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<document id="E1FE6CA15DBFE5620DCA84CF8A769C0A" ID-DOI="10.11646/zootaxa.5318.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="8158357" ID-ZooBank="161E8842-5DB1-40CA-A4B7-2287462D86E1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1689663496356" checkinUser="plazi" docAuthor="Bennett, Robb, Copley, Claudia &amp; Copley, Darren" docDate="2023" docId="03BF87ABE43A070209BEF17361C8FCF0" docLanguage="en" docName="zootaxa.5318.1.5.pdf" docOrigin="Zootaxa 5318 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5318.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rothaeina beaudini Bennett &amp; Copley &amp; Copley 2023, spec. nov." docType="treatment" docVersion="3" lastPageNumber="118" masterDocId="FF86FFD3E42907170929F7706528FFC9" masterDocTitle="Revision of the western Nearctic spider genus Cybaeina including the description of Neocybaeina gen. nov. and Rothaeina gen. nov. (Araneae: Cybaeidae: Cybaeinae)" masterLastPageNumber="129" masterPageNumber="97" pageNumber="116" updateTime="1689664496717" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7FCC07CF4FE5A39005F396818DF08394">Revision of the western Nearctic spider genus Cybaeina including the description of Neocybaeina gen. nov. and Rothaeina gen. nov. (Araneae: Cybaeidae: Cybaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="82A77CEC4C820D0826A679DD6F330851" type="personal">
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<mods:namePart id="2C187D709749B2FE9B2FB29841A0B10A">Bennett, Robb</mods:namePart>
<mods:nameIdentifier id="4A453605C91F3110E36633202AB2FCDB" type="ORCID">0000-0002-6587-7079</mods:nameIdentifier>
<mods:affiliation id="1FE22F057CDE1BE0CCC887508DFAF6B4">robb. bennett @ shaw. ca; https: // orcid. org / 0000 - 0002 - 6587 - 7079</mods:affiliation>
<mods:nameIdentifier id="940D8C577347C8136CA8E3F93DC1AD43" type="email">robb.bennett@shaw.ca</mods:nameIdentifier>
</mods:name>
<mods:name id="00D805EC2D684E8AAE1E539D21D1D9DA" type="personal">
<mods:role id="F88C076D3C2B8A01D2EAD6E7CCDEDBFF">
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<mods:namePart id="C441212860A2A205BB67AB83F81FC379">Copley, Claudia</mods:namePart>
<mods:nameIdentifier id="37C9C87D822D7A2FAFE4AC3F39467177" type="ORCID">0000-0002-8184-5819</mods:nameIdentifier>
<mods:affiliation id="793C5546FA0D86B1CAFF84AE0F82AB42">ccopley @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0002 - 8184 - 5819</mods:affiliation>
<mods:nameIdentifier id="67BFF76E0766DCEB342946014314C327" type="email">ccopley@royalbcmuseum.bc.ca</mods:nameIdentifier>
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<mods:name id="8ED39A9DA00AED06ABD7024DD18899FF" type="personal">
<mods:role id="997B2CF240240544F04B91E7F53076E4">
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<mods:namePart id="5356BE6E9B7108FACF741383D98A0B34">Copley, Darren</mods:namePart>
<mods:nameIdentifier id="CEDF56D59065BD18B99C9962336FB3E0" type="ORCID">0000-0002-1944-4272</mods:nameIdentifier>
<mods:affiliation id="D7B896D285F49F71F4D287FC3C42A6C5">dcopley @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0002 - 1944 - 4272</mods:affiliation>
<mods:nameIdentifier id="5DFBF15162453876034AF636E7D5D8ED" type="email">dcopley@royalbcmuseum.bc.ca</mods:nameIdentifier>
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<mods:title id="ECAF1369E0F86194FE39B07B11A16311">Zootaxa</mods:title>
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<mods:date id="D1578B46254C928935FA3451AE7D3969">2023</mods:date>
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<mods:number id="756C5F85DFAA1BEC9AC07FCD63D2AC6C">2023-07-18</mods:number>
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<mods:classification id="20281DE51B99F827C51983DE9C92E000">journal article</mods:classification>
<mods:identifier id="19E24562AD52C046790B438124DA0E71" type="DOI">10.11646/zootaxa.5318.1.5</mods:identifier>
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<treatment id="03BF87ABE43A070209BEF17361C8FCF0" LSID="urn:lsid:plazi:treatment:03BF87ABE43A070209BEF17361C8FCF0" httpUri="http://treatment.plazi.org/id/03BF87ABE43A070209BEF17361C8FCF0" lastPageId="21" lastPageNumber="118" pageId="19" pageNumber="116">
<subSubSection id="C30C6536E43A070409BEF1736746F9D7" box="[151,622,1539,1566]" pageId="19" pageNumber="116" type="nomenclature">
<paragraph id="8BA936BDE43A070409BEF1736746F9D7" blockId="19.[151,622,1539,1603]" box="[151,622,1539,1566]" pageId="19" pageNumber="116">
<heading id="D0E181D1E43A070409BEF1736746F9D7" bold="true" box="[151,622,1539,1566]" fontSize="11" level="1" pageId="19" pageNumber="116" reason="1">
<emphasis id="B962EAAFE43A070409BEF1736746F9D7" bold="true" box="[151,622,1539,1566]" pageId="19" pageNumber="116">
<taxonomicName id="4C164D3EE43A070409BEF17364C7F9D7" authority="Bennett" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2023" box="[151,495,1539,1566]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="116" phylum="Arthropoda" rank="species" species="beaudini" status="spec. nov.">
<emphasis id="B962EAAFE43A070409BEF17364AEF9D7" bold="true" box="[151,390,1539,1566]" italics="true" pageId="19" pageNumber="116">Rothaeina beaudini</emphasis>
Bennett
</taxonomicName>
<taxonomicNameLabel id="A25157D4E43A070408DFF1746746F9D7" box="[502,622,1540,1566]" pageId="19" pageNumber="116" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30C6536E43A070409BEF1586416F98A" box="[151,318,1576,1603]" pageId="19" pageNumber="116" type="description">
<paragraph id="8BA936BDE43A070409BEF1586416F98A" blockId="19.[151,622,1539,1603]" box="[151,318,1576,1603]" pageId="19" pageNumber="116">
<figureCitation id="132D2A38E43A070409BEF1586430F98B" box="[151,280,1576,1602]" captionStart="FIGURES 6669" captionStartId="20.[152,270,1168,1192]" captionTargetBox="[151,1435,182,1144]" captionTargetId="figure-37@20.[151,1436,181,1144]" captionTargetPageId="20" captionText="FIGURES 6669. Rothaeina beaudini spec. nov., females from California (6667 from Delta; 68 from Manzanita Lake; 69 from Lake Almanor), copulatory organ. 66 Epigyne, ventral. 6769 Vulva (6768 dorsal, 69 ventral). BG—Bennetts gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial opening (6667, 69)." figureDoi="http://doi.org/10.5281/zenodo.8158434" httpUri="https://zenodo.org/record/8158434/files/figure.png" pageId="19" pageNumber="116">Figs 6669</figureCitation>
,
<figureCitation id="132D2A38E43A0704080BF1586416F98A" box="[290,318,1576,1603]" captionStart="FIGURES 8486" captionStartId="28.[152,270,1909,1933]" captionTargetBox="[177,1411,181,1884]" captionTargetId="figure-37@28.[177,1411,181,1884]" captionTargetPageId="28" captionText="FIGURES 8486. Distributions of Neocybaeina gen. nov. and Rothaeina gen. nov. species (84 Neocybaeina burnetti spec. nov. and N. xantha comb. nov.; 85 Rothaeina beaudini spec. nov., R. jamesi spec. nov., R. mackinleyi spec. nov., and R. sequoia comb. nov.; 86 Rothaeina petersoni spec. nov.) AZ—Arizona, CA—California, ID—Idaho, MT—Montana, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington." figureDoi="http://doi.org/10.5281/zenodo.8158395" httpUri="https://zenodo.org/record/8158395/files/figure.png" pageId="19" pageNumber="116">85</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30C6536E43A070409BEF10066D4F84F" pageId="19" pageNumber="116" type="materials_examined">
<paragraph id="8BA936BDE43A070409BEF10066D4F84F" blockId="19.[151,1437,1647,1998]" pageId="19" pageNumber="116">
<emphasis id="B962EAAFE43A070409BEF1006460F943" bold="true" box="[151,328,1648,1674]" pageId="19" pageNumber="116">Type material.</emphasis>
<materialsCitation id="3B7E3CE0E43A0704087BF1006604F967" collectingDate="1961-09-19" collectionCode="AMNH" collectorName="W. Ivie &amp; W. J. Gertsch" country="United States of America" county="Tehama County" location="Lassen Volcanic National Park" pageId="19" pageNumber="116" specimenCount="1" stateProvince="California" typeStatus="holotype">
<emphasis id="B962EAAFE43A0704087BF1006624F943" bold="true" box="[338,780,1647,1674]" pageId="19" pageNumber="116">
<collectingCountry id="F301762DE43A0704087BF100648AF943" box="[338,418,1648,1674]" name="United States of America" pageId="19" pageNumber="116">U.S.A.</collectingCountry>
:
<emphasis id="B962EAAFE43A0704089AF11F6703F943" bold="true" box="[435,555,1647,1674]" italics="true" pageId="19" pageNumber="116">
<collectingRegion id="49D2F85FE43A0704089AF11F6703F943" box="[435,555,1647,1674]" country="United States of America" name="California" pageId="19" pageNumber="116">California</collectingRegion>
</emphasis>
:
<typeStatus id="54AD881FE43A07040B17F1006783F943" box="[574,683,1648,1674]" pageId="19" pageNumber="116">Holotype</typeStatus>
female.
</emphasis>
<collectingCounty id="62C84E31E43A07040A3FF10066E1F943" box="[790,969,1648,1674]" pageId="19" pageNumber="116">Tehama County</collectingCounty>
, near junction south of
<location id="8EC96066E43A07040DCAF100641BF967" LSID="urn:lsid:plazi:treatment:03BF87ABE43A070209BEF17361C8FCF0:8EC96066E43A07040DCAF100641BF967" country="United States of America" county="Tehama County" name="Lassen Volcanic National Park" pageId="19" pageNumber="116" stateProvince="California">Lassen Volcanic National Park</location>
,
<date id="FFA8107DE43A07040815F1E4649EF967" box="[316,438,1684,1710]" pageId="19" pageNumber="116" value="1961-09-19">
<collectingDate id="EFECE995E43A07040815F1E4649EF967" box="[316,438,1684,1710]" pageId="19" pageNumber="116" value="1961-09-19">19.ix.1961</collectingDate>
</date>
,
<collectorName id="26E3536BE43A07040896F1E46727F967" box="[447,527,1684,1710]" pageId="19" pageNumber="116">W. Ivie</collectorName>
&amp;
<collectorName id="26E3536BE43A07040B07F1E46793F967" box="[558,699,1684,1710]" pageId="19" pageNumber="116">W.J. Gertsch</collectorName>
(
<collectionCode id="ED07AE78E43A07040BE1F1E4660CF967" box="[712,804,1684,1710]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="19" pageNumber="116">AMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7E3CE0E43A07040A1DF1E467B0F91B" collectingDate="1966-05-05" collectionCode="CAS" collectorName="Humbug &amp; Understock Roads &amp; Dokes" country="United States of America" location="Butte" pageId="19" pageNumber="116" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
<emphasis id="B962EAAFE43A07040A1DF1E4669BF967" bold="true" box="[820,947,1684,1710]" pageId="19" pageNumber="116">
<typeStatus id="54AD881FE43A07040A1DF1E46686F967" box="[820,942,1684,1710]" pageId="19" pageNumber="116">Paratypes</typeStatus>
.
</emphasis>
<location id="8EC96066E43A07040A91F1E466DDF967" LSID="urn:lsid:plazi:treatment:03BF87ABE43A070209BEF17361C8FCF0:8EC96066E43A07040A91F1E466DDF967" box="[952,1013,1684,1710]" country="United States of America" name="Butte" pageId="19" pageNumber="116" stateProvince="California">Butte</location>
,
<specimenCount id="9D10FD34E43A07040D28F1E4610CF967" box="[1025,1060,1684,1710]" pageId="19" pageNumber="116" type="female">1♀</specimenCount>
,
<quantity id="4CEE9B58E43A07040D19F1E46155F967" box="[1072,1149,1684,1710]" metricMagnitude="3" metricUnit="m" metricValue="5.3108352" pageId="19" pageNumber="116" unit="mi" value="3.3">3.3 mi.</quantity>
from junction of
<collectorName id="26E3536BE43A07040C13F1E460B4F967" box="[1338,1436,1684,1710]" pageId="19" pageNumber="116">Humbug</collectorName>
&amp;
<collectorName id="26E3536BE43A0704099DF1C864ADF91B" box="[180,389,1720,1746]" pageId="19" pageNumber="116">Understock Roads</collectorName>
,
<date id="FFA8107DE43A070408A6F1C864DAF91A" box="[399,498,1720,1747]" pageId="19" pageNumber="116" value="1966-05-05">
<collectingDate id="EFECE995E43A070408A6F1C864DAF91A" box="[399,498,1720,1747]" pageId="19" pageNumber="116" value="1966-05-05">5.v.1966</collectingDate>
</date>
,
<collectorName id="26E3536BE43A070408D4F1C8676DF91B" box="[509,581,1720,1746]" pageId="19" pageNumber="116">Dokes</collectorName>
(
<collectionCode id="ED07AE78E43A07040B7DF1C867B8F91B" box="[596,656,1720,1746]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="19" pageNumber="116" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7E3CE0E43A07040B8AF1C860BFF91B" box="[675,1431,1720,1746]" collectingDate="1988-09-04" collectionCode="CAS" collectorName="D. Ubick" country="United States of America" location="Plumas" pageId="19" pageNumber="116" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
<location id="8EC96066E43A07040B8AF1C867DDF91B" LSID="urn:lsid:plazi:treatment:03BF87ABE43A070209BEF17361C8FCF0:8EC96066E43A07040B8AF1C867DDF91B" box="[675,757,1720,1746]" country="United States of America" name="Plumas" pageId="19" pageNumber="116" stateProvince="California">Plumas</location>
,
<specimenCount id="9D10FD34E43A07040A2DF1C8660FF91B" box="[772,807,1720,1746]" pageId="19" pageNumber="116" type="female">1♀</specimenCount>
,
<locationDeviation id="9A1B2F9FE43A07040A1CF1C86174F91B" box="[821,1116,1720,1746]" location="Bucks Lake" pageId="19" pageNumber="116">
<quantity id="4CEE9B58E43A07040A1CF1C866ABF91B" box="[821,899,1720,1746]" metricMagnitude="4" metricUnit="m" metricValue="1.48059648" pageId="19" pageNumber="116" unit="mi" value="9.2">9.2 mi.</quantity>
NE of Bucks Lake
</locationDeviation>
,
<date id="FFA8107DE43A07040D4FF1C861FBF91B" box="[1126,1235,1720,1746]" pageId="19" pageNumber="116" value="1988-09-04">
<collectingDate id="EFECE995E43A07040D4FF1C861FBF91B" box="[1126,1235,1720,1746]" pageId="19" pageNumber="116" value="1988-09-04">4.ix.1988</collectingDate>
</date>
,
<collectorName id="26E3536BE43A07040DF7F1C8606CF91B" box="[1246,1348,1720,1746]" pageId="19" pageNumber="116">D. Ubick</collectorName>
(
<collectionCode id="ED07AE78E43A07040C7DF1C860B8F91B" box="[1364,1424,1720,1746]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="19" pageNumber="116" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7E3CE0E43A070409BEF1AC66F5F93F" box="[151,989,1756,1783]" collectingDate="1959-09-05" collectionCode="AMNH" collectorName="V. D. Roth &amp; W. J. Gertsch" country="United States of America" location="Lake Almanor" pageId="19" pageNumber="116" specimenCount="7" specimenCount-female="7" stateProvince="California" typeStatus="paratype">
<specimenCount id="9D10FD34E43A070409BEF1AC6593F93F" box="[151,187,1756,1782]" pageId="19" pageNumber="116" type="female">7♀</specimenCount>
, S side of
<location id="8EC96066E43A07040805F1AC64E5F93F" LSID="urn:lsid:plazi:treatment:03BF87ABE43A070209BEF17361C8FCF0:8EC96066E43A07040805F1AC64E5F93F" box="[300,461,1756,1782]" country="United States of America" name="Lake Almanor" pageId="19" pageNumber="116" stateProvince="California">Lake Almanor</location>
,
<date id="FFA8107DE43A070408F1F1AC676BF93E" box="[472,579,1756,1783]" pageId="19" pageNumber="116" value="1959-09-05">
<collectingDate id="EFECE995E43A070408F1F1AC676BF93E" box="[472,579,1756,1783]" pageId="19" pageNumber="116" value="1959-09-05">5.ix.1959</collectingDate>
</date>
,
<collectorName id="26E3536BE43A07040B67F1AC6794F93F" box="[590,700,1756,1782]" pageId="19" pageNumber="116">V.D. Roth</collectorName>
&amp;
<collectorName id="26E3536BE43A07040BF4F1AC6643F93F" box="[733,875,1756,1782]" pageId="19" pageNumber="116">W.J. Gertsch</collectorName>
(
<collectionCode id="ED07AE78E43A07040A50F1AC66FDF93F" box="[889,981,1756,1782]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="19" pageNumber="116">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7E3CE0E43A07040ACEF1AC67A4F8D3" collectingDate="1961-09-19" collectionCode="AMNH" collectorName="W. Ivie &amp; W. J. Gertsch" country="United States of America" location="Shasta" pageId="19" pageNumber="116" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
<location id="8EC96066E43A07040ACEF1AC6118F93F" LSID="urn:lsid:plazi:treatment:03BF87ABE43A070209BEF17361C8FCF0:8EC96066E43A07040ACEF1AC6118F93F" box="[999,1072,1756,1782]" country="United States of America" name="Shasta" pageId="19" pageNumber="116" stateProvince="California">Shasta</location>
,
<specimenCount id="9D10FD34E43A07040D15F1AC6177F93F" box="[1084,1119,1756,1782]" pageId="19" pageNumber="116" type="female">1♀</specimenCount>
,
<locationDeviation id="9A1B2F9FE43A07040D45F1AC60B0F93F" box="[1132,1432,1756,1783]" location="Manzanita Lake" pageId="19" pageNumber="116">
<quantity id="4CEE9B58E43A07040D45F1AC618DF93F" box="[1132,1189,1756,1783]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="19" pageNumber="116" unit="mi" value="5.0">5 mi.</quantity>
N of Manzanita Lake
</locationDeviation>
,
<date id="FFA8107DE43A070409BEF070643AF8D3" box="[151,274,1792,1818]" pageId="19" pageNumber="116" value="1961-09-19">
<collectingDate id="EFECE995E43A070409BEF070643AF8D3" box="[151,274,1792,1818]" pageId="19" pageNumber="116" value="1961-09-19">19.ix.1961</collectingDate>
</date>
,
<collectorName id="26E3536BE43A07040832F0706444F8D3" box="[283,364,1792,1818]" pageId="19" pageNumber="116">W. Ivie</collectorName>
&amp;
<collectorName id="26E3536BE43A070408A5F0706732F8D3" box="[396,538,1792,1818]" pageId="19" pageNumber="116">W.J. Gertsch</collectorName>
(
<collectionCode id="ED07AE78E43A07040B01F07067ACF8D3" box="[552,644,1792,1818]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="19" pageNumber="116">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7E3CE0E43A07040BBFF070642BF8F7" collectingDate="1959-09-03" collectionCode="AMNH" collectorName="W. J. Gertsch &amp; V. D. Roth" country="United States of America" location="Shasta Lake" municipality="Delta" pageId="19" pageNumber="116" specimenCount="3" specimenCount-female="3" stateProvince="California" typeStatus="paratype">
<specimenCount id="9D10FD34E43A07040BBFF0706792F8D3" box="[662,698,1792,1818]" pageId="19" pageNumber="116" type="female">3♀</specimenCount>
,
<collectingMunicipality id="6BCDACC7E43A07040BEFF070662DF8D3" box="[710,773,1792,1818]" pageId="19" pageNumber="116">Delta</collectingMunicipality>
, N end of
<location id="8EC96066E43A07040A5CF07066D6F8D3" LSID="urn:lsid:plazi:treatment:03BF87ABE43A070209BEF17361C8FCF0:8EC96066E43A07040A5CF07066D6F8D3" box="[885,1022,1792,1818]" country="United States of America" municipality="Delta" name="Shasta Lake" pageId="19" pageNumber="116" stateProvince="California">Shasta Lake</location>
,
<date id="FFA8107DE43A07040D21F070615DF8D2" box="[1032,1141,1792,1819]" pageId="19" pageNumber="116" value="1959-09-03">
<collectingDate id="EFECE995E43A07040D21F070615DF8D2" box="[1032,1141,1792,1819]" pageId="19" pageNumber="116" value="1959-09-03">3.ix.1959</collectingDate>
</date>
,
<collectorName id="26E3536BE43A07040D57F0706024F8D3" box="[1150,1292,1792,1818]" pageId="19" pageNumber="116">W.J. Gertsch</collectorName>
&amp;
<collectorName id="26E3536BE43A07040C04F07060B4F8D3" box="[1325,1436,1792,1818]" pageId="19" pageNumber="116">V.D. Roth</collectorName>
(
<collectionCode id="ED07AE78E43A070409B6F05465D3F8F7" box="[159,251,1828,1854]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="19" pageNumber="116">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7E3CE0E43A07040827F054601AF8F7" box="[270,1330,1828,1855]" collectingDate="1959-09-04" collectionCode="AMNH" collectorName="V. D. Roth &amp; W. J. Gertsch" country="United States of America" location="Emigrant Ford Rd." pageId="19" pageNumber="116" specimenCount="4" specimenCount-female="4" stateProvince="California" typeStatus="paratype">
<specimenCount id="9D10FD34E43A07040827F054641BF8F7" box="[270,307,1828,1854]" pageId="19" pageNumber="116" type="female">4♀</specimenCount>
,
<location id="8EC96066E43A07040816F054673DF8F7" LSID="urn:lsid:plazi:treatment:03BF87ABE43A070209BEF17361C8FCF0:8EC96066E43A07040816F054673DF8F7" box="[319,533,1828,1854]" country="United States of America" name="Emigrant Ford Rd." pageId="19" pageNumber="116" stateProvince="California">Emigrant Ford Rd.</location>
,
<locationDeviation id="9A1B2F9FE43A07040B09F0546632F8F7" box="[544,794,1828,1854]" location="Old Station" pageId="19" pageNumber="116">
<quantity id="4CEE9B58E43A07040B09F0546773F8F7" box="[544,603,1828,1854]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="19" pageNumber="116" unit="mi" value="4.0">4 mi.</quantity>
S of Old Station
</locationDeviation>
,
<date id="FFA8107DE43A07040A0CF05466B9F8F6" box="[805,913,1828,1855]" pageId="19" pageNumber="116" value="1959-09-04">
<collectingDate id="EFECE995E43A07040A0CF05466B9F8F6" box="[805,913,1828,1855]" pageId="19" pageNumber="116" value="1959-09-04">4.ix.1959</collectingDate>
</date>
,
<collectorName id="26E3536BE43A07040AB5F0546124F8F7" box="[924,1036,1828,1854]" pageId="19" pageNumber="116">V.D. Roth</collectorName>
&amp;
<collectorName id="26E3536BE43A07040D19F0546197F8F7" box="[1072,1215,1828,1854]" pageId="19" pageNumber="116">W.J. Gertsch</collectorName>
(
<collectionCode id="ED07AE78E43A07040DE7F0546002F8F7" box="[1230,1322,1828,1854]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="19" pageNumber="116">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7E3CE0E43A07040C14F054618CF8AB" collectingDate="1968-08-08" collectionCode="UASM" collectorName="F. O. Leech" country="United States of America" location="Hwy" municipality="Tehama" pageId="19" pageNumber="116" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
<collectingMunicipality id="6BCDACC7E43A07040C14F05460BDF8F7" box="[1341,1429,1828,1854]" pageId="19" pageNumber="116">Tehama</collectingMunicipality>
,
<specimenCount id="9D10FD34E43A070409BEF0386592F8AB" box="[151,186,1864,1890]" pageId="19" pageNumber="116" type="female">1♀</specimenCount>
,
<location id="8EC96066E43A070409E3F0386428F8AB" LSID="urn:lsid:plazi:treatment:03BF87ABE43A070209BEF17361C8FCF0:8EC96066E43A070409E3F0386428F8AB" box="[202,256,1864,1890]" country="United States of America" municipality="Tehama" name="Hwy" pageId="19" pageNumber="116" stateProvince="California">Hwy</location>
89,
<locationDeviation id="9A1B2F9FE43A0704081DF038660BF8AB" box="[308,803,1864,1891]" location="Lassen Volcanic National Park" pageId="19" pageNumber="116">
<quantity id="4CEE9B58E43A0704081DF03864ACF8AB" box="[308,388,1864,1891]" metricMagnitude="3" metricUnit="m" metricValue="5.632704" pageId="19" pageNumber="116" unit="mi" value="3.5">3.5 mi.</quantity>
S of Lassen Volcanic National Park
</locationDeviation>
,
<date id="FFA8107DE43A07040A19F0386683F8AB" box="[816,939,1864,1890]" pageId="19" pageNumber="116" value="1968-08-08">
<collectingDate id="EFECE995E43A07040A19F0386683F8AB" box="[816,939,1864,1890]" pageId="19" pageNumber="116" value="1968-08-08">8.viii.1968</collectingDate>
</date>
,
<collectorName id="26E3536BE43A07040A91F038611CF8AB" box="[952,1076,1864,1890]" pageId="19" pageNumber="116">F.O. Leech</collectorName>
(
<collectionCode id="ED07AE78E43A07040D6CF03861B4F8AB" box="[1093,1180,1864,1890]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34404" lsid="urn:lsid:biocol.org:col:34404" name="University of Alberta, E.H. Strickland Entomological Museum" pageId="19" pageNumber="116">UASM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7E3CE0E43A07040D98F03866D0F84F" collectingDate="1961-09-19" collectionCode="AMNH" collectorName="W. Ivie &amp; W. J. Gertsch" country="United States of America" location="Lassen Volcanic National Park" pageId="19" pageNumber="116" specimenCount="5" specimenCount-female="5" stateProvince="California" typeStatus="paratype">
<specimenCount id="9D10FD34E43A07040D98F03861FDF8AA" box="[1201,1237,1864,1891]" pageId="19" pageNumber="116" type="female">5♀</specimenCount>
, nr. junction S of
<location id="8EC96066E43A070409BEF01C64D9F84F" LSID="urn:lsid:plazi:treatment:03BF87ABE43A070209BEF17361C8FCF0:8EC96066E43A070409BEF01C64D9F84F" box="[151,497,1900,1926]" country="United States of America" name="Lassen Volcanic National Park" pageId="19" pageNumber="116" stateProvince="California">Lassen Volcanic National Park</location>
,
<date id="FFA8107DE43A070408D4F01C675FF84F" box="[509,631,1900,1926]" pageId="19" pageNumber="116" value="1961-09-19">
<collectingDate id="EFECE995E43A070408D4F01C675FF84F" box="[509,631,1900,1926]" pageId="19" pageNumber="116" value="1961-09-19">19.ix.1961</collectingDate>
</date>
,
<collectorName id="26E3536BE43A07040BABF01C67FCF84F" box="[642,724,1900,1926]" pageId="19" pageNumber="116">W. Ivie</collectorName>
&amp;
<collectorName id="26E3536BE43A07040BDEF01C66ADF84F" box="[759,901,1900,1926]" pageId="19" pageNumber="116">W.J. Gertsch</collectorName>
(
<collectionCode id="ED07AE78E43A07040ABDF01C66D8F84F" box="[916,1008,1900,1926]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="19" pageNumber="116">AMNH</collectionCode>
)
</materialsCitation>
;
</paragraph>
</subSubSection>
<subSubSection id="C30C6536E43A070409EEF0E06673F807" pageId="19" pageNumber="116" type="etymology">
<paragraph id="8BA936BDE43A070409EEF0E06673F807" blockId="19.[151,1437,1647,1998]" pageId="19" pageNumber="116">
<emphasis id="B962EAAFE43A070409EEF0E06466F863" bold="true" box="[199,334,1936,1962]" pageId="19" pageNumber="116">Etymology.</emphasis>
The specific name is a patronym honouring Beaudin A. Bennett who was born in
<date id="FFA8107DE43A07040DC3F0E060A9F862" box="[1258,1409,1936,1963]" pageId="19" pageNumber="116" value="1985-01">January 1985</date>
at the beginning of the first authors study of Nearctic Cybaeinae.
</paragraph>
</subSubSection>
<caption id="DF696635E43D070309B1F3E065E8FADD" ID-DOI="http://doi.org/10.5281/zenodo.8158434" ID-Zenodo-Dep="8158434" httpUri="https://zenodo.org/record/8158434/files/figure.png" pageId="20" pageNumber="117" startId="20.[152,270,1168,1192]" targetBox="[151,1435,182,1144]" targetPageId="20" targetType="figure">
<paragraph id="8BA936BDE43D070309B1F3E065E8FADD" blockId="20.[151,1436,1168,1300]" pageId="20" pageNumber="117">
<emphasis id="B962EAAFE43D070309B1F3E06474FB61" bold="true" box="[152,348,1168,1192]" pageId="20" pageNumber="117">FIGURES 6669.</emphasis>
<taxonomicName id="4C164D3EE43D0703084AF3E06704FB61" authority="Bennett &amp; Copley &amp; Copley, 2023" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2023" box="[355,556,1168,1192]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="117" phylum="Arthropoda" rank="species" species="beaudini" status="spec. nov.">
<emphasis id="B962EAAFE43D0703084AF3E06704FB61" box="[355,556,1168,1192]" italics="true" pageId="20" pageNumber="117">Rothaeina beaudini</emphasis>
</taxonomicName>
<emphasis id="B962EAAFE43D07030B1AF3E067B3FB61" bold="true" box="[563,667,1168,1192]" pageId="20" pageNumber="117">
<taxonomicNameLabel id="A25157D4E43D07030B1AF3E067B3FB61" box="[563,667,1168,1192]" pageId="20" pageNumber="117" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, females from California (6667 from Delta; 68 from Manzanita Lake; 69 from Lake Almanor), copulatory organ. 66 Epigyne, ventral. 6769 Vulva (6768 dorsal, 69 ventral). BG—Bennetts gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial opening (6667, 69).
</paragraph>
</caption>
<subSubSection id="C30C6536E43D070209EEF216668FFF1C" lastPageId="21" lastPageNumber="118" pageId="20" pageNumber="117" type="diagnosis">
<paragraph id="8BA936BDE43D070309EEF21661C8FA25" blockId="20.[151,1437,1382,2021]" pageId="20" pageNumber="117">
<emphasis id="B962EAAFE43D070309EEF216646AFA49" bold="true" box="[199,322,1382,1408]" pageId="20" pageNumber="117">Diagnosis.</emphasis>
As is common among females of other Cybaeinae as well as other supraspecific taxa of spiders (
<bibRefCitation id="EF874B4CE43D070309B6F2FB6410FA6D" author="Bennett, R. G." box="[159,312,1418,1445]" pageId="20" pageNumber="117" pagination="473 - 492" refId="ref20905" refString="Bennett, R. G. (2006) Ontogeny, variation, and synonymy in North American Cybaeus spiders (Araneae: Cybaeidae). The Canadian Entomologist, 138 (4), 473 - 492. https: // doi. org / 10.4039 / n 06 - 804" type="journal article" year="2006">Bennett 2006</bibRefCitation>
), intraspecific variability and interspecific similarity of genitalic morphology characters can render the females of
<taxonomicName id="4C164D3EE43D07030839F2DE64ABFA01" authority="Bennett &amp; Copley &amp; Copley, 2023" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2023" box="[272,387,1454,1480]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="117" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B962EAAFE43D07030839F2DE64ABFA01" box="[272,387,1454,1480]" italics="true" pageId="20" pageNumber="117">Rothaeina</emphasis>
</taxonomicName>
<emphasis id="B962EAAFE43D070308A0F2DF64D8FA00" bold="true" box="[393,496,1455,1481]" pageId="20" pageNumber="117">
<taxonomicNameLabel id="A25157D4E43D070308A0F2DF64D8FA00" box="[393,496,1455,1481]" pageId="20" pageNumber="117" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
difficult to distinguish from each other. In some cases, collection with accompanying males, or geographic locality, may provide the best indication of species identification of females.
</paragraph>
<paragraph id="8BA936BDE43D070309EEF28665FBF8C5" blockId="20.[151,1437,1382,2021]" pageId="20" pageNumber="117">
The male of
<taxonomicName id="4C164D3EE43D0703087AF28764FBF9D9" authority="Bennett &amp; Copley &amp; Copley, 2023" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2023" box="[339,467,1526,1552]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="117" phylum="Arthropoda" rank="species" species="beaudini" status="spec. nov.">
<emphasis id="B962EAAFE43D0703087AF28764FBF9D9" box="[339,467,1526,1552]" italics="true" pageId="20" pageNumber="117">R. beaudini</emphasis>
</taxonomicName>
<emphasis id="B962EAAFE43D070308F0F2876762F9D8" bold="true" box="[473,586,1527,1553]" pageId="20" pageNumber="117">
<taxonomicNameLabel id="A25157D4E43D070308F0F2876762F9D8" box="[473,586,1527,1553]" pageId="20" pageNumber="117" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
is unknown; the female can usually be distinguished from its congeners by a combination of epigynal, atrial, and spermathecal characters. The atrium (
<figureCitation id="132D2A38E43D07030AEDF16A613CF9FD" box="[964,1044,1562,1588]" captionStart="FIGURES 6669" captionStartId="20.[152,270,1168,1192]" captionTargetBox="[151,1435,182,1144]" captionTargetId="figure-37@20.[151,1436,181,1144]" captionTargetPageId="20" captionText="FIGURES 6669. Rothaeina beaudini spec. nov., females from California (6667 from Delta; 68 from Manzanita Lake; 69 from Lake Almanor), copulatory organ. 66 Epigyne, ventral. 6769 Vulva (6768 dorsal, 69 ventral). BG—Bennetts gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial opening (6667, 69)." figureDoi="http://doi.org/10.5281/zenodo.8158434" httpUri="https://zenodo.org/record/8158434/files/figure.png" pageId="20" pageNumber="117">Fig. 66</figureCitation>
) is usually small but prominent and U-shaped and, although various components of the vulval ducts are visible through the integument of the uncleared epigyne, none appear decidedly ring-shaped. The vulva (
<figureCitation id="132D2A38E43D07030A3EF11266B0F9B5" box="[791,920,1634,1660]" captionStart="FIGURES 6669" captionStartId="20.[152,270,1168,1192]" captionTargetBox="[151,1435,182,1144]" captionTargetId="figure-37@20.[151,1436,181,1144]" captionTargetPageId="20" captionText="FIGURES 6669. Rothaeina beaudini spec. nov., females from California (6667 from Delta; 68 from Manzanita Lake; 69 from Lake Almanor), copulatory organ. 66 Epigyne, ventral. 6769 Vulva (6768 dorsal, 69 ventral). BG—Bennetts gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial opening (6667, 69)." figureDoi="http://doi.org/10.5281/zenodo.8158434" httpUri="https://zenodo.org/record/8158434/files/figure.png" pageId="20" pageNumber="117">Figs 6769</figureCitation>
) is relatively narrow with width (measured at widest extent of spermathecae) usually less than 3.5 times atrial height (measured from epigastric groove to atrium). Additionally, in the vulva the path of the copulatory and spermathecal ducts is relatively easily to trace from the atrium to the fertilization ducts and the medial transverse section of the spermathecae is inconspicuous in dorsal view.
</paragraph>
<paragraph id="8BA936BDE43D070209EEF067668FFF1C" blockId="20.[151,1437,1382,2021]" lastBlockId="21.[151,1437,151,825]" lastPageId="21" lastPageNumber="118" pageId="20" pageNumber="117">
In the females of the other species of
<taxonomicName id="4C164D3EE43D07030B5CF06667C0F8F9" authority="Bennett &amp; Copley &amp; Copley, 2023" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2023" box="[629,744,1814,1840]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="117" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B962EAAFE43D07030B5CF06667C0F8F9" box="[629,744,1814,1840]" italics="true" pageId="20" pageNumber="117">Rothaeina</emphasis>
</taxonomicName>
<emphasis id="B962EAAFE43D07030BD8F0676673F8F8" bold="true" box="[753,859,1815,1841]" pageId="20" pageNumber="117">
<taxonomicNameLabel id="A25157D4E43D07030BD8F0676673F8F8" box="[753,859,1815,1841]" pageId="20" pageNumber="117" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, the atrium is usually small but inconspicuous and never U-shaped (
<figureCitation id="132D2A38E43D07030872F04A64C8F89C" box="[347,480,1850,1877]" captionStart="FIGURES 7074" captionStartId="22.[152,270,1556,1580]" captionTargetBox="[158,1426,182,1532]" captionTargetId="figure-37@22.[158,1430,181,1532]" captionTargetPageId="22" captionText="FIGURES 7074. Rothaeina gen. nov. species, females (7072 R. jamesi spec. nov. from Klamath, California; 7374 R. mackinleyi spec. nov. from Charleston, Oregon), copulatory organ. 70 Epigyne, ventral. 7174 Vulva (71, 73 ventral; 72, 74 dorsal). BG—Bennetts gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial openings (7071, 74 single shafted arrows), additional loop of spermatheca (72 double shafted arrow), and medial transverse sections of spermathecae (72, 74 dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.8158438" httpUri="https://zenodo.org/record/8158438/files/figure.png" pageId="20" pageNumber="117">Figs 70, 73</figureCitation>
,
<figureCitation id="132D2A38E43D070308C7F04A6724F89C" box="[494,524,1850,1877]" captionStart="FIGURES 7579" captionStartId="25.[152,270,1849,1873]" captionTargetBox="[151,1436,418,1825]" captionTargetId="figure-112@25.[151,1436,417,1825]" captionTargetPageId="25" captionText="FIGURES 7579. Rothaeina petersoni spec. nov., females from California (75 from Kyburz; 76 from Sierra City; 77 from Tamarack Flat; 78 from Forest Hill; 79 from near Yuba Pass), copulatory organ. 75 Epigyne, ventral. 7679 Vulva, dorsal). BG—Bennetts gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial openings (7576, 78 single shafted arrows), and ring-like loops of spermathecae (7576, 79 double shafted arrows)." figureDoi="http://doi.org/10.5281/zenodo.8158389" httpUri="https://zenodo.org/record/8158389/files/figure.png" pageId="20" pageNumber="117">75</figureCitation>
,
<figureCitation id="132D2A38E43D07030B33F04B6712F89C" box="[538,570,1851,1877]" captionStart="FIGURES 8083" captionStartId="27.[152,270,1127,1151]" captionTargetBox="[151,1436,181,1104]" captionTargetId="figure-19@27.[151,1436,181,1104]" captionTargetPageId="27" captionText="FIGURES 8083. Rothaeina sequoia comb. nov. females from California (8082 from Pepperwood; 83 from Scotia), copulatory organ. 80 Epigyne, ventral. 8183 Vulva (81 ventral; 8283 dorsal). BG—Bennetts gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial openings (80, 8283 single shafted arrows), and ring-like loops of spermathecae (80 double shafted arrows)." figureDoi="http://doi.org/10.5281/zenodo.8158391" httpUri="https://zenodo.org/record/8158391/files/figure.png" pageId="20" pageNumber="117">80</figureCitation>
). Furthermore, the females of
<taxonomicName id="4C164D3EE43D07030AB1F04B6132F89D" authority="Bennett &amp; Copley &amp; Copley, 2023" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2023" box="[920,1050,1851,1876]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="117" phylum="Arthropoda" rank="species" species="petersoni" status="spec. nov.">
<emphasis id="B962EAAFE43D07030AB1F04B6132F89D" box="[920,1050,1851,1876]" italics="true" pageId="20" pageNumber="117">R petersoni</emphasis>
</taxonomicName>
<emphasis id="B962EAAFE43D07030D0AF04B61B0F89C" bold="true" box="[1059,1176,1851,1877]" pageId="20" pageNumber="117">
<taxonomicNameLabel id="A25157D4E43D07030D0AF04B61B0F89C" box="[1059,1176,1851,1877]" pageId="20" pageNumber="117" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C164D3EE43D07030DFBF04B6063F89D" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2023" baseAuthorityName="Bennett &amp; Copley &amp; Copley" baseAuthorityYear="1952" box="[1234,1355,1851,1876]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="117" phylum="Arthropoda" rank="species" species="sequoia" status="comb. nov.">
<emphasis id="B962EAAFE43D07030DFBF04B6063F89D" box="[1234,1355,1851,1876]" italics="true" pageId="20" pageNumber="117">R. sequoia</emphasis>
</taxonomicName>
<emphasis id="B962EAAFE43D07030C7DF04B65E0F8B0" bold="true" pageId="20" pageNumber="117">
<taxonomicNameLabel id="A25157D4E43D07030C7DF04B65E0F8B0" pageId="20" pageNumber="117" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
are unique among the females of
<taxonomicName id="4C164D3EE43D07030B6CF02E6790F8B1" authority="Bennett &amp; Copley &amp; Copley, 2023" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2023" box="[581,696,1886,1912]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="117" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B962EAAFE43D07030B6CF02E6790F8B1" box="[581,696,1886,1912]" italics="true" pageId="20" pageNumber="117">Rothaeina</emphasis>
</taxonomicName>
<emphasis id="B962EAAFE43D07030B96F02F6601F8B0" bold="true" box="[703,809,1887,1913]" pageId="20" pageNumber="117">
<taxonomicNameLabel id="A25157D4E43D07030B96F02F6601F8B0" box="[703,809,1887,1913]" pageId="20" pageNumber="117" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
in having the posterior-most spermathecal ducts clearly visible through the integument of the uncleared epigyne as circular, ring-like structures (
<figureCitation id="132D2A38E43D07030D5FF0F261E4F854" box="[1142,1228,1922,1949]" captionStart="FIGURES 7579" captionStartId="25.[152,270,1849,1873]" captionTargetBox="[151,1436,418,1825]" captionTargetId="figure-112@25.[151,1436,417,1825]" captionTargetPageId="25" captionText="FIGURES 7579. Rothaeina petersoni spec. nov., females from California (75 from Kyburz; 76 from Sierra City; 77 from Tamarack Flat; 78 from Forest Hill; 79 from near Yuba Pass), copulatory organ. 75 Epigyne, ventral. 7679 Vulva, dorsal). BG—Bennetts gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial openings (7576, 78 single shafted arrows), and ring-like loops of spermathecae (7576, 79 double shafted arrows)." figureDoi="http://doi.org/10.5281/zenodo.8158389" httpUri="https://zenodo.org/record/8158389/files/figure.png" pageId="20" pageNumber="117">Figs 75</figureCitation>
,
<figureCitation id="132D2A38E43D07030DF0F0F361DEF854" box="[1241,1270,1923,1949]" captionStart="FIGURES 8083" captionStartId="27.[152,270,1127,1151]" captionTargetBox="[151,1436,181,1104]" captionTargetId="figure-19@27.[151,1436,181,1104]" captionTargetPageId="27" captionText="FIGURES 8083. Rothaeina sequoia comb. nov. females from California (8082 from Pepperwood; 83 from Scotia), copulatory organ. 80 Epigyne, ventral. 8183 Vulva (81 ventral; 8283 dorsal). BG—Bennetts gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial openings (80, 8283 single shafted arrows), and ring-like loops of spermathecae (80 double shafted arrows)." figureDoi="http://doi.org/10.5281/zenodo.8158391" httpUri="https://zenodo.org/record/8158391/files/figure.png" pageId="20" pageNumber="117">80</figureCitation>
) and the vulva is relatively broad (vulval width usually 45 times atrial height) (
<figureCitation id="132D2A38E43D07030A58F0D666DBF808" box="[881,1011,1958,1985]" captionStart="FIGURES 7579" captionStartId="25.[152,270,1849,1873]" captionTargetBox="[151,1436,418,1825]" captionTargetId="figure-112@25.[151,1436,417,1825]" captionTargetPageId="25" captionText="FIGURES 7579. Rothaeina petersoni spec. nov., females from California (75 from Kyburz; 76 from Sierra City; 77 from Tamarack Flat; 78 from Forest Hill; 79 from near Yuba Pass), copulatory organ. 75 Epigyne, ventral. 7679 Vulva, dorsal). BG—Bennetts gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial openings (7576, 78 single shafted arrows), and ring-like loops of spermathecae (7576, 79 double shafted arrows)." figureDoi="http://doi.org/10.5281/zenodo.8158389" httpUri="https://zenodo.org/record/8158389/files/figure.png" pageId="20" pageNumber="117">Figs 7779</figureCitation>
,
<figureCitation id="132D2A38E43D07030AD7F0D76160F808" box="[1022,1096,1959,1985]" captionStart="FIGURES 8083" captionStartId="27.[152,270,1127,1151]" captionTargetBox="[151,1436,181,1104]" captionTargetId="figure-19@27.[151,1436,181,1104]" captionTargetPageId="27" captionText="FIGURES 8083. Rothaeina sequoia comb. nov. females from California (8082 from Pepperwood; 83 from Scotia), copulatory organ. 80 Epigyne, ventral. 8183 Vulva (81 ventral; 8283 dorsal). BG—Bennetts gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial openings (80, 8283 single shafted arrows), and ring-like loops of spermathecae (80 double shafted arrows)." figureDoi="http://doi.org/10.5281/zenodo.8158391" httpUri="https://zenodo.org/record/8158391/files/figure.png" pageId="20" pageNumber="117">8183</figureCitation>
). Finally, in the females of the remaining two species (
<taxonomicName id="4C164D3EE43D0703088AF0BB6724F82D" authority="Bennett &amp; Copley &amp; Copley, 2023" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2023" box="[419,524,1995,2020]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="117" phylum="Arthropoda" rank="species" species="jamesi" status="spec. nov.">
<emphasis id="B962EAAFE43D0703088AF0BB6724F82D" box="[419,524,1995,2020]" italics="true" pageId="20" pageNumber="117">R. jamesi</emphasis>
</taxonomicName>
<emphasis id="B962EAAFE43D07030B3AF0BB67AEF82C" bold="true" box="[531,646,1995,2021]" pageId="20" pageNumber="117">
<taxonomicNameLabel id="A25157D4E43D07030B3AF0BB67AEF82C" box="[531,646,1995,2021]" pageId="20" pageNumber="117" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C164D3EE43D07030B94F0BB667DF82D" authority="Bennett &amp; Copley &amp; Copley, 2023" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2023" box="[701,853,1994,2020]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="117" phylum="Arthropoda" rank="species" species="mackinleyi" status="spec. nov.">
<emphasis id="B962EAAFE43D07030B94F0BB667DF82D" box="[701,853,1994,2020]" italics="true" pageId="20" pageNumber="117">R. mackinleyi</emphasis>
</taxonomicName>
<emphasis id="B962EAAFE43D07030A74F0BB66F8F82C" bold="true" box="[861,976,1995,2021]" pageId="20" pageNumber="117">
<taxonomicNameLabel id="A25157D4E43D07030A74F0BB66F8F82C" box="[861,976,1995,2021]" pageId="20" pageNumber="117" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
), the vulval ducts (
<figureCitation id="132D2A38E43D07030D80F0BA6001F82C" box="[1193,1321,1994,2021]" captionStart="FIGURES 7074" captionStartId="22.[152,270,1556,1580]" captionTargetBox="[158,1426,182,1532]" captionTargetId="figure-37@22.[158,1430,181,1532]" captionTargetPageId="22" captionText="FIGURES 7074. Rothaeina gen. nov. species, females (7072 R. jamesi spec. nov. from Klamath, California; 7374 R. mackinleyi spec. nov. from Charleston, Oregon), copulatory organ. 70 Epigyne, ventral. 7174 Vulva (71, 73 ventral; 72, 74 dorsal). BG—Bennetts gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial openings (7071, 74 single shafted arrows), additional loop of spermatheca (72 double shafted arrow), and medial transverse sections of spermathecae (72, 74 dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.8158438" httpUri="https://zenodo.org/record/8158438/files/figure.png" pageId="20" pageNumber="117">Figs 72, 74</figureCitation>
) are more complex with the path from the atrium to the fertilization ducts difficult to trace and, in dorsal view, the medial transverse section of each spermatheca is prominent and conspicuous.
</paragraph>
</subSubSection>
<subSubSection id="C30C6536E43C070209EEF7AE663BFD60" pageId="21" pageNumber="118" type="description">
<paragraph id="8BA936BDE43C070209EEF7AE6777FED4" blockId="21.[151,1437,151,825]" pageId="21" pageNumber="118">
<emphasis id="B962EAAFE43C070209EEF7AE6471FF31" bold="true" box="[199,345,222,248]" pageId="21" pageNumber="118">Description.</emphasis>
As in diagnosis and description of the genus. Additional descriptive characters presented here. Abdomen pale or gray, lightly patterned.
</paragraph>
<paragraph id="8BA936BDE43C070209EEF65664AFFE88" blockId="21.[151,1437,151,825]" box="[199,391,294,321]" pageId="21" pageNumber="118">
<emphasis id="B962EAAFE43C070209EEF656642AFE88" bold="true" box="[199,258,294,321]" italics="true" pageId="21" pageNumber="118">Male</emphasis>
: Unknown.
</paragraph>
<paragraph id="8BA936BDE43C070209EEF63A66FAFE64" blockId="21.[151,1437,151,825]" pageId="21" pageNumber="118">
<emphasis id="B962EAAFE43C070209EEF63A6436FEAC" bold="true" box="[199,286,330,357]" italics="true" pageId="21" pageNumber="118">Female</emphasis>
: (n=24). Epigyne (
<figureCitation id="132D2A38E43C070208DBF63B676AFEAC" box="[498,578,331,357]" captionStart="FIGURES 6669" captionStartId="20.[152,270,1168,1192]" captionTargetBox="[151,1435,182,1144]" captionTargetId="figure-37@20.[151,1436,181,1144]" captionTargetPageId="20" captionText="FIGURES 6669. Rothaeina beaudini spec. nov., females from California (6667 from Delta; 68 from Manzanita Lake; 69 from Lake Almanor), copulatory organ. 66 Epigyne, ventral. 6769 Vulva (6768 dorsal, 69 ventral). BG—Bennetts gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial opening (6667, 69)." figureDoi="http://doi.org/10.5281/zenodo.8158434" httpUri="https://zenodo.org/record/8158434/files/figure.png" pageId="21" pageNumber="118">Fig. 66</figureCitation>
) with single, anteromedial atrium; atrium occasionally reduced or apparently absent. Vulva (
<figureCitation id="132D2A38E43C07020869F61F64EAFE40" box="[320,450,367,393]" captionStart="FIGURES 6669" captionStartId="20.[152,270,1168,1192]" captionTargetBox="[151,1435,182,1144]" captionTargetId="figure-37@20.[151,1436,181,1144]" captionTargetPageId="20" captionText="FIGURES 6669. Rothaeina beaudini spec. nov., females from California (6667 from Delta; 68 from Manzanita Lake; 69 from Lake Almanor), copulatory organ. 66 Epigyne, ventral. 6769 Vulva (6768 dorsal, 69 ventral). BG—Bennetts gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial opening (6667, 69)." figureDoi="http://doi.org/10.5281/zenodo.8158434" httpUri="https://zenodo.org/record/8158434/files/figure.png" pageId="21" pageNumber="118">Figs 6769</figureCitation>
) with copulatory ducts separated at atrium; spermathecal ducts not as convoluted as in other species; Bennetts glands within medial third of spermathecal ducts.
</paragraph>
<paragraph id="8BA936BDE43C070209EEF6C76692FE3C" blockId="21.[151,1437,151,825]" pageId="21" pageNumber="118">
Measurements (n=20). CL 1.952.45 (2.26+0.15), CW 1.481.90 (1.73+0.11), SL 1.091.33 (1.23+0.07), SW 1.031.24 (1.14+0.06).
<typeStatus id="54AD881FE43C07020888F6AB6721FE3C" box="[417,521,475,501]" pageId="21" pageNumber="118">Holotype</typeStatus>
CL 2.25, CW 1.73, SL 1.24, SW 1.17.
</paragraph>
<paragraph id="8BA936BDE43C070209EEF68F663BFD60" blockId="21.[151,1437,151,825]" pageId="21" pageNumber="118">
<emphasis id="B962EAAFE43C070209EEF68F6420FDD0" bold="true" box="[199,264,511,537]" pageId="21" pageNumber="118">Note:</emphasis>
Two species may be represented here. Specimens from the northwestern part of the distribution around Shasta Lake have more heavily pigmented abdomens and somewhat more complex spermathecal ducts (
<figureCitation id="132D2A38E43C07020C68F55360BCFDF4" box="[1345,1428,547,573]" captionStart="FIGURES 6669" captionStartId="20.[152,270,1168,1192]" captionTargetBox="[151,1435,182,1144]" captionTargetId="figure-37@20.[151,1436,181,1144]" captionTargetPageId="20" captionText="FIGURES 6669. Rothaeina beaudini spec. nov., females from California (6667 from Delta; 68 from Manzanita Lake; 69 from Lake Almanor), copulatory organ. 66 Epigyne, ventral. 6769 Vulva (6768 dorsal, 69 ventral). BG—Bennetts gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial opening (6667, 69)." figureDoi="http://doi.org/10.5281/zenodo.8158434" httpUri="https://zenodo.org/record/8158434/files/figure.png" pageId="21" pageNumber="118">Fig. 67</figureCitation>
) than specimens from southeast of Shasta Lake (
<figureCitation id="132D2A38E43C07020B8CF5376600FDA8" box="[677,808,583,609]" captionStart="FIGURES 6669" captionStartId="20.[152,270,1168,1192]" captionTargetBox="[151,1435,182,1144]" captionTargetId="figure-37@20.[151,1436,181,1144]" captionTargetPageId="20" captionText="FIGURES 6669. Rothaeina beaudini spec. nov., females from California (6667 from Delta; 68 from Manzanita Lake; 69 from Lake Almanor), copulatory organ. 66 Epigyne, ventral. 6769 Vulva (6768 dorsal, 69 ventral). BG—Bennetts gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial opening (6667, 69)." figureDoi="http://doi.org/10.5281/zenodo.8158434" httpUri="https://zenodo.org/record/8158434/files/figure.png" pageId="21" pageNumber="118">Figs 6869</figureCitation>
). Because the male of
<taxonomicName id="4C164D3EE43C07020D37F53761B6FDA8" authority="Bennett &amp; Copley &amp; Copley, 2023" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2023" box="[1054,1182,583,609]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="118" phylum="Arthropoda" rank="species" species="beaudini" status="spec. nov.">
<emphasis id="B962EAAFE43C07020D37F53761B6FDA8" box="[1054,1182,583,609]" italics="true" pageId="21" pageNumber="118">R. beaudini</emphasis>
</taxonomicName>
<emphasis id="B962EAAFE43C07020D8DF537603DFDA8" bold="true" box="[1188,1301,583,609]" pageId="21" pageNumber="118">
<taxonomicNameLabel id="A25157D4E43C07020D8DF537603DFDA8" box="[1188,1301,583,609]" pageId="21" pageNumber="118" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
is unknown and the vulval and atrial characters of females tend to be variable even within populations, all these specimens are here considered to be members of
<taxonomicName id="4C164D3EE43C07020B30F5FF67B2FD60" authority="Bennett &amp; Copley &amp; Copley, 2023" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2023" box="[537,666,655,681]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="118" phylum="Arthropoda" rank="species" species="beaudini" status="spec. nov.">
<emphasis id="B962EAAFE43C07020B30F5FF67B2FD60" box="[537,666,655,681]" italics="true" pageId="21" pageNumber="118">R. beaudini</emphasis>
</taxonomicName>
<emphasis id="B962EAAFE43C07020B88F5FF663BFD60" bold="true" box="[673,787,655,681]" pageId="21" pageNumber="118">
<taxonomicNameLabel id="A25157D4E43C07020B88F5FF663BFD60" box="[673,787,655,681]" pageId="21" pageNumber="118" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30C6536E43C070209EEF5C261C8FCF0" pageId="21" pageNumber="118" type="distribution">
<paragraph id="8BA936BDE43C070209EEF5C261C8FCF0" blockId="21.[151,1437,151,825]" pageId="21" pageNumber="118">
<emphasis id="B962EAAFE43C070209EEF5C2677DFD05" bold="true" box="[199,597,690,717]" pageId="21" pageNumber="118">Distribution and natural history.</emphasis>
(
<figureCitation id="132D2A38E43C07020B4FF5C36793FD04" box="[614,699,690,717]" captionStart="FIGURES 8486" captionStartId="28.[152,270,1909,1933]" captionTargetBox="[177,1411,181,1884]" captionTargetId="figure-37@28.[177,1411,181,1884]" captionTargetPageId="28" captionText="FIGURES 8486. Distributions of Neocybaeina gen. nov. and Rothaeina gen. nov. species (84 Neocybaeina burnetti spec. nov. and N. xantha comb. nov.; 85 Rothaeina beaudini spec. nov., R. jamesi spec. nov., R. mackinleyi spec. nov., and R. sequoia comb. nov.; 86 Rothaeina petersoni spec. nov.) AZ—Arizona, CA—California, ID—Idaho, MT—Montana, OR—Oregon, NV—Nevada, UT—Utah, WA—Washington." figureDoi="http://doi.org/10.5281/zenodo.8158395" httpUri="https://zenodo.org/record/8158395/files/figure.png" pageId="21" pageNumber="118">Fig. 85</figureCitation>
).
<taxonomicName id="4C164D3EE43C07020BFBF5C36698FD04" authority="Bennett &amp; Copley &amp; Copley, 2023" authorityName="Bennett &amp; Copley &amp; Copley" authorityYear="2023" box="[722,944,691,717]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="118" phylum="Arthropoda" rank="species" species="beaudini" status="spec. nov.">
<emphasis id="B962EAAFE43C07020BFBF5C36698FD04" box="[722,944,691,717]" italics="true" pageId="21" pageNumber="118">Rothaeina beaudini</emphasis>
</taxonomicName>
<emphasis id="B962EAAFE43C07020A90F5C36106FD04" bold="true" box="[953,1070,691,717]" pageId="21" pageNumber="118">
<taxonomicNameLabel id="A25157D4E43C07020A90F5C36106FD04" box="[953,1070,691,717]" pageId="21" pageNumber="118" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
is endemic to north central and northeastern
<collectingRegion id="49D2F85FE43C07020802F5A764B7FD38" box="[299,415,727,753]" country="United States of America" name="California" pageId="21" pageNumber="118">California</collectingRegion>
,
<collectingCountry id="F301762DE43C07020884F5A764D4FD38" box="[429,508,727,753]" name="United States of America" pageId="21" pageNumber="118">U.S.A.</collectingCountry>
, from the Shasta Lake area southeast to the northern Sierra
<collectingRegion id="49D2F85FE43C07020D9FF5A76027FD38" box="[1206,1295,727,753]" country="United States of America" name="Nevada" pageId="21" pageNumber="118">Nevada</collectingRegion>
. Within that area, females appear to have been relatively common in the 1950s and 1960s but only a single specimen has been recorded since the 1960s (in 1988) and the conservation status of this taxon is currently unknown.
</paragraph>
</subSubSection>
</treatment>
</document>