653 lines
89 KiB
XML
653 lines
89 KiB
XML
<document id="0205DF5B51C6E64EAA3878F6EBE4F668" ID-CLB-Dataset="7264" ID-DOI="10.11646/zootaxa.4952.3.3" ID-GBIF-Dataset="4c6980ee-d053-4d48-a8e3-9d9cd34d2a40" ID-ISSN="1175-5326" ID-Zenodo-Dep="4690551" ID-ZooBank="CEBF9AB5-1A6C-4AC5-AEEE-24EE2C142CED" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618444978806" checkinUser="plazi" docAuthor="Tsukamoto, Sho, Hiruta, Shimpei F., Eguchi, Katsuyuki, Liao, Jhih-Rong & Shimano, Satoshi" docDate="2021" docId="03BF87DCFFCBFFE7C891FADCFE602428" docLanguage="en" docName="zootaxa.4952.3.3.pdf" docOrigin="Zootaxa 4952 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scolopendra alcyona Tsukamoto & Hiruta & Eguchi & Liao & Shimano 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="489" masterDocId="FF86FFA4FFD9FFFFC806FFECFFDE254D" masterDocTitle="A new amphibious species of the genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae) from the Ryukyu Archipelago and Taiwan" masterLastPageNumber="494" masterPageNumber="465" pageNumber="483" updateTime="1698936722344" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="920D6E1C0E595DC63991D224AF6DC4DC">
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<mods:title id="7BBBA431E60CFB16D28AA8D19D6FC475">A new amphibious species of the genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae) from the Ryukyu Archipelago and Taiwan</mods:title>
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<mods:name id="46E8BB4084189B259E74210953F99E21" type="personal">
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<mods:roleTerm id="97925EDDCDEEEDF00AF6A1DE8602F304">Author</mods:roleTerm>
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<mods:namePart id="EDCB3E6768BDDFED4B0B4533D1682FA7">Tsukamoto, Sho</mods:namePart>
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<mods:affiliation id="FF573FBA360FBE8175B7BE048CEA85E4">Systematic Zoology Laboratory, Department of Biological Sciences, Graduate School of Science, Tokyo Metropolitan University, Minami-osawa 1 - 1 Hachioji-shi, Tokyo, 192 - 0397 Japan</mods:affiliation>
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<mods:roleTerm id="F782DCBB320FDAD9098AA99F4DE09EEA">Author</mods:roleTerm>
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<mods:namePart id="D5AE51090C35DED17B1014937A6BF9F2">Hiruta, Shimpei F.</mods:namePart>
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<mods:nameIdentifier id="EB3B73508724210701E604CD938929C8" type="ORCID">0000-0001-7890-1720</mods:nameIdentifier>
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<mods:affiliation id="13EDB6A08A3B1067FC2E115FFF381E06">Center for Molecular Biodiversity Research, National Museum of Nature and Science, Amakubo 4 - 4 - 1 Tsukuba-shi, Ibaraki, 305 - 0005 Japan. s-hiruta @ kahaku. go. jp; https: // orcid. org / 0000 - 0001 - 7890 - 1720</mods:affiliation>
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<mods:nameIdentifier id="F2777A375BB04A4865399FFA350B14D5" type="email">s-hiruta@kahaku.go.jp</mods:nameIdentifier>
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<mods:namePart id="AE12B3ED1058E59FC0D7906D747AA758">Eguchi, Katsuyuki</mods:namePart>
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<mods:nameIdentifier id="B55FA76140A2B7A0B961604E4D7902C4" type="ORCID">0000-0002-1054-1295</mods:nameIdentifier>
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<mods:affiliation id="F9CCC0D535E745E5B02F723FC80B7FA7">Systematic Zoology Laboratory, Department of Biological Sciences, Graduate School of Science, Tokyo Metropolitan University, Minami-osawa 1 - 1 Hachioji-shi, Tokyo, 192 - 0397 Japan & antist 2007 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1054 - 1295</mods:affiliation>
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<mods:nameIdentifier id="48213B6B77B02C57661E002786635499" type="email">antist2007@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B37EB034747B03E7FCF2326B42A20624">Liao, Jhih-Rong</mods:namePart>
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<mods:affiliation id="B2395C0F4B8C14CD44A2E5E0C9856FDA">Department of Entomology, National Taiwan University, Taipei City, 106332 Taiwan</mods:affiliation>
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<mods:namePart id="348F30A2EF3DC75E7D903F78A213F5C2">Shimano, Satoshi</mods:namePart>
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<mods:affiliation id="B58E87FE4E74F72D43F5EB4296A7E4EF">Science Research Center, Hosei University, Fujimi 2 - 17 - 1 Chiyoda-ku, Tokyo, 102 - 8160 Japan</mods:affiliation>
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<mods:title id="33862D9E8429A7BB24685ACA0803F368">Zootaxa</mods:title>
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<mods:part id="CA23F140B8F3BEBF21E0FB14F1AF21D6">
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<mods:date id="380A545BC16F14E3090AA5FE03A443D7">2021</mods:date>
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<mods:number id="42C9A24B2AF44FA910A818D6EEB4E9D6">2021-04-12</mods:number>
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<mods:number id="6475BC8F84442DC759045BA917ED66CB">4952</mods:number>
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<treatment id="03BF87DCFFCBFFE7C891FADCFE602428" ID-DOI="http://doi.org/10.5281/zenodo.4700694" ID-GBIF-Taxon="181198241" ID-Zenodo-Dep="4700694" LSID="urn:lsid:plazi:treatment:03BF87DCFFCBFFE7C891FADCFE602428" httpUri="http://treatment.plazi.org/id/03BF87DCFFCBFFE7C891FADCFE602428" lastPageId="24" lastPageNumber="489" pageId="18" pageNumber="483">
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<subSubSection id="C30C6541FFCBFFEDC891FADCFCF72006" box="[151,809,1328,1355]" pageId="18" pageNumber="483" type="nomenclature">
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<paragraph id="8BA936CAFFCBFFEDC891FADCFCF72006" blockId="18.[151,1437,1328,1459]" box="[151,809,1328,1355]" pageId="18" pageNumber="483">
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<heading id="D0E181A6FFCBFFEDC891FADCFCF72006" bold="true" box="[151,809,1328,1355]" fontSize="11" level="1" pageId="18" pageNumber="483" reason="1">
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<emphasis id="B962EAD8FFCBFFEDC891FADCFCF72006" bold="true" box="[151,809,1328,1355]" pageId="18" pageNumber="483">
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<taxonomicName id="4C164D49FFCBFFEDC891FADCFD612006" authority="Tsukamoto & Shimano" authorityName="Tsukamoto & Hiruta & Eguchi & Liao & Shimano" authorityYear="2021" box="[151,703,1328,1355]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="483" phylum="Arthropoda" rank="species" species="alcyona" status="sp. nov.">
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<emphasis id="B962EAD8FFCBFFEDC891FADCFE4B2006" bold="true" box="[151,405,1328,1355]" italics="true" pageId="18" pageNumber="483">Scolopendra alcyona</emphasis>
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Tsukamoto & Shimano
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25157A3FFCBFFEDCACDFADDFCF72006" box="[715,809,1329,1355]" pageId="18" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30C6541FFCBFFEDC891FAB8FCD320DC" pageId="18" pageNumber="483" type="reference_group">
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<paragraph id="8BA936CAFFCBFFEDC891FAB8FCD320DC" blockId="18.[151,1437,1328,1459]" pageId="18" pageNumber="483">
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[New Japanese name for
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<taxonomicName id="4C164D49FFCBFFEDC9B8FABFFD712020" authority="Tsukamoto & Hiruta & Eguchi & Liao & Shimano, 2021" authorityName="Tsukamoto & Hiruta & Eguchi & Liao & Shimano" authorityYear="2021" box="[446,687,1363,1389]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="483" phylum="Arthropoda" rank="species" species="alcyona" status="sp. nov.">
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<emphasis id="B962EAD8FFCBFFEDC9B8FABFFD712020" box="[446,687,1363,1389]" italics="true" pageId="18" pageNumber="483">Scolopendra alcyona</emphasis>
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</taxonomicName>
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<emphasis id="B962EAD8FFCBFFEDCABFFAB8FCCB2023" bold="true" box="[697,789,1364,1390]" pageId="18" pageNumber="483">
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<taxonomicNameLabel id="A25157A3FFCBFFEDCABFFAB8FCCB2023" box="[697,789,1364,1390]" pageId="18" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
: Ryûjin-ômukade (琉神大百足); new English name for
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<taxonomicName id="4C164D49FFCBFFEDC891FA9AFE5B20DD" authority="Tsukamoto & Hiruta & Eguchi & Liao & Shimano, 2021" authorityName="Tsukamoto & Hiruta & Eguchi & Liao & Shimano" authorityYear="2021" box="[151,389,1398,1424]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="483" phylum="Arthropoda" rank="species" species="alcyona" status="sp. nov.">
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||
<emphasis id="B962EAD8FFCBFFEDC891FA9AFE5B20DD" box="[151,389,1398,1424]" italics="true" pageId="18" pageNumber="483">Scolopendra alcyona</emphasis>
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</taxonomicName>
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||
<emphasis id="B962EAD8FFCBFFEDC98DFA9AFE3B20DD" bold="true" box="[395,485,1398,1424]" pageId="18" pageNumber="483">
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<taxonomicNameLabel id="A25157A3FFCBFFEDC98DFA9AFE3B20DD" box="[395,485,1398,1424]" pageId="18" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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: Halcyon giant centipede]
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C30C6541FFCBFFEDC891FA75FE8420FE" box="[151,346,1433,1459]" pageId="18" pageNumber="483" type="description">
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<paragraph id="8BA936CAFFCBFFEDC891FA75FE8420FE" blockId="18.[151,1437,1328,1459]" box="[151,346,1433,1459]" pageId="18" pageNumber="483">
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(
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<figureCitation id="132D2A4FFFCBFFEDC899FA75FEC520FE" box="[159,283,1433,1459]" captionStart="FIGURE 3" captionStartId="10.[151,250,1803,1827]" captionTargetBox="[154,1434,378,1782]" captionTargetId="figure-125@10.[150,1437,375,1786]" captionTargetPageId="10" captionText="FIGURE 3. A, Habitat of Scolopendra alcyona Tsukamoto & Shimano, sp. nov. in Kume-jima Island; B, C, S. alcyona Tsukamoto & Shimano, sp. nov., found underwater at the upper reach of Shirase River, Kume-jima Island (by Mr. H. Sato; specimen not collected)." figureDoi="http://doi.org/10.5281/zenodo.4690557" httpUri="https://zenodo.org/record/4690557/files/figure.png" pageId="18" pageNumber="483">Figs 3B, C</figureCitation>
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,
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<figureCitation id="132D2A4FFFCBFFEDC921FA75FE8C20FE" box="[295,338,1433,1459]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStart-4="FIGURE 8" captionStart-5="FIGURE 9" captionStartId-0="19.[151,250,1916,1940]" captionStartId-1="20.[151,250,1588,1612]" captionStartId-2="22.[151,250,1937,1961]" captionStartId-3="23.[151,250,1834,1858]" captionStartId-4="24.[151,260,1923,1949]" captionStartId-5="25.[151,250,1141,1165]" captionTargetBox-0="[198,1392,185,1895]" captionTargetBox-1="[154,1434,313,1568]" captionTargetBox-2="[360,1224,186,1914]" captionTargetBox-3="[167,1420,191,1803]" captionTargetBox-4="[167,1425,392,1896]" captionTargetBox-5="[166,1419,208,1115]" captionTargetId-0="figure-17@19.[194,1393,181,1899]" captionTargetId-1="figure-78@20.[151,1435,309,1572]" captionTargetId-2="figure-22@22.[357,1231,181,1920]" captionTargetId-3="figure-17@23.[151,1436,181,1817]" captionTargetId-4="figure-96@24.[151,1436,381,1908]" captionTargetId-5="figure-17@25.[158,1429,201,1124]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="22" captionTargetPageId-3="23" captionTargetPageId-4="24" captionTargetPageId-5="25" captionText-0="FIGURE 4. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., A, B, E, F: holotype TS-20180914-01, C: paratype TS- 20180301-01, D: paratype TS-20180522-02: A, cephalic plate, dorsal; B–D, variation of forcipular segment, ventral; E, tergites 14 and 15; F, sternites 14 and 15. Scale bars: 5 mm." captionText-1="FIGURE 5. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., holotype TS-20180914-01: A, posterior part of tergite 20, and tergite 21; B, ultimate leg-bearing segment, ventral; C, left coxopleuron, lateral; D, male genital segments, ventral; E, male genital segments, lateral. Legends: triangles = genital sternite 1; arrows = genital sternite 2; arrowheads = tergite of genital segment. Scale bars: 5 mm (A–C), 2.5 mm (D, E)." captionText-2="FIGURE 6. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., holotype TS-20180914-01: A, prefemora and femora of ultimate legs, dorsal; B, prefemora and femora of ultimate legs, ventral; C, tibiae, tarsi 1, tarsi 2, and pretarsi of ultimate legs, dorsal; D, tibiae tarsi 1, tarsi 2, and pretarsi of ultimate legs, ventral. Scale bars: 5 mm." captionText-3="FIGURE 7. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., A–C, E, F: paratype TS-20180301-01, D: holotype TS- 20180914-01: A, cephalic plate, basal antennal articles and tergite 1, dorsal; B, forcipular segment, ventral; C, D, variation of teeth on tooth-plates; E, tergites 9–11; F, sternites 9–11. Scale bars: 5 mm (A, B, E, F); 1 mm (C, D)." captionText-4="FIGURE 8. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., paratype TS-20180301-01: A, tergite 21 and prefemur of ultimate leg, dorsal; B, ultimate leg-bearing segment and prefemur of ultimate leg, ventral; C, left coxopleuron, lateral. Scale bars: 5 mm." captionText-5="FIGURE 9. Habitus and color variation of Scolopendra alcyona Tsukamoto & Shimano, sp. nov.: A, paratype TS-20180301- 01, habitus Okinawa-jima Island; B, paratype TS-20180522-02, Kume-jima Island; C, paratype TS-20180301-01, head and antennae; D, paratype TS-20180301-01, prefemur of ultimate legs, dorsal view (A, C, D by Mr. K. Tsukamoto)." figureDoi-0="http://doi.org/10.5281/zenodo.4690559" figureDoi-1="http://doi.org/10.5281/zenodo.4690561" figureDoi-2="http://doi.org/10.5281/zenodo.4690563" figureDoi-3="http://doi.org/10.5281/zenodo.4690565" figureDoi-4="http://doi.org/10.5281/zenodo.4690567" figureDoi-5="http://doi.org/10.5281/zenodo.4690569" httpUri-0="https://zenodo.org/record/4690559/files/figure.png" httpUri-1="https://zenodo.org/record/4690561/files/figure.png" httpUri-2="https://zenodo.org/record/4690563/files/figure.png" httpUri-3="https://zenodo.org/record/4690565/files/figure.png" httpUri-4="https://zenodo.org/record/4690567/files/figure.png" httpUri-5="https://zenodo.org/record/4690569/files/figure.png" pageId="18" pageNumber="483">4–9</figureCitation>
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||
)
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</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C30C6541FFCBFFEDC891FA32FF0622AF" pageId="18" pageNumber="483" type="materials_examined">
|
||
<paragraph id="8BA936CAFFCBFFEDC891FA32FA562356" blockId="18.[151,1437,1502,2018]" pageId="18" pageNumber="483">
|
||
<emphasis id="B962EAD8FFCBFFEDC891FA32FEA120B5" bold="true" box="[151,383,1502,1528]" pageId="18" pageNumber="483">Material examined.</emphasis>
|
||
<materialsCitation id="3B7E3C97FFCBFFEDC982FA32FA562356" ID-GBIF-Occurrence="3080383303" collectingDate="2018-09-14" collectorName="Hutoshi Taira." country="Japan" county="Okinawa-jima Island" latitude="26.817945" location="Male" longLatPrecision="1" longitude="128.28119" municipality="Kunigami-gun" pageId="18" pageNumber="483" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
|
||
<emphasis id="B962EAD8FFCBFFEDC982FA32FE2420B5" bold="true" box="[388,506,1502,1528]" pageId="18" pageNumber="483">
|
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<typeStatus id="54AD8868FFCBFFEDC982FA32FE2B20B5" box="[388,501,1502,1528]" pageId="18" pageNumber="483" type="holotype">Holotype</typeStatus>
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||
:
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||
</emphasis>
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||
<location id="8EC96011FFCBFFEDC9F9FA32FDE720B5" LSID="urn:lsid:plazi:treatment:03BF87DCFFCBFFE7C891FADCFE602428:8EC96011FFCBFFEDC9F9FA32FDE720B5" box="[511,569,1502,1528]" country="Japan" county="Okinawa-jima Island" latitude="26.817945" longLatPrecision="1" longitude="128.28119" municipality="Kunigami-gun" name="Male" pageId="18" pageNumber="483" stateProvince="Okinawa">Male</location>
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||
(NSMT-My 511, TS-20180914-01),
|
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<location id="8EC96011FFCBFFEDCBD7FA32FBDA20B5" LSID="urn:lsid:plazi:treatment:03BF87DCFFCBFFE7C891FADCFE602428:8EC96011FFCBFFEDCBD7FA32FBDA20B5" box="[977,1028,1502,1528]" country="Japan" county="Okinawa-jima Island" latitude="26.817945" longLatPrecision="1" longitude="128.28119" municipality="Kunigami-gun" name="Oku" pageId="18" pageNumber="483" stateProvince="Okinawa">Oku</location>
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,
|
||
<location id="8EC96011FFCBFFEDCC0BFA32FB6E20B5" LSID="urn:lsid:plazi:treatment:03BF87DCFFCBFFE7C891FADCFE602428:8EC96011FFCBFFEDCC0BFA32FB6E20B5" box="[1037,1200,1502,1528]" country="Japan" county="Okinawa-jima Island" latitude="26.817945" longLatPrecision="1" longitude="128.28119" municipality="Kunigami-gun" name="Kunigami-son" pageId="18" pageNumber="483" stateProvince="Okinawa">Kunigami-son</location>
|
||
,
|
||
<collectingMunicipality id="6BCDACB0FFCBFFEDCCBFFA32FA8120B5" box="[1209,1375,1502,1528]" pageId="18" pageNumber="483">Kunigami-gun</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62C84E46FFCBFFEDCD6FFA32FE852356" pageId="18" pageNumber="483">Okinawa-jima Island</collectingCounty>
|
||
,
|
||
<collectingRegion id="49D2F828FFCBFFEDC961F9EDFD962356" box="[359,584,1537,1563]" country="Japan" name="Okinawa" pageId="18" pageNumber="483">Okinawa prefecture</collectingRegion>
|
||
,
|
||
<collectingCountry id="F301765AFFCBFFEDCA54F9EDFD4C2356" box="[594,658,1537,1563]" name="Japan" pageId="18" pageNumber="483">Japan</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE22500DFFCBFFEDCAA7F9EDFC9D2356" box="[673,835,1537,1563]" degrees="26" direction="north" minutes="49" orientation="latitude" pageId="18" pageNumber="483" precision="1" seconds="04.6" value="26.817945">26°49’04.6”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE22500DFFCBFFEDCB49F9EDFC242351" box="[847,1018,1537,1564]" degrees="128" direction="east" minutes="16" orientation="longitude" pageId="18" pageNumber="483" precision="1" seconds="52.3" value="128.28119">128°16’52.3”E</geoCoordinate>
|
||
),
|
||
<date id="FFA8100AFFCBFFEDCC09F9EDFB7B2356" box="[1039,1189,1537,1563]" pageId="18" pageNumber="483" value="2018-09-14">
|
||
<collectingDate id="EFECE9E2FFCBFFEDCC09F9EDFB7B2356" box="[1039,1189,1537,1563]" pageId="18" pageNumber="483" value="2018-09-14">14 Sep. 2018</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="26E3531CFFCBFFEDCCEEF9EDFA562356" box="[1256,1416,1537,1563]" pageId="18" pageNumber="483">Hutoshi Taira.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFCBFFEDC8C1F9C8FED523A6" blockId="18.[151,1437,1502,2018]" pageId="18" pageNumber="483">
|
||
<materialsCitation id="3B7E3C97FFCBFFEDC8C1F9C8FBC6232D" ID-GBIF-Occurrence="3080383304" collectingDate="2018-09-12" collectorName="Katsuya Tsukamoto" country="Japan" county="Okinawajima Island" location="Yona" municipality="Kunigami-gun" pageId="18" pageNumber="483" specimenCount="1" stateProvince="Okinawa" typeStatus="paratype">
|
||
<emphasis id="B962EAD8FFCBFFEDC8C1F9C8FE962373" bold="true" box="[199,328,1572,1598]" pageId="18" pageNumber="483">
|
||
<typeStatus id="54AD8868FFCBFFEDC8C1F9C8FE9D2373" box="[199,323,1572,1598]" pageId="18" pageNumber="483" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
sex unknown (NSMT-My 512, TS-20180912-01),
|
||
<location id="8EC96011FFCBFFEDCB80F9C8FC1D2373" LSID="urn:lsid:plazi:treatment:03BF87DCFFCBFFE7C891FADCFE602428:8EC96011FFCBFFEDCB80F9C8FC1D2373" box="[902,963,1572,1598]" country="Japan" county="Okinawajima Island" municipality="Kunigami-gun" name="Yona" pageId="18" pageNumber="483" stateProvince="Okinawa">Yona</location>
|
||
,
|
||
<location id="8EC96011FFCBFFEDCBC8F9CFFBAF2370" LSID="urn:lsid:plazi:treatment:03BF87DCFFCBFFE7C891FADCFE602428:8EC96011FFCBFFEDCBC8F9CFFBAF2370" box="[974,1137,1571,1597]" country="Japan" county="Okinawajima Island" municipality="Kunigami-gun" name="Kunigami-son" pageId="18" pageNumber="483" stateProvince="Okinawa">Kunigami-son</location>
|
||
,
|
||
<collectingMunicipality id="6BCDACB0FFCBFFEDCC7AF9CFFAFC2370" box="[1148,1314,1571,1597]" pageId="18" pageNumber="483">Kunigami-gun</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62C84E46FFCBFFEDCD28F9CFFEC9232D" pageId="18" pageNumber="483">Okinawajima Island</collectingCounty>
|
||
,
|
||
<collectingRegion id="49D2F828FFCBFFEDC925F9AAFDDA232D" box="[291,516,1606,1632]" country="Japan" name="Okinawa" pageId="18" pageNumber="483">Okinawa prefecture</collectingRegion>
|
||
,
|
||
<collectingCountry id="F301765AFFCBFFEDCA08F9ABFD8F232C" box="[526,593,1607,1633]" name="Japan" pageId="18" pageNumber="483">Japan</collectingCountry>
|
||
,
|
||
<date id="FFA8100AFFCBFFEDCA5AF9AAFD2C232D" box="[604,754,1606,1632]" pageId="18" pageNumber="483" value="2018-09-12">
|
||
<collectingDate id="EFECE9E2FFCBFFEDCA5AF9AAFD2C232D" box="[604,754,1606,1632]" pageId="18" pageNumber="483" value="2018-09-12">12 Sep. 2018</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="26E3531CFFCBFFEDCB30F9ABFBC6232D" box="[822,1048,1606,1633]" pageId="18" pageNumber="483">Katsuya Tsukamoto</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B7E3C97FFCBFFEDCC25F9ABFE1F23E8" ID-GBIF-Occurrence="3080383301" collectingDate="2018-03-01" collectorName="Takeshi Sasaki" country="Japan" county="Okinawa-jima Island" location="Yona" municipality="Kunigami-gun" pageId="18" pageNumber="483" specimenCount="1" stateProvince="Okinawa" typeStatus="paratype">
|
||
sex unknown (RUMF-ZD-02001, TS-20180301-01),
|
||
<location id="8EC96011FFCBFFEDC968F985FE7523CE" LSID="urn:lsid:plazi:treatment:03BF87DCFFCBFFE7C891FADCFE602428:8EC96011FFCBFFEDC968F985FE7523CE" box="[366,427,1641,1667]" country="Japan" county="Okinawa-jima Island" municipality="Kunigami-gun" name="Yona" pageId="18" pageNumber="483" stateProvince="Okinawa">Yona</location>
|
||
,
|
||
<location id="8EC96011FFCBFFEDC9B1F985FD8423CE" LSID="urn:lsid:plazi:treatment:03BF87DCFFCBFFE7C891FADCFE602428:8EC96011FFCBFFEDC9B1F985FD8423CE" box="[439,602,1641,1667]" country="Japan" county="Okinawa-jima Island" municipality="Kunigami-gun" name="Kunigami-son" pageId="18" pageNumber="483" stateProvince="Okinawa">Kunigami-son</location>
|
||
,
|
||
<collectingMunicipality id="6BCDACB0FFCBFFEDCA60F985FCD223CE" box="[614,780,1641,1667]" pageId="18" pageNumber="483">Kunigami-gun</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62C84E46FFCBFFEDCB1FF985FBD623CE" box="[793,1032,1641,1667]" pageId="18" pageNumber="483">Okinawa-jima Island</collectingCounty>
|
||
,
|
||
<collectingRegion id="49D2F828FFCBFFEDCC13F985FB2923CE" box="[1045,1271,1641,1667]" country="Japan" name="Okinawa" pageId="18" pageNumber="483">Okinawa prefecture</collectingRegion>
|
||
,
|
||
<collectingCountry id="F301765AFFCBFFEDCD05F985FA9823CE" box="[1283,1350,1641,1667]" name="Japan" pageId="18" pageNumber="483">Japan</collectingCountry>
|
||
,
|
||
<date id="FFA8100AFFCBFFEDCD54F985FF0C23EB" pageId="18" pageNumber="483" value="2018-03-01">
|
||
<collectingDate id="EFECE9E2FFCBFFEDCD54F985FF0C23EB" pageId="18" pageNumber="483" value="2018-03-01">1 Mar. 2018</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="26E3531CFFCBFFEDC910F967FE1F23E8" box="[278,449,1675,1701]" pageId="18" pageNumber="483">Takeshi Sasaki</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B7E3C97FFCBFFEDC9CAF960FED523A6" ID-GBIF-Occurrence="3080383302" collectingDate="2018-05-22" collectorName="Humiyasu Sato." country="Japan" county="Kume-jima Island" location="Nakachi" municipality="Shimajiri-gun" pageId="18" pageNumber="483" specimenCount="1" stateProvince="Okinawa" typeStatus="paratype">
|
||
2 sex unknown (NSMT-My 513, TS-20180522-01; NSMT-My 514, TS-20180522-02),
|
||
<location id="8EC96011FFCBFFEDC891F942FF292385" LSID="urn:lsid:plazi:treatment:03BF87DCFFCBFFE7C891FADCFE602428:8EC96011FFCBFFEDC891F942FF292385" box="[151,247,1710,1736]" country="Japan" county="Kume-jima Island" municipality="Shimajiri-gun" name="Nakachi" pageId="18" pageNumber="483" stateProvince="Okinawa">Nakachi</location>
|
||
,
|
||
<location id="8EC96011FFCBFFEDC905F942FE702385" LSID="urn:lsid:plazi:treatment:03BF87DCFFCBFFE7C891FADCFE602428:8EC96011FFCBFFEDC905F942FE702385" box="[259,430,1710,1736]" country="Japan" county="Kume-jima Island" municipality="Shimajiri-gun" name="Kumejima-cho" pageId="18" pageNumber="483" stateProvince="Okinawa">Kumejima-cho</location>
|
||
,
|
||
<collectingMunicipality id="6BCDACB0FFCBFFEDC9BFF942FD862385" box="[441,600,1710,1736]" pageId="18" pageNumber="483">Shimajiri-gun</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62C84E46FFCBFFEDCA65F942FCEF2385" box="[611,817,1710,1736]" pageId="18" pageNumber="483">Kume-jima Island</collectingCounty>
|
||
,
|
||
<collectingRegion id="49D2F828FFCBFFEDCB3AF942FBC02385" box="[828,1054,1710,1736]" country="Japan" name="Okinawa" pageId="18" pageNumber="483">Okinawa prefecture</collectingRegion>
|
||
,
|
||
<collectingCountry id="F301765AFFCBFFEDCC2FF943FBB22384" box="[1065,1132,1711,1737]" name="Japan" pageId="18" pageNumber="483">Japan</collectingCountry>
|
||
,
|
||
<date id="FFA8100AFFCBFFEDCC71F942FACE2385" box="[1143,1296,1710,1737]" pageId="18" pageNumber="483" value="2018-05-22">
|
||
<collectingDate id="EFECE9E2FFCBFFEDCC71F942FACE2385" box="[1143,1296,1710,1737]" pageId="18" pageNumber="483" value="2018-05-22">22 May 2018</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="26E3531CFFCBFFEDCD55F942FED523A6" pageId="18" pageNumber="483">Humiyasu Sato.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFCBFFEDC8C1F91FFCC6227C" blockId="18.[151,1437,1502,2018]" pageId="18" pageNumber="483">
|
||
All type specimens except TS-20180301-01 are deposited at the Collection of Myriapoda, Department of Zoology,
|
||
<collectionCode id="ED07AE0FFFCBFFEDC8CBF8FAFEC2227D" box="[205,284,1814,1840]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="18" pageNumber="483">NSMT</collectionCode>
|
||
, and TS-20180301-01 is deposited at
|
||
<collectionCode id="ED07AE0FFFCBFFEDCAC2F8FBFCCB227C" box="[708,789,1815,1841]" pageId="18" pageNumber="483">RUMF</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFCBFFEDC8C1F8D5FF0622AF" blockId="18.[151,1437,1502,2018]" pageId="18" pageNumber="483">
|
||
<emphasis id="B962EAD8FFCBFFEDC8C1F8D5FE1E221E" bold="true" box="[199,448,1849,1875]" pageId="18" pageNumber="483">Non-type specimens:</emphasis>
|
||
<materialsCitation id="3B7E3C97FFCBFFEDC9C1F8D5FCD622F3" ID-GBIF-Occurrence="3080383305" collectingDate="1969-08-14" collectionCode="NTU, NSMT" collectorName="Zento Touyama & Mitsuo Takano." country="Japan" county="Nuha" location="Kunigami-gun" municipality="Ogimi-son" pageId="18" pageNumber="483" specimenCount="1" stateProvince="Pingtung">
|
||
sex unknown (NSMT-My 515, TS-2018MMDD-01),
|
||
<collectingCounty id="62C84E46FFCBFFEDCC27F8D5FBBE221E" box="[1057,1120,1849,1875]" pageId="18" pageNumber="483">Nuha</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BCDACB0FFCBFFEDCC6AF8D5FB39221E" box="[1132,1255,1849,1875]" pageId="18" pageNumber="483">Ogimi-son</collectingMunicipality>
|
||
,
|
||
<location id="8EC96011FFCBFFEDCCF4F8D5FA46221E" LSID="urn:lsid:plazi:treatment:03BF87DCFFCBFFE7C891FADCFE602428:8EC96011FFCBFFEDCCF4F8D5FA46221E" box="[1266,1432,1849,1875]" country="Japan" county="Nuha" municipality="Ogimi-son" name="Kunigami-gun" pageId="18" pageNumber="483" stateProvince="Pingtung">Kunigami-gun</location>
|
||
,
|
||
<location id="8EC96011FFCBFFEDC891F8B7FE572238" LSID="urn:lsid:plazi:treatment:03BF87DCFFCBFFE7C891FADCFE602428:8EC96011FFCBFFEDC891F8B7FE572238" box="[151,393,1883,1909]" country="Japan" county="Nuha" municipality="Ogimi-son" name="Okinawa-jima Island" pageId="18" pageNumber="483" stateProvince="Pingtung">Okinawa-jima Island</location>
|
||
, Okinawa prefecture,
|
||
<collectingCountry id="F301765AFFCBFFEDCA8FF8B0FD12223B" box="[649,716,1884,1910]" name="Japan" pageId="18" pageNumber="483">Japan</collectingCountry>
|
||
, 2018, coll.
|
||
<collectorName id="26E3531CFFCBFFEDCB59F8B0FBC8223B" box="[863,1046,1884,1910]" pageId="18" pageNumber="483">Zento Touyama</collectorName>
|
||
;
|
||
<location id="8EC96011FFCBFFEDCC25F8B7FBAB2238" LSID="urn:lsid:plazi:treatment:03BF87DCFFCBFFE7C891FADCFE602428:8EC96011FFCBFFEDCC25F8B7FBAB2238" box="[1059,1141,1883,1909]" country="Japan" county="Nuha" municipality="Ogimi-son" name="Female" pageId="18" pageNumber="483" stateProvince="Pingtung">Female</location>
|
||
(
|
||
<collectionCode id="ED07AE0FFFCBFFEDCC8EF8B0FB1D223B" box="[1160,1219,1884,1910]" pageId="18" pageNumber="483">NTU</collectionCode>
|
||
TS-19690814-01),
|
||
<location id="8EC96011FFCBFFEDC891F893FED322D4" LSID="urn:lsid:plazi:treatment:03BF87DCFFCBFFE7C891FADCFE602428:8EC96011FFCBFFEDC891F893FED322D4" box="[151,269,1919,1945]" country="Japan" county="Nuha" municipality="Ogimi-son" name="Hengchun" pageId="18" pageNumber="483" stateProvince="Pingtung">Hengchun</location>
|
||
,
|
||
<collectingRegion id="49D2F828FFCBFFEDC91EF893FE6B22D4" box="[280,437,1919,1945]" country="Taiwan" name="Pingtung" pageId="18" pageNumber="483">Pingtung City</collectingRegion>
|
||
,
|
||
<collectingCountry id="F301765AFFCBFFEDC9C6F893FDCB22D4" box="[448,533,1919,1945]" name="Taiwan" pageId="18" pageNumber="483">Taiwan</collectingCountry>
|
||
,
|
||
<date id="FFA8100AFFCBFFEDCA26F86CFD6422D4" box="[544,698,1919,1946]" pageId="18" pageNumber="483" value="1969-08-14">
|
||
<collectingDate id="EFECE9E2FFCBFFEDCA26F86CFD6422D4" box="[544,698,1919,1946]" pageId="18" pageNumber="483" value="1969-08-14">14 Aug. 1969</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="26E3531CFFCBFFEDCAFBF893FC7222D4" box="[765,940,1919,1945]" pageId="18" pageNumber="483">Mitsuo Takano.</collectorName>
|
||
TS-2018MMDD-01 is deposited at the
|
||
<location id="8EC96011FFCBFFEDCD6CF893FE5322F0" LSID="urn:lsid:plazi:treatment:03BF87DCFFCBFFE7C891FADCFE602428:8EC96011FFCBFFEDCD6CF893FE5322F0" country="Japan" county="Nuha" municipality="Ogimi-son" name="Collection of Myriapoda" pageId="18" pageNumber="483" stateProvince="Pingtung">Collection of Myriapoda</location>
|
||
,
|
||
<location id="8EC96011FFCBFFEDC99DF848FD7522F0" LSID="urn:lsid:plazi:treatment:03BF87DCFFCBFFE7C891FADCFE602428:8EC96011FFCBFFEDC99DF848FD7522F0" box="[411,683,1955,1982]" country="Japan" county="Nuha" municipality="Ogimi-son" name="Department of Zoology" pageId="18" pageNumber="483" stateProvince="Pingtung">Department of Zoology</location>
|
||
,
|
||
<collectionCode id="ED07AE0FFFCBFFEDCABFF848FCD622F3" box="[697,776,1956,1982]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="18" pageNumber="483">NSMT</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B7E3C97FFCBFFEDCB10F84FFF0A22AF" ID-GBIF-Occurrence="3080383306" collectionCode="NTU" collectorName="Taiwanese" country="Japan" pageId="18" pageNumber="483" specimenCount="1" stateProvince="Pingtung">
|
||
<collectorName id="26E3531CFFCBFFEDCB10F84FFC5222F0" box="[790,908,1955,1981]" pageId="18" pageNumber="483">Taiwanese</collectorName>
|
||
specimen (TS-19690814-01) are deposited at
|
||
<collectionCode id="ED07AE0FFFCBFFEDC891F824FF0A22AF" box="[151,212,1992,2018]" pageId="18" pageNumber="483">NTU</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF696642FFCAFFECC891F890FD722283" ID-DOI="http://doi.org/10.5281/zenodo.4690559" ID-Zenodo-Dep="4690559" httpUri="https://zenodo.org/record/4690559/files/figure.png" pageId="19" pageNumber="484" startId="19.[151,250,1916,1940]" targetBox="[198,1392,185,1895]" targetPageId="19">
|
||
<paragraph id="8BA936CAFFCAFFECC891F890FD722283" blockId="19.[151,1436,1916,1999]" pageId="19" pageNumber="484">
|
||
<emphasis id="B962EAD8FFCAFFECC891F890FECB22D9" bold="true" box="[151,277,1916,1940]" pageId="19" pageNumber="484">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C164D49FFCAFFECC91BF890FD3122D9" authority="Tsukamoto & Shimano" authorityName="Tsukamoto & Hiruta & Eguchi & Liao & Shimano" authorityYear="2021" box="[285,751,1916,1940]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="484" phylum="Arthropoda" rank="species" species="alcyona" status="sp. nov.">
|
||
<emphasis id="B962EAD8FFCAFFECC91BF890FE2822D9" box="[285,502,1916,1940]" italics="true" pageId="19" pageNumber="484">Scolopendra alcyona</emphasis>
|
||
Tsukamoto & Shimano
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B962EAD8FFCAFFECCAFDF890FC9322D9" bold="true" box="[763,845,1916,1940]" pageId="19" pageNumber="484">
|
||
<taxonomicNameLabel id="A25157A3FFCAFFECCAFDF890FC9322D9" box="[763,845,1916,1940]" pageId="19" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, A, B, E, F: holotype
|
||
<emphasis id="B962EAD8FFCAFFECCC3EF890FB3822D9" bold="true" box="[1080,1254,1916,1940]" pageId="19" pageNumber="484">TS-20180914-01</emphasis>
|
||
, C: paratype
|
||
<emphasis id="B962EAD8FFCAFFECCD73F890FECC22FC" bold="true" pageId="19" pageNumber="484">TS- 20180301-0</emphasis>
|
||
1, D: paratype
|
||
<emphasis id="B962EAD8FFCAFFECC9A0F875FD8B22FC" bold="true" box="[422,597,1945,1969]" pageId="19" pageNumber="484">TS-20180522-02</emphasis>
|
||
:
|
||
<emphasis id="B962EAD8FFCAFFECCA67F875FDAD22FC" bold="true" box="[609,627,1945,1969]" pageId="19" pageNumber="484">A</emphasis>
|
||
, cephalic plate, dorsal;
|
||
<emphasis id="B962EAD8FFCAFFECCB64F876FC4F22FF" bold="true" box="[866,913,1946,1970]" pageId="19" pageNumber="484">B–D</emphasis>
|
||
, variation of forcipular segment, ventral;
|
||
<emphasis id="B962EAD8FFCAFFECCD33F876FA9822FF" bold="true" box="[1333,1350,1946,1970]" pageId="19" pageNumber="484">E</emphasis>
|
||
, tergites 14 and 15;
|
||
<emphasis id="B962EAD8FFCAFFECC90EF85BFEC92282" bold="true" box="[264,279,1975,1999]" pageId="19" pageNumber="484">F</emphasis>
|
||
, sternites 14 and 15.
|
||
<emphasis id="B962EAD8FFCAFFECC9F6F85BFD812282" bold="true" box="[496,607,1975,1999]" pageId="19" pageNumber="484">Scale bars</emphasis>
|
||
: 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30C6541FFCDFFEBC8C1FF7BFEB92450" pageId="20" pageNumber="485" type="etymology">
|
||
<paragraph id="8BA936CAFFCDFFEBC8C1FF7BFEB92450" blockId="20.[151,1437,151,285]" pageId="20" pageNumber="485">
|
||
<emphasis id="B962EAD8FFCDFFEBC8C1FF7BFE9025FC" bold="true" box="[199,334,151,177]" pageId="20" pageNumber="485">Etymology.</emphasis>
|
||
This epithet “alcyona” derived from the name of a Greek mythological figure, “Alcyone”. In one version of the story, Alcyone was turned into a common kingfisher (halcyon bird) by Zeus (
|
||
<bibRefCitation id="EF874B3BFFCDFFEBCCA8FF57FA88259B" author="Gresseth, G. K." box="[1198,1366,187,214]" pageId="20" pageNumber="485" pagination="88 - 98" refId="ref13831" refString="Gresseth, G. K. (1964) The myth of alcyone. Transactions and Proceedings of the American Philological Association, 95, 88 - 98. https: // doi. org / 10.2307 / 283785" type="journal article" year="1964">Gresseth 1964</bibRefCitation>
|
||
). This epithet is a metaphor for halcyon bird-like jade green legs seen in the Okinawa-jima Island population, and its amphibious behavior.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF696642FFCDFFEBC891F9D8FDA423E9" ID-DOI="http://doi.org/10.5281/zenodo.4690561" ID-Zenodo-Dep="4690561" httpUri="https://zenodo.org/record/4690561/files/figure.png" pageId="20" pageNumber="485" startId="20.[151,250,1588,1612]" targetBox="[154,1434,313,1568]" targetPageId="20">
|
||
<paragraph id="8BA936CAFFCDFFEBC891F9D8FDA423E9" blockId="20.[151,1436,1588,1700]" pageId="20" pageNumber="485">
|
||
<emphasis id="B962EAD8FFCDFFEBC891F9D8FECD2301" bold="true" box="[151,275,1588,1612]" pageId="20" pageNumber="485">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C164D49FFCDFFEBC91CF9D8FD382301" authority="Tsukamoto & Shimano" authorityName="Tsukamoto & Hiruta & Eguchi & Liao & Shimano" authorityYear="2021" box="[282,742,1588,1612]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="485" phylum="Arthropoda" rank="species" species="alcyona" status="sp. nov.">
|
||
<emphasis id="B962EAD8FFCDFFEBC91CF9D8FE2F2301" box="[282,497,1588,1612]" italics="true" pageId="20" pageNumber="485">Scolopendra alcyona</emphasis>
|
||
Tsukamoto & Shimano
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B962EAD8FFCDFFEBCAF7F9D8FC9C2301" bold="true" box="[753,834,1588,1612]" pageId="20" pageNumber="485">
|
||
<taxonomicNameLabel id="A25157A3FFCDFFEBCAF7F9D8FC9C2301" box="[753,834,1588,1612]" pageId="20" pageNumber="485" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype
|
||
<emphasis id="B962EAD8FFCDFFEBCBA9F9D8FB832301" bold="true" box="[943,1117,1588,1612]" pageId="20" pageNumber="485">TS-20180914-01</emphasis>
|
||
:
|
||
<emphasis id="B962EAD8FFCDFFEBCC6DF9D8FBA32301" bold="true" box="[1131,1149,1588,1612]" pageId="20" pageNumber="485">A</emphasis>
|
||
, posterior part of tergite 20, and tergite 21;
|
||
<emphasis id="B962EAD8FFCDFFEBC928F9BEFEE12327" bold="true" box="[302,319,1618,1642]" pageId="20" pageNumber="485">B</emphasis>
|
||
, ultimate leg-bearing segment, ventral;
|
||
<emphasis id="B962EAD8FFCDFFEBCAD4F9BDFD3A2324" bold="true" box="[722,740,1617,1641]" pageId="20" pageNumber="485">C</emphasis>
|
||
, left coxopleuron, lateral;
|
||
<emphasis id="B962EAD8FFCDFFEBCBE9F9BEFBDF2327" bold="true" box="[1007,1025,1618,1642]" pageId="20" pageNumber="485">D</emphasis>
|
||
, male genital segments, ventral;
|
||
<emphasis id="B962EAD8FFCDFFEBCD48F9BEFA812327" bold="true" box="[1358,1375,1618,1642]" pageId="20" pageNumber="485">E</emphasis>
|
||
, male genital segments, lateral. Legends: triangles = genital sternite 1; arrows = genital sternite 2; arrowheads = tergite of genital segment.
|
||
<emphasis id="B962EAD8FFCDFFEBC8D1F960FE9B23E9" bold="true" box="[215,325,1676,1700]" pageId="20" pageNumber="485">Scale bars</emphasis>
|
||
: 5 mm (A–C), 2.5 mm (D, E).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30C6541FFCDFFEAC8C1F93EFC0D24C4" lastPageId="21" lastPageNumber="486" pageId="20" pageNumber="485" type="diagnosis">
|
||
<paragraph id="8BA936CAFFCDFFEAC8C1F93EFC0D24C4" blockId="20.[151,1437,1746,2025]" lastBlockId="21.[151,1437,151,1906]" lastPageId="21" lastPageNumber="486" pageId="20" pageNumber="485">
|
||
<emphasis id="B962EAD8FFCDFFEBC8C1F93EFE9C23A1" bold="true" box="[199,322,1746,1772]" pageId="20" pageNumber="485">Diagnosis.</emphasis>
|
||
17–20 antennal articles, basal 6 articles glabrous; cephalic plate with small puncta; each tooth-plate with 4–9 teeth; tergites (4, 5) 6–20 with complete paramedian sutures; tergite of ultimate leg-bearing segment without median suture; sternites 2–19 with well-defined, incomplete paramedian suture; coxopleural process with 2 apical spines and 1 subapical spine; ultimate legs long and slender with ratio of width and length of prefemur 1:3, ratio of width and length of femur 1:4 and ratios of lengths of prefemur and femur 1.06:1; femur and tibia 1.13:1; tibia and tarsus 1 1.33:1; tarsus 1 and tarsus 2 2.15:1; dorsally flattened ultimate leg prefemora with 2 VL, 1 M, 2 DM and 1–3 SP; tarsal spurs on legs 1–19; gonopods absent, penis present.
|
||
<taxonomicName id="4C164D49FFCDFFEBCBA1F846FB4A2289" authority="Tsukamoto & Hiruta & Eguchi & Liao & Shimano, 2021" authorityName="Tsukamoto & Hiruta & Eguchi & Liao & Shimano" authorityYear="2021" box="[935,1172,1962,1988]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="485" phylum="Arthropoda" rank="species" species="alcyona" status="sp. nov.">
|
||
<emphasis id="B962EAD8FFCDFFEBCBA1F846FB4A2289" box="[935,1172,1962,1988]" italics="true" pageId="20" pageNumber="485">Scolopendra alcyona</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAD8FFCDFFEBCC9CF847FB2C2288" bold="true" box="[1178,1266,1963,1989]" pageId="20" pageNumber="485">
|
||
<taxonomicNameLabel id="A25157A3FFCDFFEBCC9CF847FB2C2288" box="[1178,1266,1963,1989]" pageId="20" pageNumber="485" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to all other congeners in East and Southeast Asia (see “Morphological comparison among Japanese and Taiwanese
|
||
<emphasis id="B962EAD8FFCDFFEACD53F822FF3825FC" italics="true" lastPageId="21" lastPageNumber="486" pageId="20" pageNumber="485">Scolopendra</emphasis>
|
||
”, and
|
||
<tableCitation id="C6940371FFCCFFEAC929FF7BFE5225FC" box="[303,396,151,177]" captionStart="TABLE 6" captionStartId="15.[151,239,605,631]" captionText="TABLE 6. Morphological comparison between Scolopendra alcyona Tsukamoto & Shimano, sp. nov. and related species from East and Southeast Asia. Abbreviations: AP, apical spine; SAP, subapical spine; VL, ventro-lateral spine; VM, ventro-median spine; M, median spine; DM, dorso-median spine; SP, spine on prefemoral process; n/a, not available. Characteristics of the Chinese and Japanese populations of S. subspinipes examined by Siriwut et al. (2016) are included in the row of S. mutilans because Siriwut et al. (2016) mentioned all morphological characteristics of S. mutilans was correspond to S. subspinipes. References: 1 = Koch (1878); 2 = Kraepelin (1903); 3 = Attems (1930); 4 = Takakuwa (1940); 5 = Chao (2008); 6 = Siriwut et al. (2016)." pageId="21" pageNumber="486">Tables 6</tableCitation>
|
||
and
|
||
<tableCitation id="C6940371FFCCFFEAC9C7FF7BFE0F25FC" box="[449,465,151,177]" captionStart="TABLE 7" captionStartId="17.[151,241,468,494]" captionText="TABLE 7. Morphological comparison of Scolopendra alcyona Tsukamoto & Shimano, sp. nov. with two similar species. Abbreviations: AP, apical spine; SAP, subapical spine; VL, ventro-lateral spine; M, median spine; DM, dorso-median spine; SP, spine on prefemoral process." pageId="21" pageNumber="486">7</tableCitation>
|
||
).
|
||
<taxonomicName id="4C164D49FFCCFFEAC9E3FF7BFD8425FC" authority="Tsukamoto & Hiruta & Eguchi & Liao & Shimano, 2021" authorityName="Tsukamoto & Hiruta & Eguchi & Liao & Shimano" authorityYear="2021" box="[485,602,151,177]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="486" phylum="Arthropoda" rank="species" species="alcyona" status="sp. nov.">
|
||
<emphasis id="B962EAD8FFCCFFEAC9E3FF7BFD8425FC" box="[485,602,151,177]" italics="true" pageId="21" pageNumber="486">S. alcyona</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAD8FFCCFFEACA66FF7BFD6725FC" bold="true" box="[608,697,151,177]" pageId="21" pageNumber="486">
|
||
<taxonomicNameLabel id="A25157A3FFCCFFEACA66FF7BFD6725FC" box="[608,697,151,177]" pageId="21" pageNumber="486" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="4C164D49FFCCFFEACBE0FF7BFBBD25FD" authorityName="Koch" authorityYear="1878" box="[998,1123,151,177]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="486" phylum="Arthropoda" rank="species" species="japonica">
|
||
<emphasis id="B962EAD8FFCCFFEACBE0FF7BFBBD25FD" box="[998,1123,151,177]" italics="true" pageId="21" pageNumber="486">S. japonica</emphasis>
|
||
</taxonomicName>
|
||
by the absence gonopods in the male, and incomplete paramedian sutures.
|
||
<taxonomicName id="4C164D49FFCCFFEACA91FF57FCD52598" authority="Tsukamoto & Hiruta & Eguchi & Liao & Shimano, 2021" authorityName="Tsukamoto & Hiruta & Eguchi & Liao & Shimano" authorityYear="2021" box="[663,779,187,213]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="486" phylum="Arthropoda" rank="species" species="alcyona" status="sp. nov.">
|
||
<emphasis id="B962EAD8FFCCFFEACA91FF57FCD52598" box="[663,779,187,213]" italics="true" pageId="21" pageNumber="486">S. alcyona</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAD8FFCCFFEACB16FF57FCB62598" bold="true" box="[784,872,187,213]" pageId="21" pageNumber="486">
|
||
<taxonomicNameLabel id="A25157A3FFCCFFEACB16FF57FCB62598" box="[784,872,187,213]" pageId="21" pageNumber="486" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="4C164D49FFCCFFEACC94FF57FAC42598" authorityName="Newport" authorityYear="1844" box="[1170,1306,187,213]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="486" phylum="Arthropoda" rank="species" species="multidens">
|
||
<emphasis id="B962EAD8FFCCFFEACC94FF57FAC42598" box="[1170,1306,187,213]" italics="true" pageId="21" pageNumber="486">S. multidens</emphasis>
|
||
</taxonomicName>
|
||
by the ratio of the length of femur and tibia, and tibia and tarsus (femur and tibia 1.1:1; tibia and tarsus 1 1.3:
|
||
<quantity id="4CEE9B2FFFCCFFEACCD4FF33FB2725B4" box="[1234,1273,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="21" pageNumber="486" unit="in" value="1.0">1 in</quantity>
|
||
<taxonomicName id="4C164D49FFCCFFEACD06FF33FAAA25B4" authority="Tsukamoto & Hiruta & Eguchi & Liao & Shimano, 2021" authorityName="Tsukamoto & Hiruta & Eguchi & Liao & Shimano" authorityYear="2021" box="[1280,1396,223,249]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="486" phylum="Arthropoda" rank="species" species="alcyona" status="sp. nov.">
|
||
<emphasis id="B962EAD8FFCCFFEACD06FF33FAAA25B4" box="[1280,1396,223,249]" italics="true" pageId="21" pageNumber="486">S. alcyona</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAD8FFCCFFEACD7DFF33FF162450" bold="true" pageId="21" pageNumber="486">
|
||
<taxonomicNameLabel id="A25157A3FFCCFFEACD7DFF33FF162450" pageId="21" pageNumber="486" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; femur and tibia 1.7:1, tibia and tarsus 1 1.8:
|
||
<quantity id="4CEE9B2FFFCCFFEACABAFEEFFD3D2450" box="[700,739,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="21" pageNumber="486" unit="in" value="1.0">1 in</quantity>
|
||
<taxonomicName id="4C164D49FFCCFFEACAEFFEEFFCAC2450" authorityName="Newport" authorityYear="1844" box="[745,882,259,285]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="486" phylum="Arthropoda" rank="species" species="multidens">
|
||
<emphasis id="B962EAD8FFCCFFEACAEFFEEFFCAC2450" box="[745,882,259,285]" italics="true" pageId="21" pageNumber="486">S. multidens</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C164D49FFCCFFEACB8EFEEFFC222450" authority="Tsukamoto & Hiruta & Eguchi & Liao & Shimano, 2021" authorityName="Tsukamoto & Hiruta & Eguchi & Liao & Shimano" authorityYear="2021" box="[904,1020,259,285]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="486" phylum="Arthropoda" rank="species" species="alcyona" status="sp. nov.">
|
||
<emphasis id="B962EAD8FFCCFFEACB8EFEEFFC222450" box="[904,1020,259,285]" italics="true" pageId="21" pageNumber="486">S. alcyona</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAD8FFCCFFEACC04FEEFFB852450" bold="true" box="[1026,1115,259,285]" pageId="21" pageNumber="486">
|
||
<taxonomicNameLabel id="A25157A3FFCCFFEACC04FEEFFB852450" box="[1026,1115,259,285]" pageId="21" pageNumber="486" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="4C164D49FFCCFFEACD81FEEFFEDB240C" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="486" phylum="Arthropoda" rank="species" species="dawydoffi">
|
||
<emphasis id="B962EAD8FFCCFFEACD81FEEFFEDB240C" italics="true" pageId="21" pageNumber="486">S. dawydoffi</emphasis>
|
||
</taxonomicName>
|
||
by the tergite number on which the complete paramedian sutures start (T4–T
|
||
<quantity id="4CEE9B2FFFCCFFEACC6EFECBFB4D240C" box="[1128,1171,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="21" pageNumber="486" unit="in" value="6.0">6 in</quantity>
|
||
<taxonomicName id="4C164D49FFCCFFEACC9CFECBFAD1240C" authority="Tsukamoto & Hiruta & Eguchi & Liao & Shimano, 2021" authorityName="Tsukamoto & Hiruta & Eguchi & Liao & Shimano" authorityYear="2021" box="[1178,1295,295,321]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="486" phylum="Arthropoda" rank="species" species="alcyona" status="sp. nov.">
|
||
<emphasis id="B962EAD8FFCCFFEACC9CFECBFAD1240C" box="[1178,1295,295,321]" italics="true" pageId="21" pageNumber="486">S. alcyona</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAD8FFCCFFEACD13FECBFAB1240C" bold="true" box="[1301,1391,295,321]" pageId="21" pageNumber="486">
|
||
<taxonomicNameLabel id="A25157A3FFCCFFEACD13FECBFAB1240C" box="[1301,1391,295,321]" pageId="21" pageNumber="486" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; T2 or T
|
||
<quantity id="4CEE9B2FFFCCFFEAC8CEFEA7FF2F2428" box="[200,241,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="21" pageNumber="486" unit="in" value="3.0">3 in</quantity>
|
||
<taxonomicName id="4C164D49FFCCFFEAC8FEFEA7FE5C2428" box="[248,386,331,357]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="486" phylum="Arthropoda" rank="species" species="dawydoffi">
|
||
<emphasis id="B962EAD8FFCCFFEAC8FEFEA7FE5C2428" box="[248,386,331,357]" italics="true" pageId="21" pageNumber="486">S. dawydoffi</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C164D49FFCCFFEAC99CFEA7FDD12428" authority="Tsukamoto & Hiruta & Eguchi & Liao & Shimano, 2021" authorityName="Tsukamoto & Hiruta & Eguchi & Liao & Shimano" authorityYear="2021" box="[410,527,331,357]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="486" phylum="Arthropoda" rank="species" species="alcyona" status="sp. nov.">
|
||
<emphasis id="B962EAD8FFCCFFEAC99CFEA7FDD12428" box="[410,527,331,357]" italics="true" pageId="21" pageNumber="486">S. alcyona</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAD8FFCCFFEACA13FEA7FDB12428" bold="true" box="[533,623,331,357]" pageId="21" pageNumber="486">
|
||
<taxonomicNameLabel id="A25157A3FFCCFFEACA13FEA7FDB12428" box="[533,623,331,357]" pageId="21" pageNumber="486" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="4C164D49FFCCFFEACB99FEA7FBC52428" authorityName="Koch" authorityYear="1878" box="[927,1051,331,357]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="486" phylum="Arthropoda" rank="species" species="mutilans">
|
||
<emphasis id="B962EAD8FFCCFFEACB99FEA7FBC52428" box="[927,1051,331,357]" italics="true" pageId="21" pageNumber="486">S. mutilans</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C164D49FFCCFFEACC54FEA7FB2F2428" authorityName="Leach" authorityYear="1816" box="[1106,1265,331,357]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="486" phylum="Arthropoda" rank="species" species="subspinipes">
|
||
<emphasis id="B962EAD8FFCCFFEACC54FEA7FB2F2428" box="[1106,1265,331,357]" italics="true" pageId="21" pageNumber="486">S. subspinipes</emphasis>
|
||
</taxonomicName>
|
||
by the absence gonopods in the male, and the presence of a tarsal spur only on legs 1–19.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C6541FFCCFFEAC8C1FE7FFCAD223C" pageId="21" pageNumber="486" type="description">
|
||
<paragraph id="8BA936CAFFCCFFEAC8C1FE7FFEDB24B8" blockId="21.[151,1437,151,1906]" pageId="21" pageNumber="486">
|
||
<emphasis id="B962EAD8FFCCFFEAC8C1FE7FFC3024E1" bold="true" box="[199,1006,402,429]" pageId="21" pageNumber="486">
|
||
<typeStatus id="54AD8868FFCCFFEAC8C1FE7FFEEA24E0" box="[199,308,403,429]" pageId="21" pageNumber="486" type="holotype">Holotype</typeStatus>
|
||
description (variation of
|
||
<typeStatus id="54AD8868FFCCFFEACA58FE7FFD0A24E0" box="[606,724,403,429]" pageId="21" pageNumber="486" type="paratype">paratypes</typeStatus>
|
||
is given in parenthesis).
|
||
</emphasis>
|
||
Body length in 100% Ethanol
|
||
<quantity id="4CEE9B2FFFCCFFEACD47FE7FFA4324E0" box="[1345,1437,403,429]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="21" pageNumber="486" unit="mm" value="130.0">130 mm</quantity>
|
||
rounding off (TS-20180301-01:
|
||
<quantity id="4CEE9B2FFFCCFFEACA06FE5BFD80249C" box="[512,606,439,465]" metricMagnitude="-1" metricUnit="m" metricValue="1.9" pageId="21" pageNumber="486" unit="mm" value="190.0">190 mm</quantity>
|
||
, TS-20180912-01:
|
||
<quantity id="4CEE9B2FFFCCFFEACB30FE5BFC4B249C" box="[822,917,439,465]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="21" pageNumber="486" unit="mm" value="180.0">180 mm</quantity>
|
||
, TS-20180522-01:
|
||
<quantity id="4CEE9B2FFFCCFFEACC6AFE5BFB15249C" box="[1132,1227,439,465]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="21" pageNumber="486" unit="mm" value="140.0">140 mm</quantity>
|
||
, TS-20180522-02:
|
||
<quantity id="4CEE9B2FFFCCFFEAC891FE37FF2624B8" box="[151,248,475,501]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="21" pageNumber="486" unit="mm" value="130.0">130 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFCCFFEAC8C1FE13FAF02754" blockId="21.[151,1437,151,1906]" box="[199,1326,511,537]" pageId="21" pageNumber="486">
|
||
Antenna (
|
||
<figureCitation id="132D2A4FFFCCFFEAC933FE13FE4C2754" box="[309,402,511,537]" captionStart="FIGURE 4" captionStartId="19.[151,250,1916,1940]" captionTargetBox="[198,1392,185,1895]" captionTargetId="figure-17@19.[194,1393,181,1899]" captionTargetPageId="19" captionText="FIGURE 4. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., A, B, E, F: holotype TS-20180914-01, C: paratype TS- 20180301-01, D: paratype TS-20180522-02: A, cephalic plate, dorsal; B–D, variation of forcipular segment, ventral; E, tergites 14 and 15; F, sternites 14 and 15. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4690559" httpUri="https://zenodo.org/record/4690559/files/figure.png" pageId="21" pageNumber="486">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A4FFFCCFFEAC998FE13FE1D2754" box="[414,451,511,537]" captionStart="FIGURE 7" captionStartId="23.[151,250,1834,1858]" captionTargetBox="[167,1420,191,1803]" captionTargetId="figure-17@23.[151,1436,181,1817]" captionTargetPageId="23" captionText="FIGURE 7. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., A–C, E, F: paratype TS-20180301-01, D: holotype TS- 20180914-01: A, cephalic plate, basal antennal articles and tergite 1, dorsal; B, forcipular segment, ventral; C, D, variation of teeth on tooth-plates; E, tergites 9–11; F, sternites 9–11. Scale bars: 5 mm (A, B, E, F); 1 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.4690565" httpUri="https://zenodo.org/record/4690565/files/figure.png" pageId="21" pageNumber="486">7A</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A4FFFCCFFEAC9C8FE13FDCC2754" box="[462,530,511,537]" captionStart="FIGURE 9" captionStartId="25.[151,250,1141,1165]" captionTargetBox="[166,1419,208,1115]" captionTargetId="figure-17@25.[158,1429,201,1124]" captionTargetPageId="25" captionText="FIGURE 9. Habitus and color variation of Scolopendra alcyona Tsukamoto & Shimano, sp. nov.: A, paratype TS-20180301- 01, habitus Okinawa-jima Island; B, paratype TS-20180522-02, Kume-jima Island; C, paratype TS-20180301-01, head and antennae; D, paratype TS-20180301-01, prefemur of ultimate legs, dorsal view (A, C, D by Mr. K. Tsukamoto)." figureDoi="http://doi.org/10.5281/zenodo.4690569" httpUri="https://zenodo.org/record/4690569/files/figure.png" pageId="21" pageNumber="486">9A, C</figureCitation>
|
||
) with 18 articles (17–20 article in
|
||
<typeStatus id="54AD8868FFCCFFEACB93FE13FBDB2754" box="[917,1029,511,537]" pageId="21" pageNumber="486" type="paratype">paratypes</typeStatus>
|
||
); basal 6 articles glabrous.
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFCCFFEAC8C1FDCFFF38272C" blockId="21.[151,1437,151,1906]" pageId="21" pageNumber="486">
|
||
Cephalic plate (
|
||
<figureCitation id="132D2A4FFFCCFFEAC97FFDCFFE092770" box="[377,471,547,573]" captionStart="FIGURE 4" captionStartId="19.[151,250,1916,1940]" captionTargetBox="[198,1392,185,1895]" captionTargetId="figure-17@19.[194,1393,181,1899]" captionTargetPageId="19" captionText="FIGURE 4. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., A, B, E, F: holotype TS-20180914-01, C: paratype TS- 20180301-01, D: paratype TS-20180522-02: A, cephalic plate, dorsal; B–D, variation of forcipular segment, ventral; E, tergites 14 and 15; F, sternites 14 and 15. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4690559" httpUri="https://zenodo.org/record/4690559/files/figure.png" pageId="21" pageNumber="486">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A4FFFCCFFEAC9E2FDCFFDD72770" box="[484,521,547,573]" captionStart="FIGURE 7" captionStartId="23.[151,250,1834,1858]" captionTargetBox="[167,1420,191,1803]" captionTargetId="figure-17@23.[151,1436,181,1817]" captionTargetPageId="23" captionText="FIGURE 7. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., A–C, E, F: paratype TS-20180301-01, D: holotype TS- 20180914-01: A, cephalic plate, basal antennal articles and tergite 1, dorsal; B, forcipular segment, ventral; C, D, variation of teeth on tooth-plates; E, tergites 9–11; F, sternites 9–11. Scale bars: 5 mm (A, B, E, F); 1 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.4690565" httpUri="https://zenodo.org/record/4690565/files/figure.png" pageId="21" pageNumber="486">7A</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A4FFFCCFFEACA13FDCFFDE92770" box="[533,567,547,573]" captionStart="FIGURE 9" captionStartId="25.[151,250,1141,1165]" captionTargetBox="[166,1419,208,1115]" captionTargetId="figure-17@25.[158,1429,201,1124]" captionTargetPageId="25" captionText="FIGURE 9. Habitus and color variation of Scolopendra alcyona Tsukamoto & Shimano, sp. nov.: A, paratype TS-20180301- 01, habitus Okinawa-jima Island; B, paratype TS-20180522-02, Kume-jima Island; C, paratype TS-20180301-01, head and antennae; D, paratype TS-20180301-01, prefemur of ultimate legs, dorsal view (A, C, D by Mr. K. Tsukamoto)." figureDoi="http://doi.org/10.5281/zenodo.4690569" httpUri="https://zenodo.org/record/4690569/files/figure.png" pageId="21" pageNumber="486">9C</figureCitation>
|
||
) with small puncta entirely; short anterior median sulcus; paramedian suture absent.
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFCCFFEAC8C1FD87FC2427E4" blockId="21.[151,1437,151,1906]" pageId="21" pageNumber="486">Article 2 of second maxillary telopodite with spur. Second maxillary pretarsus 0.36 times as long as article 3 of second maxillary telopodite, rounded at the tip, with dorsal and ventral spurs.</paragraph>
|
||
<paragraph id="8BA936CAFFCCFFEAC8C1FD5FFC0C2674" blockId="21.[151,1437,151,1906]" pageId="21" pageNumber="486">
|
||
Coxosternite (
|
||
<figureCitation id="132D2A4FFFCCFFEAC963FD5FFE3D2780" box="[357,483,691,717]" captionStart="FIGURE 4" captionStartId="19.[151,250,1916,1940]" captionTargetBox="[198,1392,185,1895]" captionTargetId="figure-17@19.[194,1393,181,1899]" captionTargetPageId="19" captionText="FIGURE 4. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., A, B, E, F: holotype TS-20180914-01, C: paratype TS- 20180301-01, D: paratype TS-20180522-02: A, cephalic plate, dorsal; B–D, variation of forcipular segment, ventral; E, tergites 14 and 15; F, sternites 14 and 15. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4690559" httpUri="https://zenodo.org/record/4690559/files/figure.png" pageId="21" pageNumber="486">Figs 4B–D</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A4FFFCCFFEAC9E8FD5FFDEC2780" box="[494,562,691,717]" captionStart="FIGURE 7" captionStartId="23.[151,250,1834,1858]" captionTargetBox="[167,1420,191,1803]" captionTargetId="figure-17@23.[151,1436,181,1817]" captionTargetPageId="23" captionText="FIGURE 7. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., A–C, E, F: paratype TS-20180301-01, D: holotype TS- 20180914-01: A, cephalic plate, basal antennal articles and tergite 1, dorsal; B, forcipular segment, ventral; C, D, variation of teeth on tooth-plates; E, tergites 9–11; F, sternites 9–11. Scale bars: 5 mm (A, B, E, F); 1 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.4690565" httpUri="https://zenodo.org/record/4690565/files/figure.png" pageId="21" pageNumber="486">7B–D</figureCitation>
|
||
) surface smooth, without median suture; tooth-plates longer than wide, with 7 + 7 teeth in two groups (8 +
|
||
<quantity id="4CEE9B2FFFCCFFEAC9D4FD3BFE2327BC" box="[466,509,727,753]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="21" pageNumber="486" unit="in" value="7.0">7 in</quantity>
|
||
TS-20180301-01, 9 +
|
||
<quantity id="4CEE9B2FFFCCFFEACAFCFD3BFCFB27BC" box="[762,805,727,753]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="21" pageNumber="486" unit="in" value="8.0">8 in</quantity>
|
||
TS-20180912-01, 5 +
|
||
<quantity id="4CEE9B2FFFCCFFEACC24FD3BFB9327BC" box="[1058,1101,727,753]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="21" pageNumber="486" unit="in" value="4.0">4 in</quantity>
|
||
TS-20180522-01 and 5 +
|
||
<quantity id="4CEE9B2FFFCCFFEACD74FD3BFA4327BC" box="[1394,1437,727,753]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="21" pageNumber="486" unit="in" value="7.0">7 in</quantity>
|
||
TS-20180522-02) and with transverse basal sulci. Forcipule (
|
||
<figureCitation id="132D2A4FFFCCFFEACB4DFD17FC152658" box="[843,971,763,789]" captionStart="FIGURE 4" captionStartId="19.[151,250,1916,1940]" captionTargetBox="[198,1392,185,1895]" captionTargetId="figure-17@19.[194,1393,181,1899]" captionTargetPageId="19" captionText="FIGURE 4. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., A, B, E, F: holotype TS-20180914-01, C: paratype TS- 20180301-01, D: paratype TS-20180522-02: A, cephalic plate, dorsal; B–D, variation of forcipular segment, ventral; E, tergites 14 and 15; F, sternites 14 and 15. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4690559" httpUri="https://zenodo.org/record/4690559/files/figure.png" pageId="21" pageNumber="486">Figs 4B–D</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A4FFFCCFFEACBD1FD17FC262658" box="[983,1016,763,789]" captionStart="FIGURE 7" captionStartId="23.[151,250,1834,1858]" captionTargetBox="[167,1420,191,1803]" captionTargetId="figure-17@23.[151,1436,181,1817]" captionTargetPageId="23" captionText="FIGURE 7. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., A–C, E, F: paratype TS-20180301-01, D: holotype TS- 20180914-01: A, cephalic plate, basal antennal articles and tergite 1, dorsal; B, forcipular segment, ventral; C, D, variation of teeth on tooth-plates; E, tergites 9–11; F, sternites 9–11. Scale bars: 5 mm (A, B, E, F); 1 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.4690565" httpUri="https://zenodo.org/record/4690565/files/figure.png" pageId="21" pageNumber="486">7B</figureCitation>
|
||
) surface smooth; process of trochanteroprefemur with teeth in two groups, 1 apical tooth and 1(3) inner teeth.
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFCCFFEAC8C1FCAFFD4426A0" blockId="21.[151,1437,151,1906]" pageId="21" pageNumber="486">
|
||
Tergite (
|
||
<figureCitation id="132D2A4FFFCCFFEAC923FCAFFE5F2610" box="[293,385,835,861]" captionStart="FIGURE 4" captionStartId="19.[151,250,1916,1940]" captionTargetBox="[198,1392,185,1895]" captionTargetId="figure-17@19.[194,1393,181,1899]" captionTargetPageId="19" captionText="FIGURE 4. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., A, B, E, F: holotype TS-20180914-01, C: paratype TS- 20180301-01, D: paratype TS-20180522-02: A, cephalic plate, dorsal; B–D, variation of forcipular segment, ventral; E, tergites 14 and 15; F, sternites 14 and 15. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4690559" httpUri="https://zenodo.org/record/4690559/files/figure.png" pageId="21" pageNumber="486">Figs 4E</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A4FFFCCFFEAC98BFCAFFE702610" box="[397,430,835,861]" captionStart="FIGURE 7" captionStartId="23.[151,250,1834,1858]" captionTargetBox="[167,1420,191,1803]" captionTargetId="figure-17@23.[151,1436,181,1817]" captionTargetPageId="23" captionText="FIGURE 7. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., A–C, E, F: paratype TS-20180301-01, D: holotype TS- 20180914-01: A, cephalic plate, basal antennal articles and tergite 1, dorsal; B, forcipular segment, ventral; C, D, variation of teeth on tooth-plates; E, tergites 9–11; F, sternites 9–11. Scale bars: 5 mm (A, B, E, F); 1 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.4690565" httpUri="https://zenodo.org/record/4690565/files/figure.png" pageId="21" pageNumber="486">7E</figureCitation>
|
||
) surface smooth. Anterior margin of T1 overlapped by cephalic plate. Complete paramedian sutures on TT6–20 (paramedian sutures starting on TT
|
||
<quantity id="4CEE9B2FFFCCFFEACB28FC8BFCAD26CC" box="[814,883,871,897]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.524" metricValueMin="1.016" pageId="21" pageNumber="486" unit="in" value="5.0" valueMax="6.0" valueMin="4.0">4–6 in</quantity>
|
||
<typeStatus id="54AD8868FFCCFFEACB7FFC8BFC3726CC" box="[889,1001,871,897]" pageId="21" pageNumber="486" type="paratype">paratypes</typeStatus>
|
||
); margination on TT6–20 (margination starting on TT
|
||
<quantity id="4CEE9B2FFFCCFFEAC93CFC67FE5326E8" box="[314,397,907,933]" metricMagnitude="-1" metricUnit="m" metricValue="2.159" metricValueMax="2.794" metricValueMin="1.524" pageId="21" pageNumber="486" unit="in" value="8.5" valueMax="11.0" valueMin="6.0">6–11 in</quantity>
|
||
<typeStatus id="54AD8868FFCCFFEAC992FC67FDDB26E8" box="[404,517,907,933]" pageId="21" pageNumber="486" type="paratype">paratypes</typeStatus>
|
||
). Tergite of ultimate leg-bearing segment (
|
||
<figureCitation id="132D2A4FFFCCFFEACBE2FC67FB9C26E8" box="[996,1090,906,933]" captionStart="FIGURE 5" captionStartId="20.[151,250,1588,1612]" captionTargetBox="[154,1434,313,1568]" captionTargetId="figure-78@20.[151,1435,309,1572]" captionTargetPageId="20" captionText="FIGURE 5. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., holotype TS-20180914-01: A, posterior part of tergite 20, and tergite 21; B, ultimate leg-bearing segment, ventral; C, left coxopleuron, lateral; D, male genital segments, ventral; E, male genital segments, lateral. Legends: triangles = genital sternite 1; arrows = genital sternite 2; arrowheads = tergite of genital segment. Scale bars: 5 mm (A–C), 2.5 mm (D, E)." figureDoi="http://doi.org/10.5281/zenodo.4690561" httpUri="https://zenodo.org/record/4690561/files/figure.png" pageId="21" pageNumber="486">Figs 5A</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A4FFFCCFFEACC48FC67FBAC26E8" box="[1102,1138,907,933]" captionStart="FIGURE 8" captionStartId="24.[151,260,1923,1949]" captionTargetBox="[167,1425,392,1896]" captionTargetId="figure-96@24.[151,1436,381,1908]" captionTargetPageId="24" captionText="FIGURE 8. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., paratype TS-20180301-01: A, tergite 21 and prefemur of ultimate leg, dorsal; B, ultimate leg-bearing segment and prefemur of ultimate leg, ventral; C, left coxopleuron, lateral. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4690567" httpUri="https://zenodo.org/record/4690567/files/figure.png" pageId="21" pageNumber="486">8A</figureCitation>
|
||
) without median suture or depression; posterior margin of tergite of ultimate leg-bearing segment curved posteriorly; ratio of width and length of tergite of ultimate leg-bearing segment 1:1.
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFCCFFEAC8C1FC1BFEC92130" blockId="21.[151,1437,151,1906]" pageId="21" pageNumber="486">
|
||
Sternite (
|
||
<figureCitation id="132D2A4FFFCCFFEAC937FC1BFE52215C" box="[305,396,1015,1041]" captionStart="FIGURE 4" captionStartId="19.[151,250,1916,1940]" captionTargetBox="[198,1392,185,1895]" captionTargetId="figure-17@19.[194,1393,181,1899]" captionTargetPageId="19" captionText="FIGURE 4. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., A, B, E, F: holotype TS-20180914-01, C: paratype TS- 20180301-01, D: paratype TS-20180522-02: A, cephalic plate, dorsal; B–D, variation of forcipular segment, ventral; E, tergites 14 and 15; F, sternites 14 and 15. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4690559" httpUri="https://zenodo.org/record/4690559/files/figure.png" pageId="21" pageNumber="486">Figs 4F</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A4FFFCCFFEAC99DFC1BFE64215C" box="[411,442,1015,1041]" captionStart="FIGURE 7" captionStartId="23.[151,250,1834,1858]" captionTargetBox="[167,1420,191,1803]" captionTargetId="figure-17@23.[151,1436,181,1817]" captionTargetPageId="23" captionText="FIGURE 7. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., A–C, E, F: paratype TS-20180301-01, D: holotype TS- 20180914-01: A, cephalic plate, basal antennal articles and tergite 1, dorsal; B, forcipular segment, ventral; C, D, variation of teeth on tooth-plates; E, tergites 9–11; F, sternites 9–11. Scale bars: 5 mm (A, B, E, F); 1 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.4690565" httpUri="https://zenodo.org/record/4690565/files/figure.png" pageId="21" pageNumber="486">7F</figureCitation>
|
||
) surface smooth; SS2–19 with paramedian sutures, posteriorly incomplete, extending to approximately 65–76% length of sternite. Sternite of ultimate leg-bearing segment (
|
||
<figureCitation id="132D2A4FFFCCFFEACC7DFBF7FB062178" box="[1147,1240,1050,1077]" captionStart="FIGURE 5" captionStartId="20.[151,250,1588,1612]" captionTargetBox="[154,1434,313,1568]" captionTargetId="figure-78@20.[151,1435,309,1572]" captionTargetPageId="20" captionText="FIGURE 5. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., holotype TS-20180914-01: A, posterior part of tergite 20, and tergite 21; B, ultimate leg-bearing segment, ventral; C, left coxopleuron, lateral; D, male genital segments, ventral; E, male genital segments, lateral. Legends: triangles = genital sternite 1; arrows = genital sternite 2; arrowheads = tergite of genital segment. Scale bars: 5 mm (A–C), 2.5 mm (D, E)." figureDoi="http://doi.org/10.5281/zenodo.4690561" httpUri="https://zenodo.org/record/4690561/files/figure.png" pageId="21" pageNumber="486">Figs 5B</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A4FFFCCFFEACCE1FBF7FAD72178" box="[1255,1289,1051,1077]" captionStart="FIGURE 8" captionStartId="24.[151,260,1923,1949]" captionTargetBox="[167,1425,392,1896]" captionTargetId="figure-96@24.[151,1436,381,1908]" captionTargetPageId="24" captionText="FIGURE 8. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., paratype TS-20180301-01: A, tergite 21 and prefemur of ultimate leg, dorsal; B, ultimate leg-bearing segment and prefemur of ultimate leg, ventral; C, left coxopleuron, lateral. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4690567" httpUri="https://zenodo.org/record/4690567/files/figure.png" pageId="21" pageNumber="486">8B</figureCitation>
|
||
) with lateral margins converging posteriorly and posterior margin convex posteriorly; surface with shallow median longitudinal depression.
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFCCFFEAC8C1FB6BFECD2018" blockId="21.[151,1437,151,1906]" pageId="21" pageNumber="486">
|
||
Coxopleuron (
|
||
<figureCitation id="132D2A4FFFCCFFEAC96EFB6BFE3A21EC" box="[360,484,1158,1185]" captionStart="FIGURE 5" captionStartId="20.[151,250,1588,1612]" captionTargetBox="[154,1434,313,1568]" captionTargetId="figure-78@20.[151,1435,309,1572]" captionTargetPageId="20" captionText="FIGURE 5. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., holotype TS-20180914-01: A, posterior part of tergite 20, and tergite 21; B, ultimate leg-bearing segment, ventral; C, left coxopleuron, lateral; D, male genital segments, ventral; E, male genital segments, lateral. Legends: triangles = genital sternite 1; arrows = genital sternite 2; arrowheads = tergite of genital segment. Scale bars: 5 mm (A–C), 2.5 mm (D, E)." figureDoi="http://doi.org/10.5281/zenodo.4690561" httpUri="https://zenodo.org/record/4690561/files/figure.png" pageId="21" pageNumber="486">Figs 5B, C</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A4FFFCCFFEAC9E9FB6BFDEC21EC" box="[495,562,1159,1185]" captionStart="FIGURE 8" captionStartId="24.[151,260,1923,1949]" captionTargetBox="[167,1425,392,1896]" captionTargetId="figure-96@24.[151,1436,381,1908]" captionTargetPageId="24" captionText="FIGURE 8. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., paratype TS-20180301-01: A, tergite 21 and prefemur of ultimate leg, dorsal; B, ultimate leg-bearing segment and prefemur of ultimate leg, ventral; C, left coxopleuron, lateral. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4690567" httpUri="https://zenodo.org/record/4690567/files/figure.png" pageId="21" pageNumber="486">8B, C</figureCitation>
|
||
) with numerous pores, pore-field extending into coxopleural process covering most of its surface, except for a narrow pore-free area, dorsal margin of pore-field gradually elevated anteriorly. Coxopleural process moderately long, with 2 apical spines and 1 subapical spine (0–2 apical and 0–2 subapical spines in
|
||
<typeStatus id="54AD8868FFCCFFEAC8F9FB1FFEB12040" box="[255,367,1267,1293]" pageId="21" pageNumber="486" type="paratype">paratypes</typeStatus>
|
||
), without lateral spine; pore-free area of coxopleural process extending ventrally 79–93% length from distal part of coxopleural process to posterior margin of sternite of ultimate leg-bearing segment (73–100% in
|
||
<typeStatus id="54AD8868FFCCFFEAC891FAD7FED62018" box="[151,264,1339,1365]" pageId="21" pageNumber="486" type="paratype">paratypes</typeStatus>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFCCFFEAC8C1FAB3FD972344" blockId="21.[151,1437,151,1906]" pageId="21" pageNumber="486">
|
||
All legs without setae and tibial spur. One ventrodistal tarsal spur on tarsus 1 of legs 1–19. Ultimate legs (
|
||
<figureCitation id="132D2A4FFFCCFFEACD6CFAB3FF7920D0" captionStart="FIGURE 6" captionStartId="22.[151,250,1937,1961]" captionTargetBox="[360,1224,186,1914]" captionTargetId="figure-22@22.[357,1231,181,1920]" captionTargetPageId="22" captionText="FIGURE 6. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., holotype TS-20180914-01: A, prefemora and femora of ultimate legs, dorsal; B, prefemora and femora of ultimate legs, ventral; C, tibiae, tarsi 1, tarsi 2, and pretarsi of ultimate legs, dorsal; D, tibiae tarsi 1, tarsi 2, and pretarsi of ultimate legs, ventral. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4690563" httpUri="https://zenodo.org/record/4690563/files/figure.png" pageId="21" pageNumber="486">Figs 6</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A4FFFCCFFEAC8B2FA6FFF2420D0" box="[180,250,1411,1437]" captionStart="FIGURE 8" captionStartId="24.[151,260,1923,1949]" captionTargetBox="[167,1425,392,1896]" captionTargetId="figure-96@24.[151,1436,381,1908]" captionTargetPageId="24" captionText="FIGURE 8. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., paratype TS-20180301-01: A, tergite 21 and prefemur of ultimate leg, dorsal; B, ultimate leg-bearing segment and prefemur of ultimate leg, ventral; C, left coxopleuron, lateral. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4690567" httpUri="https://zenodo.org/record/4690567/files/figure.png" pageId="21" pageNumber="486">8A, B</figureCitation>
|
||
,
|
||
<figureCitation id="132D2A4FFFCCFFEAC901FA6FFEF520D0" box="[263,299,1411,1437]" captionStart="FIGURE 9" captionStartId="25.[151,250,1141,1165]" captionTargetBox="[166,1419,208,1115]" captionTargetId="figure-17@25.[158,1429,201,1124]" captionTargetPageId="25" captionText="FIGURE 9. Habitus and color variation of Scolopendra alcyona Tsukamoto & Shimano, sp. nov.: A, paratype TS-20180301- 01, habitus Okinawa-jima Island; B, paratype TS-20180522-02, Kume-jima Island; C, paratype TS-20180301-01, head and antennae; D, paratype TS-20180301-01, prefemur of ultimate legs, dorsal view (A, C, D by Mr. K. Tsukamoto)." figureDoi="http://doi.org/10.5281/zenodo.4690569" httpUri="https://zenodo.org/record/4690569/files/figure.png" pageId="21" pageNumber="486">9D</figureCitation>
|
||
) long and slender with ratio of width and length of prefemur 1:3, ratio of width and length of femur 1:4 and ratios of lengths of prefemur and femur 1.06:1; femur and tibia 1.13:1; tibia and tarsus 1 1.33:1; tarsus 1 and tarsus 2 2.15:1. Prefemur of ultimate leg flattened dorsally with acute blackish spines row as below: 2 VL, 1 M, 2 DM and 2–3 SP (1–3 SP in
|
||
<typeStatus id="54AD8868FFCCFFEAC9CBFA03FDE02344" box="[461,574,1519,1545]" pageId="21" pageNumber="486" type="paratype">paratypes</typeStatus>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFCCFFEAC8C1F9FFFD5023F0" blockId="21.[151,1437,151,1906]" pageId="21" pageNumber="486">
|
||
In the male
|
||
<typeStatus id="54AD8868FFCCFFEAC941F9FFFE722360" box="[327,428,1555,1581]" pageId="21" pageNumber="486" type="holotype">holotype</typeStatus>
|
||
, and a female non-type specimen (TS-19690814-01), genital segments (
|
||
<figureCitation id="132D2A4FFFCCFFEACCC2F9FFFAE22360" box="[1220,1340,1554,1581]" captionStart="FIGURE 5" captionStartId="20.[151,250,1588,1612]" captionTargetBox="[154,1434,313,1568]" captionTargetId="figure-78@20.[151,1435,309,1572]" captionTargetPageId="20" captionText="FIGURE 5. Scolopendra alcyona Tsukamoto & Shimano, sp. nov., holotype TS-20180914-01: A, posterior part of tergite 20, and tergite 21; B, ultimate leg-bearing segment, ventral; C, left coxopleuron, lateral; D, male genital segments, ventral; E, male genital segments, lateral. Legends: triangles = genital sternite 1; arrows = genital sternite 2; arrowheads = tergite of genital segment. Scale bars: 5 mm (A–C), 2.5 mm (D, E)." figureDoi="http://doi.org/10.5281/zenodo.4690561" httpUri="https://zenodo.org/record/4690561/files/figure.png" pageId="21" pageNumber="486">Figs 5D, E</figureCitation>
|
||
) everted and projected beyond distal part of coxopleural process. Sternite of genital segment 1 round and convex posteriorly. Gonopods absent. Sternite of genital segment 2 well developed in male. Tergite of genital segment small, horseshoeshaped and lacking setae. Lamina subanalis between genitalia and anal valves; lamina adanalis between anal valves and tergite of genital segment. Penis present.
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFCCFFEAC8C1F92BFCAD223C" blockId="21.[151,1437,151,1906]" pageId="21" pageNumber="486">
|
||
Coloration in living condition as in
|
||
<figureCitation id="132D2A4FFFCCFFEACA67F92BFD3C23AC" box="[609,738,1735,1761]" captionStart="FIGURE 9" captionStartId="25.[151,250,1141,1165]" captionTargetBox="[166,1419,208,1115]" captionTargetId="figure-17@25.[158,1429,201,1124]" captionTargetPageId="25" captionText="FIGURE 9. Habitus and color variation of Scolopendra alcyona Tsukamoto & Shimano, sp. nov.: A, paratype TS-20180301- 01, habitus Okinawa-jima Island; B, paratype TS-20180522-02, Kume-jima Island; C, paratype TS-20180301-01, head and antennae; D, paratype TS-20180301-01, prefemur of ultimate legs, dorsal view (A, C, D by Mr. K. Tsukamoto)." figureDoi="http://doi.org/10.5281/zenodo.4690569" httpUri="https://zenodo.org/record/4690569/files/figure.png" pageId="21" pageNumber="486">Figs 9A–D</figureCitation>
|
||
. Brownish black on cephalic plate and T1; greenish black on remaining tergites; bluish black on antenna; light brown on coxosternite and forcipule (but forcipular tarsungulum blackish); pale green on all sternites; bluish black on eupleuria, with greenish black integuments; orange on coxopleura; basal part of legs 1–20 yellow, distal part of legs 1–20 and ultimate legs greenish blue (all legs yellow in Kume-jima Island specimens); pale blue on intermediate sternite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF696642FFCFFFE9C891F87DFBD122A7" ID-DOI="http://doi.org/10.5281/zenodo.4690563" ID-Zenodo-Dep="4690563" httpUri="https://zenodo.org/record/4690563/files/figure.png" pageId="22" pageNumber="487" startId="22.[151,250,1937,1961]" targetBox="[360,1224,186,1914]" targetPageId="22">
|
||
<paragraph id="8BA936CAFFCFFFE9C891F87DFBD122A7" blockId="22.[151,1436,1937,2026]" pageId="22" pageNumber="487">
|
||
<emphasis id="B962EAD8FFCFFFE9C891F87DFECB22E4" bold="true" box="[151,277,1937,1961]" pageId="22" pageNumber="487">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C164D49FFCFFFE9C91BF87DFD3022E4" authority="Tsukamoto & Shimano" authorityName="Tsukamoto & Hiruta & Eguchi & Liao & Shimano" authorityYear="2021" box="[285,750,1937,1961]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="487" phylum="Arthropoda" rank="species" species="alcyona" status="sp. nov.">
|
||
<emphasis id="B962EAD8FFCFFFE9C91BF87DFE2B22E4" box="[285,501,1937,1961]" italics="true" pageId="22" pageNumber="487">Scolopendra alcyona</emphasis>
|
||
Tsukamoto & Shimano
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B962EAD8FFCFFFE9CAFCF87DFC9222E4" bold="true" box="[762,844,1937,1961]" pageId="22" pageNumber="487">
|
||
<taxonomicNameLabel id="A25157A3FFCFFFE9CAFCF87DFC9222E4" box="[762,844,1937,1961]" pageId="22" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype
|
||
<emphasis id="B962EAD8FFCFFFE9CBBAF87DFBB422E4" bold="true" box="[956,1130,1937,1961]" pageId="22" pageNumber="487">TS-20180914-01</emphasis>
|
||
:
|
||
<emphasis id="B962EAD8FFCFFFE9CC7CF87DFB5222E4" bold="true" box="[1146,1164,1937,1961]" pageId="22" pageNumber="487">A</emphasis>
|
||
, prefemora and femora of ultimate legs, dorsal;
|
||
<emphasis id="B962EAD8FFCFFFE9C975F842FE5A228B" bold="true" box="[371,388,1966,1990]" pageId="22" pageNumber="487">B</emphasis>
|
||
, prefemora and femora of ultimate legs, ventral;
|
||
<emphasis id="B962EAD8FFCFFFE9CB7DF842FC53228B" bold="true" box="[891,909,1966,1990]" pageId="22" pageNumber="487">C</emphasis>
|
||
, tibiae, tarsi 1, tarsi 2, and pretarsi of ultimate legs, dorsal;
|
||
<emphasis id="B962EAD8FFCFFFE9C8E4F83EFF2A22A7" bold="true" box="[226,244,2002,2026]" pageId="22" pageNumber="487">D</emphasis>
|
||
, tibiae tarsi 1, tarsi 2, and pretarsi of ultimate legs, ventral.
|
||
<emphasis id="B962EAD8FFCFFFE9CB57F83EFC1E22A7" bold="true" box="[849,960,2002,2026]" pageId="22" pageNumber="487">Scale bars</emphasis>
|
||
:
|
||
<quantity id="4CEE9B2FFFCFFFE9CBCBF83EFBD522A7" box="[973,1035,2002,2026]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="487" unit="mm" value="5.0">5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF696642FFCEFFE8C891F8C6FB7E22C9" ID-DOI="http://doi.org/10.5281/zenodo.4690565" ID-Zenodo-Dep="4690565" httpUri="https://zenodo.org/record/4690565/files/figure.png" pageId="23" pageNumber="488" startId="23.[151,250,1834,1858]" targetBox="[167,1420,191,1803]" targetPageId="23">
|
||
<paragraph id="8BA936CAFFCEFFE8C891F8C6FB7E22C9" blockId="23.[151,1436,1834,1924]" pageId="23" pageNumber="488">
|
||
<emphasis id="B962EAD8FFCEFFE8C891F8C6FECB220F" bold="true" box="[151,277,1834,1858]" pageId="23" pageNumber="488">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C164D49FFCEFFE8C91BF8C6FD2E220F" authority="Tsukamoto & Shimano" authorityName="Tsukamoto & Hiruta & Eguchi & Liao & Shimano" authorityYear="2021" box="[285,752,1834,1858]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="488" phylum="Arthropoda" rank="species" species="alcyona" status="sp. nov.">
|
||
<emphasis id="B962EAD8FFCEFFE8C91BF8C6FE28220F" box="[285,502,1834,1858]" italics="true" pageId="23" pageNumber="488">Scolopendra alcyona</emphasis>
|
||
Tsukamoto & Shimano
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B962EAD8FFCEFFE8CAFDF8C6FC90220F" bold="true" box="[763,846,1834,1858]" pageId="23" pageNumber="488">
|
||
<taxonomicNameLabel id="A25157A3FFCEFFE8CAFDF8C6FC90220F" box="[763,846,1834,1858]" pageId="23" pageNumber="488" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, A–C, E, F: paratype
|
||
<emphasis id="B962EAD8FFCEFFE8CC33F8C6FB3D220F" bold="true" box="[1077,1251,1834,1858]" pageId="23" pageNumber="488">TS-20180301-01</emphasis>
|
||
, D: holotype
|
||
<emphasis id="B962EAD8FFCEFFE8CD73F8C6FEC02212" bold="true" pageId="23" pageNumber="488">TS- 20180914-01</emphasis>
|
||
:
|
||
<emphasis id="B962EAD8FFCEFFE8C92AF8ABFEE02212" bold="true" box="[300,318,1863,1887]" pageId="23" pageNumber="488">A</emphasis>
|
||
, cephalic plate, basal antennal articles and tergite 1, dorsal;
|
||
<emphasis id="B962EAD8FFCEFFE8CBA1F8A4FC66222D" bold="true" box="[935,952,1864,1888]" pageId="23" pageNumber="488">B</emphasis>
|
||
, forcipular segment, ventral;
|
||
<emphasis id="B962EAD8FFCEFFE8CCE1F8ABFB272212" bold="true" box="[1255,1273,1863,1887]" pageId="23" pageNumber="488">C</emphasis>
|
||
,
|
||
<emphasis id="B962EAD8FFCEFFE8CD00F8A4FAC6222D" bold="true" box="[1286,1304,1864,1888]" pageId="23" pageNumber="488">D</emphasis>
|
||
, variation of teeth on tooth-plates;
|
||
<emphasis id="B962EAD8FFCEFFE8C975F880FE5A22C9" bold="true" box="[371,388,1900,1924]" pageId="23" pageNumber="488">E</emphasis>
|
||
, tergites 9–11;
|
||
<emphasis id="B962EAD8FFCEFFE8CA26F880FDF122C9" bold="true" box="[544,559,1900,1924]" pageId="23" pageNumber="488">F</emphasis>
|
||
, sternites 9–11.
|
||
<emphasis id="B962EAD8FFCEFFE8CAD0F887FC9A22C9" bold="true" box="[726,836,1899,1924]" pageId="23" pageNumber="488">Scale bars</emphasis>
|
||
: 5 mm (A, B, E, F); 1 mm (C, D).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30C6541FFC1FFE7C8C1FF7AFCA52453" pageId="24" pageNumber="489" type="distribution">
|
||
<paragraph id="8BA936CAFFC1FFE7C8C1FF7AFCA52453" blockId="24.[151,1436,150,357]" pageId="24" pageNumber="489">
|
||
<emphasis id="B962EAD8FFC1FFE7C8C1FF7AFEBD25FD" bold="true" box="[199,355,150,176]" pageId="24" pageNumber="489">Distribution:</emphasis>
|
||
Ryukyu Archipelago (Okinawa-jima Island, Kume-jima Island) and
|
||
<collectingCountry id="F301765AFFC1FFE7CC60FF7BFB6225FC" box="[1126,1212,151,177]" name="Taiwan" pageId="24" pageNumber="489">Taiwan</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C164D49FFC1FFE7CCC1FF7BFAE225FC" authority="Tsukamoto & Hiruta & Eguchi & Liao & Shimano, 2021" authorityName="Tsukamoto & Hiruta & Eguchi & Liao & Shimano" authorityYear="2021" box="[1223,1340,151,177]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="24" pageNumber="489" phylum="Arthropoda" rank="species" species="alcyona" status="sp. nov.">
|
||
<emphasis id="B962EAD8FFC1FFE7CCC1FF7BFAE225FC" box="[1223,1340,151,177]" italics="true" pageId="24" pageNumber="489">S. alcyona</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAD8FFC1FFE7CD45FF7BFA4225FC" bold="true" box="[1347,1436,151,177]" pageId="24" pageNumber="489">
|
||
<taxonomicNameLabel id="A25157A3FFC1FFE7CD45FF7BFA4225FC" box="[1347,1436,151,177]" pageId="24" pageNumber="489" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
specimens examined in this study were collected in Okinawa-jima Island, Kume-jima Island, and
|
||
<collectingCountry id="F301765AFFC1FFE7CCFEFF57FA932598" box="[1272,1357,187,213]" name="Taiwan" pageId="24" pageNumber="489">Taiwan</collectingCountry>
|
||
, but
|
||
<taxonomicName id="4C164D49FFC1FFE7CD81FF57FF2E25B4" authority="Tsukamoto & Hiruta & Eguchi & Liao & Shimano, 2021" authorityName="Tsukamoto & Hiruta & Eguchi & Liao & Shimano" authorityYear="2021" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="24" pageNumber="489" phylum="Arthropoda" rank="species" species="alcyona" status="sp. nov.">
|
||
<emphasis id="B962EAD8FFC1FFE7CD81FF57FF2E25B4" italics="true" pageId="24" pageNumber="489">S. alcyona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25157A3FFC1FFE7C8F1FF33FE8D25B4" box="[247,339,223,249]" pageId="24" pageNumber="489" rank="species">
|
||
<emphasis id="B962EAD8FFC1FFE7C8F1FF33FE9225B4" bold="true" box="[247,332,223,249]" pageId="24" pageNumber="489">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
could be also distributed in Tokashiki-jima Island, Ishigaki-jima Island, and Iriomote-jima Island (Masashi Sugimoto and Taku Shimada, personal communication).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C6541FFC1FFE7C8C1FECAFE602428" pageId="24" pageNumber="489" type="biology_ecology">
|
||
<paragraph id="8BA936CAFFC1FFE7C8C1FECAFE602428" blockId="24.[151,1436,150,357]" pageId="24" pageNumber="489">
|
||
<emphasis id="B962EAD8FFC1FFE7C8C1FECAFEF2240D" bold="true" box="[199,300,294,320]" pageId="24" pageNumber="489">Habitat:</emphasis>
|
||
<taxonomicName id="4C164D49FFC1FFE7C935FECBFDFE240C" authority="Tsukamoto & Hiruta & Eguchi & Liao & Shimano, 2021" authorityName="Tsukamoto & Hiruta & Eguchi & Liao & Shimano" authorityYear="2021" box="[307,544,295,321]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="24" pageNumber="489" phylum="Arthropoda" rank="species" species="alcyona" status="sp. nov.">
|
||
<emphasis id="B962EAD8FFC1FFE7C935FECBFDFE240C" box="[307,544,295,321]" italics="true" pageId="24" pageNumber="489">Scolopendra alcyona</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAD8FFC1FFE7CA21FECBFD5E240C" bold="true" box="[551,640,295,321]" pageId="24" pageNumber="489">
|
||
<taxonomicNameLabel id="A25157A3FFC1FFE7CA21FECBFD5E240C" box="[551,640,295,321]" pageId="24" pageNumber="489" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
lives along streams that flow through the forest. It is often found under stones nearby the streams.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |