Purushothaman, Abhilash, Kumar & Lal, 2021, Papua New Guinea; F, Alpheus bellulus Miya & Miyake, 1969, Sulawesi, Indonesia. Photographs courtesy of Andrey Ryanskiy (A–C, E, F) and Yury Ivanov (D)." figureDoi="http://doi.org/10.5281/zenodo.5881121" httpUri="https://zenodo.org/record/5881121/files/figure.png" pageId="12" pageNumber="285">8A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA936FD05170513B4E18987FD4D1A0A" blockId="12.[151,1437,151,1870]" pageId="12" pageNumber="285">
|
||
The non-type male specimen of
|
||
<taxonomicName id="4C164D7E05170513B6158987FD401B4E" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[563,662,1519,1545]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="285" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
|
||
<emphasis id="B962EAEF05170513B6158987FD401B4E" box="[563,662,1519,1545]" italics="true" pageId="12" pageNumber="285">A. sciolii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05170513B6B88987FD2E1B4E" bold="true" box="[670,760,1519,1545]" pageId="12" pageNumber="285">
|
||
<taxonomicNameLabel id="A251579405170513B6B88987FD2E1B4E" box="[670,760,1519,1545]" pageId="12" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from Nha Trang,
|
||
<collectingCountry id="F301766D05170513B7E48987FBF21B4E" box="[962,1060,1519,1545]" name="Vietnam" pageId="12" pageNumber="285">Vietnam</collectingCountry>
|
||
, has noticeably larger and stouter chelipeds, with a markedly stronger balaeniceps condition on the minor chela fingers, compared to that of the complete male type specimens from the
|
||
<collectingCountry id="F301766D05170513B6788A5FFCCC1B16" box="[606,794,1591,1617]" name="Solomon Islands" pageId="12" pageNumber="285">Solomon Islands</collectingCountry>
|
||
and
|
||
<collectingCountry id="F301766D05170513B7768A5FFC6C1B16" box="[848,954,1591,1617]" name="Australia" pageId="12" pageNumber="285">Australia</collectingCountry>
|
||
, (
|
||
<figureCitation id="132D2A7805170513B7E88A5FFB9E1B16" box="[974,1096,1591,1617]" captionStart="FIGURE 6" captionStartId="11.[151,250,1665,1689]" captionTargetBox="[190,1400,792,1619]" captionTargetId="figure-271@11.[151,1436,777,1642]" captionTargetPageId="11" captionText="FIGURE 6. Alpheus sciolii sp. nov., non-type male (cl 14.5 mm) from Nha Trang Bay, Vietnam (MNHN-IU-2018-5671):A, left (major) cheliped, carpus and chela, lateral; B, right (minor) cheliped, chela, lateral; C, telson, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881113" httpUri="https://zenodo.org/record/5881113/files/figure.png" pageId="12" pageNumber="285">Fig. 6A, B</figureCitation>
|
||
), which may be simply due to its larger size (cl
|
||
<quantity id="4CEE9B1805170513B57C8A32FE691B32" box="[346,447,1626,1653]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="12" pageNumber="285" unit="mm" value="14.5">14.5 mm</quantity>
|
||
<emphasis id="B962EAEF05170513B5E08A33FE0B1B33" box="[454,477,1627,1652]" italics="true" pageId="12" pageNumber="285">vs</emphasis>
|
||
.
|
||
<quantity id="4CEE9B1805170513B5CA8A33FDAE1B32" box="[492,632,1627,1653]" metricMagnitude="-3" metricUnit="m" metricValue="7.95" metricValueMax="9.3" metricValueMin="6.6" pageId="12" pageNumber="285" unit="mm" value="7.95" valueMax="9.3" valueMin="6.6">6.6–9.3 mm</quantity>
|
||
). A large
|
||
<typeStatus id="54AD885F05170513B6C28A33FC921B32" box="[740,836,1627,1653]" pageId="12" pageNumber="285" type="paratype">paratype</typeStatus>
|
||
male of similar size (cl 16.0 mm) from Heron Island is unfortunately lacking both of its chelipeds (see
|
||
<bibRefCitation id="EF874B0C05170513B6E38A17FC081BDE" author="Anker, A. & De Grave, S." box="[709,990,1663,1689]" pageId="12" pageNumber="285" pagination="145 - 150" refId="ref10292" refString="Anker, A. & De Grave, S. (2019) Further records of burrow-associated palaemonid shrimps (Decapoda: Palaemonidae). Zootaxa, 4612 (1), 145 - 150. https: // doi. org / 10.11646 / zootaxa. 4612.1.13" type="journal article" year="2019">Anker & De Grave 2019</bibRefCitation>
|
||
: fig. 4C). The Nha Trang specimen also seems to have a darker colour (see
|
||
<bibRefCitation id="EF874B0C05170513B6008ACBFD781BFA" author="Marin, I." box="[550,686,1699,1725]" pageId="12" pageNumber="285" pagination="375 - 390" refId="ref11334" refString="Marin, I. (2008) Description of two new species from the genera Palaemonella Dana, 1852 and Vir Holthuis, 1952 (Crustacea: Caridea: Palaemonidae: Pontoniinae). Zoologische Mededelingen, 82, 375 - 390." type="journal article" year="2008">Marin 2008</bibRefCitation>
|
||
: fig. 9b), although it is partly covered by detritus and sediment. In addition, a colour pattern very similar to that of
|
||
<taxonomicName id="4C164D7E05170513B6988AAFFCF41BA6" authorityName="Anker" authorityYear="2022" box="[702,802,1735,1761]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="285" phylum="Arthropoda" rank="species" species="sciolii">
|
||
<emphasis id="B962EAEF05170513B6988AAFFCF41BA6" box="[702,802,1735,1761]" italics="true" pageId="12" pageNumber="285">A. sciolii</emphasis>
|
||
</taxonomicName>
|
||
exists in shrimps associated with different gobies, such as
|
||
<taxonomicName id="4C164D7E05170513B4918A83FD9D1A41" authority="(Klausewitz)" baseAuthorityName="Klausewitz" baseAuthorityYear="1974" box="[183,587,1771,1798]" class="Actinopterygii" family="Gobiidae" genus="Amblyeleotris" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="285" phylum="Chordata" rank="species" species="steinitzi">
|
||
<emphasis id="B962EAEF05170513B4918A83FE641A43" box="[183,434,1771,1797]" italics="true" pageId="12" pageNumber="285">Amblyeleotris steinitzi</emphasis>
|
||
(Klausewitz)
|
||
</taxonomicName>
|
||
(see
|
||
<figureCitation id="132D2A7805170513B6AE8A83FD341A42" box="[648,738,1771,1797]" captionStart="FIGURE 8" captionStartId="14.[151,250,1545,1569]" captionTargetBox="[158,1430,181,1521]" captionTargetId="figure-17@14.[158,1430,181,1521]" captionTargetPageId="14" captionText="FIGURE 8. Some goby-associated species of Alpheus mentioned in the text, photographed in situ: A, Alpheus thompsoni sp. nov. associated with Amblyeleotris wheeleri, Anilao, Philippines; B, same, associated with Amblyeleotris guttata, Sulawesi, Indonesia; C, Alpheus sciolii sp. nov. associated with Cryptocentrus inexplicatus, Papua New Guinea; D, Alpheus sp. aff. A. sciolii sp. nov. associated with Amblyeleotris steinitzi, Bali, Indonesia; E, Alpheus sp. cf. mannarensis Purushothaman, Abhilash, Kumar & Lal, 2021, Papua New Guinea; F, Alpheus bellulus Miya & Miyake, 1969, Sulawesi, Indonesia. Photographs courtesy of Andrey Ryanskiy (A–C, E, F) and Yury Ivanov (D)." figureDoi="http://doi.org/10.5281/zenodo.5881121" httpUri="https://zenodo.org/record/5881121/files/figure.png" pageId="12" pageNumber="285">Fig. 8D</figureCitation>
|
||
). Therefore, collection of additional material and a molecular comparison between the specimens of
|
||
<taxonomicName id="4C164D7E05170513B6698B67FD621A6E" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[591,692,1807,1833]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="285" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
|
||
<emphasis id="B962EAEF05170513B6698B67FD621A6E" box="[591,692,1807,1833]" italics="true" pageId="12" pageNumber="285">A. sciolii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05170513B69B8B67FCCE1A6E" bold="true" box="[701,792,1807,1833]" pageId="12" pageNumber="285">
|
||
<taxonomicNameLabel id="A251579405170513B69B8B67FCCE1A6E" box="[701,792,1807,1833]" pageId="12" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from
|
||
<collectingCountry id="F301766D05170513B7798B67FBCA1A6E" box="[863,1052,1807,1833]" name="Solomon Islands" pageId="12" pageNumber="285">Solomon Islands</collectingCountry>
|
||
/
|
||
<collectingCountry id="F301766D05170513B0158B67FB4C1A6E" box="[1075,1178,1807,1833]" name="Australia" pageId="12" pageNumber="285">Australia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F301766D05170513B0F48B67FAE71A6E" box="[1234,1329,1807,1833]" name="Vietnam" pageId="12" pageNumber="285">Vietnam</collectingCountry>
|
||
(or other parts of South-East Asia) are highly desirable.
|
||
</paragraph>
|
||
<caption id="DF69667505160512B4B18B30FD2C1AFF" ID-DOI="http://doi.org/10.5281/zenodo.5881119" ID-Zenodo-Dep="5881119" httpUri="https://zenodo.org/record/5881119/files/figure.png" pageId="13" pageNumber="286" startId="13.[151,250,1880,1904]" targetBox="[151,1435,181,1856]" targetPageId="13">
|
||
<paragraph id="8BA936FD05160512B4B18B30FD2C1AFF" blockId="13.[151,1436,1880,1976]" pageId="13" pageNumber="286">
|
||
<emphasis id="B962EAEF05160512B4B18B30FEDB1A37" bold="true" box="[151,269,1880,1904]" pageId="13" pageNumber="286">FIGURE 7</emphasis>
|
||
.
|
||
<taxonomicName id="4C164D7E05160512B53C8B30FE661A37" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[282,432,1880,1904]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="286" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
|
||
<emphasis id="B962EAEF05160512B53C8B30FE661A37" box="[282,432,1880,1904]" italics="true" pageId="13" pageNumber="286">Alpheus sciolii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05160512B5908B30FDD11A37" bold="true" box="[438,519,1880,1904]" pageId="13" pageNumber="286">
|
||
<taxonomicNameLabel id="A251579405160512B5908B30FDD11A37" box="[438,519,1880,1904]" pageId="13" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (cl 8.6 mm) from New Georgia, Solomon Islands (MNHN-IU-2018-5667) [A, B]; paratype female (cl 10.7 mm) from Heron Island, Queensland, Australia (MNHN-IU-2018-5668) [C]: A, C, living shrimps, dorsal; B, same, lateral. Photographs by the author.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF69667505150511B4B18A61FCCD1B92" ID-DOI="http://doi.org/10.5281/zenodo.5881121" ID-Zenodo-Dep="5881121" httpUri="https://zenodo.org/record/5881121/files/figure.png" pageId="14" pageNumber="287" startId="14.[151,250,1545,1569]" targetBox="[158,1430,181,1521]" targetPageId="14">
|
||
<paragraph id="8BA936FD05150511B4B18A61FCCD1B92" blockId="14.[151,1437,1545,1750]" pageId="14" pageNumber="287">
|
||
<emphasis id="B962EAEF05150511B4B18A61FED81B66" bold="true" box="[151,270,1545,1569]" pageId="14" pageNumber="287">FIGURE 8</emphasis>
|
||
. Some goby-associated species of
|
||
<taxonomicName id="4C164D7E05150511B6528A61FD101B66" authorityName="Fabricius" authorityYear="1798" box="[628,710,1545,1569]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAEF05150511B6528A61FD101B66" box="[628,710,1545,1569]" italics="true" pageId="14" pageNumber="287">Alpheus</emphasis>
|
||
</taxonomicName>
|
||
mentioned in the text, photographed
|
||
<emphasis id="B962EAEF05150511B0618A62FB511B66" box="[1095,1159,1546,1569]" italics="true" pageId="14" pageNumber="287">in situ</emphasis>
|
||
: A,
|
||
<taxonomicName id="4C164D7E05150511B0928A61FAA11B66" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[1204,1399,1545,1569]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="thompsoni" status="sp. nov.">
|
||
<emphasis id="B962EAEF05150511B0928A61FAA11B66" box="[1204,1399,1545,1569]" italics="true" pageId="14" pageNumber="287">Alpheus thompsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05150511B1588A61FF151B02" bold="true" pageId="14" pageNumber="287">
|
||
<taxonomicNameLabel id="A251579405150511B1588A61FF151B02" pageId="14" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
associated with
|
||
<taxonomicName id="4C164D7E05150511B5558A45FD601B02" authority=", Anilao" authorityName="Anilao" box="[371,694,1581,1605]" class="Actinopterygii" family="Gobiidae" genus="Amblyeleotris" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="287" phylum="Chordata" rank="species" species="wheeleri">
|
||
<emphasis id="B962EAEF05150511B5558A45FDB61B02" box="[371,608,1581,1605]" italics="true" pageId="14" pageNumber="287">Amblyeleotris wheeleri</emphasis>
|
||
, Anilao
|
||
</taxonomicName>
|
||
, Philippines; B, same, associated with
|
||
<taxonomicName id="4C164D7E05150511B0688A45FAFD1B02" baseAuthorityName="Fowler" baseAuthorityYear="1938" box="[1102,1323,1581,1605]" class="Actinopterygii" family="Gobiidae" genus="Amblyeleotris" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="287" phylum="Chordata" rank="species" species="guttata">
|
||
<emphasis id="B962EAEF05150511B0688A45FAFD1B02" box="[1102,1323,1581,1605]" italics="true" pageId="14" pageNumber="287">Amblyeleotris guttata</emphasis>
|
||
</taxonomicName>
|
||
, Sulawesi, Indonesia; C,
|
||
<taxonomicName id="4C164D7E05150511B50D8A39FE131B2E" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[299,453,1617,1641]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
|
||
<emphasis id="B962EAEF05150511B50D8A39FE131B2E" box="[299,453,1617,1641]" italics="true" pageId="14" pageNumber="287">Alpheus sciolii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05150511B5E98A39FDF41B2E" bold="true" box="[463,546,1617,1641]" pageId="14" pageNumber="287">
|
||
<taxonomicNameLabel id="A251579405150511B5E98A39FDF41B2E" box="[463,546,1617,1641]" pageId="14" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
associated with
|
||
<taxonomicName id="4C164D7E05150511B6F38A39FC3C1B2E" baseAuthorityName="Herre" baseAuthorityYear="1934" box="[725,1002,1617,1641]" class="Actinopterygii" family="Gobiidae" genus="Cryptocentrus" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="287" phylum="Chordata" rank="species" species="inexplicatus">
|
||
<emphasis id="B962EAEF05150511B6F38A39FC3C1B2E" box="[725,1002,1617,1641]" italics="true" pageId="14" pageNumber="287">Cryptocentrus inexplicatus</emphasis>
|
||
</taxonomicName>
|
||
, Papua New Guinea; D,
|
||
<taxonomicName id="4C164D7E05150511B0DE8A39FAA71B2E" box="[1272,1393,1617,1641]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B962EAEF05150511B0DE8A39FA9C1B2E" box="[1272,1354,1617,1641]" italics="true" pageId="14" pageNumber="287">Alpheus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
aff.
|
||
<taxonomicName id="4C164D7E05150511B4B18A1DFF231BCA" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[151,245,1653,1677]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
|
||
<emphasis id="B962EAEF05150511B4B18A1DFF231BCA" box="[151,245,1653,1677]" italics="true" pageId="14" pageNumber="287">A. sciolii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05150511B4D88A1DFE841BCA" bold="true" box="[254,338,1653,1677]" pageId="14" pageNumber="287">
|
||
<taxonomicNameLabel id="A251579405150511B4D88A1DFE841BCA" box="[254,338,1653,1677]" pageId="14" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
associated with
|
||
<taxonomicName id="4C164D7E05150511B6238A1DFD3D1BCA" baseAuthorityName="Klausewitz" baseAuthorityYear="1974" box="[517,747,1653,1677]" class="Actinopterygii" family="Gobiidae" genus="Amblyeleotris" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="287" phylum="Chordata" rank="species" species="steinitzi">
|
||
<emphasis id="B962EAEF05150511B6238A1DFD3D1BCA" box="[517,747,1653,1677]" italics="true" pageId="14" pageNumber="287">Amblyeleotris steinitzi</emphasis>
|
||
</taxonomicName>
|
||
, Bali, Indonesia; E,
|
||
<taxonomicName id="4C164D7E05150511B7EE8A1DFB971BCA" box="[968,1089,1653,1677]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B962EAEF05150511B7EE8A1DFBCC1BCA" box="[968,1050,1653,1677]" italics="true" pageId="14" pageNumber="287">Alpheus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
<taxonomicName id="4C164D7E05150511B06C8A1DFE131BF6" authority="Purushothaman, Abhilash, Kumar & Lal, 2021" authorityName="Purushothaman, Abhilash, Kumar & Lal" authorityYear="2021" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="mannarensis">
|
||
cf.
|
||
<emphasis id="B962EAEF05150511B0488A1EFB391BCA" box="[1134,1263,1654,1677]" italics="true" pageId="14" pageNumber="287">mannarensis</emphasis>
|
||
<bibRefCitation id="EF874B0C05150511B0DE8A1DFE131BF6" author="Purushothaman, P. & Abhilash, C. P. & Ajith Kumar, T. T. & Lal, K. K." pageId="14" pageNumber="287" pagination="127 - 135" refId="ref11627" refString="Purushothaman, P., Abhilash, C. P., Ajith Kumar, T. T. & Lal, K. K. (2021) A new alpheid shrimp, Alpheus mannarensis sp. nov. (Crustacea; Decapoda; Alpheidae) from Gulf of Mannar, southern India. Zootaxa, 5026 (1), 127 - 135. https: // doi. org / 10.11646 / zootaxa. 5026.1.5" type="journal article" year="2021">Purushothaman, Abhilash, Kumar & Lal, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
, Papua New Guinea; F,
|
||
<taxonomicName id="4C164D7E05150511B6948AF1FB961BF6" authority="Miya & Miyake, 1969" authorityName="Miya & Miyake" authorityYear="1969" box="[690,1088,1689,1713]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="bellulus">
|
||
<emphasis id="B962EAEF05150511B6948AF1FC8F1BF6" box="[690,857,1689,1713]" italics="true" pageId="14" pageNumber="287">Alpheus bellulus</emphasis>
|
||
<bibRefCitation id="EF874B0C05150511B7788AF1FB961BF6" author="Miya, Y. & Miyake, S." box="[862,1088,1689,1713]" pageId="14" pageNumber="287" pagination="307 - 314" refId="ref11409" refString="Miya, Y. & Miyake, S. (1969) Description of Alpheus bellulus sp. nov. associated with gobies from Japan (Crustacea, Decapoda, Alpheidae). Publications of the Seto Marine Biological Laboratory, 16, 307 - 314. https: // doi. org / 10.5134 / 175556" type="journal article" year="1969">Miya & Miyake, 1969</bibRefCitation>
|
||
</taxonomicName>
|
||
, Sulawesi, Indonesia. Photographs courtesy of Andrey Ryanskiy (A–C, E, F) and Yury Ivanov (D).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA936FD05150510B4E18B6BFDA71C22" blockId="14.[151,1436,1795,2038]" lastBlockId="15.[151,1437,151,1041]" lastPageId="15" lastPageNumber="288" pageId="14" pageNumber="287">
|
||
The separation of
|
||
<taxonomicName id="4C164D7E05150511B5B58B6CFE211A5A" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[403,503,1795,1822]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
|
||
<emphasis id="B962EAEF05150511B5B58B6CFE211A5A" box="[403,503,1795,1822]" italics="true" pageId="14" pageNumber="287">A. sciolii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05150511B5D98B6BFD8F1A5A" bold="true" box="[511,601,1795,1821]" pageId="14" pageNumber="287">
|
||
<taxonomicNameLabel id="A251579405150511B5D98B6BFD8F1A5A" box="[511,601,1795,1821]" pageId="14" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from
|
||
<taxonomicName id="4C164D7E05150511B6B88B6CFCE21A5A" box="[670,820,1795,1822]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="djeddensis">
|
||
<emphasis id="B962EAEF05150511B6B88B6CFCE21A5A" box="[670,820,1795,1822]" italics="true" pageId="14" pageNumber="287">A. djeddensis</emphasis>
|
||
</taxonomicName>
|
||
is based on the same criteria as those used by
|
||
<bibRefCitation id="EF874B0C05150511B1678B6CFF341A06" author="De Man, J. G." pageId="14" pageNumber="287" pagination="99 - 125" refId="ref10774" refString="De Man, J. G. (1909) Diagnoses of new species of macrurous decapod Crustacea from the " Siboga-Expedition ". Tijdschrift der Nederlandse Dierkundige Vereeniging, 2, 99 - 125." type="journal article" year="1909">De Man (1909)</bibRefCitation>
|
||
to distinguish
|
||
<taxonomicName id="4C164D7E05150511B5B28B40FDE81A06" authorityName="De Man" authorityYear="1909" box="[404,574,1831,1858]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="djiboutensis">
|
||
<emphasis id="B962EAEF05150511B5B28B40FDE81A06" box="[404,574,1831,1858]" italics="true" pageId="14" pageNumber="287">A. djiboutensis</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C164D7E05150511B6AD8B40FCF21A06" box="[651,804,1831,1858]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="djeddensis">
|
||
<emphasis id="B962EAEF05150511B6AD8B40FCF21A06" box="[651,804,1831,1858]" italics="true" pageId="14" pageNumber="287">A. djeddensis</emphasis>
|
||
</taxonomicName>
|
||
, viz. the scaphocerite blade reaching well beyond the adjacent distolateral tooth in both
|
||
<taxonomicName id="4C164D7E05150511B6368B24FDA41A22" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[528,626,1867,1894]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
|
||
<emphasis id="B962EAEF05150511B6368B24FDA41A22" box="[528,626,1867,1894]" italics="true" pageId="14" pageNumber="287">A. sciolii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05150511B65E8B23FD071A22" bold="true" box="[632,721,1867,1893]" pageId="14" pageNumber="287">
|
||
<taxonomicNameLabel id="A251579405150511B65E8B23FD071A22" box="[632,721,1867,1893]" pageId="14" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C164D7E05150511B7238B24FC7C1A22" authorityName="De Man" authorityYear="1909" box="[773,938,1867,1894]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="djiboutensis">
|
||
<emphasis id="B962EAEF05150511B7238B24FC7C1A22" box="[773,938,1867,1894]" italics="true" pageId="14" pageNumber="287">A. djiboutensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B962EAEF05150511B79C8B24FC071A22" box="[954,977,1868,1893]" italics="true" pageId="14" pageNumber="287">vs</emphasis>
|
||
. not reaching beyond it in
|
||
<taxonomicName id="4C164D7E05150511B0DE8B24FA5A1A22" box="[1272,1420,1867,1894]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="djeddensis">
|
||
<emphasis id="B962EAEF05150511B0DE8B24FA5A1A22" box="[1272,1420,1867,1894]" italics="true" pageId="14" pageNumber="287">A. djeddensis</emphasis>
|
||
</taxonomicName>
|
||
), and the first subarticle of the second pereiopod carpus noticeably longer than the second in
|
||
<taxonomicName id="4C164D7E05150511B0838B18FADF1ACE" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[1189,1289,1903,1930]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
|
||
<emphasis id="B962EAEF05150511B0838B18FADF1ACE" box="[1189,1289,1903,1930]" italics="true" pageId="14" pageNumber="287">A. sciolii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05150511B1378B07FABA1ACE" bold="true" box="[1297,1388,1903,1929]" pageId="14" pageNumber="287">
|
||
<taxonomicNameLabel id="A251579405150511B1378B07FABA1ACE" box="[1297,1388,1903,1929]" pageId="14" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C164D7E05150511B4B18BFCFEE81AEA" authorityName="De Man" authorityYear="1909" box="[151,318,1939,1966]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="djiboutensis">
|
||
<emphasis id="B962EAEF05150511B4B18BFCFEE81AEA" box="[151,318,1939,1966]" italics="true" pageId="14" pageNumber="287">A. djiboutensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B962EAEF05150511B5698BFCFEB01AEA" box="[335,358,1940,1965]" italics="true" pageId="14" pageNumber="287">vs</emphasis>
|
||
. as long as the second in
|
||
<taxonomicName id="4C164D7E05150511B6A18BFCFCCA1AEA" box="[647,796,1939,1966]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="djeddensis">
|
||
<emphasis id="B962EAEF05150511B6A18BFCFCCA1AEA" box="[647,796,1939,1966]" italics="true" pageId="14" pageNumber="287">A. djeddensis</emphasis>
|
||
</taxonomicName>
|
||
). On the other hand,
|
||
<taxonomicName id="4C164D7E05150511B02B8BFCFBA61AEA" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[1037,1136,1939,1966]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
|
||
<emphasis id="B962EAEF05150511B02B8BFCFBA61AEA" box="[1037,1136,1939,1966]" italics="true" pageId="14" pageNumber="287">A. sciolii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05150511B05E8BFBFB041AEA" bold="true" box="[1144,1234,1939,1965]" pageId="14" pageNumber="287">
|
||
<taxonomicNameLabel id="A251579405150511B05E8BFBFB041AEA" box="[1144,1234,1939,1965]" pageId="14" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
may be separated from
|
||
<taxonomicName id="4C164D7E05150511B4F38BD0FEAA1A96" authorityName="De Man" authorityYear="1909" box="[213,380,1975,2002]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="djiboutensis">
|
||
<emphasis id="B962EAEF05150511B4F38BD0FEAA1A96" box="[213,380,1975,2002]" italics="true" pageId="14" pageNumber="287">A. djiboutensis</emphasis>
|
||
</taxonomicName>
|
||
by the much broader telson, which is around 1.8 times as long as maximal width in
|
||
<taxonomicName id="4C164D7E05150511B11E8BD0FA4A1A96" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[1336,1436,1975,2002]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
|
||
<emphasis id="B962EAEF05150511B11E8BD0FA4A1A96" box="[1336,1436,1975,2002]" italics="true" pageId="14" pageNumber="287">A. sciolii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05150511B4B18BB3FF261AB2" bold="true" box="[151,240,2011,2037]" pageId="14" pageNumber="287">
|
||
<taxonomicNameLabel id="A251579405150511B4B18BB3FF261AB2" box="[151,240,2011,2037]" pageId="14" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="132D2A7805150511B4D98BB3FEB61AB2" box="[255,352,2011,2037]" captionStart="FIGURE 4" captionStartId="9.[151,250,1839,1863]" captionTargetBox="[169,1413,191,1801]" captionTargetId="figure-15@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="14" pageNumber="287">Figs. 4C</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A7805150511B54D8BB3FE5B1AB1" box="[363,397,2011,2038]" captionStart="FIGURE 6" captionStartId="11.[151,250,1665,1689]" captionTargetBox="[190,1400,792,1619]" captionTargetId="figure-271@11.[151,1436,777,1642]" captionTargetPageId="11" captionText="FIGURE 6. Alpheus sciolii sp. nov., non-type male (cl 14.5 mm) from Nha Trang Bay, Vietnam (MNHN-IU-2018-5671):A, left (major) cheliped, carpus and chela, lateral; B, right (minor) cheliped, chela, lateral; C, telson, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881113" httpUri="https://zenodo.org/record/5881113/files/figure.png" pageId="14" pageNumber="287">6C</figureCitation>
|
||
)
|
||
<emphasis id="B962EAEF05150511B5BD8BB4FE6C1AB2" box="[411,442,2012,2037]" italics="true" pageId="14" pageNumber="287">vs.</emphasis>
|
||
2.1 times in
|
||
<taxonomicName id="4C164D7E05150511B66E8BB4FD381AB2" authorityName="De Man" authorityYear="1909" box="[584,750,2011,2038]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="djiboutensis">
|
||
<emphasis id="B962EAEF05150511B66E8BB4FD381AB2" box="[584,750,2011,2038]" italics="true" pageId="14" pageNumber="287">A. djiboutensis</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EF874B0C05150511B70E8BB4FC121AB2" author="De Man, J. G." box="[808,964,2011,2038]" pageId="14" pageNumber="287" pagination="99 - 125" refId="ref10774" refString="De Man, J. G. (1909) Diagnoses of new species of macrurous decapod Crustacea from the " Siboga-Expedition ". Tijdschrift der Nederlandse Dierkundige Vereeniging, 2, 99 - 125." type="journal article" year="1909">De Man 1909</bibRefCitation>
|
||
: fig. 18). In fact, the telson of
|
||
<taxonomicName id="4C164D7E05150511B1348BB4FAA21AB2" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[1298,1396,2011,2038]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="287" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
|
||
<emphasis id="B962EAEF05150511B1348BB4FAA21AB2" box="[1298,1396,2011,2038]" italics="true" pageId="14" pageNumber="287">A. sciolii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05150510B15D8BB3FF1E1DF6" bold="true" lastPageId="15" lastPageNumber="288" pageId="14" pageNumber="287">
|
||
<taxonomicNameLabel id="A251579405150510B15D8BB3FF1E1DF6" lastPageId="15" lastPageNumber="288" pageId="14" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
appears to be the broadest among the presently known species of the
|
||
<emphasis id="B962EAEF05140510B7F08CFFFAFB1DF6" box="[982,1325,151,177]" italics="true" pageId="15" pageNumber="288">
|
||
<taxonomicName id="4C164D7E05140510B7F08CFFFBBD1DF6" box="[982,1131,151,177]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="djeddensis">A. djeddensis</taxonomicName>
|
||
–
|
||
<taxonomicName id="4C164D7E05140510B0A18CFFFAFB1DF6" authorityName="De Man" authorityYear="1909" box="[1159,1325,151,177]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="djiboutensis">A. djiboutensis</taxonomicName>
|
||
</emphasis>
|
||
complex. Also noteworthy is that no snapping shrimp with the colour pattern of
|
||
<taxonomicName id="4C164D7E05140510B7E48CD3FBF11D92" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[962,1063,187,213]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
|
||
<emphasis id="B962EAEF05140510B7E48CD3FBF11D92" box="[962,1063,187,213]" italics="true" pageId="15" pageNumber="288">A. sciolii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05140510B0168CD3FB5D1D92" bold="true" box="[1072,1163,187,213]" pageId="15" pageNumber="288">
|
||
<taxonomicNameLabel id="A251579405140510B0168CD3FB5D1D92" box="[1072,1163,187,213]" pageId="15" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known from the
|
||
<collectingRegion id="49D2F81F05140510B1498CD3FF691DBE" country="Sudan" name="Al Bahr al Ahmar" pageId="15" pageNumber="288">Red Sea</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EF874B0C05140510B4F68CB7FE791DBE" author="Karplus, I. & Szlep, R. & Tsurnamal, M." box="[208,431,223,249]" pageId="15" pageNumber="288" pagination="1 - 19" refId="ref11166" refString="Karplus, I., Szlep, R. & Tsurnamal, M. (1981) Goby-shrimp partner specificity. I. Distribution in the northern Red Sea and partner specificity. Journal of Experimental Marine Biology and Ecology, 51, 1 - 19. https: // doi. org / 10.1016 / 0022 - 0981 (81) 90151 - 9" type="journal article" year="1981">
|
||
Karplus
|
||
<emphasis id="B962EAEF05140510B5158CB7FEBB1DBE" box="[307,365,223,249]" italics="true" pageId="15" pageNumber="288">et al.</emphasis>
|
||
1981
|
||
</bibRefCitation>
|
||
) or western Indian Ocean (
|
||
<bibRefCitation id="EF874B0C05140510B6CE8CB7FBC71DBE" author="Polunin, N. V. C. & Lubbock, R." box="[744,1041,223,249]" pageId="15" pageNumber="288" pagination="63 - 101" refId="ref11525" refString="Polunin, N. V. C. & Lubbock, R. (1977) Prawn-associated gobies (Teleostei: Gobiidae) from the Seychelles, western Indian Ocean: systematics and ecology. Journal of Zoology, London, 183, 63 - 101. https: // doi. org / 10.1111 / j. 1469 - 7998.1977. tb 04174. x" type="journal article" year="1977">Polunin & Lubbock 1977</bibRefCitation>
|
||
), from where
|
||
<taxonomicName id="4C164D7E05140510B0958CB7FA9C1DBE" box="[1203,1354,223,249]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="djeddensis">
|
||
<emphasis id="B962EAEF05140510B0958CB7FA9C1DBE" box="[1203,1354,223,249]" italics="true" pageId="15" pageNumber="288">A. djeddensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C164D7E05140510B1A28CB7FEC81C5A" authorityName="De Man" authorityYear="1909" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="djiboutensis">
|
||
<emphasis id="B962EAEF05140510B1A28CB7FEC81C5A" italics="true" pageId="15" pageNumber="288">A. djiboutensis</emphasis>
|
||
</taxonomicName>
|
||
were origially described. The Australian material reported as
|
||
<taxonomicName id="4C164D7E05140510B7D28D6BFB481C5A" authorityName="De Man" authorityYear="1909" box="[1012,1182,259,285]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="djiboutensis">
|
||
<emphasis id="B962EAEF05140510B7D28D6BFB481C5A" box="[1012,1182,259,285]" italics="true" pageId="15" pageNumber="288">A. djiboutensis</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF874B0C05140510B0E98D6BFF341C06" author="Banner, D. M. & Banner, A. H." pageId="15" pageNumber="288" pagination="1 - 357" refId="ref10443" refString="Banner, D. M. & Banner, A. H. (1982) The alpheid shrimp of Australia, part III: the remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner & Banner (1982)</bibRefCitation>
|
||
appears to contain more than one species (possibly including the true
|
||
<taxonomicName id="4C164D7E05140510B7D58D4FFB4F1C06" authorityName="De Man" authorityYear="1909" box="[1011,1177,295,321]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="djiboutensis">
|
||
<emphasis id="B962EAEF05140510B7D58D4FFB4F1C06" box="[1011,1177,295,321]" italics="true" pageId="15" pageNumber="288">A. djiboutensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C164D7E05140510B0E98D4FFAE41C06" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[1231,1330,295,321]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
|
||
<emphasis id="B962EAEF05140510B0E98D4FFAE41C06" box="[1231,1330,295,321]" italics="true" pageId="15" pageNumber="288">A. sciolii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05140510B11C8D4FFA451C06" bold="true" box="[1338,1427,295,321]" pageId="15" pageNumber="288">
|
||
<taxonomicNameLabel id="A251579405140510B11C8D4FFA451C06" box="[1338,1427,295,321]" pageId="15" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) and will need to be carefully re-examined.
|
||
</paragraph>
|
||
<paragraph id="8BA936FD05140510B4E18D07FCF31956" blockId="15.[151,1437,151,1041]" pageId="15" pageNumber="288">
|
||
<taxonomicName id="4C164D7E05140510B4E18D07FEBB1CCE" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[199,365,367,393]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="sciolii" status="sp. nov.">
|
||
<emphasis id="B962EAEF05140510B4E18D07FEBB1CCE" box="[199,365,367,393]" italics="true" pageId="15" pageNumber="288">Alpheus sciolii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05140510B5538D07FE061CCE" bold="true" box="[373,464,367,393]" pageId="15" pageNumber="288">
|
||
<taxonomicNameLabel id="A251579405140510B5538D07FE061CCE" box="[373,464,367,393]" pageId="15" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished morphologically from
|
||
<taxonomicName id="4C164D7E05140510B7E28D07FBEB1CCE" authorityName="Miya & Miyake" authorityYear="1969" box="[964,1085,367,393]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="bellulus">
|
||
<emphasis id="B962EAEF05140510B7E28D07FBEB1CCE" box="[964,1085,367,393]" italics="true" pageId="15" pageNumber="288">A. bellulus</emphasis>
|
||
</taxonomicName>
|
||
by the rostrum much narrower and longer (
|
||
<emphasis id="B962EAEF05140510B5058DFBFEEC1CEB" box="[291,314,403,428]" italics="true" pageId="15" pageNumber="288">vs</emphasis>
|
||
. shorter and subtriangular in
|
||
<taxonomicName id="4C164D7E05140510B6A88DFBFCDE1CEA" authorityName="Miya & Miyake" authorityYear="1969" box="[654,776,403,429]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="bellulus">
|
||
<emphasis id="B962EAEF05140510B6A88DFBFCDE1CEA" box="[654,776,403,429]" italics="true" pageId="15" pageNumber="288">A. bellulus</emphasis>
|
||
</taxonomicName>
|
||
), the ventral margin of the major chela almost straight to slightly convex (
|
||
<emphasis id="B962EAEF05140510B5778DDFFEBE1C97" box="[337,360,439,464]" italics="true" pageId="15" pageNumber="288">vs</emphasis>
|
||
. somewhat concave in
|
||
<taxonomicName id="4C164D7E05140510B64F8DDFFD091C96" authorityName="Miya & Miyake" authorityYear="1969" box="[617,735,439,465]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="bellulus">
|
||
<emphasis id="B962EAEF05140510B64F8DDFFD091C96" box="[617,735,439,465]" italics="true" pageId="15" pageNumber="288">A. bellulus</emphasis>
|
||
</taxonomicName>
|
||
), and the telson 1.8 times as long as broad (
|
||
<emphasis id="B962EAEF05140510B0998DDFFB001C97" box="[1215,1238,439,464]" italics="true" pageId="15" pageNumber="288">vs</emphasis>
|
||
. 2.5 times as long as broad) (see
|
||
<bibRefCitation id="EF874B0C05140510B51B8DB3FDE11CB2" author="Miya, Y. & Miyake, S." box="[317,567,475,501]" pageId="15" pageNumber="288" pagination="307 - 314" refId="ref11409" refString="Miya, Y. & Miyake, S. (1969) Description of Alpheus bellulus sp. nov. associated with gobies from Japan (Crustacea, Decapoda, Alpheidae). Publications of the Seto Marine Biological Laboratory, 16, 307 - 314. https: // doi. org / 10.5134 / 175556" type="journal article" year="1969">Miya & Miyake 1969</bibRefCitation>
|
||
: figs. 1, 2B, C); from
|
||
<taxonomicName id="4C164D7E05140510B7158DB3FC011CB2" authorityName="Chace" authorityYear="1988" box="[819,983,475,501]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="macellarius">
|
||
<emphasis id="B962EAEF05140510B7158DB3FC011CB2" box="[819,983,475,501]" italics="true" pageId="15" pageNumber="288">A. macellarius</emphasis>
|
||
</taxonomicName>
|
||
by the distolateral tooth of the antennal scaphocerite pointing straight-forward (
|
||
<emphasis id="B962EAEF05140510B6758D97FDBC1F5F" box="[595,618,511,536]" italics="true" pageId="15" pageNumber="288">vs</emphasis>
|
||
. curved mesially in
|
||
<taxonomicName id="4C164D7E05140510B76D8D97FC3B1F5E" authorityName="Chace" authorityYear="1988" box="[843,1005,511,537]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="macellarius">
|
||
<emphasis id="B962EAEF05140510B76D8D97FC3B1F5E" box="[843,1005,511,537]" italics="true" pageId="15" pageNumber="288">A. macellarius</emphasis>
|
||
</taxonomicName>
|
||
), the much stouter minor chela (which is very slender in
|
||
<taxonomicName id="4C164D7E05140510B57A8E4BFE281F7A" authorityName="Chace" authorityYear="1988" box="[348,510,547,573]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="macellarius">
|
||
<emphasis id="B962EAEF05140510B57A8E4BFE281F7A" box="[348,510,547,573]" italics="true" pageId="15" pageNumber="288">A. macellarius</emphasis>
|
||
</taxonomicName>
|
||
), and the much stouter third pereiopod merus, about four times as long as wide (vs. almost six times in
|
||
<taxonomicName id="4C164D7E05140510B5518E2FFDCA1F26" authorityName="Chace" authorityYear="1988" box="[375,540,583,609]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="macellarius">
|
||
<emphasis id="B962EAEF05140510B5518E2FFDCA1F26" box="[375,540,583,609]" italics="true" pageId="15" pageNumber="288">A. macellarius</emphasis>
|
||
</taxonomicName>
|
||
) (see
|
||
<bibRefCitation id="EF874B0C05140510B6458E2FFD381F26" author="Chace, F. A. Jr." box="[611,750,583,609]" pageId="15" pageNumber="288" pagination="1 - 99" refId="ref10577" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, 1 - 99. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1988">Chace 1988</bibRefCitation>
|
||
: fig. 6e, f, i); from
|
||
<taxonomicName id="4C164D7E05140510B7EA8E2FFBED1F26" authorityName="Bruce" authorityYear="1994" box="[972,1083,583,609]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="fenneri">
|
||
<emphasis id="B962EAEF05140510B7EA8E2FFBED1F26" box="[972,1083,583,609]" italics="true" pageId="15" pageNumber="288">A. fenneri</emphasis>
|
||
</taxonomicName>
|
||
by the distolateral tooth of the scaphocerite reaching far beyond the blade (
|
||
<emphasis id="B962EAEF05140510B6A38E03FD4A1FC3" box="[645,668,619,644]" italics="true" pageId="15" pageNumber="288">vs</emphasis>
|
||
. not or very slightly overreaching the blade in
|
||
<taxonomicName id="4C164D7E05140510B0808E03FAC51FC2" authorityName="Bruce" authorityYear="1994" box="[1190,1299,619,645]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="fenneri">
|
||
<emphasis id="B962EAEF05140510B0808E03FAC51FC2" box="[1190,1299,619,645]" italics="true" pageId="15" pageNumber="288">A. fenneri</emphasis>
|
||
</taxonomicName>
|
||
), the mesial surface of the major and minor chelae not granulated (
|
||
<emphasis id="B962EAEF05140510B7358EE7FCFC1FEF" box="[787,810,655,680]" italics="true" pageId="15" pageNumber="288">vs</emphasis>
|
||
. distinctly granulated in
|
||
<taxonomicName id="4C164D7E05140510B0688EE7FB691FEE" authorityName="Bruce" authorityYear="1994" box="[1102,1215,655,681]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="fenneri">
|
||
<emphasis id="B962EAEF05140510B0688EE7FB691FEE" box="[1102,1215,655,681]" italics="true" pageId="15" pageNumber="288">A. fenneri</emphasis>
|
||
</taxonomicName>
|
||
), and the posterior margin of the telson unarmed between the mesial pair of the posterolateral spiniform setae (
|
||
<emphasis id="B962EAEF05140510B0F78EDBFB3E1F8B" box="[1233,1256,691,716]" italics="true" pageId="15" pageNumber="288">vs</emphasis>
|
||
. with a row of spiniform setae in
|
||
<taxonomicName id="4C164D7E05140510B5428EBFFE041FB6" authorityName="Bruce" authorityYear="1994" box="[356,466,727,753]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="fenneri">
|
||
<emphasis id="B962EAEF05140510B5428EBFFE041FB6" box="[356,466,727,753]" italics="true" pageId="15" pageNumber="288">A. fenneri</emphasis>
|
||
</taxonomicName>
|
||
) (see
|
||
<bibRefCitation id="EF874B0C05140510B6338EBFFD4C1FB6" author="Bruce, A. J." box="[533,666,727,753]" pageId="15" pageNumber="288" pagination="15 - 28" refId="ref10508" refString="Bruce, A. J. (1994) Alpheus fenneri sp. nov. and A. williamsi sp. nov., two new Indo-West Pacific alpheid shrimps of the brevirostris species group. The Beagle, Records of the Museums and Art Galleries of the Northern Territory, 11, 15 - 28. https: // doi. org / 10.5962 / p. 264023" type="journal article" year="1994">Bruce 1994</bibRefCitation>
|
||
: figs. 1C, M, N, 2C); from
|
||
<taxonomicName id="4C164D7E05140510B7E38EBFFBA41FB7" authorityName="Purushothaman, Abhilash, Kumar & Lal" authorityYear="2021" box="[965,1138,727,753]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="mannarensis">
|
||
<emphasis id="B962EAEF05140510B7E38EBFFBA41FB7" box="[965,1138,727,753]" italics="true" pageId="15" pageNumber="288">A. mannarensis</emphasis>
|
||
</taxonomicName>
|
||
by the distolateral tooth of the scaphocerite reaching far beyond the blade (
|
||
<emphasis id="B962EAEF05140510B68D8E93FD141E53" box="[683,706,763,788]" italics="true" pageId="15" pageNumber="288">vs</emphasis>
|
||
. not reaching the anterior margin of the blade in
|
||
<taxonomicName id="4C164D7E05140510B0C68E93FA5A1E53" authorityName="Purushothaman, Abhilash, Kumar & Lal" authorityYear="2021" box="[1248,1420,763,789]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="mannarensis">
|
||
<emphasis id="B962EAEF05140510B0C68E93FA5A1E53" box="[1248,1420,763,789]" italics="true" pageId="15" pageNumber="288">A. mannarensis</emphasis>
|
||
</taxonomicName>
|
||
), the second pereiopod carpus with the first subarticle slightly longer than the second (
|
||
<emphasis id="B962EAEF05140510B0748F77FBBF1E7F" box="[1106,1129,799,824]" italics="true" pageId="15" pageNumber="288">vs</emphasis>
|
||
. with the second subarticle noticeably longer than the first in
|
||
<taxonomicName id="4C164D7E05140510B6308F2BFD121E1B" authorityName="Purushothaman, Abhilash, Kumar & Lal" authorityYear="2021" box="[534,708,835,861]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="mannarensis">
|
||
<emphasis id="B962EAEF05140510B6308F2BFD121E1B" box="[534,708,835,861]" italics="true" pageId="15" pageNumber="288">A. mannarensis</emphasis>
|
||
</taxonomicName>
|
||
), the posterior margin of the telson unarmed between the mesial pair of the posterolateral spiniform setae (
|
||
<emphasis id="B962EAEF05140510B64A8F0FFD551EC7" box="[620,643,871,896]" italics="true" pageId="15" pageNumber="288">vs</emphasis>
|
||
. with a row of spiniform setae in
|
||
<taxonomicName id="4C164D7E05140510B7DB8F0FFB7C1EC7" authorityName="Purushothaman, Abhilash, Kumar & Lal" authorityYear="2021" box="[1021,1194,871,897]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="mannarensis">
|
||
<emphasis id="B962EAEF05140510B7DB8F0FFB7C1EC7" box="[1021,1194,871,897]" italics="true" pageId="15" pageNumber="288">A. mannarensis</emphasis>
|
||
</taxonomicName>
|
||
) (see
|
||
<bibRefCitation id="EF874B0C05140510B0C88F0FFEC71EE2" author="Purushothaman, P. & Abhilash, C. P. & Ajith Kumar, T. T. & Lal, K. K." pageId="15" pageNumber="288" pagination="127 - 135" refId="ref11627" refString="Purushothaman, P., Abhilash, C. P., Ajith Kumar, T. T. & Lal, K. K. (2021) A new alpheid shrimp, Alpheus mannarensis sp. nov. (Crustacea; Decapoda; Alpheidae) from Gulf of Mannar, southern India. Zootaxa, 5026 (1), 127 - 135. https: // doi. org / 10.11646 / zootaxa. 5026.1.5" type="journal article" year="2021">
|
||
Purushothaman
|
||
<emphasis id="B962EAEF05140510B4B18FE3FF1E1EE2" box="[151,200,907,933]" italics="true" pageId="15" pageNumber="288">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
: figs. 2B, E, 3G); and from the above-described
|
||
<taxonomicName id="4C164D7E05140510B70F8FE3FC6A1EE2" authority="Anker, 2022" authorityName="Anker" authorityYear="2022" box="[809,956,907,933]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="288" phylum="Arthropoda" rank="species" species="thompsoni" status="sp. nov.">
|
||
<emphasis id="B962EAEF05140510B70F8FE3FC6A1EE2" box="[809,956,907,933]" italics="true" pageId="15" pageNumber="288">A. thompsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAEF05140510B7E48FE3FBCD1EE2" bold="true" box="[962,1051,907,933]" pageId="15" pageNumber="288">
|
||
<taxonomicNameLabel id="A251579405140510B7E48FE3FBCD1EE2" box="[962,1051,907,933]" pageId="15" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the relative development of the distolateral tooth of the scaphocerite (cf.
|
||
<figureCitation id="132D2A7805140510B67B8FC7FD681E8E" box="[605,702,943,969]" captionStart="FIGURE 1" captionStartId="3.[151,250,1814,1838]" captionTargetBox="[153,1420,187,1767]" captionTargetId="figure-15@3.[151,1436,181,1790]" captionTargetPageId="3" captionText="FIGURE 1. Alpheus thompsoni sp. nov., holotype, female (cl 13.3 mm) from Madang, Papua New Guinea (MNHN-2013- 16813): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal portion of propodus and dactylus, ventrolateral, slender setae on propodus omitted; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventromesial; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881097" httpUri="https://zenodo.org/record/5881097/files/figure.png" pageId="15" pageNumber="288">Figs. 1A</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A7805140510B6EF8FC7FD391E8E" box="[713,751,943,969]" captionStart="FIGURE 4" captionStartId="9.[151,250,1839,1863]" captionTargetBox="[169,1413,191,1801]" captionTargetId="figure-15@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="15" pageNumber="288">4A</figureCitation>
|
||
), the length-width proportions of the telson (cf.
|
||
<figureCitation id="132D2A7805140510B1268FC7FA891E8E" box="[1280,1375,943,969]" captionStart="FIGURE 1" captionStartId="3.[151,250,1814,1838]" captionTargetBox="[153,1420,187,1767]" captionTargetId="figure-15@3.[151,1436,181,1790]" captionTargetPageId="3" captionText="FIGURE 1. Alpheus thompsoni sp. nov., holotype, female (cl 13.3 mm) from Madang, Papua New Guinea (MNHN-2013- 16813): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal portion of propodus and dactylus, ventrolateral, slender setae on propodus omitted; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventromesial; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881097" httpUri="https://zenodo.org/record/5881097/files/figure.png" pageId="15" pageNumber="288">Figs. 1C</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A7805140510B14D8FC7FA591E8E" box="[1387,1423,943,969]" captionStart="FIGURE 4" captionStartId="9.[151,250,1839,1863]" captionTargetBox="[169,1413,191,1801]" captionTargetId="figure-15@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="15" pageNumber="288">4C</figureCitation>
|
||
), the much stouter third maxilliped, without extremely elongate setae on the pendultimate article (cf.
|
||
<figureCitation id="132D2A7805140510B1268FBBFA891EAA" box="[1280,1375,979,1005]" captionStart="FIGURE 1" captionStartId="3.[151,250,1814,1838]" captionTargetBox="[153,1420,187,1767]" captionTargetId="figure-15@3.[151,1436,181,1790]" captionTargetPageId="3" captionText="FIGURE 1. Alpheus thompsoni sp. nov., holotype, female (cl 13.3 mm) from Madang, Papua New Guinea (MNHN-2013- 16813): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal portion of propodus and dactylus, ventrolateral, slender setae on propodus omitted; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventromesial; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881097" httpUri="https://zenodo.org/record/5881097/files/figure.png" pageId="15" pageNumber="288">Figs. 1E</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A7805140510B14B8FBBFA591EAA" box="[1389,1423,979,1005]" captionStart="FIGURE 4" captionStartId="9.[151,250,1839,1863]" captionTargetBox="[169,1413,191,1801]" captionTargetId="figure-15@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="15" pageNumber="288">4E</figureCitation>
|
||
), and the much stouter walking legs (cf.
|
||
<figureCitation id="132D2A7805140510B66D8F9FFD121956" box="[587,708,1015,1041]" captionStart="FIGURE 1" captionStartId="3.[151,250,1814,1838]" captionTargetBox="[153,1420,187,1767]" captionTargetId="figure-15@3.[151,1436,181,1790]" captionTargetPageId="3" captionText="FIGURE 1. Alpheus thompsoni sp. nov., holotype, female (cl 13.3 mm) from Madang, Papua New Guinea (MNHN-2013- 16813): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, same, distal portion of propodus and dactylus, ventrolateral, slender setae on propodus omitted; I, fifth pereiopod, lateral; J, same, propodus and dactylus, ventromesial; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881097" httpUri="https://zenodo.org/record/5881097/files/figure.png" pageId="15" pageNumber="288">Figs. 1G, I</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A7805140510B6F68F9FFCCF1956" box="[720,793,1015,1041]" captionStart="FIGURE 4" captionStartId="9.[151,250,1839,1863]" captionTargetBox="[169,1413,191,1801]" captionTargetId="figure-15@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 4. Alpheus sciolii sp. nov., paratype, male (cl 16.0 mm) from Heron Island, Australia (MNHN-IU-2018-5525): A, frontal region, dorsal; B, same, lateral; C, telson, dorsal, plumose setae omitted; D, tooth on ventromesial carina of first article of antennular peduncle, lateral; E, third maxilliped, lateral; F, second pereiopod, lateral; G, third pereiopod, lateral; H, fifth pereiopod, lateral; I, same, propodus and dactylus, ventromesial; J, second pleopod, detail of appendices masculina and interna, anterior; K, uropod, dorsal, plumose setae omitted." figureDoi="http://doi.org/10.5281/zenodo.5881105" httpUri="https://zenodo.org/record/5881105/files/figure.png" pageId="15" pageNumber="288">4G, H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |