treatments-xml/data/03/BF/87/03BF87A3C84DFFB9FF3DFBBCFF4DA481.xml
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<document id="4FCDE8DC2118EDED2FDF70F320236B46" ID-CLB-Dataset="8059" ID-DOI="10.11646/zootaxa.4933.2.1" ID-GBIF-Dataset="1a2ba1ca-028f-4b95-8dda-5ddec4e63524" ID-ISSN="1175-5326" ID-Zenodo-Dep="4550146" ID-ZooBank="5D1D20A5-0F44-4AEF-AF5F-A758FC37D076" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613716226573" checkinUser="plazi" docAuthor="Anistratenko, Vitaliy V., Neubauer, Thomas A., Anistratenko, Olga Yu., Kijashko, Pavel V. &amp; Wesselingh, Frank P." docDate="2021" docId="03BF87A3C84DFFB9FF3DFBBCFF4DA481" docLanguage="en" docName="zootaxa.4933.2.1.pdf" docOrigin="Zootaxa 4933 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Andrusovia cyrensis Anistratenko &amp; Neubauer &amp; Anistratenko &amp; Kijashko &amp; Wesselingh 2021, sp. nov." docType="treatment" docUuid="3A413153-1A98-47A7-A93A-B00CC7CE8E04" docUuidSource="ZooBank" docVersion="8" lastPageNumber="165" masterDocId="FF86FFDBC841FFB7FFAAFFDEFFA5A577" masterDocTitle="A revision of the Pontocaspian gastropods of the subfamily Caspiinae (Caenogastropoda: Hydrobiidae)" masterLastPageNumber="197" masterPageNumber="151" pageNumber="163" updateTime="1698930299740" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D2D96E74158B17346252BE73B8E83FBF">A revision of the Pontocaspian gastropods of the subfamily Caspiinae (Caenogastropoda: Hydrobiidae)</mods:title>
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<mods:namePart id="6943458B5E31B2C14288E53C8CC4D7CC">Anistratenko, Vitaliy V.</mods:namePart>
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<mods:affiliation id="D7407FC25051B724A87D9856BFB1F59B">Department of Animal Ecology and Systematics, Justus Liebig University, Heinrich-Buff-Ring 26 - 32 IFZ, 35392 Giessen, Germany. tneub @ zo. jlug. de; https: // orcid. org / 0000 - 0002 - 1398 - 9941 &amp; Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, The Netherlands.</mods:affiliation>
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<mods:affiliation id="0D59417672F891E75AFF79BAC43B8066">Department of Invertebrate Fauna and Systematics, Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, B. Khmelnytsky Str. 15, 01030 Kiev, Ukraine. &amp; Department of Cainozoic Deposits, Institute of Geological Sciences, National Academy of Sciences of Ukraine, O. Gontchar Str. 55 b, 01054 Kiev, Ukraine.</mods:affiliation>
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<paragraph id="8BA936B5C84DFFBBFF6DFA87FDCDA0E0" blockId="12.[151,1437,1261,2043]" pageId="12" pageNumber="163">
<materialsCitation id="3B7E3CE8C84DFFBBFF6DFA87FDC1A0E0" ID-GBIF-Occurrence="3042095327" collectionCode="WGS" country="Azerbaijan" latitude="39.286983" location="South Caspian Basin" longLatPrecision="1" longitude="49.602116" municipality="Kura" pageId="12" pageNumber="163" specimenCode="WGS84" specimenCount="1" typeStatus="holotype">
<emphasis id="B962EAA7C84DFFBBFF6DFA87FEC2A004" bold="true" box="[199,359,1369,1395]" pageId="12" pageNumber="163">
<typeStatus id="54AD8817C84DFFBBFF6DFA87FEA7A004" box="[199,258,1369,1395]" pageId="12" pageNumber="163">Type</typeStatus>
locality.
</emphasis>
<location id="8EC9606EC84DFFBBFEDAFA87FDF8A004" LSID="urn:lsid:plazi:treatment:03BF87A3C84DFFB9FF3DFBBCFF4DA481:8EC9606EC84DFFBBFEDAFA87FDF8A004" box="[368,605,1369,1395]" country="Azerbaijan" latitude="39.286983" longLatPrecision="1" longitude="49.602116" municipality="Kura" name="South Caspian Basin" pageId="12" pageNumber="163">South Caspian Basin</location>
off
<collectingMunicipality id="6BCDACCFC84DFFBBFD24FA87FD63A004" box="[654,710,1369,1395]" pageId="12" pageNumber="163">Kura</collectingMunicipality>
delta,
<collectingCountry id="F3017625C84DFFBBFCBEFA87FC2AA004" box="[788,911,1369,1395]" name="Azerbaijan" pageId="12" pageNumber="163">Azerbaijan</collectingCountry>
(locality 25f, sample 14-1); GPS coordinates:
<geoCoordinate id="EE225072C84DFFBBFF3DFAA3FEE3A0E0" box="[151,326,1405,1431]" degrees="39" direction="north" minutes="17" orientation="latitude" pageId="12" pageNumber="163" precision="1" seconds="13.14" value="39.286983">39°1713.14”N</geoCoordinate>
,
<geoCoordinate id="EE225072C84DFFBBFEFBFAA3FE49A0E0" box="[337,492,1405,1431]" degrees="49" direction="east" minutes="36" orientation="longitude" pageId="12" pageNumber="163" precision="1" seconds="7.62" value="49.602116">49°367.62”E</geoCoordinate>
(
<specimenCode id="DBB09ECEC84DFFBBFE51FAA3FDF9A0E0" box="[507,604,1405,1431]" collectionCode="WGS" pageId="12" pageNumber="163">WGS84</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA936B5C84DFFBBFF6DFA7FFCEBA0CB" blockId="12.[151,1437,1261,2043]" box="[199,846,1441,1468]" pageId="12" pageNumber="163">
<emphasis id="B962EAA7C84DFFBBFF6DFA7FFF58A0CC" bold="true" box="[199,253,1441,1467]" pageId="12" pageNumber="163">Age.</emphasis>
Subfossil, late Holocene (
<bibRefCitation id="EF874B44C84DFFBBFD89FA7FFCE7A0CB" author="Hoogendoorn, R. M. &amp; Boels, J. F. &amp; Kroonenberg, S. B. &amp; Simmons, M. D. &amp; Aliyeva, E. &amp; Babazadeh, A. D. &amp; Huseynov, D." box="[547,834,1441,1468]" pageId="12" pageNumber="163" pagination="359 - 380" refId="ref31193" refString="Hoogendoorn, R. M., Boels, J. F., Kroonenberg, S. B., Simmons, M. D., Aliyeva, E., Babazadeh, A. D. &amp; Huseynov, D. (2005) Development of the Kura delta, Azerbaijan; a record of Holocene Caspian sea-level changes. Marine Geology, 222 - 223, 359 - 380. https: // doi. org / 10.1016 / j. margeo. 2005.06.007" type="journal article" year="2005">
Hoogendoorn
<emphasis id="B962EAA7C84DFFBBFD6CFA7CFD52A0CC" box="[710,759,1441,1467]" italics="true" pageId="12" pageNumber="163">et al</emphasis>
. 2005
</bibRefCitation>
).
</paragraph>
<paragraph id="8BA936B5C84DFFBBFF6DFA1BFB10A097" blockId="12.[151,1437,1261,2043]" box="[199,1205,1477,1504]" pageId="12" pageNumber="163">
<emphasis id="B962EAA7C84DFFBBFF6DFA1BFE24A0A8" bold="true" box="[199,385,1477,1503]" pageId="12" pageNumber="163">Other material.</emphasis>
Three specimens from localities 25d and 25f (
<collectionCode id="ED07AE70C84DFFBBFC20FA1BFC69A0A8" box="[906,972,1477,1503]" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="163" type="Museum">RGM</collectionCode>
13102961310298).
</paragraph>
</subSubSection>
<subSubSection id="C30C653EC84DFFBBFF6DFA37FCC6A374" box="[199,867,1513,1539]" pageId="12" pageNumber="163" type="etymology">
<paragraph id="8BA936B5C84DFFBBFF6DFA37FCC6A374" blockId="12.[151,1437,1261,2043]" box="[199,867,1513,1539]" pageId="12" pageNumber="163">
<emphasis id="B962EAA7C84DFFBBFF6DFA37FEEBA374" bold="true" box="[199,334,1513,1539]" pageId="12" pageNumber="163">Etymology.</emphasis>
From Cyrus, the Latin name of the Kura River.
</paragraph>
</subSubSection>
<subSubSection id="C30C653EC84DFFBBFF6DF9D3FCC3A3E4" pageId="12" pageNumber="163" type="diagnosis">
<paragraph id="8BA936B5C84DFFBBFF6DF9D3FCC3A3E4" blockId="12.[151,1437,1261,2043]" pageId="12" pageNumber="163">
<emphasis id="B962EAA7C84DFFBBFF6DF9D3FEE7A350" bold="true" box="[199,322,1549,1575]" pageId="12" pageNumber="163">Diagnosis.</emphasis>
Low trochiform, conical, small
<taxonomicName id="4C164D36C84DFFBBFD05F9D3FC95A350" box="[687,816,1549,1575]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="12" pageNumber="163" phylum="Ciliophora" rank="genus">
<emphasis id="B962EAA7C84DFFBBFD05F9D3FC95A350" box="[687,816,1549,1575]" italics="true" pageId="12" pageNumber="163">Andrusovia</emphasis>
</taxonomicName>
with up to three convex, slightly stepped whorls, wide umbilicus, broadly ovoid aperture, weakly sinuate and weakly thickened peristome, strongly wrinkled protoconch with faint spiral threads and teleoconch microsculpture consisting of fine, densely spaced, slightly irregular, raised spiral threads, which are typically dissected into elongate beads.
</paragraph>
</subSubSection>
<subSubSection id="C30C653EC84DFFBBFF6DF943FC3FA28C" pageId="12" pageNumber="163" type="description">
<paragraph id="8BA936B5C84DFFBBFF6DF943FC3FA28C" blockId="12.[151,1437,1261,2043]" pageId="12" pageNumber="163">
<emphasis id="B962EAA7C84DFFBBFF6DF943FEFCA3C0" bold="true" box="[199,345,1693,1719]" pageId="12" pageNumber="163">Description.</emphasis>
Small shell, slightly broader as high, with up to three whorls with stepped appearance. Protoconch low domical, cap-like, consisting of 1 whorl that measures c.
<quantity id="4CEE9B50C84DFFBBFCE4F91FFC45A3AC" box="[846,992,1729,1756]" metricMagnitude="-1" metricUnit="m" metricValue="2.975" metricValueMax="3.05" metricValueMin="2.9" pageId="12" pageNumber="163" unit="mm" value="297.5" valueMax="305.0" valueMin="290.0">290305 µm</quantity>
; protoconch covered with densely malleate sculpture that may become wider meshed and spikier on second half; faint traces of up to four spiral threads appear on first half near lower suture in some specimens; final triangular portion reaching from lower suture to end of protoconch, starting at c. 0.75 whorls, devoid of malleate sculpture but bearing growth rims that increase in strength towards transition to teleoconch; transition marked by growth stop and onset of fine prosocyrt growth lines. Teleoconch whorls covered with dense microsculpture consisting of fine, densely spaced, slightly irregular, raised spiral threads, which are typically dissected into elongate beads. Suture incised and deep. Body whorl attains c. 80% of total shell height, forms shallow base. Aperture wide, ovoid, with weak notch at contact to penultimate whorl; umbilicus circular, wide, deep. Peristome slightly thickened, sinuate.
</paragraph>
</subSubSection>
<caption id="DF69663DC84CFFBAFF3DF899FD92A298" ID-DOI="http://doi.org/10.5281/zenodo.4550160" ID-Zenodo-Dep="4550160" httpUri="https://zenodo.org/record/4550160/files/figure.png" pageId="13" pageNumber="164" startId="13.[151,250,1863,1887]" targetBox="[176,1411,181,1839]" targetPageId="13">
<paragraph id="8BA936B5C84CFFBAFF3DF899FD92A298" blockId="13.[151,1437,1863,2032]" pageId="13" pageNumber="164">
<emphasis id="B962EAA7C84CFFBAFF3DF899FEB6A228" bold="true" box="[151,275,1863,1887]" pageId="13" pageNumber="164">FIGURE 5.</emphasis>
<taxonomicName id="4C164D36C84CFFBAFEB3F899FE43A228" authority="Anistratenko &amp; Neubauer &amp; Anistratenko &amp; Kijashko &amp; Wesselingh, 2021" authorityName="Anistratenko &amp; Neubauer &amp; Anistratenko &amp; Kijashko &amp; Wesselingh" authorityYear="2021" box="[281,486,1863,1887]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="13" pageNumber="164" phylum="Ciliophora" rank="species" species="cyrensis" status="sp. nov.">
<emphasis id="B962EAA7C84CFFBAFEB3F899FE43A228" box="[281,486,1863,1887]" italics="true" pageId="13" pageNumber="164">Andrusovia cyrensis</emphasis>
</taxonomicName>
<emphasis id="B962EAA7C84CFFBAFE46F899FD98A228" bold="true" box="[492,573,1863,1887]" pageId="13" pageNumber="164">
<taxonomicNameLabel id="A25157DCC84CFFBAFE46F899FD98A228" box="[492,573,1863,1887]" pageId="13" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from the late Holocene of the South Caspian Sea off the Kura delta, Azerbaijan. (a, j, k, r) Holotype, locality 25f (RGM 1310292). (b, m) Paratype, locality 25d (RGM 1310293). (c, d, l, q, s) Paratype, locality 25f (RGM 1310294). (e, f, i, p, t) Paratype, locality 25f (RGM 1310295). (g) Specimen from locality 25d (RGM 1310296). (h, n) Specimen from locality 25f (RGM 1310297). (o) Specimen from locality 25f (RGM 1310298). Scale bars = 500 µm (ah), 100 µm (i, j, lq, s, t), 50 µm (k), 10 µm (r).
</paragraph>
</caption>
<subSubSection id="C30C653EC84FFFB9FF6DFF49FDFAA4DA" pageId="14" pageNumber="165" type="discussion">
<paragraph id="8BA936B5C84FFFB9FF6DFF49FDFAA4DA" blockId="14.[151,1437,151,502]" pageId="14" pageNumber="165">
<emphasis id="B962EAA7C84FFFB9FF6DFF49FE9EA5C6" bold="true" box="[199,315,151,177]" pageId="14" pageNumber="165">Remarks.</emphasis>
The new species resembles
<taxonomicName id="4C164D36C84FFFB9FDDDFF49FCA3A5C6" authorityName="Brusina" authorityYear="1902" box="[631,774,151,177]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="14" pageNumber="165" phylum="Ciliophora" rank="species" species="dybowskii">
<emphasis id="B962EAA7C84FFFB9FDDDFF49FCA3A5C6" box="[631,774,151,177]" italics="true" pageId="14" pageNumber="165">A. dybowskii</emphasis>
</taxonomicName>
superficially concerning the low trochiform shape, but it is distinctly smaller, has a lower spire, a smaller protoconch (
<quantity id="4CEE9B50C84FFFB9FC84FF64FC1AA5A2" box="[814,959,186,213]" metricMagnitude="-1" metricUnit="m" metricValue="2.975" metricValueMax="3.05" metricValueMin="2.9" pageId="14" pageNumber="165" unit="mm" value="297.5" valueMax="305.0" valueMin="290.0">290305 µm</quantity>
versus
<quantity id="4CEE9B50C84FFFB9FBBFFF64FB03A5A2" box="[1045,1190,186,213]" metricMagnitude="-1" metricUnit="m" metricValue="3.975" metricValueMax="4.15" metricValueMin="3.8" pageId="14" pageNumber="165" unit="mm" value="397.5" valueMax="415.0" valueMin="380.0">380415 µm</quantity>
) at approximately the same number of whorls, more convex whorls and a teleoconch microsculpture that is so far unknown in any other
<taxonomicName id="4C164D36C84FFFB9FF3DFEDDFEBDA46A" box="[151,280,259,285]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="14" pageNumber="165" phylum="Ciliophora" rank="genus">
<emphasis id="B962EAA7C84FFFB9FF3DFEDDFEBDA46A" box="[151,280,259,285]" italics="true" pageId="14" pageNumber="165">Andrusovia</emphasis>
</taxonomicName>
species. Only specimens formerly attributed to
<taxonomicName id="4C164D36C84FFFB9FC81FEDDFB87A46A" box="[811,1058,259,285]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="14" pageNumber="165" phylum="Ciliophora" rank="species" species="andrusovi">
<emphasis id="B962EAA7C84FFFB9FC81FEDDFB87A46A" box="[811,1058,259,285]" italics="true" pageId="14" pageNumber="165">Andrusovia andrusovi</emphasis>
</taxonomicName>
(here considered a synonym of
<taxonomicName id="4C164D36C84FFFB9FA2EFEDDFEA2A436" authorityName="Brusina" authorityYear="1902" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="14" pageNumber="165" phylum="Ciliophora" rank="species" species="dybowskii">
<emphasis id="B962EAA7C84FFFB9FA2EFEDDFEA2A436" italics="true" pageId="14" pageNumber="165">A. dybowskii</emphasis>
</taxonomicName>
; see above) more closely match
<taxonomicName id="4C164D36C84FFFB9FDD3FEF9FD56A437" authority="Anistratenko &amp; Neubauer &amp; Anistratenko &amp; Kijashko &amp; Wesselingh, 2021" authorityName="Anistratenko &amp; Neubauer &amp; Anistratenko &amp; Kijashko &amp; Wesselingh" authorityYear="2021" box="[633,755,295,321]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="14" pageNumber="165" phylum="Ciliophora" rank="species" species="cyrensis" status="sp. nov.">
<emphasis id="B962EAA7C84FFFB9FDD3FEF9FD56A437" box="[633,755,295,321]" italics="true" pageId="14" pageNumber="165">A. cyrensis</emphasis>
</taxonomicName>
<emphasis id="B962EAA7C84FFFB9FD51FEF9FCF0A436" bold="true" box="[763,853,295,321]" pageId="14" pageNumber="165">
<taxonomicNameLabel id="A25157DCC84FFFB9FD51FEF9FCF0A436" box="[763,853,295,321]" pageId="14" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in terms of the depressed shell, but we consider the larger size, larger protoconch, more thickened shell and (apparent) lack of teleoconch microsculpture (
<figureCitation id="132D2A30C84FFFB9FAA0FE95FAE9A412" box="[1290,1356,331,357]" captionStart="FIGURE 2" captionStartId="9.[151,250,1721,1745]" captionTargetBox="[324,1262,181,1697]" captionTargetId="figure-17@9.[324,1262,181,1697]" captionTargetPageId="9" captionText="FIGURE 2. Andrusovia dybowskii Brusina in Westerlund, 1903. (a, b, e) Lectotype, locality 26 (CNHM ZMKJ 50). (c, d, f) Paralectotype, locality 26 (CNHM ZMKJ 51). (g, h) Paratype of A. brusinai, locality 22 (MSU Lc-24598). (ik) Specimen from Baer collection, locality 26 (ZB-M W.Dyb. 70). (l, m, q) Specimen from Baer collection, locality 26 (ZB-M W.Dyb. 28). (np) Specimen from Baer collection, locality 26 (ZB-M W.Dyb. 28). (r, v, w) Specimen from Baer collection, locality 26 (ZB-M W.Dyb. 116). (su) Specimen from Baer collection, locality 26 (ZB-M W.Dyb. 70). (x) Specimen from Baer collection, locality 26 (ZB-M W.Dyb. 70). (y) Paratype of H. marina, locality 22 (MSU Lc-24614c). (z, ad) Paratype of H. marina, locality 22 (MSU Lc-24614a). (aa) Paratype of H. marina, locality 22 (MSU Lc-24614b). (ab) Specimen from Baer collection, locality 26 (ZB-M W.Dyb. 70). (ac) Specimen from Baer collection, locality 26 (ZB-M W.Dyb. 28). (ae) Paratype of A. andrusovi, locality 18 (MSU Lc-24601). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4550152" httpUri="https://zenodo.org/record/4550152/files/figure.png" pageId="14" pageNumber="165">Fig. 2</figureCitation>
ae), 3c, h) sufficient to distinguish them on the species level. The Pleistocene
<taxonomicName id="4C164D36C84FFFB9FC0BFEB1FB05A4FF" authority="Anistratenko &amp; Neubauer &amp; Anistratenko &amp; Kijashko &amp; Wesselingh, 2021" authorityName="Anistratenko &amp; Neubauer &amp; Anistratenko &amp; Kijashko &amp; Wesselingh" authorityYear="2021" box="[929,1184,367,393]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="14" pageNumber="165" phylum="Ciliophora" rank="species" species="antecessor" status="sp. nov.">
<emphasis id="B962EAA7C84FFFB9FC0BFEB1FB05A4FF" box="[929,1184,367,393]" italics="true" pageId="14" pageNumber="165">Andrusovia antecessor</emphasis>
</taxonomicName>
<emphasis id="B962EAA7C84FFFB9FB0DFEB1FAA5A4FE" bold="true" box="[1191,1280,367,393]" pageId="14" pageNumber="165">
<taxonomicNameLabel id="A25157DCC84FFFB9FB0DFEB1FAA5A4FE" box="[1191,1280,367,393]" pageId="14" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is even larger and has a higher spire with more whorls.
</paragraph>
</subSubSection>
<subSubSection id="C30C653EC84FFFB9FF6DFE68FF4DA481" pageId="14" pageNumber="165" type="distribution">
<paragraph id="8BA936B5C84FFFB9FF6DFE68FF4DA481" blockId="14.[151,1437,151,502]" pageId="14" pageNumber="165">
<emphasis id="B962EAA7C84FFFB9FF6DFE68FEC4A4A7" bold="true" box="[199,353,438,464]" pageId="14" pageNumber="165">Distribution.</emphasis>
Caspian Sea. So far only known from the late Holocene of the Kura delta deposits (South Caspian Basin).
</paragraph>
</subSubSection>
</treatment>
</document>