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<document id="568663B29AF44EBD166CE8FFB46B0C6D" ID-DOI="10.11646/zootaxa.5351.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="8391329" ID-ZooBank="08CAFDFE-D4CF-4E5A-A173-3DA9F19F5FA5" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695989053433" checkinUser="plazi" docAuthor="Jałoszyński, Paweł &amp; Wakimura, Ryotaro" docDate="2023" docId="03BF87BFFFA8FFE6A7E0FF75FD2E89F2" docLanguage="en" docName="zootaxa.5351.1.5.pdf" docOrigin="Zootaxa 5351 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5351.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tohlezkus uminchu Jaloszynski 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="136" masterDocId="FF86FFC7FFA3FFE8A777FFE2FFCA8A57" masterDocTitle="A new genus, new species and new records of Eucinetidae in Japan and Taiwan (Coleoptera)" masterLastPageNumber="138" masterPageNumber="122" pageNumber="133" updateTime="1695989500669" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="BEAF7DAC0BC6A652C19C6A8F1F0A783E">
<mods:title id="66B0099B03B77485EB963C190596376B">A new genus, new species and new records of Eucinetidae in Japan and Taiwan (Coleoptera)</mods:title>
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<mods:namePart id="389161C87EEF404B2A1101D89065C018">Jałoszyński, Paweł</mods:namePart>
<mods:affiliation id="7CD0C839FEAEA39C469D673125F9B2C2">Museum of Natural History, University of Wrocław, Sienkiewicza 21, 50 - 335 Wrocław, Poland</mods:affiliation>
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<mods:namePart id="D5E8A1C83CA3848E1A84F5DF50ACE28B">Wakimura, Ryotaro</mods:namePart>
<mods:nameIdentifier id="5127379232B3A83D11AB9D54994D4F22" type="ORCID">0009-0000-8410-0567</mods:nameIdentifier>
<mods:affiliation id="06ED0E79FA449C63A05823C66749AB74">Department of Biology, School of Biological Sciences, Tokai University, Hokkaido, 005 - 8601, Japan https: // orcid. org / 0009 - 0000 - 8410 - 0567</mods:affiliation>
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<mods:date id="2E37BFCB4AFCD31F567EB3E113C2B83D">2023</mods:date>
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<mods:number id="297AEE3A2AB6085BF4A761D327115C03">2023-09-25</mods:number>
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<mods:number id="4B5E463C03EFEFE671B6C2E34873A54F">5351</mods:number>
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<mods:identifier id="F34382453651CDA92204D8285A189B9E" type="DOI">10.11646/zootaxa.5351.1.5</mods:identifier>
<mods:identifier id="FDC90D28D437C2568E521C4DEA9298D3" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03BF87BFFFA8FFE6A7E0FF75FD2E89F2" LSID="urn:lsid:plazi:treatment:03BF87BFFFA8FFE6A7E0FF75FD2E89F2" httpUri="http://treatment.plazi.org/id/03BF87BFFFA8FFE6A7E0FF75FD2E89F2" lastPageId="14" lastPageNumber="136" pageId="11" pageNumber="133">
<subSubSection id="C30C6522FFA8FFE3A7E0FF75FDAF8AE6" box="[151,613,150,178]" pageId="11" pageNumber="133" type="nomenclature">
<paragraph id="8BA936A9FFA8FFE3A7E0FF75FDAF8AE6" blockId="11.[151,613,150,214]" box="[151,613,150,178]" pageId="11" pageNumber="133">
<heading id="D0E181C5FFA8FFE3A7E0FF75FDAF8AE6" bold="true" box="[151,613,150,178]" fontSize="11" level="1" pageId="11" pageNumber="133" reason="1">
<emphasis id="B962EABBFFA8FFE3A7E0FF75FDAF8AE6" bold="true" box="[151,613,150,178]" pageId="11" pageNumber="133">
<taxonomicName id="4C164D2AFFA8FFE3A7E0FF75FDD18AE6" authority="Jaloszynski" authorityName="Jaloszynski" authorityYear="2023" box="[151,539,150,178]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="133" phylum="Arthropoda" rank="species" species="uminchu" status="sp. n.">
<emphasis id="B962EABBFFA8FFE3A7E0FF75FE4F8AE5" bold="true" box="[151,389,151,178]" italics="true" pageId="11" pageNumber="133">Tohlezkus uminchu</emphasis>
Jałoszyński
</taxonomicName>
<taxonomicNameLabel id="A25157C0FFA8FFE3A554FF75FDAF8AE6" box="[547,613,151,177]" pageId="11" pageNumber="133" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30C6522FFA8FFE3A7E0FF59FE6F8A82" box="[151,421,187,214]" pageId="11" pageNumber="133" type="description">
<paragraph id="8BA936A9FFA8FFE3A7E0FF59FE6F8A82" blockId="11.[151,613,150,214]" box="[151,421,187,214]" pageId="11" pageNumber="133">
(
<figureCitation id="132D2A2CFFA8FFE3A7E8FF59FEEA8A82" box="[159,288,187,213]" captionStart-0="FIGURES 23‒24" captionStart-1="FIGURES 25‒29" captionStart-2="FIGURES 30‒32" captionStart-3="FIGURES 33‒34" captionStartId-0="9.[151,264,1856,1880]" captionStartId-1="10.[151,264,1726,1750]" captionStartId-2="11.[151,264,1096,1120]" captionStartId-3="11.[151,264,1899,1923]" captionTargetBox-0="[151,1436,648,1832]" captionTargetBox-1="[151,1435,181,1702]" captionTargetBox-2="[151,1435,381,1073]" captionTargetBox-3="[151,1435,1181,1876]" captionTargetId-0="figure-539@9.[151,1436,648,1832]" captionTargetId-1="figure-17@10.[151,1435,181,1702]" captionTargetId-2="figure-75@11.[151,1435,381,1073]" captionTargetId-3="figure-111@11.[151,1435,1181,1876]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="11" captionText-0="FIGURES 23‒24. Tohlezkus uminchu sp. n. Dorsal (23) and ventral (24) habitus." captionText-1="FIGURES 25‒29. Tohlezkus uminchu sp. n. Head and pronotum in anterodorsal view (25); head in dorsal (26) and ventral (27) views with suctorial mouthparts tilted ventrally; labrum and prelabium directed ventrally (28); left antenna in ventral view (29)." captionText-2="FIGURES 30‒32. Tohlezkus uminchu sp. n. Head and thorax in ventral view (30); middle leg in ventral view (31); apical spurs of mesotibia in ventral view (32)." captionText-3="FIGURES 33‒34. Metaventral features to distinguish Tohlezkus uminchu sp. n. (33) from T. rufus (M. Sakai) (34). Arrowheads indicate (from left to right): anterior marginal line of anterior metaventral process; shape of median region of metaventrite exposed between metacoxal plates; and shape of lateral margin of metacoxal plate." figureDoi-0="http://doi.org/10.5281/zenodo.8391341" figureDoi-1="http://doi.org/10.5281/zenodo.8391343" figureDoi-2="http://doi.org/10.5281/zenodo.8391347" figureDoi-3="http://doi.org/10.5281/zenodo.8391349" httpUri-0="https://zenodo.org/record/8391341/files/figure.png" httpUri-1="https://zenodo.org/record/8391343/files/figure.png" httpUri-2="https://zenodo.org/record/8391347/files/figure.png" httpUri-3="https://zenodo.org/record/8391349/files/figure.png" pageId="11" pageNumber="133">Figs 23–33</figureCitation>
,
<figureCitation id="132D2A2CFFA8FFE3A65BFF59FEBE8A81" box="[300,372,187,214]" captionStart="FIGURES 35‒36" captionStartId="12.[151,264,1251,1275]" captionTargetBox="[151,1435,181,1227]" captionTargetId="figure-17@12.[151,1435,181,1227]" captionTargetPageId="12" captionText="FIGURES 35‒36. Tohlezkus uminchu sp. n. Aedeagus (35) and ovipositor (36) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.8391351" httpUri="https://zenodo.org/record/8391351/files/figure.png" pageId="11" pageNumber="133">35–36</figureCitation>
,
<figureCitation id="132D2A2CFFA8FFE3A6F7FF59FE578A82" box="[384,413,187,213]" captionStart="FIGURES 37‒40" captionStartId="13.[151,264,1195,1219]" captionTargetBox="[163,1426,190,1160]" captionTargetId="figure-22@13.[151,1435,181,1171]" captionTargetPageId="13" captionText="FIGURES 37‒40. Aedeagus. Bisayodes hure gen. &amp; sp. n. (37); Tohlezkus uminchu sp. n. (38); Tohlezkus orientalis Vít (39); Tohlezkus rufus (M. Sakai) (40)." figureDoi="http://doi.org/10.5281/zenodo.8391357" httpUri="https://zenodo.org/record/8391357/files/figure.png" pageId="11" pageNumber="133">38</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30C6522FFA8FFE3A7E0FEE1FB6C8B32" pageId="11" pageNumber="133" type="materials_examined">
<paragraph id="8BA936A9FFA8FFE3A7E0FEE1FB6C8B32" blockId="11.[151,1437,259,358]" pageId="11" pageNumber="133">
<emphasis id="B962EABBFFA8FFE3A7E0FEE1FE6D8B4A" bold="true" box="[151,423,259,285]" pageId="11" pageNumber="133">Type material studied.</emphasis>
<materialsCitation id="3B7E3CF4FFA8FFE3A6C7FEE1FD4A8B32" collectingDate="2023-02-26" collectionCode="NSMT" country="Japan" latitude="26.75436" location="Jashiki" longLatPrecision="1" longitude="128.24579" municipality="Ohkuni-rindo" pageId="11" pageNumber="133" specimenCount="2" stateProvince="Okinawa" typeStatus="holotype">
<emphasis id="B962EABBFFA8FFE3A6C7FEE1FD518B4A" bold="true" box="[432,667,259,285]" pageId="11" pageNumber="133">
<typeStatus id="54AD880BFFA8FFE3A6C7FEE1FDD78B4A" box="[432,541,259,285]" pageId="11" pageNumber="133">Holotype</typeStatus>
(
<collectingCountry id="F3017639FFA8FFE3A55AFEE1FD448B4A" box="[557,654,259,285]" name="Japan" pageId="11" pageNumber="133">JAPAN</collectingCountry>
):
</emphasis>
♁, two labels: “JAPAN,
<collectingRegion id="49D2F84BFFA8FFE3A4C8FEE1FB4A8B4A" box="[959,1152,259,285]" country="Japan" name="Okinawa" pageId="11" pageNumber="133">OKINAWA Pref.</collectingRegion>
/ OKINAWA-jima,
<date id="FFA81069FFA8FFE3A21FFEE1FF058B16" pageId="11" pageNumber="133" value="2023-02-26">
<collectingDate id="EFECE981FFA8FFE3A21FFEE1FF058B16" pageId="11" pageNumber="133" value="2023-02-26">26 ii 2023</collectingDate>
</date>
/
<collectingMunicipality id="6BCDACD3FFA8FFE3A794FEC5FEB48B16" box="[227,382,295,321]" pageId="11" pageNumber="133">Ohkuni-rindo</collectingMunicipality>
,
<location id="8EC96072FFA8FFE3A6FFFEC5FE1C8B16" LSID="urn:lsid:plazi:treatment:03BF87BFFFA8FFE6A7E0FF75FD2E89F2:8EC96072FFA8FFE3A6FFFEC5FE1C8B16" box="[392,470,295,321]" country="Japan" latitude="26.75436" longLatPrecision="1" longitude="128.24579" municipality="Ohkuni-rindo" name="Jashiki" pageId="11" pageNumber="133" stateProvince="Okinawa">Jashiki</location>
/
<geoCoordinate id="EE22506EFFA8FFE3A69DFEC5FD4B8B15" box="[490,641,295,322]" degrees="26" direction="north" minutes="45" orientation="latitude" pageId="11" pageNumber="133" precision="1" seconds="15.7" value="26.75436">26°45'15.7''N</geoCoordinate>
/
<geoCoordinate id="EE22506EFFA8FFE3A5FDFEC5FCE18B15" box="[650,811,295,322]" degrees="128" direction="east" minutes="14" orientation="longitude" pageId="11" pageNumber="133" precision="1" seconds="44.8" value="128.24579">128°14'44.8''E</geoCoordinate>
/ leg. P. JAŁOSZYŃSKI”, “
<emphasis id="B962EABBFFA8FFE3A31EFEC5FAB38B16" box="[1129,1401,295,322]" italics="true" pageId="11" pageNumber="133">
<taxonomicName id="4C164D2AFFA8FFE3A31EFEC5FACC8B16" box="[1129,1286,295,321]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="133" phylum="Arthropoda" rank="genus">TOHLEZKUS</taxonomicName>
/
<taxonomicName id="4C164D2AFFA8FFE3A26EFEC5FAB38B16" authorityName="Jaloszynski" authorityYear="2023" box="[1305,1401,295,321]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="133" phylum="Arthropoda" rank="species" species="uminchu">uminchu</taxonomicName>
</emphasis>
m. / Jałoszyński ‘23/
<typeStatus id="54AD880BFFA8FFE3A615FEA9FDC18B32" box="[354,523,331,357]" pageId="11" pageNumber="133">HOLOTYPUS</typeStatus>
” (
<collectionCode id="ED07AE6CFFA8FFE3A553FEA9FDB28B32" box="[548,632,331,357]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="11" pageNumber="133">NSMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7E3CF4FFA8FFE3A5FDFEA9FB6B8B32" box="[650,1185,331,358]" collectingDate="2023-02-26" collectionCode="NSMT" country="Japan" latitude="26.75436" location="Jashiki" longLatPrecision="1" longitude="128.24579" municipality="Ohkuni-rindo" pageId="11" pageNumber="133" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="paratype">
<emphasis id="B962EABBFFA8FFE3A5FDFEA9FCCB8B32" bold="true" box="[650,769,331,357]" pageId="11" pageNumber="133">
<typeStatus id="54AD880BFFA8FFE3A5FDFEA9FD318B32" box="[650,763,331,357]" pageId="11" pageNumber="133">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D10FD20FFA8FFE3A47FFEA9FCF88B31" box="[776,818,331,358]" pageId="11" pageNumber="133" type="female">1 ♀</specimenCount>
, same data as for holotype (cPJ)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF696621FFA8FFE3A7E0FBAAFE208ED2" ID-DOI="http://doi.org/10.5281/zenodo.8391347" ID-Zenodo-Dep="8391347" httpUri="https://zenodo.org/record/8391347/files/figure.png" pageId="11" pageNumber="133" startId="11.[151,264,1096,1120]" targetBox="[151,1435,381,1073]" targetPageId="11" targetType="figure">
<paragraph id="8BA936A9FFA8FFE3A7E0FBAAFE208ED2" blockId="11.[151,1436,1096,1157]" pageId="11" pageNumber="133">
<emphasis id="B962EABBFFA8FFE3A7E0FBAAFE9E8E37" bold="true" box="[151,340,1096,1120]" pageId="11" pageNumber="133">FIGURES 30‒32.</emphasis>
<taxonomicName id="4C164D2AFFA8FFE3A62EFBAAFDD38E37" authority="Jałoszyński &amp; Wakimura, 2023" authorityName="Jałoszyński &amp; Wakimura" authorityYear="2023" box="[345,537,1096,1120]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="133" phylum="Arthropoda" rank="species" species="uminchu" status="sp. n.">
<emphasis id="B962EABBFFA8FFE3A62EFBAAFDD38E37" box="[345,537,1096,1120]" italics="true" pageId="11" pageNumber="133">Tohlezkus uminchu</emphasis>
</taxonomicName>
<emphasis id="B962EABBFFA8FFE3A568FBABFD928E36" bold="true" box="[543,600,1097,1121]" pageId="11" pageNumber="133">
<taxonomicNameLabel id="A25157C0FFA8FFE3A568FBABFD928E36" box="[543,600,1097,1121]" pageId="11" pageNumber="133" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Head and thorax in ventral view (
<emphasis id="B962EABBFFA8FFE3A4C7FBAAFC038E37" bold="true" box="[944,969,1096,1120]" pageId="11" pageNumber="133">30</emphasis>
); middle leg in ventral view (
<emphasis id="B962EABBFFA8FFE3A381FBAAFAC58E37" bold="true" box="[1270,1295,1096,1120]" pageId="11" pageNumber="133">31</emphasis>
); apical spurs of mesotibia in ventral view (
<emphasis id="B962EABBFFA8FFE3A6B5FB8EFE118ED3" bold="true" box="[450,475,1132,1156]" pageId="11" pageNumber="133">32</emphasis>
).
</paragraph>
</caption>
<caption id="DF696621FFA8FFE3A7E0F889FC148D9B" ID-DOI="http://doi.org/10.5281/zenodo.8391349" ID-Zenodo-Dep="8391349" httpUri="https://zenodo.org/record/8391349/files/figure.png" pageId="11" pageNumber="133" startId="11.[151,264,1899,1923]" targetBox="[151,1435,1181,1876]" targetPageId="11" targetType="figure">
<paragraph id="8BA936A9FFA8FFE3A7E0F889FC148D9B" blockId="11.[151,1437,1899,1996]" pageId="11" pageNumber="133">
<emphasis id="B962EABBFFA8FFE3A7E0F889FE9E8DD3" bold="true" box="[151,340,1899,1924]" pageId="11" pageNumber="133">FIGURES 33‒34.</emphasis>
Metaventral features to distinguish
<taxonomicName id="4C164D2AFFA8FFE3A5CAF88EFCB78DD3" authority="Jałoszyński &amp; Wakimura, 2023" authorityName="Jałoszyński &amp; Wakimura" authorityYear="2023" box="[701,893,1900,1924]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="133" phylum="Arthropoda" rank="species" species="uminchu" status="sp. n.">
<emphasis id="B962EABBFFA8FFE3A5CAF88EFCB78DD3" box="[701,893,1900,1924]" italics="true" pageId="11" pageNumber="133">Tohlezkus uminchu</emphasis>
</taxonomicName>
<emphasis id="B962EABBFFA8FFE3A4F4F88EFC718DD3" bold="true" box="[899,955,1900,1924]" pageId="11" pageNumber="133">
<taxonomicNameLabel id="A25157C0FFA8FFE3A4F4F88EFC718DD3" box="[899,955,1900,1924]" pageId="11" pageNumber="133" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<emphasis id="B962EABBFFA8FFE3A4BDF88EFC298DD3" bold="true" box="[970,995,1900,1924]" pageId="11" pageNumber="133">33</emphasis>
) from
<taxonomicName id="4C164D2AFFA8FFE3A35FF88FFB2C8DD3" authority="(M. Sakai)" authorityName="" baseAuthorityName="M. Sakai" baseAuthorityYear="1980" box="[1064,1254,1900,1924]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="133" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="B962EABBFFA8FFE3A35FF88FFBBE8DD3" box="[1064,1140,1900,1924]" italics="true" pageId="11" pageNumber="133">T. rufus</emphasis>
(M. Sakai)
</taxonomicName>
(
<emphasis id="B962EABBFFA8FFE3A383F88EFAC78DD3" bold="true" box="[1268,1293,1900,1924]" pageId="11" pageNumber="133">34</emphasis>
). Arrowheads indicate (from left to right): anterior marginal line of anterior metaventral process; shape of median region of metaventrite exposed between metacoxal plates; and shape of lateral margin of metacoxal plate.
</paragraph>
</caption>
<caption id="DF696621FFAFFFE4A7E0FB01FBAA8EAC" ID-DOI="http://doi.org/10.5281/zenodo.8391351" ID-Zenodo-Dep="8391351" box="[151,1120,1251,1275]" httpUri="https://zenodo.org/record/8391351/files/figure.png" pageId="12" pageNumber="134" startId="12.[151,264,1251,1275]" targetBox="[151,1435,181,1227]" targetPageId="12" targetType="figure">
<paragraph id="8BA936A9FFAFFFE4A7E0FB01FBAA8EAC" blockId="12.[151,1120,1251,1275]" box="[151,1120,1251,1275]" pageId="12" pageNumber="134">
<emphasis id="B962EABBFFAFFFE4A7E0FB01FE9E8EAC" bold="true" box="[151,340,1251,1275]" pageId="12" pageNumber="134">FIGURES 35‒36.</emphasis>
<taxonomicName id="4C164D2AFFAFFFE4A62DFB01FDD08EAC" authority="Jałoszyński &amp; Wakimura, 2023" authorityName="Jałoszyński &amp; Wakimura" authorityYear="2023" box="[346,538,1251,1275]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="134" phylum="Arthropoda" rank="species" species="uminchu" status="sp. n.">
<emphasis id="B962EABBFFAFFFE4A62DFB01FDD08EAC" box="[346,538,1251,1275]" italics="true" pageId="12" pageNumber="134">Tohlezkus uminchu</emphasis>
</taxonomicName>
<emphasis id="B962EABBFFAFFFE4A556FB01FD908EAC" bold="true" box="[545,602,1251,1275]" pageId="12" pageNumber="134">
<taxonomicNameLabel id="A25157C0FFAFFFE4A556FB01FD908EAC" box="[545,602,1251,1275]" pageId="12" pageNumber="134" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Aedeagus (
<emphasis id="B962EABBFFAFFFE4A5A5FB01FD218EAC" bold="true" box="[722,747,1251,1275]" pageId="12" pageNumber="134">35</emphasis>
) and ovipositor (
<emphasis id="B962EABBFFAFFFE4A4EDFB01FC798EAC" bold="true" box="[922,947,1251,1275]" pageId="12" pageNumber="134">36</emphasis>
) in ventral view.
</paragraph>
</caption>
<subSubSection id="C30C6522FFAFFFE4A7B0FAAFFA4C8F84" pageId="12" pageNumber="134" type="diagnosis">
<paragraph id="8BA936A9FFAFFFE4A7B0FAAFFA4C8F84" blockId="12.[151,1437,1357,2031]" pageId="12" pageNumber="134">
<emphasis id="B962EABBFFAFFFE4A7B0FAAFFE8A8F30" bold="true" box="[199,320,1357,1383]" pageId="12" pageNumber="134">Diagnosis.</emphasis>
Apical rod-like region of maxillary palpomere 4 shorter than 1/4 length of entire palpomere; posterior marginal carina of anterior metaventral process forming inversely U-shaped anterior expansion at middle; triangular median region of metaventrite about as long as 1/3 length of metacoxal plate; lateral margin of metacoxal plate barely noticeably rounded and in anterior half straight; each paramere with distinctly concave mesal margin in distal 1/3.
</paragraph>
</subSubSection>
<subSubSection id="C30C6522FFAFFFE6A7B0FA3FFB178BA2" lastPageId="14" lastPageNumber="136" pageId="12" pageNumber="134" type="description">
<paragraph id="8BA936A9FFAFFFE4A7B0FA3FFA478C4B" blockId="12.[151,1437,1357,2031]" pageId="12" pageNumber="134">
<emphasis id="B962EABBFFAFFFE4A7B0FA3FFE938FA0" bold="true" box="[199,345,1501,1527]" pageId="12" pageNumber="134">Description.</emphasis>
Body of male (
<figureCitation id="132D2A2CFFAFFFE4A564FA3FFD948FA0" box="[531,606,1501,1527]" captionStart="FIGURES 1‒4" captionStartId="1.[151,264,1474,1498]" captionTargetBox="[157,1429,860,1439]" captionTargetId="figure-191@1.[151,1435,849,1451]" captionTargetPageId="1" captionText="FIGURES 1‒4. Dorsal habitus of males. Bisayodes hure gen. &amp; sp. n. (1); Tohlezkus uminchu sp. n. (2); Tohlezkus orientalis Vít (3); Tohlezkus rufus (M. Sakai) (4)." figureDoi="http://doi.org/10.5281/zenodo.8391331" httpUri="https://zenodo.org/record/8391331/files/figure.png" pageId="12" pageNumber="134">Figs 2</figureCitation>
,
<figureCitation id="132D2A2CFFAFFFE4A51AFA3FFD7F8FA0" box="[621,693,1501,1527]" captionStart="FIGURES 23‒24" captionStartId="9.[151,264,1856,1880]" captionTargetBox="[151,1436,648,1832]" captionTargetId="figure-539@9.[151,1436,648,1832]" captionTargetPageId="9" captionText="FIGURES 23‒24. Tohlezkus uminchu sp. n. Dorsal (23) and ventral (24) habitus." figureDoi="http://doi.org/10.5281/zenodo.8391341" httpUri="https://zenodo.org/record/8391341/files/figure.png" pageId="12" pageNumber="134">23–24</figureCitation>
) elongate oval, strongly convex dorsally and flattened ventrally, brown with umbra hue, appendages indistinctly lighter than body, setae slightly lighter than cuticle; BL
<quantity id="4CEE9B4CFFAFFFE4A255F9E3FA438C4B" box="[1314,1417,1537,1564]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" pageId="12" pageNumber="134" unit="mm" value="1.18">1.18 mm</quantity>
.
</paragraph>
<paragraph id="8BA936A9FFAFFFE4A7B0F9C7FD7D8C87" blockId="12.[151,1437,1357,2031]" pageId="12" pageNumber="134">
Head (
<figureCitation id="132D2A2CFFAFFFE4A666F9C7FE5B8C17" box="[273,401,1573,1600]" captionStart="FIGURES 25‒29" captionStartId="10.[151,264,1726,1750]" captionTargetBox="[151,1435,181,1702]" captionTargetId="figure-17@10.[151,1435,181,1702]" captionTargetPageId="10" captionText="FIGURES 25‒29. Tohlezkus uminchu sp. n. Head and pronotum in anterodorsal view (25); head in dorsal (26) and ventral (27) views with suctorial mouthparts tilted ventrally; labrum and prelabium directed ventrally (28); left antenna in ventral view (29)." figureDoi="http://doi.org/10.5281/zenodo.8391343" httpUri="https://zenodo.org/record/8391343/files/figure.png" pageId="12" pageNumber="134">Figs 25–28</figureCitation>
) broadest at eyes, HL
<quantity id="4CEE9B4CFFAFFFE4A5FAF9C7FD398C17" box="[653,755,1573,1600]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="134" unit="mm" value="0.13">0.13 mm</quantity>
, HW
<quantity id="4CEE9B4CFFAFFFE4A443F9C7FC518C17" box="[820,923,1573,1600]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="12" pageNumber="134" unit="mm" value="0.26">0.26 mm</quantity>
; lateral margins of head behind eyes in dorsal view straight, vertex and frons together weakly convex, strongly transverse; punctures on dorsum small, sharply marked and evenly distributed, separated by spaces 2–3 × as wide as diameters of punctures, microsculpture of frons and vertex scale-like; setae short and sparse, weakly suberect, directed anteriorly. Genae (
<figureCitation id="132D2A2CFFAFFFE4A3A6F973FAE98CFC" box="[1233,1315,1681,1707]" captionStart="FIGURES 25‒29" captionStartId="10.[151,264,1726,1750]" captionTargetBox="[151,1435,181,1702]" captionTargetId="figure-17@10.[151,1435,181,1702]" captionTargetPageId="10" captionText="FIGURES 25‒29. Tohlezkus uminchu sp. n. Head and pronotum in anterodorsal view (25); head in dorsal (26) and ventral (27) views with suctorial mouthparts tilted ventrally; labrum and prelabium directed ventrally (28); left antenna in ventral view (29)." figureDoi="http://doi.org/10.5281/zenodo.8391343" httpUri="https://zenodo.org/record/8391343/files/figure.png" pageId="12" pageNumber="134">Fig. 27</figureCitation>
) and short postgenae asetose; gula and submentum asetose.
</paragraph>
<paragraph id="8BA936A9FFAFFFE4A7B0F938FE698D6C" blockId="12.[151,1437,1357,2031]" pageId="12" pageNumber="134">
Antennae (
<figureCitation id="132D2A2CFFAFFFE4A633F93BFE528CA4" box="[324,408,1753,1779]" captionStart="FIGURES 25‒29" captionStartId="10.[151,264,1726,1750]" captionTargetBox="[151,1435,181,1702]" captionTargetId="figure-17@10.[151,1435,181,1702]" captionTargetPageId="10" captionText="FIGURES 25‒29. Tohlezkus uminchu sp. n. Head and pronotum in anterodorsal view (25); head in dorsal (26) and ventral (27) views with suctorial mouthparts tilted ventrally; labrum and prelabium directed ventrally (28); left antenna in ventral view (29)." figureDoi="http://doi.org/10.5281/zenodo.8391343" httpUri="https://zenodo.org/record/8391343/files/figure.png" pageId="12" pageNumber="134">Fig. 29</figureCitation>
) distinctly longer than length of entire head capsule, AnL
<quantity id="4CEE9B4CFFAFFFE4A349F93BFB628CA3" box="[1086,1192,1753,1780]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="134" unit="mm" value="0.25">0.25 mm</quantity>
; scape narrower than pedicel, transverse, pedicel subquadrate, antennomeres 3–10 each distinctly transverse, 11 subquadrate, nearly as long as 9–10 combined.
</paragraph>
<paragraph id="8BA936A9FFAFFFE4A7B0F8A4FE4B8DB8" blockId="12.[151,1437,1357,2031]" pageId="12" pageNumber="134">
Pronotum (
<figureCitation id="132D2A2CFFAFFFE4A631F8A7FE5B8D08" box="[326,401,1861,1887]" captionStart="FIGURES 1‒4" captionStartId="1.[151,264,1474,1498]" captionTargetBox="[157,1429,860,1439]" captionTargetId="figure-191@1.[151,1435,849,1451]" captionTargetPageId="1" captionText="FIGURES 1‒4. Dorsal habitus of males. Bisayodes hure gen. &amp; sp. n. (1); Tohlezkus uminchu sp. n. (2); Tohlezkus orientalis Vít (3); Tohlezkus rufus (M. Sakai) (4)." figureDoi="http://doi.org/10.5281/zenodo.8391331" httpUri="https://zenodo.org/record/8391331/files/figure.png" pageId="12" pageNumber="134">Figs 2</figureCitation>
,
<figureCitation id="132D2A2CFFAFFFE4A6E8F8A7FE778D08" box="[415,445,1861,1887]" captionStart="FIGURES 23‒24" captionStartId="9.[151,264,1856,1880]" captionTargetBox="[151,1436,648,1832]" captionTargetId="figure-539@9.[151,1436,648,1832]" captionTargetPageId="9" captionText="FIGURES 23‒24. Tohlezkus uminchu sp. n. Dorsal (23) and ventral (24) habitus." figureDoi="http://doi.org/10.5281/zenodo.8391341" httpUri="https://zenodo.org/record/8391341/files/figure.png" pageId="12" pageNumber="134">23</figureCitation>
,
<figureCitation id="132D2A2CFFAFFFE4A6BDF8A7FE2D8D37" box="[458,487,1861,1888]" captionStart="FIGURES 25‒29" captionStartId="10.[151,264,1726,1750]" captionTargetBox="[151,1435,181,1702]" captionTargetId="figure-17@10.[151,1435,181,1702]" captionTargetPageId="10" captionText="FIGURES 25‒29. Tohlezkus uminchu sp. n. Head and pronotum in anterodorsal view (25); head in dorsal (26) and ventral (27) views with suctorial mouthparts tilted ventrally; labrum and prelabium directed ventrally (28); left antenna in ventral view (29)." figureDoi="http://doi.org/10.5281/zenodo.8391343" httpUri="https://zenodo.org/record/8391343/files/figure.png" pageId="12" pageNumber="134">25</figureCitation>
) less convex than elytra, broadest at posterior corners, PL
<quantity id="4CEE9B4CFFAFFFE4A3FEF8A7FB3A8D37" box="[1161,1264,1861,1888]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="12" pageNumber="134" unit="mm" value="0.23">0.23 mm</quantity>
, PW
<quantity id="4CEE9B4CFFAFFFE4A247F8A7FA538D37" box="[1328,1433,1861,1888]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="12" pageNumber="134" unit="mm" value="0.43">0.43 mm</quantity>
. In dorsal view, anterior and lateral margins strongly and evenly rounded; posterior corners strongly projecting posteriorly and sharp-angled; posterior margin bisinuate, medially strongly and evenly convex. Punctures and setae on pronotal disc similar to those on frons and vertex, but surface not microsculptured. Hypomera asetose and virtually impunctate.
</paragraph>
<caption id="DF696621FFAEFFE5A7E0FB49FE148EBF" ID-DOI="http://doi.org/10.5281/zenodo.8391357" ID-Zenodo-Dep="8391357" httpUri="https://zenodo.org/record/8391357/files/figure.png" pageId="13" pageNumber="135" startId="13.[151,264,1195,1219]" targetBox="[163,1426,190,1160]" targetPageId="13" targetType="figure">
<paragraph id="8BA936A9FFAEFFE5A7E0FB49FE148EBF" blockId="13.[151,1436,1195,1256]" pageId="13" pageNumber="135">
<emphasis id="B962EABBFFAEFFE5A7E0FB49FE9F8E94" bold="true" box="[151,341,1195,1219]" pageId="13" pageNumber="135">FIGURES 37‒40.</emphasis>
Aedeagus.
<taxonomicName id="4C164D2AFFAEFFE5A6BBFB49FDAC8E94" authorityName="Wakimura &amp; Jaloszynski" authorityYear="2023" box="[460,614,1195,1219]" class="Insecta" family="Eucinetidae" genus="Bisayodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="135" phylum="Arthropoda" rank="species" species="hure">
<emphasis id="B962EABBFFAEFFE5A6BBFB49FDAC8E94" box="[460,614,1195,1219]" italics="true" pageId="13" pageNumber="135">Bisayodes hure</emphasis>
</taxonomicName>
<emphasis id="B962EABBFFAEFFE5A51AFB4EFD508E93" bold="true" box="[621,666,1196,1220]" pageId="13" pageNumber="135">gen.</emphasis>
&amp;
<emphasis id="B962EABBFFAEFFE5A5CCFB4EFD3E8E93" bold="true" box="[699,756,1196,1220]" pageId="13" pageNumber="135">sp. n.</emphasis>
(
<emphasis id="B962EABBFFAEFFE5A473FB49FCD78E94" bold="true" box="[772,797,1195,1219]" pageId="13" pageNumber="135">37</emphasis>
);
<taxonomicName id="4C164D2AFFAEFFE5A444FB49FC3E8E94" authority="Jałoszyński &amp; Wakimura, 2023" authorityName="Jałoszyński &amp; Wakimura" authorityYear="2023" box="[819,1012,1195,1219]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="135" phylum="Arthropoda" rank="species" species="uminchu" status="sp. n.">
<emphasis id="B962EABBFFAEFFE5A444FB49FC3E8E94" box="[819,1012,1195,1219]" italics="true" pageId="13" pageNumber="135">Tohlezkus uminchu</emphasis>
</taxonomicName>
<emphasis id="B962EABBFFAEFFE5A48CFB4EFBFE8E93" bold="true" box="[1019,1076,1196,1220]" pageId="13" pageNumber="135">
<taxonomicNameLabel id="A25157C0FFAEFFE5A48CFB4EFBFE8E93" box="[1019,1076,1196,1220]" pageId="13" pageNumber="135" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<emphasis id="B962EABBFFAEFFE5A334FB49FB968E94" bold="true" box="[1091,1116,1195,1219]" pageId="13" pageNumber="135">38</emphasis>
);
<taxonomicName id="4C164D2AFFAEFFE5A304FB49FAAE8E93" authority="Vit" authorityName="Vit" authorityYear="1981" box="[1139,1380,1195,1220]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="135" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B962EABBFFAEFFE5A304FB49FAF48E94" box="[1139,1342,1195,1219]" italics="true" pageId="13" pageNumber="135">Tohlezkus orientalis</emphasis>
Vít
</taxonomicName>
(
<emphasis id="B962EABBFFAEFFE5A204FB49FA468E94" bold="true" box="[1395,1420,1195,1219]" pageId="13" pageNumber="135">39</emphasis>
);
<taxonomicName id="4C164D2AFFAEFFE5A7E0FB2DFE6D8EB0" authority="(M. Sakai)" authorityName="" baseAuthorityName="M. Sakai" baseAuthorityYear="1980" box="[151,423,1231,1256]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="135" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="B962EABBFFAEFFE5A7E0FB2DFEFF8EB0" box="[151,309,1231,1255]" italics="true" pageId="13" pageNumber="135">Tohlezkus rufus</emphasis>
(M. Sakai)
</taxonomicName>
(
<emphasis id="B962EABBFFAEFFE5A6C1FB2DFE058EB0" bold="true" box="[438,463,1231,1255]" pageId="13" pageNumber="135">40</emphasis>
).
</paragraph>
</caption>
<caption id="DF696621FFAEFFE5A7E0F914FAB48D58" ID-DOI="http://doi.org/10.5281/zenodo.8391359" ID-Zenodo-Dep="8391359" box="[151,1406,1782,1809]" httpUri="https://zenodo.org/record/8391359/files/figure.png" pageId="13" pageNumber="135" startId="13.[151,264,1782,1806]" targetBox="[151,1435,1280,1759]" targetPageId="13" targetType="figure">
<paragraph id="8BA936A9FFAEFFE5A7E0F914FAB48D58" blockId="13.[151,1406,1782,1809]" box="[151,1406,1782,1809]" pageId="13" pageNumber="135">
<emphasis id="B962EABBFFAEFFE5A7E0F914FE9E8D58" bold="true" box="[151,340,1782,1807]" pageId="13" pageNumber="135">FIGURES 41‒42.</emphasis>
Collecting sites of
<taxonomicName id="4C164D2AFFAEFFE5A56EF915FD798D58" authorityName="Wakimura &amp; Jaloszynski" authorityYear="2023" box="[537,691,1783,1807]" class="Insecta" family="Eucinetidae" genus="Bisayodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="135" phylum="Arthropoda" rank="species" species="hure">
<emphasis id="B962EABBFFAEFFE5A56EF915FD798D58" box="[537,691,1783,1807]" italics="true" pageId="13" pageNumber="135">Bisayodes hure</emphasis>
</taxonomicName>
<emphasis id="B962EABBFFAEFFE5A5CDF915FD2C8D58" bold="true" box="[698,742,1783,1807]" pageId="13" pageNumber="135">gen.</emphasis>
&amp;
<emphasis id="B962EABBFFAEFFE5A471F915FCF58D58" bold="true" box="[774,831,1783,1807]" pageId="13" pageNumber="135">sp. n.</emphasis>
in Hiroo-chô, Bitatanunke, Hokkaidô (© R. Wakimura).
</paragraph>
</caption>
<paragraph id="8BA936A9FFAEFFE5A7B0F883FD468D94" blockId="13.[151,1437,1889,2023]" pageId="13" pageNumber="135">
Elytra (
<figureCitation id="132D2A2CFFAEFFE5A66BF883FEAD8D2C" box="[284,359,1889,1915]" captionStart="FIGURES 1‒4" captionStartId="1.[151,264,1474,1498]" captionTargetBox="[157,1429,860,1439]" captionTargetId="figure-191@1.[151,1435,849,1451]" captionTargetPageId="1" captionText="FIGURES 1‒4. Dorsal habitus of males. Bisayodes hure gen. &amp; sp. n. (1); Tohlezkus uminchu sp. n. (2); Tohlezkus orientalis Vít (3); Tohlezkus rufus (M. Sakai) (4)." figureDoi="http://doi.org/10.5281/zenodo.8391331" httpUri="https://zenodo.org/record/8391331/files/figure.png" pageId="13" pageNumber="135">Figs 2</figureCitation>
,
<figureCitation id="132D2A2CFFAEFFE5A603F883FE5B8D2C" box="[372,401,1889,1915]" captionStart="FIGURES 23‒24" captionStartId="9.[151,264,1856,1880]" captionTargetBox="[151,1436,648,1832]" captionTargetId="figure-539@9.[151,1436,648,1832]" captionTargetPageId="9" captionText="FIGURES 23‒24. Tohlezkus uminchu sp. n. Dorsal (23) and ventral (24) habitus." figureDoi="http://doi.org/10.5281/zenodo.8391341" httpUri="https://zenodo.org/record/8391341/files/figure.png" pageId="13" pageNumber="135">23</figureCitation>
) together oval, broadest near anterior fourth and strongly narrowing posteriorly to broadly subtriangular apex; EL
<quantity id="4CEE9B4CFFAEFFE5A6D7F867FDCD8DF7" box="[416,519,1925,1952]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="13" pageNumber="135" unit="mm" value="0.83">0.83 mm</quantity>
, EW
<quantity id="4CEE9B4CFFAEFFE5A530F867FD648DF7" box="[583,686,1925,1952]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="13" pageNumber="135" unit="mm" value="0.58">0.58 mm</quantity>
, EI 1.42. Punctures and setae similar to those on frons and vertex, surface with weak scale-like microsculpture.
</paragraph>
<paragraph id="8BA936A9FFAEFFE5A7B0F82FFE7D8DB0" blockId="13.[151,1437,1889,2023]" box="[199,439,1997,2023]" pageId="13" pageNumber="135">Hind wings vestigial.</paragraph>
<paragraph id="8BA936A9FFADFFE6A7B0FF75FD588AAE" blockId="14.[151,1437,151,933]" pageId="14" pageNumber="136">
Legs (
<figureCitation id="132D2A2CFFADFFE6A679FF75FEAC8AE6" box="[270,358,151,177]" captionStart="FIGURES 23‒24" captionStartId="9.[151,264,1856,1880]" captionTargetBox="[151,1436,648,1832]" captionTargetId="figure-539@9.[151,1436,648,1832]" captionTargetPageId="9" captionText="FIGURES 23‒24. Tohlezkus uminchu sp. n. Dorsal (23) and ventral (24) habitus." figureDoi="http://doi.org/10.5281/zenodo.8391341" httpUri="https://zenodo.org/record/8391341/files/figure.png" pageId="14" pageNumber="136">Figs 24</figureCitation>
,
<figureCitation id="132D2A2CFFADFFE6A604FF75FE718AE6" box="[371,443,151,177]" captionStart="FIGURES 30‒32" captionStartId="11.[151,264,1096,1120]" captionTargetBox="[151,1435,381,1073]" captionTargetId="figure-75@11.[151,1435,381,1073]" captionTargetPageId="11" captionText="FIGURES 30‒32. Tohlezkus uminchu sp. n. Head and thorax in ventral view (30); middle leg in ventral view (31); apical spurs of mesotibia in ventral view (32)." figureDoi="http://doi.org/10.5281/zenodo.8391347" httpUri="https://zenodo.org/record/8391347/files/figure.png" pageId="14" pageNumber="136">30–32</figureCitation>
) short, robust, flattened dorsoventrally. Profemora with row of dense, thick setae along anterior margin in basal half of ventral surface; remaining legs unmodified. Apical spurs on mesotibiae (
<figureCitation id="132D2A2CFFADFFE6A236FF59FA5E8A82" box="[1345,1428,187,213]" captionStart="FIGURES 30‒32" captionStartId="11.[151,264,1096,1120]" captionTargetBox="[151,1435,381,1073]" captionTargetId="figure-75@11.[151,1435,381,1073]" captionTargetPageId="11" captionText="FIGURES 30‒32. Tohlezkus uminchu sp. n. Head and thorax in ventral view (30); middle leg in ventral view (31); apical spurs of mesotibia in ventral view (32)." figureDoi="http://doi.org/10.5281/zenodo.8391347" httpUri="https://zenodo.org/record/8391347/files/figure.png" pageId="14" pageNumber="136">Fig. 32</figureCitation>
) equal in length, sinuate, with rounded apices.
</paragraph>
<paragraph id="8BA936A9FFADFFE6A7B0FEE1FD818BDE" blockId="14.[151,1437,151,933]" pageId="14" pageNumber="136">
Aedeagus (
<figureCitation id="132D2A2CFFADFFE6A63EFEE1FE6F8B4A" box="[329,421,258,285]" captionStart="FIGURES 35‒36" captionStartId="12.[151,264,1251,1275]" captionTargetBox="[151,1435,181,1227]" captionTargetId="figure-17@12.[151,1435,181,1227]" captionTargetPageId="12" captionText="FIGURES 35‒36. Tohlezkus uminchu sp. n. Aedeagus (35) and ovipositor (36) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.8391351" httpUri="https://zenodo.org/record/8391351/files/figure.png" pageId="14" pageNumber="136">Figs 35</figureCitation>
,
<figureCitation id="132D2A2CFFADFFE6A6C1FEE1FE198B4A" box="[438,467,259,285]" captionStart="FIGURES 37‒40" captionStartId="13.[151,264,1195,1219]" captionTargetBox="[163,1426,190,1160]" captionTargetId="figure-22@13.[151,1435,181,1171]" captionTargetPageId="13" captionText="FIGURES 37‒40. Aedeagus. Bisayodes hure gen. &amp; sp. n. (37); Tohlezkus uminchu sp. n. (38); Tohlezkus orientalis Vít (39); Tohlezkus rufus (M. Sakai) (40)." figureDoi="http://doi.org/10.5281/zenodo.8391357" httpUri="https://zenodo.org/record/8391357/files/figure.png" pageId="14" pageNumber="136">38</figureCitation>
) strongly elongate, AeL
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; penis tapering towards subtriangular apex with membranous, inflatable preputial sac, lateral margins of penis straight; in sole studied male penis with exserted slender flagellum; parameres massive, at base as broad as penis, tapering towards rounded apices, with mesal margins clearly concave in distal third.
</paragraph>
<paragraph id="8BA936A9FFADFFE6A7B0FE71FB178BA2" blockId="14.[151,1437,151,933]" pageId="14" pageNumber="136">
<emphasis id="B962EABBFFADFFE6A7B0FE71FEEF8BFA" bold="true" box="[199,293,403,429]" pageId="14" pageNumber="136">Female.</emphasis>
Externally indistinguishable from male. BL
<quantity id="4CEE9B4CFFADFFE6A46CFE71FC488BFA" box="[795,898,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="14" pageNumber="136" unit="mm" value="1.23">1.23 mm</quantity>
; HL
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, HW
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, AnL
<quantity id="4CEE9B4CFFADFFE6A27DFE71FABB8BFA" box="[1290,1393,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="14" pageNumber="136" unit="mm" value="0.28">0.28 mm</quantity>
; PL
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, PW
<quantity id="4CEE9B4CFFADFFE6A64CFE55FE688B86" box="[315,418,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="14" pageNumber="136" unit="mm" value="0.48">0.48 mm</quantity>
; EL
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, EW
<quantity id="4CEE9B4CFFADFFE6A50BFE55FD298B86" box="[636,739,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="136" unit="mm" value="0.6">0.60 mm</quantity>
, EI 1.42. Gonocoxites (
<figureCitation id="132D2A2CFFADFFE6A49AFE55FBF58B86" box="[1005,1087,439,465]" captionStart="FIGURES 35‒36" captionStartId="12.[151,264,1251,1275]" captionTargetBox="[151,1435,181,1227]" captionTargetId="figure-17@12.[151,1435,181,1227]" captionTargetPageId="12" captionText="FIGURES 35‒36. Tohlezkus uminchu sp. n. Aedeagus (35) and ovipositor (36) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.8391351" httpUri="https://zenodo.org/record/8391351/files/figure.png" pageId="14" pageNumber="136">Fig. 36</figureCitation>
) subtriangular elongate, setose on outer margins, each with minute subconical and pointed gonostylus bearing several long setae.
</paragraph>
</subSubSection>
<subSubSection id="C30C6522FFADFFE6A7B0FE1CFCC4884E" box="[199,782,510,537]" pageId="14" pageNumber="136" type="distribution">
<paragraph id="8BA936A9FFADFFE6A7B0FE1CFCC4884E" blockId="14.[151,1437,151,933]" box="[199,782,510,537]" pageId="14" pageNumber="136">
<emphasis id="B962EABBFFADFFE6A7B0FE1CFEAB884F" bold="true" box="[199,353,510,536]" pageId="14" pageNumber="136">Distribution.</emphasis>
Southwestern
<collectingCountry id="F3017639FFADFFE6A570FE1DFD81884E" box="[519,587,511,537]" name="Japan" pageId="14" pageNumber="136">Japan</collectingCountry>
:
<collectingRegion id="49D2F84BFFADFFE6A522FE1DFD70884E" box="[597,698,511,537]" country="Japan" name="Okinawa" pageId="14" pageNumber="136">Okinawa</collectingRegion>
Island.
</paragraph>
</subSubSection>
<subSubSection id="C30C6522FFADFFE6A7B0FDC1FF3A88D2" pageId="14" pageNumber="136" type="etymology">
<paragraph id="8BA936A9FFADFFE6A7B0FDC1FF3A88D2" blockId="14.[151,1437,151,933]" pageId="14" pageNumber="136">
<emphasis id="B962EABBFFADFFE6A7B0FDC1FE84886A" bold="true" box="[199,334,547,573]" pageId="14" pageNumber="136">Etymology.</emphasis>
The noun in apposition
<taxonomicName id="4C164D2AFFADFFE6A51FFDC1FD02886A" authorityName="Jałoszyński &amp; Wakimura" authorityYear="2023" box="[616,712,547,573]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="136" phylum="Arthropoda" rank="species" species="uminchu">
<emphasis id="B962EABBFFADFFE6A51FFDC1FD02886A" box="[616,712,547,573]" italics="true" pageId="14" pageNumber="136">uminchu</emphasis>
</taxonomicName>
is taken from the Uchinaaguchi language of Okinawa-jima; its meaning is “a person of the sea”, or, in a narrower sense, a fisherman, referring to the islandic origin of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C30C6522FFADFFE6A7B0FD6DFD2E89F2" pageId="14" pageNumber="136" type="discussion">
<paragraph id="8BA936A9FFADFFE6A7B0FD6DFA5988FE" blockId="14.[151,1437,151,933]" box="[199,1427,655,681]" pageId="14" pageNumber="136">
<emphasis id="B962EABBFFADFFE6A7B0FD6DFEF288FE" bold="true" box="[199,312,655,681]" pageId="14" pageNumber="136">Remarks.</emphasis>
Specimens used in this study were collected by sifting leaf litter and rotten wood in subtropical forest.
</paragraph>
<paragraph id="8BA936A9FFADFFE6A7B0FD51FD2E89F2" blockId="14.[151,1437,151,933]" pageId="14" pageNumber="136">
Adults of
<taxonomicName id="4C164D2AFFADFFE6A649FD51FDDC889A" authorityName="Jałoszyński &amp; Wakimura" authorityYear="2023" box="[318,534,691,717]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="136" phylum="Arthropoda" rank="species" species="uminchu">
<emphasis id="B962EABBFFADFFE6A649FD51FDDC889A" box="[318,534,691,717]" italics="true" pageId="14" pageNumber="136">Tohlezkus uminchu</emphasis>
</taxonomicName>
are the smallest among species known to occur in
<collectingCountry id="F3017639FFADFFE6A303FD51FB7E889A" box="[1140,1204,691,717]" name="Japan" pageId="14" pageNumber="136">Japan</collectingCountry>
and
<collectingCountry id="F3017639FFADFFE6A386FD51FA89889A" box="[1265,1347,691,717]" name="Taiwan" pageId="14" pageNumber="136">Taiwan</collectingCountry>
(
<figureCitation id="132D2A2CFFADFFE6A221FD51FA56889A" box="[1366,1436,691,717]" captionStart="FIGURES 1‒4" captionStartId="1.[151,264,1474,1498]" captionTargetBox="[157,1429,860,1439]" captionTargetId="figure-191@1.[151,1435,849,1451]" captionTargetPageId="1" captionText="FIGURES 1‒4. Dorsal habitus of males. Bisayodes hure gen. &amp; sp. n. (1); Tohlezkus uminchu sp. n. (2); Tohlezkus orientalis Vít (3); Tohlezkus rufus (M. Sakai) (4)." figureDoi="http://doi.org/10.5281/zenodo.8391331" httpUri="https://zenodo.org/record/8391331/files/figure.png" pageId="14" pageNumber="136">Fig. 2</figureCitation>
vs. 3–4). They resemble
<taxonomicName id="4C164D2AFFADFFE6A6C0FD35FD8A88A6" authorityName="Vit" authorityYear="1981" box="[439,576,727,753]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="136" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B962EABBFFADFFE6A6C0FD35FD8A88A6" box="[439,576,727,753]" italics="true" pageId="14" pageNumber="136">T. orientalis</emphasis>
</taxonomicName>
in the umbra body pigmentation (distinctly reddish in
<taxonomicName id="4C164D2AFFADFFE6A3CAFD35FADF88A6" authorityName="" baseAuthorityName="M. Sakai" baseAuthorityYear="1980" box="[1213,1301,727,753]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="136" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="B962EABBFFADFFE6A3CAFD35FADF88A6" box="[1213,1301,727,753]" italics="true" pageId="14" pageNumber="136">T. rufus</emphasis>
</taxonomicName>
).
<taxonomicName id="4C164D2AFFADFFE6A258FD35FEC88942" authorityName="Vit" authorityYear="1981" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="136" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B962EABBFFADFFE6A258FD35FEC88942" italics="true" pageId="14" pageNumber="136">Tohlezkus orientalis</emphasis>
</taxonomicName>
, known to occur in
<collectingCountry id="F3017639FFADFFE6A687FD19FD8F8942" box="[496,581,763,789]" name="Taiwan" pageId="14" pageNumber="136">Taiwan</collectingCountry>
, apart from having a proportionally much larger aedeagus with parameres longer in relation to the penis and with less concave mesal margins, has a much longer apical rod-like portion of the maxillary palpomere 4 (illustrated in Vít (1981): figs 1, 2). The Japanese
<taxonomicName id="4C164D2AFFADFFE6A4C8FCA1FBD9890A" authorityName="" baseAuthorityName="M. Sakai" baseAuthorityYear="1980" box="[959,1043,835,861]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="136" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="B962EABBFFADFFE6A4C8FCA1FBD9890A" box="[959,1043,835,861]" italics="true" pageId="14" pageNumber="136">T. rufus</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C164D2AFFADFFE6A3AAFCA1FA92890A" authorityName="Jałoszyński &amp; Wakimura" authorityYear="2023" box="[1245,1368,835,861]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="136" phylum="Arthropoda" rank="species" species="uminchu">
<emphasis id="B962EABBFFADFFE6A3AAFCA1FA92890A" box="[1245,1368,835,861]" italics="true" pageId="14" pageNumber="136">T. uminchu</emphasis>
</taxonomicName>
in the relative length and structure of the aedeagus, but clearly differs in several ventral characters compared in
<figureCitation id="132D2A2CFFADFFE6A25EFC85FAB289D6" box="[1321,1400,871,897]" captionStart="FIGURES 33‒34" captionStartId="11.[151,264,1899,1923]" captionTargetBox="[151,1435,1181,1876]" captionTargetId="figure-111@11.[151,1435,1181,1876]" captionTargetPageId="11" captionText="FIGURES 33‒34. Metaventral features to distinguish Tohlezkus uminchu sp. n. (33) from T. rufus (M. Sakai) (34). Arrowheads indicate (from left to right): anterior marginal line of anterior metaventral process; shape of median region of metaventrite exposed between metacoxal plates; and shape of lateral margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.8391349" httpUri="https://zenodo.org/record/8391349/files/figure.png" pageId="14" pageNumber="136">Fig. 33</figureCitation>
(
<taxonomicName id="4C164D2AFFADFFE6A2FFFC85FF3D89F2" authorityName="Jałoszyński &amp; Wakimura" authorityYear="2023" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="136" phylum="Arthropoda" rank="species" species="uminchu">
<emphasis id="B962EABBFFADFFE6A2FFFC85FF3D89F2" italics="true" pageId="14" pageNumber="136">T. uminchu</emphasis>
</taxonomicName>
) and
<figureCitation id="132D2A2CFFADFFE6A640FC69FE4D89F2" box="[311,391,907,933]" captionStart="FIGURES 33‒34" captionStartId="11.[151,264,1899,1923]" captionTargetBox="[151,1435,1181,1876]" captionTargetId="figure-111@11.[151,1435,1181,1876]" captionTargetPageId="11" captionText="FIGURES 33‒34. Metaventral features to distinguish Tohlezkus uminchu sp. n. (33) from T. rufus (M. Sakai) (34). Arrowheads indicate (from left to right): anterior marginal line of anterior metaventral process; shape of median region of metaventrite exposed between metacoxal plates; and shape of lateral margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.8391349" httpUri="https://zenodo.org/record/8391349/files/figure.png" pageId="14" pageNumber="136">Fig. 34</figureCitation>
(
<taxonomicName id="4C164D2AFFADFFE6A6E0FC69FE2789F2" authorityName="" baseAuthorityName="M. Sakai" baseAuthorityYear="1980" box="[407,493,907,933]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="136" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="B962EABBFFADFFE6A6E0FC69FE2789F2" box="[407,493,907,933]" italics="true" pageId="14" pageNumber="136">T. rufus</emphasis>
</taxonomicName>
) and listed in the key.
</paragraph>
</subSubSection>
</treatment>
</document>
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