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<document id="061ECA77ACF02BF884573AA8B3B39830" ID-DOI="10.5281/zenodo.186683" ID-GBIF-Dataset="f72ec086-e1a2-4abd-b53e-860754f33f3a" ID-ISSN="1175-5326" ID-Zenodo-Dep="186683" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461055588076" checkinUser="plazi" docAuthor="Yao, Gang, Yang, Ding &amp; Evenhuis, Neal L." docDate="2009" docId="03BF878E1125FFFDF7E7FF3AA3840556" docLanguage="en" docName="zt02055p060.pdf" docOrigin="Zootaxa 2055" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Villa aspros" docType="treatment" docVersion="9" lastPageNumber="54" masterDocId="FF86FFF61121FFF8F770FFADA1490036" masterDocTitle="Four new species and a new record of Villa Lioy, 1864 from China (Diptera: Bombyliidae)" masterLastPageNumber="60" masterPageNumber="49" pageNumber="53" updateTime="1698581030920" updateUser="plazi">
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<mods:title id="9EC60B35B84373F5B2B758F4D79F3D12">Four new species and a new record of Villa Lioy, 1864 from China (Diptera: Bombyliidae)</mods:title>
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<mods:namePart id="77174AF2C81CD2A94BFFC2C47F402548">Yao, Gang</mods:namePart>
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<mods:namePart id="32B1DCC0F58B825D3389F7B4514799F1">Yang, Ding</mods:namePart>
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<mods:namePart id="0792D194FABA779D392D792D922979BD">Evenhuis, Neal L.</mods:namePart>
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<mods:date id="63F17E6E2783A40EC9A2A82A36BFF24E">2009</mods:date>
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<treatment id="03BF878E1125FFFDF7E7FF3AA3840556" ID-DOI="http://doi.org/10.5281/zenodo.6222185" ID-GBIF-Taxon="119612953" ID-Zenodo-Dep="6222185" LSID="urn:lsid:plazi:treatment:03BF878E1125FFFDF7E7FF3AA3840556" httpUri="http://treatment.plazi.org/id/03BF878E1125FFFDF7E7FF3AA3840556" lastPageId="5" lastPageNumber="54" pageId="4" pageNumber="53">
<subSubSection id="C30C65131125FFFCF7E7FF3AA01B00EE" pageId="4" pageNumber="53" type="nomenclature">
<paragraph id="8BA936981125FFFCF7E7FF3AA0C30087" blockId="4.[151,394,151,216]" box="[151,394,151,177]" pageId="4" pageNumber="53">
<heading id="D0E181F41125FFFCF7E7FF3AA0C30087" bold="true" box="[151,394,151,177]" fontSize="11" level="1" pageId="4" pageNumber="53" reason="1">
<emphasis id="B962EA8A1125FFFCF7E7FF3AA0C30087" bold="true" box="[151,394,151,177]" pageId="4" pageNumber="53">
<emphasis id="B962EA8A1125FFFCF7E7FF3AA07E0087" bold="true" box="[151,311,151,177]" italics="true" pageId="4" pageNumber="53">
<taxonomicName id="4C164D1B1125FFFCF7E7FF3AA06C0087" ID-CoL="8FLKM" box="[151,293,151,177]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="aspros">Villa aspros</taxonomicName>
s
</emphasis>
p. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA936981125FFFCF7E7FF13A01B00EE" blockId="4.[151,394,151,216]" box="[151,338,190,216]" pageId="4" pageNumber="53">
(
<figureCitation id="132D2A1D1125FFFCF7EFFF13A1BA00EE" box="[159,243,190,216]" captionStart="FIGURES 1 5" captionStartId="1.[151,269,1973,1997]" captionTargetBox="[174,1389,1223,1950]" captionTargetId="figure@1.[165,1409,1212,1968]" captionTargetPageId="1" captionText="FIGURES 1 5. Villa wings. 1. Villa aquila sp. nov.; 2. Villa aspros sp. nov.; 3. Villa cingulata Meigen; 4. Villa flavidus sp. nov.; 5. Villa obtusus sp. nov.." httpUri="https://zenodo.org/record/186684/files/figure.png" pageId="4" pageNumber="53">Figs. 2</figureCitation>
,
<figureCitation id="132D2A1D1125FFFCF78FFF13A00000EE" box="[255,329,190,216]" captionStart="FIGURES 12 17" captionStartId="4.[151,273,1615,1639]" captionTargetBox="[168,1358,516,1535]" captionTargetId="figure@4.[151,1436,506,1590]" captionTargetPageId="4" captionText="FIGURES 12 17. Villa aspros sp. nov. male genitalia. 12. epandrium and cercus, dorsal view; 13. epandrium and cercus, lateral view; 14. gonocoxa and gonostylus, lateral view; 15. phallus, dorsal view; 16. phallus, lateral view; 17. gonocoxa and gonostylus, ventral view." httpUri="https://zenodo.org/record/186686/files/figure.png" pageId="4" pageNumber="53">1217</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30C65131125FFFCF7E7FEA6A06C01F6" pageId="4" pageNumber="53" type="diagnosis">
<paragraph id="8BA936981125FFFCF7E7FEA6A06C01F6" blockId="4.[151,1436,267,448]" pageId="4" pageNumber="53">
<emphasis id="B962EA8A1125FFFCF7E7FEA6A05E0113" bold="true" box="[151,279,267,293]" pageId="4" pageNumber="53">Diagnosis.</emphasis>
Tergites 2 and 4 with white recumbent scales anteriorly and tergite 3 with white recumbent scales antero-laterally, tergites 56 with white recumbent scales posteriorly. Gonocoxa distinctly narrowing apically in ventral view, with numerous black bristle-like hairs apically; gonostylus with an acute tip and narrowing from base to tip in lateral view; epiphallus subquadrate, narrowing from base to tip, with an irregular tip in dorsal view.
</paragraph>
</subSubSection>
<caption id="DF6966101125FFFCF7E7F9E2A374069D" httpUri="https://zenodo.org/record/186686/files/figure.png" pageId="4" pageNumber="53" targetBox="[168,1358,516,1535]" targetPageId="4">
<paragraph id="8BA936981125FFFCF7E7F9E2A374069D" blockId="4.[151,1436,1615,1707]" pageId="4" pageNumber="53">
<emphasis id="B962EA8A1125FFFCF7E7F9E2A02C0651" bold="true" box="[151,357,1615,1639]" pageId="4" pageNumber="53">FIGURES 1217.</emphasis>
<taxonomicName id="4C164D1B1125FFFCF61DF9E2A0BB0650" box="[365,498,1615,1638]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="aspros" status="sp. nov.">
<emphasis id="B962EA8A1125FFFCF61DF9E2A0BB0650" box="[365,498,1615,1638]" italics="true" pageId="4" pageNumber="53">Villa aspros</emphasis>
</taxonomicName>
<emphasis id="B962EA8A1125FFFCF68BF9E2A31C0651" bold="true" box="[507,597,1615,1639]" pageId="4" pageNumber="53">
<taxonomicNameLabel id="A25157F11125FFFCF68BF9E2A31C0651" box="[507,597,1615,1639]" pageId="4" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male genitalia. 12. epandrium and cercus, dorsal view; 13. epandrium and cercus, lateral view; 14. gonocoxa and gonostylus, lateral view; 15. phallus, dorsal view; 16. phallus, lateral view; 17. gonocoxa and gonostylus, ventral view.
</paragraph>
</caption>
<subSubSection id="C30C65131125FFFDF7B5F974A47403A6" lastPageId="5" lastPageNumber="54" pageId="4" pageNumber="53" type="description">
<paragraph id="8BA936981125FFFCF7B5F974A2DD06C2" blockId="4.[151,1437,1753,2012]" box="[197,916,1753,1780]" pageId="4" pageNumber="53">
<emphasis id="B962EA8A1125FFFCF7B5F974A01406C5" bold="true" box="[197,349,1753,1779]" pageId="4" pageNumber="53">Description.</emphasis>
Male. Body length
<quantity id="4CEE9B7D1125FFFCF536F977A3D506C2" box="[582,668,1754,1780]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="53" unit="mm" value="13.0">13 mm</quantity>
, wing length
<quantity id="4CEE9B7D1125FFFCF44AF977A2D906C2" box="[826,912,1754,1780]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="53" unit="mm" value="12.0">12 mm</quantity>
.
</paragraph>
<paragraph id="8BA936981125FFFCF7B5F8ACA37807EA" blockId="4.[151,1437,1753,2012]" pageId="4" pageNumber="53">Head black with grey pollen; ocellar tubercle reddish brown. Hairs on head black and white; frons with dense black erect hairs and sparse yellow erect scale-like hairs, face with dense black erect hairs; occiput with sparse short black erect hairs and white scales close to eyes, a row of yellow erect hairs on edge. Antenna black; scape short and thick, slightly longer than wide, with dense black hairs; pedicel nearly as long as wide, almost bare; first flagellomere cone-shaped, bare. Antennal ratio: 3:1:6. Proboscis dark brown with brown hairs; palpus dark with black hairs.</paragraph>
<paragraph id="8BA936981124FFFDF7B5FF35A43001D1" blockId="5.[151,1437,152,1376]" pageId="5" pageNumber="54">
Thorax black with grey pollen. Hairs on thorax mostly yellow, bristles yellow and black; postpronotal lobe with long yellow hairs, mesonotum with row of long yellow hairs at anterior margin and three long yellow lateral bristles near base of wing, postalar callus with four yellow bristles; thorax with black scales on back. Scutellum with scales-like yellow hairs on posterior edge and black scales on back. Legs dark brown with black scales. Hairs on legs mostly black, bristles black. Femora with long black hairs; tibiae and tarsi dense short black hairs. Mid femur with four
<emphasis id="B962EA8A1124FFFDF5DAFEF4A38F0144" box="[682,710,345,370]" italics="true" pageId="5" pageNumber="54">av</emphasis>
apically; hind femur with six
<emphasis id="B962EA8A1124FFFDF356FEF4A50B0144" box="[1062,1090,345,370]" italics="true" pageId="5" pageNumber="54">av</emphasis>
apically. Mid tibia with eight
<emphasis id="B962EA8A1124FFFDF7E7FE2DA1FD01AF" box="[151,180,384,409]" italics="true" pageId="5" pageNumber="54">ad</emphasis>
, nine
<emphasis id="B962EA8A1124FFFDF78EFE2DA05201AF" box="[254,283,384,409]" italics="true" pageId="5" pageNumber="54">pd</emphasis>
, seven
<emphasis id="B962EA8A1124FFFDF607FE2DA0DA01AF" box="[375,403,384,409]" italics="true" pageId="5" pageNumber="54">av</emphasis>
and 11
<emphasis id="B962EA8A1124FFFDF682FE2DA34701AF" box="[498,526,384,409]" italics="true" pageId="5" pageNumber="54">pv</emphasis>
; hind tibia with nine
<emphasis id="B962EA8A1124FFFDF47FFE2DA26001AF" box="[783,809,384,409]" italics="true" pageId="5" pageNumber="54">av</emphasis>
, and dense bristle-like hairs all round. Wing (
<figureCitation id="132D2A1D1124FFFDF23BFE2DA4DB01AC" box="[1355,1426,384,410]" captionStart="FIGURES 1 5" captionStartId="1.[151,269,1973,1997]" captionTargetBox="[174,1389,1223,1950]" captionTargetId="figure@1.[165,1409,1212,1968]" captionTargetPageId="1" captionText="FIGURES 1 5. Villa wings. 1. Villa aquila sp. nov.; 2. Villa aspros sp. nov.; 3. Villa cingulata Meigen; 4. Villa flavidus sp. nov.; 5. Villa obtusus sp. nov.." httpUri="https://zenodo.org/record/186684/files/figure.png" pageId="5" pageNumber="54">Fig. 2</figureCitation>
) hyaline entirely, with metalline reflecting. Base of vein C with brush-like long black hairs, tegula almost bare. Wing with basicosta strong; vein r-m nearly 2/5 length of discal cell apically. Halteres brown with tip pale.
</paragraph>
<paragraph id="8BA936981124FFFDF7B5FE59A51702B4" blockId="5.[151,1437,152,1376]" pageId="5" pageNumber="54">Abdomen black. Hairs on abdomen black and white; dorsum with dense black recumbent scales except tergites 2 and 4 with white recumbent scales anteriorly and tergite 3 with white recumbent scales anterolaterally, tergites 56 with white recumbent scales posteriorly. Sternites with sparse long white erect hairs and dense black recumbent scales except sternite 4 with dense white recumbent scales.</paragraph>
<paragraph id="8BA936981124FFFDF7B5FD23A2EF035C" blockId="5.[151,1437,152,1376]" pageId="5" pageNumber="54">
Male genitalia (
<figureCitation id="132D2A1D1124FFFDF6F6FD23A355029E" box="[390,540,654,680]" captionStart="FIGURES 12 17" captionStartId="4.[151,273,1615,1639]" captionTargetBox="[168,1358,516,1535]" captionTargetId="figure@4.[151,1436,506,1590]" captionTargetPageId="4" captionText="FIGURES 12 17. Villa aspros sp. nov. male genitalia. 12. epandrium and cercus, dorsal view; 13. epandrium and cercus, lateral view; 14. gonocoxa and gonostylus, lateral view; 15. phallus, dorsal view; 16. phallus, lateral view; 17. gonocoxa and gonostylus, ventral view." httpUri="https://zenodo.org/record/186686/files/figure.png" pageId="5" pageNumber="54">Figs. 1217</figureCitation>
). Epandrium subquadrate, nearly twice longer than high, with dense long black bristles apically, cercus well exposed in lateral view, epandrium about as long as wide, slightly narrowed from base to tip in dorsal view; gonocoxa with numerous black bristle-like hairs apically, distinctly narrowing apically in ventral view; gonostylus subquadrate, its tip acute and narrowed from base to tip in lateral view; epiphallus subquadrate, narrowing from base to tip, with an irregular tip in dorsal view, distiphallus rather narrow, long and curved apically in lateral view.
</paragraph>
<paragraph id="8BA936981124FFFDF7B5FCDBA47403A6" blockId="5.[151,1437,152,1376]" box="[197,1341,886,912]" pageId="5" pageNumber="54">
Female. Body length
<quantity id="4CEE9B7D1124FFFDF6B1FCDBA35E03A6" box="[449,535,886,912]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="54" unit="mm" value="14.0">14 mm</quantity>
, wing length
<quantity id="4CEE9B7D1124FFFDF5C5FCDBA24203A6" box="[693,779,886,912]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="54" unit="mm" value="12.0">12 mm</quantity>
. Similar to male, but face with long white hairs.
</paragraph>
</subSubSection>
<subSubSection id="C30C65131124FFFDF7B5FC30A072041D" pageId="5" pageNumber="54" type="materials_examined">
<paragraph id="8BA936981124FFFDF7B5FC30A072041D" blockId="5.[151,1437,152,1376]" pageId="5" pageNumber="54">
<emphasis id="B962EA8A1124FFFDF7B5FC30A0C90381" bold="true" box="[197,384,925,951]" pageId="5" pageNumber="54">
<typeStatus id="54AD883A1124FFFDF7B5FC30A04B0381" box="[197,258,925,951]" pageId="5" pageNumber="54">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54AD883A1124FFFDF6FBFC30A0B50381" box="[395,508,925,951]" pageId="5" pageNumber="54" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="F30176081124FFFDF53FFC30A3FA0381" box="[591,691,925,951]" name="China" pageId="5" pageNumber="54">CHINA</collectingCountry>
: Henan, Jiyuan Wangwushan (
<geoCoordinate id="EE22505F1124FFFDF35AFC30A5E20381" box="[1066,1195,925,951]" direction="north" orientation="latitude" pageId="5" pageNumber="54" precision="925" value="112.166664">N 112°10</geoCoordinate>
<geoCoordinate id="EE22505F1124FFFDF3C3FC30A46A0381" box="[1203,1315,925,951]" direction="east" orientation="longitude" pageId="5" pageNumber="54" precision="925" value="35.266666">E 35°16</geoCoordinate>
),
<date id="FFA810581124FFFDF24BFC30A19D03E8" pageId="5" pageNumber="54" value="2007-07-27">27. VII. 2007</date>
, Juncao Wang.
<typeStatus id="54AD883A1124FFFDF6FCFC69A0B703E8" box="[396,510,964,990]" pageId="5" pageNumber="54" type="paratype">Paratypes</typeStatus>
<specimenCount id="9D10FD111124FFFDF576FC69A31103E8" box="[518,600,964,990]" pageId="5" pageNumber="54" type="male">1 male</specimenCount>
,
<collectingCountry id="F30176081124FFFDF514FC69A38C03E8" box="[612,709,964,990]" name="China" pageId="5" pageNumber="54">CHINA</collectingCountry>
: Beijing, Mentougou Longmenjian (
<geoCoordinate id="EE22505F1124FFFDF301FC69A5A403E8" box="[1137,1261,964,990]" direction="north" orientation="latitude" pageId="5" pageNumber="54" precision="925" value="114.666664">N 114°40</geoCoordinate>
<geoCoordinate id="EE22505F1124FFFDF385FC69A42803E8" box="[1269,1377,964,990]" direction="east" orientation="longitude" pageId="5" pageNumber="54" precision="925" value="38.783333">E 38°47</geoCoordinate>
),
<date id="FFA810581124FFFDF207FC69A05E0432" pageId="5" pageNumber="54" value="2007-08-15">15. VIII. 2007</date>
, Gang Yao;
<specimenCount id="9D10FD111124FFFDF6D9FC47A35A0432" box="[425,531,1002,1028]" pageId="5" pageNumber="54" type="female">1 female</specimenCount>
,
<collectingCountry id="F30176081124FFFDF550FC47A3CB0432" box="[544,642,1002,1028]" name="China" pageId="5" pageNumber="54">CHINA</collectingCountry>
: Henan, Jiyuan Wangwushan (
<geoCoordinate id="EE22505F1124FFFDF49CFC47A5200432" box="[1004,1129,1002,1028]" direction="north" orientation="latitude" pageId="5" pageNumber="54" precision="925" value="112.166664">N 112°10</geoCoordinate>
<geoCoordinate id="EE22505F1124FFFDF301FC47A5940432" box="[1137,1245,1002,1028]" direction="east" orientation="longitude" pageId="5" pageNumber="54" precision="925" value="35.266666">E 35°16</geoCoordinate>
),
<date id="FFA810581124FFFDF384FC47A4D10432" box="[1268,1432,1002,1028]" pageId="5" pageNumber="54" value="2007-07-28">28. VII. 2007</date>
, Juncao Wang.
</paragraph>
</subSubSection>
<subSubSection id="C30C65131124FFFDF7B5FB9AA3CE0464" box="[197,647,1079,1106]" pageId="5" pageNumber="54" type="distribution">
<paragraph id="8BA936981124FFFDF7B5FB9AA3CE0464" blockId="5.[151,1437,152,1376]" box="[197,647,1079,1106]" pageId="5" pageNumber="54">
<emphasis id="B962EA8A1124FFFDF7B5FB9AA02C0467" bold="true" box="[197,357,1079,1105]" pageId="5" pageNumber="54">Distribution.</emphasis>
<collectingCountry id="F30176081124FFFDF61CFB95A0FB0464" box="[364,434,1080,1106]" name="China" pageId="5" pageNumber="54">China</collectingCountry>
(Beijing, Henan).
</paragraph>
</subSubSection>
<subSubSection id="C30C65131124FFFDF7B5FBF3A0EE04A9" pageId="5" pageNumber="54" type="etymology">
<paragraph id="8BA936981124FFFDF7B5FBF3A0EE04A9" blockId="5.[151,1437,152,1376]" pageId="5" pageNumber="54">
<emphasis id="B962EA8A1124FFFDF7B5FBF3A01B044E" bold="true" box="[197,338,1118,1144]" pageId="5" pageNumber="54">Etymology.</emphasis>
The species epithet derives from the Latin “
<taxonomicName id="4C164D1B1124FFFDF42CFBF3A2E5044E" box="[860,940,1118,1144]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="aspros">aspros</taxonomicName>
” means “white diaper”; referring to white scales on the abdomen.
</paragraph>
</subSubSection>
<subSubSection id="C30C65131124FFFDF7B5FB06A3840556" pageId="5" pageNumber="54" type="discussion">
<paragraph id="8BA936981124FFFDF7B5FB06A3840556" blockId="5.[151,1437,152,1376]" pageId="5" pageNumber="54">
<emphasis id="B962EA8A1124FFFDF7B5FB06A07704F3" bold="true" box="[197,318,1195,1221]" pageId="5" pageNumber="54">Remarks.</emphasis>
The new species is similar to
<taxonomicName id="4C164D1B1124FFFDF5ECFB01A21604F0" authority="Rossi" authorityName="Rossi" box="[668,863,1196,1222]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="panisca">
<emphasis id="B962EA8A1124FFFDF5ECFB01A25D04F3" box="[668,788,1196,1221]" italics="true" pageId="5" pageNumber="54">V. panisca</emphasis>
Rossi
</taxonomicName>
, but it can be separated from the latter by tergites 2 and 4 with white recumbent scales anteriorly, tergite 3 with white recumbent scales antero-laterally, tergites 56 with white recumbent scales posteriorly; sternite 4 with the dense white recumbent scales. In
<taxonomicName id="4C164D1B1124FFFDF26CFB54A4DD0524" box="[1308,1428,1273,1298]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="panisca">
<emphasis id="B962EA8A1124FFFDF26CFB54A4DD0524" box="[1308,1428,1273,1298]" italics="true" pageId="5" pageNumber="54">V. panisca</emphasis>
</taxonomicName>
, tergites 3 and 4 have yellow hairs laterally, and tergites 56 have black hair laterally; the sternites have the black erect hairs and yellowish recumbent hairs.
</paragraph>
</subSubSection>
</treatment>
</document>