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<document id="CBB4EA80876084EA1E626F1CAA6BF262" ID-DOI="10.11646/zootaxa.5351.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="8391329" ID-ZooBank="08CAFDFE-D4CF-4E5A-A173-3DA9F19F5FA5" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695989053433" checkinUser="plazi" docAuthor="Jałoszyński, Paweł &amp; Wakimura, Ryotaro" docDate="2023" docId="03BF87BFFFA1FFEFA7E0FF3DFF248B32" docLanguage="en" docName="zootaxa.5351.1.5.pdf" docOrigin="Zootaxa 5351 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5351.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bisayodes Wakimura &amp; Jaloszynski 2023, gen. n." docType="treatment" docVersion="2" lastPageNumber="128" masterDocId="FF86FFC7FFA3FFE8A777FFE2FFCA8A57" masterDocTitle="A new genus, new species and new records of Eucinetidae in Japan and Taiwan (Coleoptera)" masterLastPageNumber="138" masterPageNumber="122" pageNumber="124" updateTime="1695989500669" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3B298292BD933EA0E3CA676D1008B330">A new genus, new species and new records of Eucinetidae in Japan and Taiwan (Coleoptera)</mods:title>
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<mods:namePart id="B8716B4CF04FEA08EE08737EBBBF2358">Jałoszyński, Paweł</mods:namePart>
<mods:affiliation id="6055FEC6F67991B9F05730D0E9AD339F">Museum of Natural History, University of Wrocław, Sienkiewicza 21, 50 - 335 Wrocław, Poland</mods:affiliation>
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<mods:namePart id="DC0A6C90F544DD7BE244F39DB8FF641C">Wakimura, Ryotaro</mods:namePart>
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<mods:affiliation id="7AE2AFF389448B5FCB169CB93F08B90A">Department of Biology, School of Biological Sciences, Tokai University, Hokkaido, 005 - 8601, Japan https: // orcid. org / 0009 - 0000 - 8410 - 0567</mods:affiliation>
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<paragraph id="8BA936A9FFA1FFEAA7E0FF3DFCC78AAE" blockId="2.[151,781,222,250]" box="[151,781,222,250]" pageId="2" pageNumber="124">
<heading id="D0E181C5FFA1FFEAA7E0FF3DFCC78AAE" bold="true" box="[151,781,222,250]" fontSize="11" level="1" pageId="2" pageNumber="124" reason="1">
<emphasis id="B962EABBFFA1FFEAA7E0FF3DFCC78AAE" bold="true" box="[151,781,222,250]" pageId="2" pageNumber="124">
Genus 
<taxonomicName id="4C164D2AFFA1FFEAA799FF3DFD678AAE" authority="Wakimura &amp; Jaloszynski" authorityName="Wakimura &amp; Jaloszynski" authorityYear="2023" box="[238,685,222,250]" class="Insecta" family="Eucinetidae" genus="Bisayodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="124" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis id="B962EABBFFA1FFEAA799FF3DFEAD8AAD" bold="true" box="[238,359,223,250]" italics="true" pageId="2" pageNumber="124">Bisayodes</emphasis>
Wakimura &amp; Jałoszyński
</taxonomicName>
, 
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</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30C6522FFA1FFEAA7E0FEC6FCC18B6B" box="[151,779,292,316]" pageId="2" pageNumber="124" type="materials_examined">
<paragraph id="8BA936A9FFA1FFEAA7E0FEC6FCC18B6B" blockId="2.[151,779,292,316]" box="[151,779,292,316]" pageId="2" pageNumber="124">
<typeStatus id="54AD880BFFA1FFEAA7E0FEC6FF038B6B" box="[151,201,292,316]" pageId="2" pageNumber="124">Type</typeStatus>
species: 
<taxonomicName id="4C164D2AFFA1FFEAA651FEC6FD0D8B6B" authority="Wakimura &amp; Jaloszynski" authorityName="Wakimura &amp; Jaloszynski" authorityYear="2023" box="[294,711,292,316]" class="Insecta" family="Eucinetidae" genus="Bisayodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="124" phylum="Arthropoda" rank="species" species="hure" status="sp. n.">
<emphasis id="B962EABBFFA1FFEAA651FEC6FE0A8B6B" box="[294,448,292,316]" italics="true" pageId="2" pageNumber="124">Bisayodes hure</emphasis>
Wakimura &amp; Jałoszyński
</taxonomicName>
, 
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<taxonomicNameLabel id="A25157C0FFA1FFEAA5A5FEC6FCC18B6B" box="[722,779,292,316]" pageId="2" pageNumber="124" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
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<subSubSection id="C30C6522FFA1FFEAA7E0FE8BFC758947" pageId="2" pageNumber="124" type="diagnosis">
<paragraph id="8BA936A9FFA1FFEAA7E0FE8BFC758947" blockId="2.[151,1437,361,821]" pageId="2" pageNumber="124">
<emphasis id="B962EABBFFA1FFEAA7E0FE8BFED88BD4" bold="true" box="[151,274,361,387]" pageId="2" pageNumber="124">Diagnosis.</emphasis>
Body small, not reaching 
<quantity id="4CEE9B4CFFA1FFEAA548FE88FD4B8BD3" box="[575,641,362,388]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="124" unit="mm" value="2.0">2 mm</quantity>
in length, short oval; head subpentagonal with antennal insertions not exposed in dorsal view; frons not constricted in front of eyes; frontoclypeal margin carinate; clypeus exposed in dorsal view, about 6 × wide as long at middle; mouthparts semisuctorial, with labrum and prelabium about twice as long as broad and truncate apically; mentum as long as wide; labial palpomere 2 pedunculate; labial palpomere 3 fusiform and rapidly narrowing in distal third; galea vestigial, lacinia longer than stipes and with several conspicuously short and widely spaced thick distomesal setae; maxillary palpomere 1 subtriangular and shorter than palpomere 2; maxillary palpomere 2 subcylindrical; maxillary palpomere 3 fusiform, nearly gradually narrowing distally; antenna loosely assembled, with slender and elongate proximal antennomeres and clearly delimited trimerous club; mesoventral process broadly subtriangular and not separating mesocoxae; metaventrite behind mesocoxae with transverse carina; median region of metaventrite broadly subtriangular and posteriorly reaching behind half length of metacoxal plates; apical mesotibial spurs of subequal length and apically rounded; elytra lacking adsutural striae and transverse strigulae; parameres shorter than penis and lacking setae.
</paragraph>
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<subSubSection id="C30C6522FFA1FFEEA7B0FCFBFB9589BA" lastPageId="6" lastPageNumber="128" pageId="2" pageNumber="124" type="description">
<paragraph id="8BA936A9FFA1FFEAA7B0FCFBFA8F8963" blockId="2.[151,1437,361,821]" box="[199,1349,793,821]" pageId="2" pageNumber="124">
<emphasis id="B962EABBFFA1FFEAA7B0FCFBFE938964" bold="true" box="[199,345,793,819]" pageId="2" pageNumber="124">Description.</emphasis>
Body (
<figureCitation id="132D2A2CFFA1FFEAA6DCFCF8FE388963" box="[427,498,794,820]" captionStart="FIGURES 1‒4" captionStartId="1.[151,264,1474,1498]" captionTargetBox="[157,1429,860,1439]" captionTargetId="figure-191@1.[151,1435,849,1451]" captionTargetPageId="1" captionText="FIGURES 1‒4. Dorsal habitus of males. Bisayodes hure gen. &amp; sp. n. (1); Tohlezkus uminchu sp. n. (2); Tohlezkus orientalis Vít (3); Tohlezkus rufus (M. Sakai) (4)." figureDoi="http://doi.org/10.5281/zenodo.8391331" httpUri="https://zenodo.org/record/8391331/files/figure.png" pageId="2" pageNumber="124">Figs 1</figureCitation>
, 
<figureCitation id="132D2A2CFFA1FFEAA577FCF8FDE18962" box="[512,555,794,821]" captionStart="FIGURES 5‒6" captionStartId="2.[151,264,1898,1922]" captionTargetBox="[151,1435,844,1875]" captionTargetId="figure-233@2.[151,1435,844,1875]" captionTargetPageId="2" captionText="FIGURES 5‒6. Bisayodes hure gen. &amp; sp. n. Dorsal (5) and ventral (6) habitus." figureDoi="http://doi.org/10.5281/zenodo.8391333" httpUri="https://zenodo.org/record/8391333/files/figure.png" pageId="2" pageNumber="124">5–6</figureCitation>
) short oval and strongly convex dorsally, not reaching 
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in length.
</paragraph>
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<paragraph id="8BA936A9FFA1FFEAA7E0F888FC0D8DD4" blockId="2.[151,967,1898,1923]" box="[151,967,1898,1923]" pageId="2" pageNumber="124">
<emphasis id="B962EABBFFA1FFEAA7E0F888FEF18DD5" bold="true" box="[151,315,1898,1922]" pageId="2" pageNumber="124">FIGURES 5‒6.</emphasis>
<taxonomicName id="4C164D2AFFA1FFEAA636F889FE118DD4" authorityName="Wakimura &amp; Jaloszynski" authorityYear="2023" box="[321,475,1899,1923]" class="Insecta" family="Eucinetidae" genus="Bisayodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="124" phylum="Arthropoda" rank="species" species="hure">
<emphasis id="B962EABBFFA1FFEAA636F889FE118DD4" box="[321,475,1899,1923]" italics="true" pageId="2" pageNumber="124">Bisayodes hure</emphasis>
</taxonomicName>
<emphasis id="B962EABBFFA1FFEAA695F889FDC48DD4" bold="true" box="[482,526,1899,1923]" pageId="2" pageNumber="124">gen.</emphasis>
&amp; 
<emphasis id="B962EABBFFA1FFEAA559F889FDAD8DD4" bold="true" box="[558,615,1899,1923]" pageId="2" pageNumber="124">sp. n.</emphasis>
Dorsal (
<emphasis id="B962EABBFFA1FFEAA5B7F889FD078DD4" bold="true" box="[704,717,1899,1923]" pageId="2" pageNumber="124">5</emphasis>
) and ventral (
<emphasis id="B962EABBFFA1FFEAA42BF888FCA38DD5" bold="true" box="[860,873,1898,1922]" pageId="2" pageNumber="124">6</emphasis>
) habitus.
</paragraph>
</caption>
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<paragraph id="8BA936A9FFA0FFEBA7E0F8E3FE6F8D6A" blockId="3.[151,1436,1793,1853]" pageId="3" pageNumber="125">
<emphasis id="B962EABBFFA0FFEBA7E0F8E3FE8D8D4E" bold="true" box="[151,327,1793,1817]" pageId="3" pageNumber="125">FIGURES 7‒12.</emphasis>
<taxonomicName id="4C164D2AFFA0FFEBA63AF8E3FE2D8D4E" authorityName="Wakimura &amp; Jaloszynski" authorityYear="2023" box="[333,487,1793,1817]" class="Insecta" family="Eucinetidae" genus="Bisayodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="125" phylum="Arthropoda" rank="species" species="hure">
<emphasis id="B962EABBFFA0FFEBA63AF8E3FE2D8D4E" box="[333,487,1793,1817]" italics="true" pageId="3" pageNumber="125">Bisayodes hure</emphasis>
</taxonomicName>
<emphasis id="B962EABBFFA0FFEBA69AF8E3FDD08D4E" bold="true" box="[493,538,1793,1817]" pageId="3" pageNumber="125">gen.</emphasis>
&amp; 
<emphasis id="B962EABBFFA0FFEBA54EF8E3FDB88D4E" bold="true" box="[569,626,1793,1817]" pageId="3" pageNumber="125">sp. n.</emphasis>
Head in dorsal (
<emphasis id="B962EABBFFA0FFEBA46DF8E3FC818D4E" bold="true" box="[794,843,1793,1817]" pageId="3" pageNumber="125">7, 11</emphasis>
), ventral (
<emphasis id="B962EABBFFA0FFEBA4C2F8E3FC2D8D4E" bold="true" box="[949,999,1793,1817]" pageId="3" pageNumber="125">8, 12</emphasis>
) and anteroventral (
<emphasis id="B962EABBFFA0FFEBA3C3F8E3FB0B8D4E" bold="true" box="[1204,1217,1793,1817]" pageId="3" pageNumber="125">9</emphasis>
) views; right antenna in anteroventral view (
<emphasis id="B962EABBFFA0FFEBA60AF8C7FE5C8D6A" bold="true" box="[381,406,1829,1853]" pageId="3" pageNumber="125">10</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BA936A9FFA0FFECA7B0F86DFF2A8AAE" blockId="3.[151,1436,1935,2034]" lastBlockId="4.[151,1436,151,789]" lastPageId="4" lastPageNumber="126" pageId="3" pageNumber="125">
Head (
<figureCitation id="132D2A2CFFA0FFEBA663F86DFE428DFE" box="[276,392,1935,1961]" captionStart="FIGURES 7‒12" captionStartId="3.[151,264,1793,1817]" captionTargetBox="[151,1435,181,1769]" captionTargetId="figure-22@3.[151,1435,181,1769]" captionTargetPageId="3" captionText="FIGURES 7‒12. Bisayodes hure gen. &amp; sp. n. Head in dorsal (7, 11), ventral (8, 12) and anteroventral (9) views; right antenna in anteroventral view (10)." figureDoi="http://doi.org/10.5281/zenodo.8391335" httpUri="https://zenodo.org/record/8391335/files/figure.png" pageId="3" pageNumber="125">Figs 7–12</figureCitation>
) flattened and transverse, subpentagonal with tempora strongly converging posteriorly and sides of frons strongly converging anteriorly. Vertex and frons confluent; anterior margin of frons demarcated from clypeus by arcuate transverse carina laterally extending along mesal margins of eyes; clypeus visible in dorsal view, arcuate, about 6 × as wide as long. Eyes convex. Each gena with infraocular carina. Antennal sulcus present at each side of head, extending ventrally from antennal fossa to lateral margin of submentum, with carinate outer edge (
<figureCitation id="132D2A2CFFA7FFECA21DFF59FF1E8AAE" captionStart="FIGURES 7‒12" captionStartId="3.[151,264,1793,1817]" captionTargetBox="[151,1435,181,1769]" captionTargetId="figure-22@3.[151,1435,181,1769]" captionTargetPageId="3" captionText="FIGURES 7‒12. Bisayodes hure gen. &amp; sp. n. Head in dorsal (7, 11), ventral (8, 12) and anteroventral (9) views; right antenna in anteroventral view (10)." figureDoi="http://doi.org/10.5281/zenodo.8391335" httpUri="https://zenodo.org/record/8391335/files/figure.png" pageId="4" pageNumber="126">Figs 8–10</figureCitation>
).
</paragraph>
<paragraph id="8BA936A9FFA7FFECA7B0FEE1FC038B4A" blockId="4.[151,1436,151,789]" box="[199,969,259,285]" pageId="4" pageNumber="126">Gular plate subrectangular, slightly transverse; gular sutures distinct.</paragraph>
<paragraph id="8BA936A9FFA7FFECA7B0FEC5FB208942" blockId="4.[151,1436,151,789]" pageId="4" pageNumber="126">
Mouthparts (
<figureCitation id="132D2A2CFFA7FFECA62FFEC5FE128B16" box="[344,472,295,321]" captionStart="FIGURES 13‒19" captionStartId="4.[151,264,1942,1966]" captionTargetBox="[151,1435,813,1919]" captionTargetId="figure-282@4.[151,1435,813,1919]" captionTargetPageId="4" captionText="FIGURES 13‒19. Bisayodes hure gen. &amp; sp. n. Labrum in dorsal view (13); mandible in dorsal view (14); right maxilla in dorsal (15) and ventral (16, 17) views; labium (excl. submentum) in ventral view (18); mentum, prelabium and maxilla in ventral view (19)." figureDoi="http://doi.org/10.5281/zenodo.8391337" httpUri="https://zenodo.org/record/8391337/files/figure.png" pageId="4" pageNumber="126">Figs 13–19</figureCitation>
) semisuctorial. Labrum (
<figureCitation id="132D2A2CFFA7FFECA583FEC5FC8E8B16" box="[756,836,295,321]" captionStart="FIGURES 13‒19" captionStartId="4.[151,264,1942,1966]" captionTargetBox="[151,1435,813,1919]" captionTargetId="figure-282@4.[151,1435,813,1919]" captionTargetPageId="4" captionText="FIGURES 13‒19. Bisayodes hure gen. &amp; sp. n. Labrum in dorsal view (13); mandible in dorsal view (14); right maxilla in dorsal (15) and ventral (16, 17) views; labium (excl. submentum) in ventral view (18); mentum, prelabium and maxilla in ventral view (19)." figureDoi="http://doi.org/10.5281/zenodo.8391337" httpUri="https://zenodo.org/record/8391337/files/figure.png" pageId="4" pageNumber="126">Fig. 13</figureCitation>
) trapezoidal, elongate, with truncate anterior margin, together with prelabium forming tubular sucking apparatus (
<figureCitation id="132D2A2CFFA7FFECA45BFEA9FCA48B32" box="[812,878,331,357]" captionStart="FIGURES 7‒12" captionStartId="3.[151,264,1793,1817]" captionTargetBox="[151,1435,181,1769]" captionTargetId="figure-22@3.[151,1435,181,1769]" captionTargetPageId="3" captionText="FIGURES 7‒12. Bisayodes hure gen. &amp; sp. n. Head in dorsal (7, 11), ventral (8, 12) and anteroventral (9) views; right antenna in anteroventral view (10)." figureDoi="http://doi.org/10.5281/zenodo.8391335" httpUri="https://zenodo.org/record/8391335/files/figure.png" pageId="4" pageNumber="126">Fig. 9</figureCitation>
). Mandibles (
<figureCitation id="132D2A2CFFA7FFECA372FEA9FB9E8B32" box="[1029,1108,331,357]" captionStart="FIGURES 13‒19" captionStartId="4.[151,264,1942,1966]" captionTargetBox="[151,1435,813,1919]" captionTargetId="figure-282@4.[151,1435,813,1919]" captionTargetPageId="4" captionText="FIGURES 13‒19. Bisayodes hure gen. &amp; sp. n. Labrum in dorsal view (13); mandible in dorsal view (14); right maxilla in dorsal (15) and ventral (16, 17) views; labium (excl. submentum) in ventral view (18); mentum, prelabium and maxilla in ventral view (19)." figureDoi="http://doi.org/10.5281/zenodo.8391337" httpUri="https://zenodo.org/record/8391337/files/figure.png" pageId="4" pageNumber="126">Fig. 14</figureCitation>
) short, elongate subtriangular and flattened, with weakly sclerotized apical subtriangular and pointed tooth directed anteriorly, mesal margin sharp, with short and broad preapical tooth and lacking prostheca and mola. Maxillae (
<figureCitation id="132D2A2CFFA7FFECA367FE71FB458BFA" box="[1040,1167,402,429]" captionStart="FIGURES 13‒19" captionStartId="4.[151,264,1942,1966]" captionTargetBox="[151,1435,813,1919]" captionTargetId="figure-282@4.[151,1435,813,1919]" captionTargetPageId="4" captionText="FIGURES 13‒19. Bisayodes hure gen. &amp; sp. n. Labrum in dorsal view (13); mandible in dorsal view (14); right maxilla in dorsal (15) and ventral (16, 17) views; labium (excl. submentum) in ventral view (18); mentum, prelabium and maxilla in ventral view (19)." figureDoi="http://doi.org/10.5281/zenodo.8391337" httpUri="https://zenodo.org/record/8391337/files/figure.png" pageId="4" pageNumber="126">Figs 15–17</figureCitation>
) with slender stipes and palpifer, galea vestigial, presumably represented by narrow and short slender projection at base of maxillary palp; lacinia slender and weakly curved, slightly longer than stipes, with a few short and sparse distomesal setae. Maxillary palp tetramerous, palpomere 1 smallest, elongate, asetose; palpomere 2 subtriangular, weakly elongate and shorter than 3, with only a few setae; palpomere 3 subcylindrical with oblique distal margin, sparsely setose; palpomere 4 largest, fusiform and nearly gradually tapering apically, sparsely setose on entire surface. Labium (
<figureCitation id="132D2A2CFFA7FFECA381FDA5FAF58836" box="[1270,1343,583,609]" captionStart="FIGURES 7‒12" captionStartId="3.[151,264,1793,1817]" captionTargetBox="[151,1435,181,1769]" captionTargetId="figure-22@3.[151,1435,181,1769]" captionTargetPageId="3" captionText="FIGURES 7‒12. Bisayodes hure gen. &amp; sp. n. Head in dorsal (7, 11), ventral (8, 12) and anteroventral (9) views; right antenna in anteroventral view (10)." figureDoi="http://doi.org/10.5281/zenodo.8391335" httpUri="https://zenodo.org/record/8391335/files/figure.png" pageId="4" pageNumber="126">Figs 8</figureCitation>
, 
<figureCitation id="132D2A2CFFA7FFECA23AFDA5FA5E8836" box="[1357,1428,583,609]" captionStart="FIGURES 13‒19" captionStartId="4.[151,264,1942,1966]" captionTargetBox="[151,1435,813,1919]" captionTargetId="figure-282@4.[151,1435,813,1919]" captionTargetPageId="4" captionText="FIGURES 13‒19. Bisayodes hure gen. &amp; sp. n. Labrum in dorsal view (13); mandible in dorsal view (14); right maxilla in dorsal (15) and ventral (16, 17) views; labium (excl. submentum) in ventral view (18); mentum, prelabium and maxilla in ventral view (19)." figureDoi="http://doi.org/10.5281/zenodo.8391337" httpUri="https://zenodo.org/record/8391337/files/figure.png" pageId="4" pageNumber="126">18–19</figureCitation>
) with strongly transverse and short, slightly impressed submentum, trapezoidal mentum with lateral and anterior marginal carinae, and elongate, subtrapezoidal prelabium truncate at apex; premental sclerites absent. Labial palps inserted at base of prelabium, trimerous, palpomere 1 vestigial, subtriangular, asetose; palpomere 2 pedunculate, strongly broadening distally to middle and then strongly narrowing, covered with sparse setae; palpomere 3 about as long and about half as wide as 2, fusiform with rapidly narrowing apical region, sparsely setose.
</paragraph>
<caption id="DF696621FFA7FFECA7E0F874FF358DA0" ID-DOI="http://doi.org/10.5281/zenodo.8391337" ID-Zenodo-Dep="8391337" httpUri="https://zenodo.org/record/8391337/files/figure.png" pageId="4" pageNumber="126" startId="4.[151,264,1942,1966]" targetBox="[151,1435,813,1919]" targetPageId="4" targetType="figure">
<paragraph id="8BA936A9FFA7FFECA7E0F874FF358DA0" blockId="4.[151,1437,1942,2039]" pageId="4" pageNumber="126">
<emphasis id="B962EABBFFA7FFECA7E0F874FE9C8DF8" bold="true" box="[151,342,1942,1967]" pageId="4" pageNumber="126">FIGURES 13‒19.</emphasis>
<taxonomicName id="4C164D2AFFA7FFECA629F875FE338DF8" authorityName="Wakimura &amp; Jaloszynski" authorityYear="2023" box="[350,505,1943,1967]" class="Insecta" family="Eucinetidae" genus="Bisayodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="126" phylum="Arthropoda" rank="species" species="hure">
<emphasis id="B962EABBFFA7FFECA629F875FE338DF8" box="[350,505,1943,1967]" italics="true" pageId="4" pageNumber="126">Bisayodes hure</emphasis>
</taxonomicName>
<emphasis id="B962EABBFFA7FFECA575F875FDE58DF8" bold="true" box="[514,559,1943,1967]" pageId="4" pageNumber="126">gen.</emphasis>
&amp; 
<emphasis id="B962EABBFFA7FFECA525F875FD478DF8" bold="true" box="[594,653,1943,1967]" pageId="4" pageNumber="126">sp. n.</emphasis>
Labrum in dorsal view (
<emphasis id="B962EABBFFA7FFECA4E6F875FC608DF8" bold="true" box="[913,938,1943,1967]" pageId="4" pageNumber="126">13</emphasis>
); mandible in dorsal view (
<emphasis id="B962EABBFFA7FFECA3BBF875FB2F8DF8" bold="true" box="[1228,1253,1943,1967]" pageId="4" pageNumber="126">14</emphasis>
); right maxilla in dorsal (
<emphasis id="B962EABBFFA7FFECA794F859FF368D84" bold="true" box="[227,252,1979,2003]" pageId="4" pageNumber="126">15</emphasis>
) and ventral (
<emphasis id="B962EABBFFA7FFECA6F0F859FE0F8D84" bold="true" box="[391,453,1979,2003]" pageId="4" pageNumber="126">16, 17</emphasis>
) views; labium (excl. submentum) in ventral view (
<emphasis id="B962EABBFFA7FFECA4BDF859FC298D84" bold="true" box="[970,995,1979,2003]" pageId="4" pageNumber="126">18</emphasis>
); mentum, prelabium and maxilla in ventral view (
<emphasis id="B962EABBFFA7FFECA7A0F83DFF3A8DA0" bold="true" box="[215,240,2015,2039]" pageId="4" pageNumber="126">19</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BA936A9FFA6FFEDA7B0FF75FB6F8A82" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="127">
Antenna (
<figureCitation id="132D2A2CFFA6FFEDA64EFF75FE778AE6" box="[313,445,151,177]" captionStart="FIGURES 7‒12" captionStartId="3.[151,264,1793,1817]" captionTargetBox="[151,1435,181,1769]" captionTargetId="figure-22@3.[151,1435,181,1769]" captionTargetPageId="3" captionText="FIGURES 7‒12. Bisayodes hure gen. &amp; sp. n. Head in dorsal (7, 11), ventral (8, 12) and anteroventral (9) views; right antenna in anteroventral view (10)." figureDoi="http://doi.org/10.5281/zenodo.8391335" httpUri="https://zenodo.org/record/8391335/files/figure.png" pageId="5" pageNumber="127">Figs 10–12</figureCitation>
) slender, with enlarged scape, elongate pedicel and elongate proximal flagellomeres, distally thickened to 
<taxonomicName id="4C164D2AFFA6FFEDA6F6FF59FDDB8A82" box="[385,529,187,213]" form="distinct" pageId="5" pageNumber="127" rank="form">form distinct</taxonomicName>
trimerous club. All antennomeres covered with long setae.
</paragraph>
<paragraph id="8BA936A9FFA6FFEDA7B0FF3DFCB18BDE" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="127">
Pronotum (
<figureCitation id="132D2A2CFFA6FFEDA63FFF3DFE478AAE" box="[328,397,222,249]" captionStart="FIGURES 5‒6" captionStartId="2.[151,264,1898,1922]" captionTargetBox="[151,1435,844,1875]" captionTargetId="figure-233@2.[151,1435,844,1875]" captionTargetPageId="2" captionText="FIGURES 5‒6. Bisayodes hure gen. &amp; sp. n. Dorsal (5) and ventral (6) habitus." figureDoi="http://doi.org/10.5281/zenodo.8391333" httpUri="https://zenodo.org/record/8391333/files/figure.png" pageId="5" pageNumber="127">Fig. 5</figureCitation>
) strongly transverse and strongly convex, with anterior and lateral margins confluent and broadly rounded, posterior margin strongly arcuate and posteriorly convex in middle, laterally strongly emarginate, so that posterior pronotal corners are projecting posterolaterally. Lateral pronotal edges visible in ventral view; hypomera concave; prosternum short, in front of procoxae nearly carinate; prosternal process narrow, rounded at apex, not projecting ventrally beyond ventral surface of procoxae.
</paragraph>
<paragraph id="8BA936A9FFA6FFEDA7B0FE71FD308BFA" blockId="5.[151,1437,151,645]" box="[199,762,403,429]" pageId="5" pageNumber="127">Mesoscutellar shield not exposed in intact beetles.</paragraph>
<paragraph id="8BA936A9FFA6FFEDA7B0FE55FC1E8BA2" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="127">
Elytra (
<figureCitation id="132D2A2CFFA6FFEDA656FE55FEA38B86" box="[289,361,438,465]" captionStart="FIGURES 5‒6" captionStartId="2.[151,264,1898,1922]" captionTargetBox="[151,1435,844,1875]" captionTargetId="figure-233@2.[151,1435,844,1875]" captionTargetPageId="2" captionText="FIGURES 5‒6. Bisayodes hure gen. &amp; sp. n. Dorsal (5) and ventral (6) habitus." figureDoi="http://doi.org/10.5281/zenodo.8391333" httpUri="https://zenodo.org/record/8391333/files/figure.png" pageId="5" pageNumber="127">Fig. 5</figureCitation>
) strongly convex, oval, lacking fine transverse strigulae and lacking adsutural striae, with subhorizontal epipleurae (
<figureCitation id="132D2A2CFFA6FFEDA6CCFE39FE348BA2" box="[443,510,475,501]" captionStart="FIGURES 5‒6" captionStartId="2.[151,264,1898,1922]" captionTargetBox="[151,1435,844,1875]" captionTargetId="figure-233@2.[151,1435,844,1875]" captionTargetPageId="2" captionText="FIGURES 5‒6. Bisayodes hure gen. &amp; sp. n. Dorsal (5) and ventral (6) habitus." figureDoi="http://doi.org/10.5281/zenodo.8391333" httpUri="https://zenodo.org/record/8391333/files/figure.png" pageId="5" pageNumber="127">Fig. 6</figureCitation>
) weakly, gradually narrowing posteriorly.
</paragraph>
<paragraph id="8BA936A9FFA6FFEDA7B0FE1DFCAC88D2" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="127">
Mesoventrite (
<figureCitation id="132D2A2CFFA6FFEDA61BFE1DFE0A884E" box="[364,448,511,537]" captionStart="FIGURES 20‒22" captionStartId="5.[151,264,1947,1971]" captionTargetBox="[151,1435,669,1924]" captionTargetId="figure-196@5.[151,1435,669,1924]" captionTargetPageId="5" captionText="FIGURES 20‒22. Bisayodes hure gen. &amp; sp. n. Pterothorax in ventral view (20); fore femur, tibia and tarsus in ventral view (21); left mesotarsus and apex of mesotibia in ventral view (22)." figureDoi="http://doi.org/10.5281/zenodo.8391339" httpUri="https://zenodo.org/record/8391339/files/figure.png" pageId="5" pageNumber="127">Fig. 20</figureCitation>
) short, anteriorly concave and posteriorly convex, with impressed and broad procoxal rests in its anterior region; posteriorly, median region of mesoventrite forming short and broadly subtriangular mesoventral process entirely situated in front of mesocoxae; median region of mesoventrite forming demarcated elongate rhomboidal plate with median longitudinal impression.
</paragraph>
<caption id="DF696621FFA6FFEDA7E0F879FCEB8D8F" ID-DOI="http://doi.org/10.5281/zenodo.8391339" ID-Zenodo-Dep="8391339" httpUri="https://zenodo.org/record/8391339/files/figure.png" pageId="5" pageNumber="127" startId="5.[151,264,1947,1971]" targetBox="[151,1435,669,1924]" targetPageId="5" targetType="figure">
<paragraph id="8BA936A9FFA6FFEDA7E0F879FCEB8D8F" blockId="5.[151,1436,1947,2008]" pageId="5" pageNumber="127">
<emphasis id="B962EABBFFA6FFEDA7E0F879FE9F8DE4" bold="true" box="[151,341,1947,1971]" pageId="5" pageNumber="127">FIGURES 20‒22.</emphasis>
<taxonomicName id="4C164D2AFFA6FFEDA62BF879FE3D8DE4" authorityName="Wakimura &amp; Jaloszynski" authorityYear="2023" box="[348,503,1947,1971]" class="Insecta" family="Eucinetidae" genus="Bisayodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="species" species="hure">
<emphasis id="B962EABBFFA6FFEDA62BF879FE3D8DE4" box="[348,503,1947,1971]" italics="true" pageId="5" pageNumber="127">Bisayodes hure</emphasis>
</taxonomicName>
<emphasis id="B962EABBFFA6FFEDA688F87EFDE68DE3" bold="true" box="[511,556,1948,1972]" pageId="5" pageNumber="127">gen.</emphasis>
&amp; 
<emphasis id="B962EABBFFA6FFEDA539F87EFD428DE3" bold="true" box="[590,648,1948,1972]" pageId="5" pageNumber="127">sp. n.</emphasis>
Pterothorax in ventral view (
<emphasis id="B962EABBFFA6FFEDA4C1F879FC058DE4" bold="true" box="[950,975,1947,1971]" pageId="5" pageNumber="127">20</emphasis>
); fore femur, tibia and tarsus in ventral view (
<emphasis id="B962EABBFFA6FFEDA7D7F85DFF738D80" bold="true" box="[160,185,1983,2007]" pageId="5" pageNumber="127">21</emphasis>
); left mesotarsus and apex of mesotibia in ventral view (
<emphasis id="B962EABBFFA6FFEDA58DF85DFCD98D80" bold="true" box="[762,787,1983,2007]" pageId="5" pageNumber="127">22</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BA936A9FFA5FFEEA7B0FF75FD0F8B32" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="128">
Metaventrite (
<figureCitation id="132D2A2CFFA5FFEEA611FF75FE728AE6" box="[358,440,151,177]" captionStart="FIGURES 20‒22" captionStartId="5.[151,264,1947,1971]" captionTargetBox="[151,1435,669,1924]" captionTargetId="figure-196@5.[151,1435,669,1924]" captionTargetPageId="5" captionText="FIGURES 20‒22. Bisayodes hure gen. &amp; sp. n. Pterothorax in ventral view (20); fore femur, tibia and tarsus in ventral view (21); left mesotarsus and apex of mesotibia in ventral view (22)." figureDoi="http://doi.org/10.5281/zenodo.8391339" httpUri="https://zenodo.org/record/8391339/files/figure.png" pageId="6" pageNumber="128">Fig. 20</figureCitation>
) with median region broadly subtriangular and bearing anterior transverse carina behind mesocoxae; posterior tip of median region of metaventrite reaching distinctly behind middle of metacoxal plates; metanepisterna not exposed in ventral view. Large metacoxal plates meeting along midline behind median region of metaventrite, with posterior margin of each plate forming two angles: outer rounded and more posterior and inner (mesal) more subtriangular and less projecting posteriorly; lateral margin of each metacoxal plate weakly rounded in anterior and posterior thirds, at middle straight.
</paragraph>
<paragraph id="8BA936A9FFA5FFEEA7B0FE8DFAED886A" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="128">
Fore legs (
<figureCitation id="132D2A2CFFA5FFEEA636FE8DFE418BDE" box="[321,395,367,393]" captionStart="FIGURES 5‒6" captionStartId="2.[151,264,1898,1922]" captionTargetBox="[151,1435,844,1875]" captionTargetId="figure-233@2.[151,1435,844,1875]" captionTargetPageId="2" captionText="FIGURES 5‒6. Bisayodes hure gen. &amp; sp. n. Dorsal (5) and ventral (6) habitus." figureDoi="http://doi.org/10.5281/zenodo.8391333" httpUri="https://zenodo.org/record/8391333/files/figure.png" pageId="6" pageNumber="128">Figs 6</figureCitation>
, 
<figureCitation id="132D2A2CFFA5FFEEA6EEFE8DFE7C8BDE" box="[409,438,367,393]" captionStart="FIGURES 20‒22" captionStartId="5.[151,264,1947,1971]" captionTargetBox="[151,1435,669,1924]" captionTargetId="figure-196@5.[151,1435,669,1924]" captionTargetPageId="5" captionText="FIGURES 20‒22. Bisayodes hure gen. &amp; sp. n. Pterothorax in ventral view (20); fore femur, tibia and tarsus in ventral view (21); left mesotarsus and apex of mesotibia in ventral view (22)." figureDoi="http://doi.org/10.5281/zenodo.8391339" httpUri="https://zenodo.org/record/8391339/files/figure.png" pageId="6" pageNumber="128">21</figureCitation>
) with strongly transverse procoxae, each bearing ventral lateral carina on entire length; protrochanters small, subtriangular; profemora (
<figureCitation id="132D2A2CFFA5FFEEA5CDFE71FCC78BFA" box="[698,781,403,429]" captionStart="FIGURES 20‒22" captionStartId="5.[151,264,1947,1971]" captionTargetBox="[151,1435,669,1924]" captionTargetId="figure-196@5.[151,1435,669,1924]" captionTargetPageId="5" captionText="FIGURES 20‒22. Bisayodes hure gen. &amp; sp. n. Pterothorax in ventral view (20); fore femur, tibia and tarsus in ventral view (21); left mesotarsus and apex of mesotibia in ventral view (22)." figureDoi="http://doi.org/10.5281/zenodo.8391339" httpUri="https://zenodo.org/record/8391339/files/figure.png" pageId="6" pageNumber="128">Fig. 21</figureCitation>
) flattened laterally, slightly narrowing distally, each with anteroventral row of dense thick and slightly unevenly distributed setae in basal half; protibiae (
<figureCitation id="132D2A2CFFA5FFEEA3ACFE55FAE48B86" box="[1243,1326,439,465]" captionStart="FIGURES 20‒22" captionStartId="5.[151,264,1947,1971]" captionTargetBox="[151,1435,669,1924]" captionTargetId="figure-196@5.[151,1435,669,1924]" captionTargetPageId="5" captionText="FIGURES 20‒22. Bisayodes hure gen. &amp; sp. n. Pterothorax in ventral view (20); fore femur, tibia and tarsus in ventral view (21); left mesotarsus and apex of mesotibia in ventral view (22)." figureDoi="http://doi.org/10.5281/zenodo.8391339" httpUri="https://zenodo.org/record/8391339/files/figure.png" pageId="6" pageNumber="128">Fig. 21</figureCitation>
) about as long as profemora, broadening distally and flattened dorsoventrally, lacking apical spurs, with ventral and dorsal apical rows of densely set thick trichia; protarsi (
<figureCitation id="132D2A2CFFA5FFEEA5B9FE1DFCE8884E" box="[718,802,511,537]" captionStart="FIGURES 20‒22" captionStartId="5.[151,264,1947,1971]" captionTargetBox="[151,1435,669,1924]" captionTargetId="figure-196@5.[151,1435,669,1924]" captionTargetPageId="5" captionText="FIGURES 20‒22. Bisayodes hure gen. &amp; sp. n. Pterothorax in ventral view (20); fore femur, tibia and tarsus in ventral view (21); left mesotarsus and apex of mesotibia in ventral view (22)." figureDoi="http://doi.org/10.5281/zenodo.8391339" httpUri="https://zenodo.org/record/8391339/files/figure.png" pageId="6" pageNumber="128">Fig. 21</figureCitation>
) longer than half length of tibiae, with tarsomeres 1–4 subequal in lengths, subcylindrical, tarsomere 5 elongate, subcylindrical, with pair of unmodified claws.
</paragraph>
<paragraph id="8BA936A9FFA5FFEEA7B0FDA5FD84896E" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="128">
Middle legs (
<figureCitation id="132D2A2CFFA5FFEEA621FDA5FE1B8836" box="[342,465,583,609]" captionStart="FIGURES 20‒22" captionStartId="5.[151,264,1947,1971]" captionTargetBox="[151,1435,669,1924]" captionTargetId="figure-196@5.[151,1435,669,1924]" captionTargetPageId="5" captionText="FIGURES 20‒22. Bisayodes hure gen. &amp; sp. n. Pterothorax in ventral view (20); fore femur, tibia and tarsus in ventral view (21); left mesotarsus and apex of mesotibia in ventral view (22)." figureDoi="http://doi.org/10.5281/zenodo.8391339" httpUri="https://zenodo.org/record/8391339/files/figure.png" pageId="6" pageNumber="128">Figs 20, 22</figureCitation>
) with strongly transverse mesocoxae slightly longer than procoxae and each also bearing posteroventral carina, distal coxal region above coxo-trochanteral articulation projecting mesally as subtriangular tooth; mesotrochanters small, subtriangular; mesofemora flattened dorsoventrally and narrowing distally; mesotibiae indistinctly shorter than mesofemora, flattened dorsoventrally and broadening distally, each with apical transverse row of dense and thick trichia along ventral and dorsal margin, and with two apical mesal spurs (
<figureCitation id="132D2A2CFFA5FFEEA3A7FD35FAEB88A6" box="[1232,1313,727,753]" captionStart="FIGURES 20‒22" captionStartId="5.[151,264,1947,1971]" captionTargetBox="[151,1435,669,1924]" captionTargetId="figure-196@5.[151,1435,669,1924]" captionTargetPageId="5" captionText="FIGURES 20‒22. Bisayodes hure gen. &amp; sp. n. Pterothorax in ventral view (20); fore femur, tibia and tarsus in ventral view (21); left mesotarsus and apex of mesotibia in ventral view (22)." figureDoi="http://doi.org/10.5281/zenodo.8391339" httpUri="https://zenodo.org/record/8391339/files/figure.png" pageId="6" pageNumber="128">Fig. 22</figureCitation>
) of similar lengths, slightly curved in apical regions and blunt; mesotarsi (
<figureCitation id="132D2A2CFFA5FFEEA42AFD19FC658942" box="[861,943,763,789]" captionStart="FIGURES 20‒22" captionStartId="5.[151,264,1947,1971]" captionTargetBox="[151,1435,669,1924]" captionTargetId="figure-196@5.[151,1435,669,1924]" captionTargetPageId="5" captionText="FIGURES 20‒22. Bisayodes hure gen. &amp; sp. n. Pterothorax in ventral view (20); fore femur, tibia and tarsus in ventral view (21); left mesotarsus and apex of mesotibia in ventral view (22)." figureDoi="http://doi.org/10.5281/zenodo.8391339" httpUri="https://zenodo.org/record/8391339/files/figure.png" pageId="6" pageNumber="128">Fig. 22</figureCitation>
) shorter than mesotibiae, similar to protarsi but with tarsomere 1 strongly elongate.
</paragraph>
<paragraph id="8BA936A9FFA5FFEEA7B0FCA1FC0489F2" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="128">
Hind legs (
<figureCitation id="132D2A2CFFA5FFEEA639FCA1FE6F890A" box="[334,421,835,861]" captionStart="FIGURES 20‒22" captionStartId="5.[151,264,1947,1971]" captionTargetBox="[151,1435,669,1924]" captionTargetId="figure-196@5.[151,1435,669,1924]" captionTargetPageId="5" captionText="FIGURES 20‒22. Bisayodes hure gen. &amp; sp. n. Pterothorax in ventral view (20); fore femur, tibia and tarsus in ventral view (21); left mesotarsus and apex of mesotibia in ventral view (22)." figureDoi="http://doi.org/10.5281/zenodo.8391339" httpUri="https://zenodo.org/record/8391339/files/figure.png" pageId="6" pageNumber="128">Fig. 20</figureCitation>
) with coxae hidden under metacoxal plates; metatrochanters small and subtriangular; metafemora, metatibiae and metatarsi similar to those in middle legs, but femora with apical spurs gradually narrowing to pointed apices, and metatarsi slightly longer than mesotarsi.
</paragraph>
<paragraph id="8BA936A9FFA5FFEEA7B0FC4DFA91899E" blockId="6.[151,1437,151,2013]" box="[199,1371,943,969]" pageId="6" pageNumber="128">
Abdomen (
<figureCitation id="132D2A2CFFA5FFEEA632FC4DFE42899E" box="[325,392,943,969]" captionStart="FIGURES 5‒6" captionStartId="2.[151,264,1898,1922]" captionTargetBox="[151,1435,844,1875]" captionTargetId="figure-233@2.[151,1435,844,1875]" captionTargetPageId="2" captionText="FIGURES 5‒6. Bisayodes hure gen. &amp; sp. n. Dorsal (5) and ventral (6) habitus." figureDoi="http://doi.org/10.5281/zenodo.8391333" httpUri="https://zenodo.org/record/8391333/files/figure.png" pageId="6" pageNumber="128">Fig. 6</figureCitation>
) with five visible ventrites, the first longest, but largely hidden under metacoxal plates.
</paragraph>
<paragraph id="8BA936A9FFA5FFEEA7B0FC31FB9589BA" blockId="6.[151,1437,151,2013]" box="[199,1119,979,1005]" pageId="6" pageNumber="128">
Aedeagus (
<figureCitation id="132D2A2CFFA5FFEEA632FC31FE5C89BA" box="[325,406,979,1005]" captionStart="FIGURES 37‒40" captionStartId="13.[151,264,1195,1219]" captionTargetBox="[163,1426,190,1160]" captionTargetId="figure-22@13.[151,1435,181,1171]" captionTargetPageId="13" captionText="FIGURES 37‒40. Aedeagus. Bisayodes hure gen. &amp; sp. n. (37); Tohlezkus uminchu sp. n. (38); Tohlezkus orientalis Vít (39); Tohlezkus rufus (M. Sakai) (40)." figureDoi="http://doi.org/10.5281/zenodo.8391357" httpUri="https://zenodo.org/record/8391357/files/figure.png" pageId="6" pageNumber="128">Fig. 37</figureCitation>
) with parameres devoid of setae and shorter than slender penis.
</paragraph>
</subSubSection>
<subSubSection id="C30C6522FFA5FFEEA7B0FC14FBAE8E46" box="[199,1124,1014,1041]" pageId="6" pageNumber="128" type="distribution">
<paragraph id="8BA936A9FFA5FFEEA7B0FC14FBAE8E46" blockId="6.[151,1437,151,2013]" box="[199,1124,1014,1041]" pageId="6" pageNumber="128">
<emphasis id="B962EABBFFA5FFEEA7B0FC14FDE68E47" bold="true" box="[199,556,1014,1041]" pageId="6" pageNumber="128">Distribution and composition.</emphasis>
One species inhabiting northern 
<collectingCountry id="F3017639FFA5FFEEA4E8FC15FC298E46" box="[927,995,1015,1041]" name="Japan" pageId="6" pageNumber="128">Japan</collectingCountry>
: 
<collectingRegion id="49D2F84BFFA5FFEEA499FC15FBAA8E46" box="[1006,1120,1015,1041]" country="Japan" name="Hokkaido" pageId="6" pageNumber="128">Hokkaidô</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C30C6522FFA5FFEEA7B0FBF9FB528E62" box="[199,1176,1051,1077]" pageId="6" pageNumber="128" type="etymology">
<paragraph id="8BA936A9FFA5FFEEA7B0FBF9FB528E62" blockId="6.[151,1437,151,2013]" box="[199,1176,1051,1077]" pageId="6" pageNumber="128">
<emphasis id="B962EABBFFA5FFEEA7B0FBF9FE848E62" bold="true" box="[199,334,1051,1077]" pageId="6" pageNumber="128">Etymology.</emphasis>
The name 
<taxonomicName id="4C164D2AFFA5FFEEA6BDFBF9FDF68E62" authorityName="Wakimura &amp; Jaloszynski" authorityYear="2023" box="[458,572,1051,1077]" class="Insecta" family="Eucinetidae" genus="Bisayodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA6BDFBF9FDF68E62" box="[458,572,1051,1077]" italics="true" pageId="6" pageNumber="128">Bisayodes</emphasis>
</taxonomicName>
(gender masculine) is derived from the genus 
<taxonomicName id="4C164D2AFFA5FFEEA332FBF9FB5B8E63" authorityName="Reitter" authorityYear="1884" box="[1093,1169,1051,1076]" class="Insecta" family="Eucinetidae" genus="Bisaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA332FBF9FB5B8E63" box="[1093,1169,1051,1076]" italics="true" pageId="6" pageNumber="128">Bisaya</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30C6522FFA5FFEFA7B0FBDDFF248B32" lastPageId="7" lastPageNumber="129" pageId="6" pageNumber="128" type="discussion">
<paragraph id="8BA936A9FFA5FFEFA7B0FBDDFF248B32" blockId="6.[151,1437,151,2013]" lastBlockId="7.[151,1436,151,357]" lastPageId="7" lastPageNumber="129" pageId="6" pageNumber="128">
<emphasis id="B962EABBFFA5FFEEA7B0FBDDFEF18E0E" bold="true" box="[199,315,1087,1113]" pageId="6" pageNumber="128">Remarks.</emphasis>
<taxonomicName id="4C164D2AFFA5FFEEA632FBDDFE7D8E0E" authorityName="Wakimura &amp; Jaloszynski" authorityYear="2023" box="[325,439,1087,1113]" class="Insecta" family="Eucinetidae" genus="Bisayodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA632FBDDFE7D8E0E" box="[325,439,1087,1113]" italics="true" pageId="6" pageNumber="128">Bisayodes</emphasis>
</taxonomicName>
can be distinguished from all other extant genera of 
<taxonomicName id="4C164D2AFFA5FFEEA352FBDDFB638E0E" authorityName="Lacordaire" authorityYear="1857" box="[1061,1193,1087,1113]" class="Insecta" family="Eucinetidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="family">Eucinetidae</taxonomicName>
by the following set of characters: semisuctorial mouthparts, antennae slender with trimerous clubs, antennal fossae not exposed in dorsal view, and the carinate frontoclypeal border demarcating a relatively long, dorsally clearly visible clypeus (see identification key for genera of 
<taxonomicName id="4C164D2AFFA5FFEEA54EFB49FD778E92" authorityName="Lacordaire" authorityYear="1857" box="[569,701,1195,1221]" class="Insecta" family="Eucinetidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="family">Eucinetidae</taxonomicName>
given below). Among the genera with suctorial or semisuctorial mouthparts, 
<taxonomicName id="4C164D2AFFA5FFEEA654FB2DFE5F8EBE" authorityName="Wakimura &amp; Jaloszynski" authorityYear="2023" box="[291,405,1231,1257]" class="Insecta" family="Eucinetidae" genus="Bisayodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA654FB2DFE5F8EBE" box="[291,405,1231,1257]" italics="true" pageId="6" pageNumber="128">Bisayodes</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="4C164D2AFFA5FFEEA513FB2DFD7A8EBF" authorityName="Reitter" authorityYear="1884" box="[612,688,1231,1256]" class="Insecta" family="Eucinetidae" genus="Bisaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA513FB2DFD7A8EBF" box="[612,688,1231,1256]" italics="true" pageId="6" pageNumber="128">Bisaya</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C164D2AFFA5FFEEA591FB2DFC998EBE" box="[742,851,1231,1257]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA591FB2DFC998EBE" box="[742,851,1231,1257]" italics="true" pageId="6" pageNumber="128">Tohlezkus</emphasis>
</taxonomicName>
. It differs from 
<taxonomicName id="4C164D2AFFA5FFEEA370FB2DFB998EBF" authorityName="Reitter" authorityYear="1884" box="[1031,1107,1231,1256]" class="Insecta" family="Eucinetidae" genus="Bisaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA370FB2DFB998EBF" box="[1031,1107,1231,1256]" italics="true" pageId="6" pageNumber="128">Bisaya</emphasis>
</taxonomicName>
in: (1) head pentagonal, with anterior margin strongly convex (in 
<taxonomicName id="4C164D2AFFA5FFEEA543FB11FD4A8F5B" authorityName="Reitter" authorityYear="1884" box="[564,640,1267,1292]" class="Insecta" family="Eucinetidae" genus="Bisaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA543FB11FD4A8F5B" box="[564,640,1267,1292]" italics="true" pageId="6" pageNumber="128">Bisaya</emphasis>
</taxonomicName>
hexagonal, with anterior margin truncate); (2) head not constricted at antennal insertions, hence antennal fossae not exposed in dorsal view (in 
<taxonomicName id="4C164D2AFFA5FFEEA485FAF5FBF48F67" authorityName="Reitter" authorityYear="1884" box="[1010,1086,1303,1328]" class="Insecta" family="Eucinetidae" genus="Bisaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA485FAF5FBF48F67" box="[1010,1086,1303,1328]" italics="true" pageId="6" pageNumber="128">Bisaya</emphasis>
</taxonomicName>
head strongly constricted and antennal fossae exposed in dorsal view); (3) frontoclypeal border carinate (in 
<taxonomicName id="4C164D2AFFA5FFEEA492FAD9FBFB8F03" authorityName="Reitter" authorityYear="1884" box="[997,1073,1339,1364]" class="Insecta" family="Eucinetidae" genus="Bisaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA492FAD9FBFB8F03" box="[997,1073,1339,1364]" italics="true" pageId="6" pageNumber="128">Bisaya</emphasis>
</taxonomicName>
frons and clypeus confluent); (4) labial palpomere 3 fusiform, broadest near proximal third and strongly narrowing distally, with strongly sinuate outer margin, about half as wide as palpomere 2 and as long as palpomere 2 (in 
<taxonomicName id="4C164D2AFFA5FFEEA4BDFA61FBDC8FCB" authorityName="Reitter" authorityYear="1884" box="[970,1046,1411,1436]" class="Insecta" family="Eucinetidae" genus="Bisaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA4BDFA61FBDC8FCB" box="[970,1046,1411,1436]" italics="true" pageId="6" pageNumber="128">Bisaya</emphasis>
</taxonomicName>
labial palpomere 3 nearly filiform, very slender, with outer margin nearly straight, as wide as 1/4 width of palpomere 2 and longer than palpomere 2); (5) maxillary palpomere 1 subtriangular and shorter than palpomere 2 (in 
<taxonomicName id="4C164D2AFFA5FFEEA4A0FA29FBE98FB3" authorityName="Reitter" authorityYear="1884" box="[983,1059,1483,1508]" class="Insecta" family="Eucinetidae" genus="Bisaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA4A0FA29FBE98FB3" box="[983,1059,1483,1508]" italics="true" pageId="6" pageNumber="128">Bisaya</emphasis>
</taxonomicName>
subcylindrical and clearly longer than 2); (6) galea vestigial, lacinia much longer than maxillary palpomere 1 and 2 combined, with a few widely spaced and conspicuously short distomesal setae (in 
<taxonomicName id="4C164D2AFFA5FFEEA595F9F1FCE48C7B" authorityName="Reitter" authorityYear="1884" box="[738,814,1555,1580]" class="Insecta" family="Eucinetidae" genus="Bisaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA595F9F1FCE48C7B" box="[738,814,1555,1580]" italics="true" pageId="6" pageNumber="128">Bisaya</emphasis>
</taxonomicName>
galea distinct, lacinia clearly shorter than palpomeres 1 and 2 combined, with distomesal group of dense long setae); (7) postcoxal carina or line on metaventrite at middle straight (in 
<taxonomicName id="4C164D2AFFA5FFEEA66CF9B9FEAD8C23" authorityName="Reitter" authorityYear="1884" box="[283,359,1627,1652]" class="Insecta" family="Eucinetidae" genus="Bisaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA66CF9B9FEAD8C23" box="[283,359,1627,1652]" italics="true" pageId="6" pageNumber="128">Bisaya</emphasis>
</taxonomicName>
forming subtriangular expansion between mesocoxae projecting anteriorly); (8) median region of metaventrite with its apex posteriorly reaching far behind middle length of metacoxal plates (in 
<taxonomicName id="4C164D2AFFA5FFEEA383F99DFA8A8CCF" authorityName="Reitter" authorityYear="1884" box="[1268,1344,1663,1688]" class="Insecta" family="Eucinetidae" genus="Bisaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA383F99DFA8A8CCF" box="[1268,1344,1663,1688]" italics="true" pageId="6" pageNumber="128">Bisaya</emphasis>
</taxonomicName>
apex of median region of metaventrite not reaching middle of metacoxal plates). 
<taxonomicName id="4C164D2AFFA5FFEEA4A1F941FB828CEA" authorityName="Wakimura &amp; Jaloszynski" authorityYear="2023" box="[982,1096,1699,1725]" class="Insecta" family="Eucinetidae" genus="Bisayodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA4A1F941FB828CEA" box="[982,1096,1699,1725]" italics="true" pageId="6" pageNumber="128">Bisayodes</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C164D2AFFA5FFEEA397F941FA878CEA" box="[1248,1357,1699,1725]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA397F941FA878CEA" box="[1248,1357,1699,1725]" italics="true" pageId="6" pageNumber="128">Tohlezkus</emphasis>
</taxonomicName>
in: (1) antennae loosely assembled with antennomeres 1–5 elongate and well-defined trimerous club (in 
<taxonomicName id="4C164D2AFFA5FFEEA3B1F925FAF98CB6" box="[1222,1331,1735,1761]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA3B1F925FAF98CB6" box="[1222,1331,1735,1761]" italics="true" pageId="6" pageNumber="128">Tohlezkus</emphasis>
</taxonomicName>
antennae massive, compact and with antennomeres 2–10 transverse); (2) mouthparts semisuctorial, with labrum and prelabium about twice as long as broad and with truncate anterior margins (in 
<taxonomicName id="4C164D2AFFA5FFEEA4F6F8EDFC248D7E" box="[897,1006,1807,1833]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA4F6F8EDFC248D7E" box="[897,1006,1807,1833]" italics="true" pageId="6" pageNumber="128">Tohlezkus</emphasis>
</taxonomicName>
mouthparts suctorial, with labrum and prelabium more than twice as long as broad and apically pointed); (3) maxillary palpomere 2 subtriangular, about 1.5 × as long as broad and clearly shorter than palpomere 3 (in 
<taxonomicName id="4C164D2AFFA5FFEEA414F8B5FC1A8D26" box="[867,976,1879,1905]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA414F8B5FC1A8D26" box="[867,976,1879,1905]" italics="true" pageId="6" pageNumber="128">Tohlezkus</emphasis>
</taxonomicName>
pipe-shaped or weakly clavate, 2–3 × as long as broad and clearly longer than palpomere 3); (4) maxillary palpomere 4 almost gradually narrowing distally (in 
<taxonomicName id="4C164D2AFFA5FFEEA7C8F87DFEE68DEE" box="[191,300,1951,1977]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA7C8F87DFEE68DEE" box="[191,300,1951,1977]" italics="true" pageId="6" pageNumber="128">Tohlezkus</emphasis>
</taxonomicName>
rapidly narrowing so that apical region forms a nearly rod-like projection); (5) postcoxal carina on metaventrite at middle straight (in 
<taxonomicName id="4C164D2AFFA5FFEEA54FF821FD6F8D8A" box="[568,677,1987,2013]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA5FFEEA54FF821FD6F8D8A" box="[568,677,1987,2013]" italics="true" pageId="6" pageNumber="128">Tohlezkus</emphasis>
</taxonomicName>
forming subtriangular expansion between mesocoxae projecting anteriorly); (6) median region of metaventrite broadly subtriangular, in posterior half forming a nearly equilateral triangle (in 
<taxonomicName id="4C164D2AFFA4FFEFA66DFF59FE4D8A82" box="[282,391,187,213]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA4FFEFA66DFF59FE4D8A82" box="[282,391,187,213]" italics="true" pageId="7" pageNumber="129">Tohlezkus</emphasis>
</taxonomicName>
median region of metaventrite has strongly concave sides, so that its anterior region is broad but posterior very narrow, nearly needle-shaped, posterior half forming strongly elongate triangle with narrowly sharp posterior apical angle); (7) apical mesotibial spurs of subequal lengths and both subparallel up to rounded apices (in 
<taxonomicName id="4C164D2AFFA4FFEFA678FEC5FEB68B16" box="[271,380,295,321]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B962EABBFFA4FFEFA678FEC5FEB68B16" box="[271,380,295,321]" italics="true" pageId="7" pageNumber="129">Tohlezkus</emphasis>
</taxonomicName>
mesotibial spurs are of strongly unequal lengths and both narrowing from bases to pointed apices).
</paragraph>
</subSubSection>
</treatment>
</document>