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<mods:title id="A6488A4846013A0324F27A5A7DAB9F2C">Redescription and lectotype designation of Chasmogenus abnormalis (Sharp), with notes on its distribution</mods:title>
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<heading id="D0E17A6FF06C4A60C7F7368FBEC1907B" bold="true" box="[151,642,1535,1562]" fontSize="11" level="1" pageId="0" pageNumber="296" reason="1">
<taxonomicName id="4C16B680F06C4A60C7F7368FBEC1907B" authority="Sharp, 1890" authorityName="Sharp" authorityYear="1890" box="[151,642,1535,1562]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="296" phylum="Arthropoda" rank="species" species="abnormalis">
<emphasis id="B9621111F06C4A60C7F7368FBEC1907B" bold="true" box="[151,642,1535,1562]" pageId="0" pageNumber="296">
<emphasis id="B9621111F06C4A60C7F7368FBD919078" bold="true" box="[151,466,1535,1561]" italics="true" pageId="0" pageNumber="296">Chasmogenus abnormalis</emphasis>
(
<bibRefCitation id="EF87B0F2F06C4A60C6813570BE39907B" author="Sharp" box="[481,634,1536,1562]" pageId="0" pageNumber="296" refString="Sharp, D. (1890) On some aquatic Coleoptera from Ceylon. Transactions of the Entomological Society of London, 1890, 339 - 359. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1890. tb 03026. x" type="journal article" year="1890">Sharp, 1890</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA9CD03F06C4A60C7F73552BD509058" blockId="0.[151,1437,1535,1831]" box="[151,275,1570,1593]" pageId="0" pageNumber="296">
(
<figureCitation id="132DD186F06C4A60C7FE3552BD499058" box="[158,266,1570,1593]" captionStart-0="FIGURES 1 6" captionStart-1="FIGURES 7 13. 7 8" captionStartId-0="1.[151,258,1622,1644]" captionStartId-1="2.[151,258,1886,1908]" captionTargetBox-0="[150,1429,213,1598]" captionTargetBox-1="[160,1435,365,1858]" captionTargetId-0="figure@1.[150,1436,194,1599]" captionTargetId-1="figure@2.[151,1436,365,1865]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 1 6. Lectotype of Philydrus abnormalis Sharp, 1890 (= Chasmogenus abnormalis). 1: general habitus, dorsal view; 2: detail of head and the shape of clypeus; 3 5: aedeagus (3: ventral, 4: lateral, 5: dorsal); 6: labels of the lectotype." captionText-1="FIGURES 7 13. 7 8: habitus of alive specimen of Chasmogenus abnormalis (Sharp) from Manipur (7: dorsal, 8: ventral); 9: known distribution of C. abnormalis (green: localities of the examined specimens, blue: literature data, T: type locality). 10 13: Morphology of the aedeagus and detail of apex of parameres of examined specimens (10: lectotype; 11: specimen from India: Manipur; 12: specimen from Laos: Vientiane; 13: specimen from Japan: Ryukyu Islands; 10 12: ventral, 13 dorsal)." httpUri-0="https://zenodo.org/record/261466/files/figure.png" httpUri-1="https://zenodo.org/record/261467/files/figure.png" pageId="0" pageNumber="296">Figs 113</figureCitation>
)
</paragraph>
</subSubSection>
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<materialsCitation id="3B7EC75EF06C4A60C7F73535BF31903B" ID-GBIF-Occurrence="1288103071" box="[151,882,1605,1626]" country="Sri Lanka" location="Sri Lanka" pageId="0" pageNumber="296" specimenCount="1" typeStatus="holotype">
<treatmentCitationGroup id="AB06EA2DF06C4A60C7F73535BF31903B" box="[151,882,1605,1626]" pageId="0" pageNumber="296">
<taxonomicName id="4C16B680F06C4A60C7F73535BE56903B" ID-CoL="4GDQW" authority="Sharp, 1890: 351" authorityName="Sharp" authorityPageNumber="351" authorityYear="1890" box="[151,533,1605,1626]" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="296" phylum="Arthropoda" rank="species" species="abnormalis">
<emphasis id="B9621111F06C4A60C7F73535BD24903B" box="[151,359,1605,1626]" italics="true" pageId="0" pageNumber="296">Philydrus abnormalis</emphasis>
<treatmentCitation id="0AB7EB12F06C4A60C60D3535BE56903B" author="Sharp" box="[365,533,1605,1626]" page="351" pageId="0" pageNumber="296" year="1890">
<bibRefCitation id="EF87B0F2F06C4A60C60D3535BDA6903B" author="Sharp" box="[365,485,1605,1626]" pageId="0" pageNumber="296" refString="Sharp, D. (1890) On some aquatic Coleoptera from Ceylon. Transactions of the Entomological Society of London, 1890, 339 - 359. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1890. tb 03026. x" type="journal article" year="1890">Sharp, 1890</bibRefCitation>
: 351
</treatmentCitation>
</taxonomicName>
.
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locality:
<collectingCountry id="F3018D93F06C4A60C5C83535BF44903B" box="[680,775,1605,1626]" name="Sri Lanka" pageId="0" pageNumber="296">Sri Lanka</collectingCountry>
, Colombo.
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</materialsCitation>
</paragraph>
<paragraph id="8BA9CD03F06C4A60C7F73512BD8C90F5" blockId="0.[151,1437,1535,1831]" pageId="0" pageNumber="296">
<materialsCitation id="3B7EC75EF06C4A60C7F73512BD8C90F5" ID-GBIF-Occurrence="1288103064" country="Vietnam" county="Cochinchine" location="Synonymized" municipality="My Tho" pageId="0" pageNumber="296" specimenCount="1" stateProvince="Sumatra" typeStatus="holotype">
<emphasis id="B9621111F06C4A60C7F73512BD2E9016" box="[151,365,1634,1655]" italics="true" pageId="0" pageNumber="296">Phylhydrus ferrugatus</emphasis>
<bibRefCitation id="EF87B0F2F06C4A60C6133512BE559016" author="Regimbart" box="[371,534,1634,1655]" pageId="0" pageNumber="296" refString="Regimbart, M. (1903) Contribution a la faune Indo-Chinoise. 19 e memoire. Annales de la Societe Entomologique de France, 72, 52 - 64." type="journal article" year="1903">Régimbart, 1903</bibRefCitation>
: 57.
<typeStatus id="54AD73A1F06C4A60C5223512BE329016" box="[578,625,1634,1655]" pageId="0" pageNumber="296">Type</typeStatus>
localities:
<collectingCounty id="62C8B58FF06C4A60C5BA3512BF109016" box="[730,851,1634,1655]" pageId="0" pageNumber="296">Cochinchine</collectingCounty>
[=
<collectingCountry id="F3018D93F06C4A60C4143512BF899016" box="[884,970,1634,1655]" name="Vietnam" pageId="0" pageNumber="296">Vietnam</collectingCountry>
],
<collectingMunicipality id="6BCD5779F06C4A60C4B83512B8659016" box="[984,1062,1634,1655]" pageId="0" pageNumber="296">My Tho</collectingMunicipality>
and
<collectingCountry id="F3018D93F06C4A60C3333512B8F79016" box="[1107,1204,1634,1655]" name="Indonesia" pageId="0" pageNumber="296">Indonesia</collectingCountry>
:
<collectingRegion id="49D203E1F06C4A60C3DC3512B94C9016" box="[1212,1295,1634,1655]" country="Indonesia" name="Sumatera Utara" pageId="0" pageNumber="296">Sumatra</collectingRegion>
.
<location id="8EC99BD8F06C4A60C2773512B9D89016" LSID="urn:lsid:plazi:treatment:03BF7C15F06C4A64C7F7368FB9CD94F6:8EC99BD8F06C4A60C2773512B9D89016" box="[1303,1435,1634,1655]" country="Vietnam" county="Cochinchine" municipality="My Tho" name="Synonymized" pageId="0" pageNumber="296" stateProvince="Sumatra">Synonymized</location>
by
<bibRefCitation id="EF87B0F2F06C4A60C785350FBD8890F5" author="Orchymont" box="[229,459,1663,1684]" pageId="0" pageNumber="296" refString="Orchymont, A. dʼ (1939) Revision des especes du sous-genre Crephelochares dʼ Helochares. Bulletin et Annales de la Societe Entomologique de Belgique, 79, 154 - 166." type="journal article" year="1939">
<location id="8EC99BD8F06C4A60C785350FBD1190F5" LSID="urn:lsid:plazi:treatment:03BF7C15F06C4A64C7F7368FB9CD94F6:8EC99BD8F06C4A60C785350FBD1190F5" box="[229,338,1663,1684]" country="Vietnam" county="Cochinchine" municipality="My Tho" name="Orchymont" pageId="0" pageNumber="296" stateProvince="Sumatra">Orchymont</location>
(1939: 159)
</bibRefCitation>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA9CD03F06C4A60C7F735EDBE1A90AE" blockId="0.[151,1437,1535,1831]" pageId="0" pageNumber="296">
<treatmentCitationGroup id="AB06EA2DF06C4A60C7F735EDBE1A90AE" pageId="0" pageNumber="296">
<materialsCitation id="3B7EC75EF06C4A60C7F735EDBF9090D3" ID-GBIF-Occurrence="1288103073" box="[151,979,1693,1714]" country="Vietnam" location="My Tho" pageId="0" pageNumber="296" specimenCount="1" stateProvince="Saigon" typeStatus="holotype">
<taxonomicName id="4C16B680F06C4A60C7F735EDBE6B90D3" authority="Regimbart 1903: 57" authorityName="Regimbart" authorityPageNumber="57" authorityYear="1903" box="[151,552,1693,1714]" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="296" phylum="Arthropoda" rank="species" species="nigritulus">
<emphasis id="B9621111F06C4A60C7F735EDBD1A90D3" box="[151,345,1693,1714]" italics="true" pageId="0" pageNumber="296">Philydrus nigritulus</emphasis>
<treatmentCitation id="0AB7EB12F06C4A60C60135EDBE6B90D3" author="Regimbart" box="[353,552,1693,1714]" page="57" pageId="0" pageNumber="296" year="1903">
<bibRefCitation id="EF87B0F2F06C4A60C60135EDBE4190D3" author="Regimbart" box="[353,514,1693,1714]" pageId="0" pageNumber="296" refString="Regimbart, M. (1903) Contribution a la faune Indo-Chinoise. 19 e memoire. Annales de la Societe Entomologique de France, 72, 52 - 64." type="journal article" year="1903">Régimbart 1903</bibRefCitation>
: 57
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54AD73A1F06C4A60C55335EDBE2090D3" box="[563,611,1693,1714]" pageId="0" pageNumber="296">Type</typeStatus>
localities:
<collectingCountry id="F3018D93F06C4A60C5B035EDBF6690D3" box="[720,805,1693,1714]" name="Vietnam" pageId="0" pageNumber="296">Vietnam</collectingCountry>
:
<collectingRegion id="49D203E1F06C4A60C45035EDBF3690D3" box="[816,885,1693,1714]" country="Vietnam" name="Ho Chi Minh City" pageId="0" pageNumber="296">Saigon</collectingRegion>
,
<location id="8EC99BD8F06C4A60C4E035EDBF9090D3" LSID="urn:lsid:plazi:treatment:03BF7C15F06C4A64C7F7368FB9CD94F6:8EC99BD8F06C4A60C4E035EDBF9090D3" box="[896,979,1693,1714]" country="Vietnam" name="My Tho" pageId="0" pageNumber="296" stateProvince="Saigon">My Tho</location>
</materialsCitation>
; Cambodia, Phnom Penh;
<materialsCitation id="3B7EC75EF06C4A60C3BB35EDBE1A90AE" ID-GBIF-Occurrence="1288103068" country="Indonesia" location="Sumatra" pageId="0" pageNumber="296" specimenCount="1" stateProvince="Sumatra" typeStatus="holotype">
<collectingCountry id="F3018D93F06C4A60C3BB35EDB97890D3" box="[1243,1339,1693,1714]" name="Indonesia" pageId="0" pageNumber="296">Indonesia</collectingCountry>
,
<collectingRegion id="49D203E1F06C4A60C22635EDB9DA90D3" box="[1350,1433,1693,1714]" country="Indonesia" name="Sumatera Utara" pageId="0" pageNumber="296">Sumatra</collectingRegion>
. Synonymized by
<treatmentCitation id="0AB7EB12F06C4A60C60F35CABE1790AE" author="Orchymont" box="[367,596,1722,1743]" page="159" pageId="0" pageNumber="296" year="1939">
<bibRefCitation id="EF87B0F2F06C4A60C60F35CABE1790AE" author="Orchymont" box="[367,596,1722,1743]" pageId="0" pageNumber="296" refString="Orchymont, A. dʼ (1939) Revision des especes du sous-genre Crephelochares dʼ Helochares. Bulletin et Annales de la Societe Entomologique de Belgique, 79, 154 - 166." type="journal article" year="1939">Orchymont (1939: 159)</bibRefCitation>
</treatmentCitation>
.
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA9CD03F06C4A60C7F735A7BFF2908D" blockId="0.[151,1437,1535,1831]" box="[151,945,1751,1772]" pageId="0" pageNumber="296">
<taxonomicName id="4C16B680F06C4A60C7F735A7BE74908D" authority="Sharp" authorityName="Sharp" box="[151,567,1751,1772]" class="Insecta" family="Hydrophilidae" genus="Enochrus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="296" phylum="Arthropoda" rank="species" species="abnormicollis" subGenus="Lumetus">
<emphasis id="B9621111F06C4A60C7F735A7BCB0908D" box="[151,243,1751,1772]" italics="true" pageId="0" pageNumber="296">Enochrus</emphasis>
(
<emphasis id="B9621111F06C4A60C66135A7BD11908D" box="[257,338,1751,1772]" italics="true" pageId="0" pageNumber="296">Lumetus</emphasis>
)
<emphasis id="B9621111F06C4A60C60035A7BDA6908D" box="[352,485,1751,1772]" italics="true" pageId="0" pageNumber="296">abnormicollis</emphasis>
(Sharp)
</taxonomicName>
:
<bibRefCitation id="EF87B0F2F06C4A60C55F35A7BFEE908D" author="Zaitzev" box="[575,941,1751,1772]" pageId="0" pageNumber="296" refString="Zaitzev, F. A. (1908) Catalogue des Coleopteres aquatiques des familles Dryopidae, Georyssidae, Cyathoceridae, Heteroceridae et Hydrophilidae. Horae Societatis Entomologicae Rossicae, 38, 283 - 420." type="journal article" year="1908">Zaitzev (1908: 385, misspelled name)</bibRefCitation>
.
</paragraph>
<paragraph id="8BA9CD03F06C4A60C7F73585BDF29146" blockId="0.[151,1437,1535,1831]" pageId="0" pageNumber="296">
<treatmentCitationGroup id="AB06EA2DF06C4A60C7F73585BDF29146" pageId="0" pageNumber="296">
<taxonomicName id="4C16B680F06C4A60C7F73585BE89916B" authority="Knisch, 1924: 195" authorityName="Knisch" authorityPageNumber="195" authorityYear="1924" box="[151,714,1781,1802]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="296" phylum="Arthropoda" rank="species" species="regimbarti" subGenus="Chasmogenus">
<emphasis id="B9621111F06C4A60C7F73585BD46916B" box="[151,261,1781,1802]" italics="true" pageId="0" pageNumber="296">Helochares</emphasis>
(
<emphasis id="B9621111F06C4A60C6743585BDD9916B" box="[276,410,1781,1802]" italics="true" pageId="0" pageNumber="296">Chasmogenus</emphasis>
)
<emphasis id="B9621111F06C4A60C6CA3585BE4C916B" box="[426,527,1781,1802]" italics="true" pageId="0" pageNumber="296">regimbarti</emphasis>
<treatmentCitation id="0AB7EB12F06C4A60C5773585BE89916B" author="Knisch" box="[535,714,1781,1802]" page="195" pageId="0" pageNumber="296" year="1924">
<bibRefCitation id="EF87B0F2F06C4A60C5773585BEDF916B" author="Knisch" box="[535,668,1781,1802]" pageId="0" pageNumber="296" refString="Knisch, A. (1924) Hydrophilidae. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Vol. 14, part 79. Junk, W., Berlin, 1 - 306." type="journal article" year="1924">Knisch, 1924</bibRefCitation>
: 195
</treatmentCitation>
</taxonomicName>
(replacement name for
<taxonomicName id="4C16B680F06C4A60C4D53585B8AB916B" authority="Regimbart" authorityName="Regimbart" box="[949,1256,1781,1802]" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="296" phylum="Arthropoda" rank="species" species="nigritulus">
<emphasis id="B9621111F06C4A60C4D53585B835916B" box="[949,1142,1781,1802]" italics="true" pageId="0" pageNumber="296">Philydrus nigritulus</emphasis>
Régimbart
</taxonomicName>
). Synonymized by
<treatmentCitation id="0AB7EB12F06C4A60C7A73462BDEE9146" author="Orchymont" box="[199,429,1810,1831]" page="159" pageId="0" pageNumber="296" year="1939">
<bibRefCitation id="EF87B0F2F06C4A60C7A73462BDEE9146" author="Orchymont" box="[199,429,1810,1831]" pageId="0" pageNumber="296" refString="Orchymont, A. dʼ (1939) Revision des especes du sous-genre Crephelochares dʼ Helochares. Bulletin et Annales de la Societe Entomologique de Belgique, 79, 154 - 166." type="journal article" year="1939">Orchymont (1939: 159)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30C9E88F06C4A62C7F73422B95A9758" lastPageId="2" lastPageNumber="298" pageId="0" pageNumber="296" type="materials_examined">
<paragraph id="8BA9CD03F06C4A60C7F73422BDE89195" blockId="0.[151,1436,1874,2036]" pageId="0" pageNumber="296">
<materialsCitation id="3B7EC75EF06C4A60C7F73422BD7C91ED" ID-GBIF-Occurrence="1288103065" collectingDate="82-04-17" collectionCode="BMNH" collectorName="D. Sharp" country="Sri Lanka" location="Ceylon" pageId="0" pageNumber="296" specimenCount="2" typeStatus="lectotype">
<emphasis id="B9621111F06C4A60C7F73422BE6C9108" bold="true" box="[151,559,1874,1897]" pageId="0" pageNumber="296">
<typeStatus id="54AD73A1F06C4A60C7F73422BC8C9108" box="[151,207,1874,1897]" pageId="0" pageNumber="296">Type</typeStatus>
material examined.
<typeStatus id="54AD73A1F06C4A60C6DF3422BE6C9108" box="[447,559,1874,1897]" pageId="0" pageNumber="296" type="lectotype">Lectotype</typeStatus>
</emphasis>
(hereby designated,
<figureCitation id="132DD186F06C4A60C4763422BF3B9108" box="[790,888,1874,1897]" captionStart="FIGURES 1 6" captionStartId="1.[151,258,1622,1644]" captionTargetBox="[150,1429,213,1598]" captionTargetId="figure@1.[150,1436,194,1599]" captionTargetPageId="1" captionText="FIGURES 1 6. Lectotype of Philydrus abnormalis Sharp, 1890 (= Chasmogenus abnormalis). 1: general habitus, dorsal view; 2: detail of head and the shape of clypeus; 3 5: aedeagus (3: ventral, 4: lateral, 5: dorsal); 6: labels of the lectotype." httpUri="https://zenodo.org/record/261466/files/figure.png" pageId="0" pageNumber="296">Figs 16</figureCitation>
): male (
<collectionCode id="ED0755C6F06C4A60C4B43422B8689108" box="[980,1067,1874,1897]" pageId="0" pageNumber="296">BMNH</collectionCode>
): “
<taxonomicName id="4C16B680F06C4A60C3283422B8F39108" box="[1096,1200,1874,1897]" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="297" phylum="Arthropoda" rank="genus">Philydrus</taxonomicName>
/
<taxonomicName id="4C16B680F06C4A60C3AB3422B9019108" box="[1227,1346,1874,1897]" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="296" phylum="Arthropoda" rank="species" species="abnormalis">abnormalis</taxonomicName>
/
<collectorName id="26E3A8D5F06C4A60C23B3422B9C89108" box="[1371,1419,1874,1897]" name="D. Sharp" pageId="0" pageNumber="296">D.S.</collectorName>
/
<collectingCountry id="F3018D93F06C4A60C7F73405BCA791ED" box="[151,228,1909,1932]" name="Sri Lanka" pageId="0" pageNumber="296">Ceylon</collectingCountry>
<date id="FFA8EBC3F06C4A60C78C3405BD7C91ED" box="[236,319,1909,1932]" pageId="0" pageNumber="296" value="82-04-17">
<collectingDate id="EFEC122BF06C4A60C78C3405BD7C91ED" box="[236,319,1909,1932]" pageId="0" pageNumber="296" value="82-04-17">17.4.82</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B7EC75EF06C4A60C62A3405BCA991B0" ID-GBIF-Occurrence="1288103072" collectingDate="1982-04-07" collectingDateMax="1982-04-27" collectingDateMin="1982-04-07" collectorName="G. Lewis" country="Sri Lanka" location="Ceylon" pageId="0" pageNumber="296" specimenCount="1" typeStatus="holotype">
/ Lewis // [male symbol] // Colombo / On coast level /
<date id="FFA8EBC3F06C4A60C4C13405B85991ED" box="[929,1050,1909,1932]" pageId="0" pageNumber="296" value="1982-04-07" valueMax="1982-04-27" valueMin="1982-04-07">
<collectingDate id="EFEC122BF06C4A60C4C13405B85991ED" box="[929,1050,1909,1932]" pageId="0" pageNumber="296" value="1982-04-07" valueMax="1982-04-27" valueMin="1982-04-07">727.IV.82</collectingDate>
</date>
, //
<collectingCountry id="F3018D93F06C4A60C35C3405B8CA91ED" box="[1084,1161,1909,1932]" name="Sri Lanka" pageId="0" pageNumber="296">Ceylon</collectingCountry>
/
<collectorName id="26E3A8D5F06C4A60C3C13405B94391ED" box="[1185,1280,1909,1932]" pageId="0" pageNumber="296">G. Lewis</collectorName>
/ 1910-320. // Co- /
<typeStatus id="54AD73A1F06C4A60C7BC34E8BD4991CE" box="[220,266,1944,1967]" pageId="0" pageNumber="296">type</typeStatus>
// A. dʼOrchymont vid. 1924 // 1936 / A. dʼOrchymont det. /
<taxonomicName id="4C16B680F06C4A60C4AE34E8B94B91CE" authority="Crephelochares" authorityName="Crephelochares" box="[974,1288,1944,1967]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="296" phylum="Arthropoda" rank="genus">Helochares (Crephelochares)</taxonomicName>
/
<taxonomicName id="4C16B680F06C4A60C24534E8BCA991B0" authority="Sharp" authorityName="Sharp" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="296" phylum="Arthropoda" rank="species" species="abnormalis">abnormalis (Sharp)</taxonomicName>
</materialsCitation>
.
<materialsCitation id="3B7EC75EF06C4A60C79434CABDE89195" ID-GBIF-Occurrence="1288103074" collectingDate="82-04-17" collectionCode="BMNH" collectorName="Type D. S." country="Sri Lanka" location="Ceylon" pageId="0" pageNumber="296" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<emphasis id="B9621111F06C4A60C79434CABDD991B0" bold="true" box="[244,410,1978,2001]" pageId="0" pageNumber="296">
<typeStatus id="54AD73A1F06C4A60C79434CABDD691B0" box="[244,405,1978,2001]" pageId="0" pageNumber="296" type="paralectotype">Paralectotype</typeStatus>
:
</emphasis>
<specimenCount id="9D10068AF06C4A60C6C034CABDBE91B0" box="[416,509,1978,2001]" pageId="0" pageNumber="296" type="female">1 female</specimenCount>
(
<collectionCode id="ED0755C6F06C4A60C56B34CABE2191B0" box="[523,610,1978,2001]" pageId="0" pageNumber="296">BMNH</collectionCode>
):
<taxonomicName id="4C16B680F06C4A60C51334CABE9A91B0" box="[627,729,1978,2001]" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="297" phylum="Arthropoda" rank="genus">Philydrus</taxonomicName>
/
<taxonomicName id="4C16B680F06C4A60C58E34CABF2791B0" box="[750,868,1978,2001]" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="296" phylum="Arthropoda" rank="species" species="abnormalis">abnormalis</taxonomicName>
/
<collectorName id="26E3A8D5F06C4A60C41934CABFA791B0" box="[889,996,1978,2001]" pageId="0" pageNumber="296">
<typeStatus id="54AD73A1F06C4A60C41934CABFED91B0" box="[889,942,1978,2001]" pageId="0" pageNumber="296">Type</typeStatus>
D.S.
</collectorName>
/
<collectingCountry id="F3018D93F06C4A60C49934CAB80591B0" box="[1017,1094,1978,2001]" name="Sri Lanka" pageId="0" pageNumber="296">Ceylon</collectingCountry>
<date id="FFA8EBC3F06C4A60C32D34CAB8E391B0" box="[1101,1184,1978,2001]" pageId="0" pageNumber="296" value="82-04-17">
<collectingDate id="EFEC122BF06C4A60C32D34CAB8E391B0" box="[1101,1184,1978,2001]" pageId="0" pageNumber="296" value="82-04-17">17.4.82</collectingDate>
</date>
. / Lewis //
<typeStatus id="54AD73A1F06C4A60C27534CAB90991B0" box="[1301,1354,1978,2001]" pageId="0" pageNumber="296">Type</typeStatus>
/ H.T. / / Sharp Coll. / 1905-313.”
</materialsCitation>
</paragraph>
<caption id="DF699D8BF06D4A61C7F73526B89390E8" httpUri="https://zenodo.org/record/261466/files/figure.png" pageId="1" pageNumber="297" targetBox="[150,1429,213,1598]" targetPageId="1">
<paragraph id="8BA9CD03F06D4A61C7F73526B89390E8" blockId="1.[151,1436,1622,1673]" pageId="1" pageNumber="297">
<emphasis id="B9621111F06D4A61C7F73526BD77900D" bold="true" box="[151,308,1622,1644]" pageId="1" pageNumber="297">FIGURES 16.</emphasis>
Lectotype of
<taxonomicName id="4C16B680F06D4A61C6A13527BF51900D" authority="Sharp, 1890" authorityName="Sharp" authorityYear="1890" box="[449,786,1623,1644]" class="Insecta" family="Hydrophilidae" genus="Philydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="297" phylum="Arthropoda" rank="species" species="abnormalis">
<emphasis id="B9621111F06D4A61C6A13527BED0900D" box="[449,659,1623,1644]" italics="true" pageId="1" pageNumber="297">Philydrus abnormalis</emphasis>
Sharp, 1890
</taxonomicName>
(=
<taxonomicName id="4C16B680F06D4A61C4583527B877900D" box="[824,1076,1623,1644]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="297" phylum="Arthropoda" rank="species" species="abnormalis">
<emphasis id="B9621111F06D4A61C4583527B877900D" box="[824,1076,1623,1644]" italics="true" pageId="1" pageNumber="297">Chasmogenus abnormalis</emphasis>
</taxonomicName>
). 1: general habitus, dorsal view; 2: detail of head and the shape of clypeus; 35: aedeagus (3: ventral, 4: lateral, 5: dorsal); 6: labels of the lectotype.
</paragraph>
</caption>
<paragraph id="8BA9CD03F06D4A62C7A735DFB95A9758" blockId="1.[151,1436,1711,2011]" lastBlockId="2.[151,1436,152,313]" lastPageId="2" lastPageNumber="298" pageId="1" pageNumber="297">
<emphasis id="B9621111F06D4A61C7A735DFBE5E90A7" bold="true" box="[199,541,1711,1734]" pageId="1" pageNumber="297">Additional material examined</emphasis>
(17 specimens).
<materialsCitation id="3B7EC75EF06D4A61C5B435DFBF1B914F" ID-GBIF-Occurrence="1288103067" collectingDate="1982-04-07" collectingDateMax="1982-04-27" collectingDateMin="1982-04-07" collectionCode="BMNH" collectorName="Colombo. &amp; On &amp; Ceylon &amp; G. Lewis &amp; Considered &amp; Orchymont" country="Sri Lanka" determinerName="Orchymont" location="SRI LANKA" pageId="1" pageNumber="297" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B9621111F06D4A61C5B435DFBF3290A7" bold="true" box="[724,881,1711,1734]" pageId="1" pageNumber="297">
<collectingCountry id="F3018D93F06D4A61C5B435DFBF2890A7" box="[724,875,1711,1734]" name="Sri Lanka" pageId="1" pageNumber="297">SRI LANKA</collectingCountry>
:
</emphasis>
<specimenCount id="9D10068AF06D4A61C41835DFBF8390A7" box="[888,960,1711,1734]" pageId="1" pageNumber="297" type="male">1 male</specimenCount>
(
<collectionCode id="ED0755C6F06D4A61C4AF35DFB86690A7" box="[975,1061,1711,1734]" pageId="1" pageNumber="297">BMNH</collectionCode>
): “[male symbol] //
<collectorName id="26E3A8D5F06D4A61C39C35DFB92590A7" box="[1276,1382,1711,1734]" pageId="1" pageNumber="297">Colombo.</collectorName>
/
<collectorName id="26E3A8D5F06D4A61C21B35DFB9DF90A7" box="[1403,1436,1711,1734]" pageId="1" pageNumber="297">On</collectorName>
coast level /
<date id="FFA8EBC3F06D4A61C67E35A1BDD49089" box="[286,407,1745,1768]" pageId="1" pageNumber="297" value="1982-04-07" valueMax="1982-04-27" valueMin="1982-04-07">
<collectingDate id="EFEC122BF06D4A61C67E35A1BDD49089" box="[286,407,1745,1768]" pageId="1" pageNumber="297" value="1982-04-07" valueMax="1982-04-27" valueMin="1982-04-07">727.IV.82</collectingDate>
</date>
. //
<collectorName id="26E3A8D5F06D4A61C6D735A1BE479089" box="[439,516,1745,1768]" pageId="1" pageNumber="297">Ceylon</collectorName>
/
<collectorName id="26E3A8D5F06D4A61C57A35A1BE3B9089" box="[538,632,1745,1768]" pageId="1" pageNumber="297">G. Lewis</collectorName>
/ 1910-320. // 1936 / A. d´
<determinerName id="7E7521C7F06D4A61C4F635A1B8519089" box="[918,1042,1745,1768]" pageId="1" pageNumber="297">Orchymont</determinerName>
det. /
<taxonomicName id="4C16B680F06D4A61C33735A1B9CE9089" authority="Crephelochares" authorityName="Crephelochares" box="[1111,1421,1745,1768]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="297" phylum="Arthropoda" rank="genus">Helochares (Crephelochares)</taxonomicName>
/
<taxonomicName id="4C16B680F06D4A61C7F73584BD29916A" authority="Sharp" authorityName="Sharp" box="[151,362,1780,1803]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="297" phylum="Arthropoda" rank="species" species="abnormalis">abnormalis (Sharp)</taxonomicName>
<collectorName id="26E3A8D5F06D4A61C61E3584BDBB916A" box="[382,504,1780,1803]" pageId="1" pageNumber="297">Considered</collectorName>
as part of
<typeStatus id="54AD73A1F06D4A61C5193584BEE4916A" box="[633,679,1780,1803]" pageId="1" pageNumber="297">type</typeStatus>
series by
<bibRefCitation id="EF87B0F2F06D4A61C47E3584BFAF916A" author="Orchymont" box="[798,1004,1780,1803]" pageId="1" pageNumber="297" refString="Orchymont, A. dʼ (1939) Revision des especes du sous-genre Crephelochares dʼ Helochares. Bulletin et Annales de la Societe Entomologique de Belgique, 79, 154 - 166." type="journal article" year="1939">
<collectorName id="26E3A8D5F06D4A61C47E3584BFD4916A" box="[798,919,1780,1803]" pageId="1" pageNumber="297">Orchymont</collectorName>
(1939)
</bibRefCitation>
, but the label data do not corresponds completely with the original description (see below for details)
</materialsCitation>
.
<materialsCitation id="3B7EC75EF06D4A61C4063467BDAC9112" ID-GBIF-Occurrence="1288103069" collectingDate="2013-11-21" collectionCode="IARI" collectorName="Imphal &amp; Phubala &amp; M. Bhubaneshwari &amp; O. Sandhyarani &amp; S. Dineshwar" country="India" elevation="763" location="Loktak Lake" pageId="1" pageNumber="297" specimenCount="2" specimenCount-male="2" stateProvince="Manipur" typeStatus="holotype">
<emphasis id="B9621111F06D4A61C4063467B874914F" bold="true" box="[870,1079,1815,1838]" pageId="1" pageNumber="297">
<collectingCountry id="F3018D93F06D4A61C4063467BFFA914F" box="[870,953,1815,1838]" name="India" pageId="1" pageNumber="297">INDIA</collectingCountry>
:
<collectingRegion id="49D203E1F06D4A61C4A83467B871914F" box="[968,1074,1815,1838]" country="India" name="Manipur" pageId="1" pageNumber="297">Manipur</collectingRegion>
:
</emphasis>
<specimenCount id="9D10068AF06D4A61C3223467B8DB914F" box="[1090,1176,1815,1838]" pageId="1" pageNumber="297" type="male">2 males</specimenCount>
(
<collectionCode id="ED0755C6F06D4A61C3CA3467B8A4914F" box="[1194,1255,1815,1838]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13272" name="Indian Agricultural Research Institute" pageId="1" pageNumber="297">IARI</collectionCode>
):
<quantity id="4CEE60E6F06D4A61C39D3467B907914F" box="[1277,1348,1815,1838]" metricMagnitude="4" metricUnit="m" metricValue="3.9" pageId="1" pageNumber="297" unit="km" value="39.0">39 km</quantity>
S from
<collectorName id="26E3A8D5F06D4A61C7F7344ABCA69130" box="[151,229,1850,1873]" pageId="1" pageNumber="297">Imphal</collectorName>
,
<collectorName id="26E3A8D5F06D4A61C791344ABD099130" box="[241,330,1850,1873]" pageId="1" pageNumber="297">Phubala</collectorName>
,
<location id="8EC99BD8F06D4A61C634344ABDA39130" LSID="urn:lsid:plazi:treatment:03BF7C15F06C4A64C7F7368FB9CD94F6:8EC99BD8F06D4A61C634344ABDA39130" box="[340,480,1850,1873]" country="India" name="Loktak Lake" pageId="1" pageNumber="297" stateProvince="Manipur">Loktak Lake</location>
, wetland, 24°27.327ʹN 93º51.295ʹE,
<elevation id="003B2A30F06D4A61C40D344ABFA89130" box="[877,1003,1850,1873]" metricMagnitude="2" metricUnit="m" metricValue="7.63" pageId="1" pageNumber="297" unit="m" value="763.0">
<quantity id="4CEE60E6F06D4A61C40D344ABFF29130" box="[877,945,1850,1873]" metricMagnitude="2" metricUnit="m" metricValue="7.63" pageId="1" pageNumber="297" unit="m" value="763.0">763 m</quantity>
a.s.l.
</elevation>
,
<date id="FFA8EBC3F06D4A61C499344AB82E9130" box="[1017,1133,1850,1873]" pageId="1" pageNumber="297" value="2013-11-21">
<collectingDate id="EFEC122BF06D4A61C499344AB82E9130" box="[1017,1133,1850,1873]" pageId="1" pageNumber="297" value="2013-11-21">21.xi.2013</collectingDate>
</date>
, leg.
<collectorName id="26E3A8D5F06D4A61C3C8344AB9349130" box="[1192,1399,1850,1873]" pageId="1" pageNumber="297">M. Bhubaneshwari</collectorName>
,
<collectorName id="26E3A8D5F06D4A61C2E1344ABD5F9112" pageId="1" pageNumber="297">O. Sandhyarani</collectorName>
and
<collectorName id="26E3A8D5F06D4A61C638342CBDAC9112" box="[344,495,1884,1907]" pageId="1" pageNumber="297">S. Dineshwar</collectorName>
</materialsCitation>
;
<materialsCitation id="3B7EC75EF06D4A61C69E342CB97F91F7" ID-GBIF-Occurrence="1288103066" collectingDate="2014-02-07" collectionCode="DCIM" collectorName="Imphal &amp; Longum &amp; M. Bhubaneswari &amp; O. Sandhyrani &amp; S. Dineshwar" country="India" elevation="822" location="Loktak Lake" pageId="1" pageNumber="297" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Manipur" typeStatus="holotype">
<specimenCount id="9D10068AF06D4A61C69E342CBE0E9112" box="[510,589,1884,1907]" pageId="1" pageNumber="297" type="male">1 male</specimenCount>
,
<specimenCount id="9D10068AF06D4A61C53B342CBEFF9112" box="[603,700,1884,1907]" pageId="1" pageNumber="297" type="female">1 female</specimenCount>
(
<collectionCode id="ED0755C6F06D4A61C5AE342CBF599112" box="[718,794,1884,1907]" pageId="1" pageNumber="297">DCIM</collectionCode>
):
<quantity id="4CEE60E6F06D4A61C450342CBF3B9112" box="[816,888,1884,1907]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="297" unit="km" value="45.0">45 km</quantity>
S from
<collectorName id="26E3A8D5F06D4A61C4BB342CB86A9112" box="[987,1065,1884,1907]" pageId="1" pageNumber="297">Imphal</collectorName>
,
<collectorName id="26E3A8D5F06D4A61C358342CB8D69112" box="[1080,1173,1884,1907]" pageId="1" pageNumber="297">Longum</collectorName>
,
<location id="8EC99BD8F06D4A61C3C4342CB9719112" LSID="urn:lsid:plazi:treatment:03BF7C15F06C4A64C7F7368FB9CD94F6:8EC99BD8F06D4A61C3C4342CB9719112" box="[1188,1330,1884,1907]" country="India" name="Loktak Lake" pageId="1" pageNumber="297" stateProvince="Manipur">Loktak Lake</location>
, wetland, 24º31.011ʹN 93º49.066ʹE,
<elevation id="003B2A30F06D4A61C6DB340FBE0291F7" box="[443,577,1919,1942]" metricMagnitude="2" metricUnit="m" metricValue="8.22" pageId="1" pageNumber="297" unit="m" value="822.0">
<quantity id="4CEE60E6F06D4A61C6DB340FBE4091F7" box="[443,515,1919,1942]" metricMagnitude="2" metricUnit="m" metricValue="8.22" pageId="1" pageNumber="297" unit="m" value="822.0">822 m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="EFEC122BF06D4A61C532340FBEF091F7" box="[594,691,1919,1942]" pageId="1" pageNumber="297" value="2014-02-07">
<date id="FFA8EBC3F06D4A61C532340FBEF091F7" box="[594,691,1919,1942]" pageId="1" pageNumber="297" value="2014-02-07">7.ii.2014</date>
</collectingDate>
, leg.
<collectorName id="26E3A8D5F06D4A61C595340FBFF991F7" box="[757,954,1919,1942]" pageId="1" pageNumber="297">M. Bhubaneswari</collectorName>
,
<collectorName id="26E3A8D5F06D4A61C4A9340FB82B91F7" box="[969,1128,1919,1942]" pageId="1" pageNumber="297">O. Sandhyrani</collectorName>
and
<collectorName id="26E3A8D5F06D4A61C3C5340FB97F91F7" box="[1189,1340,1919,1942]" pageId="1" pageNumber="297">S. Dineshwar</collectorName>
</materialsCitation>
.
<materialsCitation id="3B7EC75EF06D4A61C22A340FB9DB91D8" ID-GBIF-Occurrence="1288103063" collectingDate="2004-05-01" collectingDateMax="2004-05-04" collectingDateMin="2004-05-01" collectionCode="NMPC" collectorName="J. Bezdek" country="Laos" elevation="200" location="Vientiane" pageId="1" pageNumber="297" specimenCount="1" specimenCount-male="1" stateProvince="Vientiane" typeStatus="holotype">
<emphasis id="B9621111F06D4A61C22A340FBD4F91D8" bold="true" pageId="1" pageNumber="297">
<collectingCountry id="F3018D93F06D4A61C22A340FB9D491F7" box="[1354,1431,1919,1942]" name="Laos" pageId="1" pageNumber="297">LAOS</collectingCountry>
:
<collectingRegion id="49D203E1F06D4A61C7F734D2BD4B91D8" box="[151,264,1954,1977]" country="Laos" name="Viangchan" pageId="1" pageNumber="297">Vientiane</collectingRegion>
:
</emphasis>
<specimenCount id="9D10068AF06D4A61C67634D2BD2791D8" box="[278,356,1954,1977]" pageId="1" pageNumber="297" type="male">1 male</specimenCount>
, 1 unsexed specimen (
<collectionCode id="ED0755C6F06D4A61C50334D2BEF691D8" box="[611,693,1954,1977]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="297">NMPC</collectionCode>
):
<quantity id="4CEE60E6F06D4A61C5AA34D2BF5291D8" box="[714,785,1954,1977]" metricMagnitude="4" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="297" unit="km" value="55.0">55 km</quantity>
NE of
<collectingRegion id="49D203E1F06D4A61C40A34D2BF9091D8" box="[874,979,1954,1977]" country="Laos" name="Viangchan" pageId="1" pageNumber="297">Vientiane</collectingRegion>
,
<elevation id="003B2A30F06D4A61C48134D2B82691D8" box="[993,1125,1954,1977]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="297" unit="m" value="200.0">
<quantity id="4CEE60E6F06D4A61C48134D2B86B91D8" box="[993,1064,1954,1977]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="297" unit="m" value="200.0">200 m</quantity>
a.s.l.
</elevation>
,
<date id="FFA8EBC3F06D4A61C31534D2B8B791D8" box="[1141,1268,1954,1977]" pageId="1" pageNumber="297" value="2004-05-01" valueMax="2004-05-04" valueMin="2004-05-01">
<collectingDate id="EFEC122BF06D4A61C31534D2B8B791D8" box="[1141,1268,1954,1977]" pageId="1" pageNumber="297" value="2004-05-01" valueMax="2004-05-04" valueMin="2004-05-01">1.4.v.2004</collectingDate>
</date>
,
<collectorName id="26E3A8D5F06D4A61C26334D2B92D91D8" box="[1283,1390,1954,1977]" pageId="1" pageNumber="297">J. Bezděk</collectorName>
lgt
</materialsCitation>
.
<materialsCitation id="3B7EC75EF06D4A62C7F734B4BD2F96CE" ID-GBIF-Occurrence="1288103070" collectingDate="2005-02-25" collectionCode="KMNH" collectorName="Amagi-cho &amp; Tokunoshima I. &amp; Y. Minoshima" country="Japan" lastPageId="2" lastPageNumber="298" location="Ryukyu Islands" pageId="1" pageNumber="297" specimenCount="1" stateProvince="Kagoshima" typeStatus="holotype">
<emphasis id="B9621111F06D4A61C7F734B4BECE91BA" bold="true" box="[151,653,1988,2011]" pageId="1" pageNumber="297">
<collectingCountry id="F3018D93F06D4A61C7F734B4BCB291BA" box="[151,241,1988,2011]" name="Japan" pageId="1" pageNumber="297">JAPAN</collectingCountry>
:
<location id="8EC99BD8F06D4A61C79E34B4BDF691BA" LSID="urn:lsid:plazi:treatment:03BF7C15F06C4A64C7F7368FB9CD94F6:8EC99BD8F06D4A61C79E34B4BDF691BA" box="[254,437,1988,2011]" country="Japan" name="Ryukyu Islands" pageId="1" pageNumber="297" stateProvince="Kagoshima">Ryukyu Islands</location>
:
<collectingRegion id="49D203E1F06D4A61C6A434B4BE0691BA" box="[452,581,1988,2011]" country="Japan" name="Kagoshima" pageId="1" pageNumber="297">Kagoshima</collectingRegion>
Pref.:
</emphasis>
1 ex. (
<collectionCode id="ED0755C6F06D4A61C5BC34B4BF7091BA" box="[732,819,1988,2011]" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13440" name="Kitakyushu Museum and Institute of Natural History" pageId="1" pageNumber="297">KMNH</collectionCode>
),
<collectorName id="26E3A8D5F06D4A61C42734B4BF8191BA" box="[839,962,1988,2011]" pageId="1" pageNumber="297">Amagi-chô</collectorName>
,
<collectorName id="26E3A8D5F06D4A61C4AE34B4B83A91BA" box="[974,1145,1988,2011]" pageId="1" pageNumber="297">Tokunoshima I.</collectorName>
,
<date id="FFA8EBC3F06D4A61C3E734B4B8B691BA" box="[1159,1269,1988,2011]" pageId="1" pageNumber="297" value="2005-02-25">
<collectingDate id="EFEC122BF06D4A61C3E734B4B8B691BA" box="[1159,1269,1988,2011]" pageId="1" pageNumber="297" value="2005-02-25">25.ii.2005</collectingDate>
</date>
,
<collectorName id="26E3A8D5F06D4A61C26334B4B9DF91BA" box="[1283,1436,1988,2011]" pageId="1" pageNumber="297">Y. Minoshima</collectorName>
leg.
<emphasis id="B9621111F06E4A62C7A633E8BD2F96CE" bold="true" box="[198,364,152,175]" pageId="2" pageNumber="298">
<collectingRegion id="49D203E1F06E4A62C7A633E8BD6D96CE" box="[198,302,152,175]" country="Japan" name="Okinawa" pageId="2" pageNumber="298">Okinawa</collectingRegion>
Pref
</emphasis>
</materialsCitation>
.: 1 male (KMNH): Kita, Minamidaitô-jima I., Daitô Is., 25º51.006ʹN 131º14.338ʹE,
<date id="FFA8EBC3F06E4A62C39D33E8B93A96CE" box="[1277,1401,152,175]" pageId="2" pageNumber="298" value="2005-12-12">12.xii.2005</date>
, Y. Minoshima leg.; 1 female (KMNH): near Daitô-jinja, Minamidaitô-jima I., Daitô Is., 25º50.535ʹN 131º13.702ʹE,
<date id="FFA8EBC3F06E4A62C7F733ADBD489695" box="[151,267,221,244]" pageId="2" pageNumber="298" value="2005-03-11">11.iii.2005</date>
, Y. Minoshima leg.; 1 ex. (KMNH), Nagura, Ishigaki-jima I., 24º24.48ʹN 124º09.62ʹE,
<date id="FFA8EBC3F06E4A62C39E33ADB9379695" box="[1278,1396,221,244]" pageId="2" pageNumber="298" value="2005-11-18">18.xi.2005</date>
; Y. Minoshima leg.; 6 exs. (KMNH), Urauchi, Iriomote-jima I., 24º24.78ʹN 123º46.78ʹE,
<date id="FFA8EBC3F06E4A62C3523270B8EE9776" box="[1074,1197,256,279]" pageId="2" pageNumber="298" value="2005-12-31">31.xii.2005</date>
, Y. Minoshima leg.; 1 ex. (KMNH), Minami-bokujô, Yonaguni-jima I., 24º26.682ʹN 122º57.593ʹE,
<date id="FFA8EBC3F06E4A62C4AD3252B80A9758" box="[973,1097,290,313]" pageId="2" pageNumber="298" value="2005-12-25">25.xii.2005</date>
, Y. Minoshima leg.
</paragraph>
</subSubSection>
<caption id="DF699D8BF06E4A62C7F7342EB82F91AA" httpUri="https://zenodo.org/record/261467/files/figure.png" pageId="2" pageNumber="298" targetBox="[160,1435,365,1858]" targetPageId="2">
<paragraph id="8BA9CD03F06E4A62C7F7342EB82F91AA" blockId="2.[151,1436,1886,1995]" pageId="2" pageNumber="298">
<emphasis id="B9621111F06E4A62C7F7342EBD7E9115" bold="true" box="[151,317,1886,1908]" pageId="2" pageNumber="298">FIGURES 713.</emphasis>
78: habitus of alive specimen of
<taxonomicName id="4C16B680F06E4A62C5EC342EBF979112" authority="Sharp" authorityName="Sharp" box="[652,980,1886,1907]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="298" phylum="Arthropoda" rank="species" species="abnormalis">
<emphasis id="B9621111F06E4A62C5EC342EBFC59112" box="[652,902,1886,1907]" italics="true" pageId="2" pageNumber="298">Chasmogenus abnormalis</emphasis>
(Sharp)
</taxonomicName>
from Manipur (7: dorsal, 8: ventral); 9: known distribution of
<taxonomicName id="4C16B680F06E4A62C646340CBDEC91F0" box="[294,431,1916,1937]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="298" phylum="Arthropoda" rank="species" species="abnormalis">
<emphasis id="B9621111F06E4A62C646340CBDEC91F0" box="[294,431,1916,1937]" italics="true" pageId="2" pageNumber="298">C. abnormalis</emphasis>
</taxonomicName>
(green: localities of the examined specimens, blue: literature data, T: type locality). 1013: Morphology of the aedeagus and detail of apex of parameres of examined specimens (10: lectotype; 11: specimen from India: Manipur; 12: specimen from Laos: Vientiane; 13: specimen from Japan: Ryukyu Islands; 1012: ventral, 13 dorsal).
</paragraph>
</caption>
<subSubSection id="C30C9E88F06F4A63C7A733E8BEAC930D" pageId="3" pageNumber="299" type="description">
<paragraph id="8BA9CD03F06F4A63C7A733E8BD4597C0" blockId="3.[151,1436,152,2012]" pageId="3" pageNumber="299">
<emphasis id="B9621111F06F4A63C7A733E8BE1896CE" bold="true" box="[199,603,152,175]" pageId="3" pageNumber="299">
Redescription.
<emphasis id="B9621111F06F4A63C61433E8BE1896CE" bold="true" box="[372,603,152,175]" italics="true" pageId="3" pageNumber="299">Form and coloration</emphasis>
</emphasis>
: Body elongate oval but rather depressed in lateral view, medium size, body length
<quantity id="4CEE60E6F06F4A63C78333CABD2596B0" box="[227,358,186,209]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" metricValueMax="4.4" metricValueMin="3.3" pageId="3" pageNumber="299" unit="mm" value="3.85" valueMax="4.4" valueMin="3.3">3.34.4 mm</quantity>
(
<typeStatus id="54AD73A1F06F4A63C61633CABD9E96B0" box="[374,477,186,209]" pageId="3" pageNumber="299" type="lectotype">lectotype</typeStatus>
:
<quantity id="4CEE60E6F06F4A63C68A33CABE0296B0" box="[490,577,186,209]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="299" unit="mm" value="3.5">3.5 mm</quantity>
), body width
<quantity id="4CEE60E6F06F4A63C5BB33CABF1E96B0" box="[731,861,186,209]" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="3" pageNumber="299" unit="mm" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1.71.9 mm</quantity>
(
<typeStatus id="54AD73A1F06F4A63C40E33CABF9696B0" box="[878,981,186,209]" pageId="3" pageNumber="299" type="lectotype">lectotype</typeStatus>
:
<quantity id="4CEE60E6F06F4A63C48233CAB87996B0" box="[994,1082,186,209]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="299" unit="mm" value="1.8">1.8 mm</quantity>
). General dorsal coloration dark brown (in
<typeStatus id="54AD73A1F06F4A63C66E33ADBD329695" box="[270,369,221,244]" pageId="3" pageNumber="299" type="lectotype">lectotype</typeStatus>
) to black, shiny. Head shiny, dark reddish, with nearly whole clypeus (in
<typeStatus id="54AD73A1F06F4A63C3D233ADB9569695" box="[1202,1301,221,244]" pageId="3" pageNumber="299" type="lectotype">lectotype</typeStatus>
) or only its anterolateral portions slightly paler; pronotum of the same color as head, with anterior, lateral and posterolateral margins slightly paler. Elytra dark brown, of the same color as or slightly darker than head and pronotum. Ventral face of head brown, mentum and mouthparts slightly paler; mouthparts and antennae yellowish. Ventral face of thorax brown to dark brown. Abdominal ventrites dark brown with posterior margin of each ventrite slightly paler. Legs brown, tarsi yellowish.
</paragraph>
<paragraph id="8BA9CD03F06F4A63C7A732DDB90694F5" blockId="3.[151,1436,152,2012]" pageId="3" pageNumber="299">
<emphasis id="B9621111F06F4A63C7A732DDBD4897A5" bold="true" box="[199,267,429,452]" italics="true" pageId="3" pageNumber="299">Head:</emphasis>
Punctation on clypeus moderately coarse but sparse, slightly finer on frons, with distinct larger systematic punctures on clypeus and at inner margin of each eye; interstices smooth, without microsculpture. Anterior margin of clypeus deeply excised, with the median straight portion rather narrow. Labrum strongly sclerotized, largely exposed in front of clypeus, intersegmentary membrane between clypeus and labrum narrowly exposed medially. Eyes large both dorsally and ventrally, dorsally divided by 2.3× the width of one eye. Maxillary palps long, palpomere 4 slightly shorter than palpomere 3, palpomere 3 slightly shorter than palpomere 2. Antennae with 9 antennomeres. Mentum coarsely punctuate, almost rugose; with distinct notch on anteromedial margin and depressed on anteromedial two-thirds.
</paragraph>
<paragraph id="8BA9CD03F06F4A63C7A731D0B95895E6" blockId="3.[151,1436,152,2012]" pageId="3" pageNumber="299">
<emphasis id="B9621111F06F4A63C7A731D0BD6294D6" box="[199,289,672,695]" italics="true" pageId="3" pageNumber="299">
<emphasis id="B9621111F06F4A63C7A731D0BD5A94D6" bold="true" box="[199,281,672,695]" italics="true" pageId="3" pageNumber="299">Thorax</emphasis>
:
</emphasis>
Punctation on pronotum slightly coarser than on frons, with slightly larger systematic punctures present anterolaterally and posterolaterally; interstices smooth. Prosternum rather long in front of procoxae, with median carina present only extremely posteriorly, and without transverse groove. Mesoventrite with evenly rounded low carina, not bearing any long setae. Metaventrite with a narrow posteromedial glabrous area comprising ca. a half the total length of metaventrite. All femora densely pubescent in basal 0.800.85. Elytra with distinct sutural stria on posterior 0.75 to 0.80 of elytral length; punctation on elytral disc similar to pronotum, finer and sparser laterally and posteriorly; elytra without series of punctures, only with rather unorganized longitudinal groups of slightly larger systematic punctures.
</paragraph>
<paragraph id="8BA9CD03F06F4A63C7A730E2BCA8958E" blockId="3.[151,1436,152,2012]" pageId="3" pageNumber="299">
<emphasis id="B9621111F06F4A63C7A730E2BD7A95C8" box="[199,313,914,937]" italics="true" pageId="3" pageNumber="299">
<emphasis id="B9621111F06F4A63C7A730E2BD7395C8" bold="true" box="[199,304,914,937]" italics="true" pageId="3" pageNumber="299">Abdomen</emphasis>
:
</emphasis>
Abdominal ventrites densely pubescent, slightly more dense on posterior half, especially on ventrites 2 5. Fifth ventrite with small medial emargination on posterior margin, ca. twice as long as wide and lined with yellow bristles.
</paragraph>
<paragraph id="8BA9CD03F06F4A63C7A7308ABEAC930D" blockId="3.[151,1436,152,2012]" pageId="3" pageNumber="299">
<emphasis id="B9621111F06F4A63C7A7308ABD379270" box="[199,372,1018,1041]" italics="true" pageId="3" pageNumber="299">
<emphasis id="B9621111F06F4A63C7A7308ABD289270" bold="true" box="[199,363,1018,1041]" italics="true" pageId="3" pageNumber="299">Male genitalia</emphasis>
:
</emphasis>
Aedeagus
<quantity id="4CEE60E6F06F4A63C695308ABED69270" box="[501,661,1018,1041]" metricMagnitude="-4" metricUnit="m" metricValue="9.299999999999999" metricValueMax="9.8" metricValueMin="8.8" pageId="3" pageNumber="299" unit="mm" value="0.9299999999999999" valueMax="0.98" valueMin="0.88">0.880.98 mm</quantity>
(
<typeStatus id="54AD73A1F06F4A63C5C8308ABF489270" box="[680,779,1018,1041]" pageId="3" pageNumber="299" type="lectotype">lectotype</typeStatus>
<quantity id="4CEE60E6F06F4A63C477308ABFC39270" box="[791,896,1018,1041]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="3" pageNumber="299" unit="mm" value="0.98">0.98 mm</quantity>
). Median lobe of aedeagus deeply inserted into phallobase, rather narrow on base, slightly widening up to apical third, then abruptly narrowing, with two subapical constrictions making subapical portion of lateral margin strongly sinuate; apex abruptly widened, truncate; internal sack with complex system on internal sclerites visible as dark internal structures; ventral portion with a strongly sclerotized median projection arising ca. at midlength of median lobe and reaching nearly the level of apical constriction of median lobe; base of the dorsal projection wide, then narrowing apicad into very narrow but apically truncate apex. Parameres only indistinctly longer than phallobase ventrally, but with much longer dorsal portion, inner margin of combined parameres V-shaped ventrally and widely U-shaped dorsally; inner margin of parameres weakly convex ventrally, strongly sinuate dorsally; outer margin of parameres evenly arcuate from base to apex; apical portion of paramere widened, inner margin of the apex with two small teeth (one dorsal and one ventral). Phallobase rather short, symmetrical, slightly narrowed basally, rounded at base.
</paragraph>
</subSubSection>
<subSubSection id="C30C9E88F06F4A63C7A73608B95A93EE" box="[199,1305,1400,1423]" pageId="3" pageNumber="299" type="biology_ecology">
<paragraph id="8BA9CD03F06F4A63C7A73608B95A93EE" blockId="3.[151,1436,152,2012]" box="[199,1305,1400,1423]" pageId="3" pageNumber="299">
<emphasis id="B9621111F06F4A63C7A73608BD5893EE" bold="true" box="[199,283,1400,1423]" pageId="3" pageNumber="299">Biology</emphasis>
. Aquatic species; the Indian specimens were collected in along densely vegetated lake margins.
</paragraph>
</subSubSection>
<subSubSection id="C30C9E88F06F4A63C7A736EABFAF9078" pageId="3" pageNumber="299" type="distribution">
<paragraph id="8BA9CD03F06F4A63C7A736EABFAF9078" blockId="3.[151,1436,152,2012]" pageId="3" pageNumber="299">
<emphasis id="B9621111F06F4A63C7A736EABD1193D0" bold="true" box="[199,338,1434,1457]" pageId="3" pageNumber="299">Distribution</emphasis>
. Widely distributed in the Oriental region (
<collectingCountry id="F3018D93F06F4A63C44F36EABF2993D0" box="[815,874,1434,1457]" name="India" pageId="3" pageNumber="299">India</collectingCountry>
,
<collectingCountry id="F3018D93F06F4A63C41736EABFA793D0" box="[887,996,1434,1457]" name="Sri Lanka" pageId="3" pageNumber="299">
<collectingRegion id="49D203E1F06F4A63C41736EABFD593D0" box="[887,918,1434,1457]" country="United Kingdom" name="Surrey" pageId="3" pageNumber="299">Sri</collectingRegion>
Lanka
</collectingCountry>
,
<collectingCountry id="F3018D93F06F4A63C49136EAB81193D0" box="[1009,1106,1434,1457]" name="Thailand" pageId="3" pageNumber="299">Thailand</collectingCountry>
,
<collectingCountry id="F3018D93F06F4A63C30036EAB89393D0" box="[1120,1232,1434,1457]" name="Cambodia" pageId="3" pageNumber="299">Cambodia</collectingCountry>
,
<collectingCountry id="F3018D93F06F4A63C3BD36EAB97993D0" box="[1245,1338,1434,1457]" name="Vietnam" pageId="3" pageNumber="299">Vietnam</collectingCountry>
,
<collectingCountry id="F3018D93F06F4A63C22736EAB9DB93D0" box="[1351,1432,1434,1457]" name="Taiwan" pageId="3" pageNumber="299">Taiwan</collectingCountry>
,
<collectingCountry id="F3018D93F06F4A63C7F736CDBD4593B5" box="[151,262,1469,1492]" name="Singapore" pageId="3" pageNumber="299">Singapore</collectingCountry>
,
<collectingRegion id="49D203E1F06F4A63C67036CDBD2F93B5" box="[272,364,1469,1492]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="299">Sumatra</collectingRegion>
,
<collectingRegion id="49D203E1F06F4A63C61736CDBDEA93B5" box="[375,425,1469,1492]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="299">Java</collectingRegion>
,
<collectingRegion id="49D203E1F06F4A63C6D336CDBE5593B5" box="[435,534,1469,1492]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="299">Sulawesi</collectingRegion>
, Borneo,
<collectingCountry id="F3018D93F06F4A63C51D36CDBEBB93B5" box="[637,760,1469,1492]" name="Philippines" pageId="3" pageNumber="299">Philippines</collectingCountry>
, Ryukyu Islands). Although we confirm here that the species is widely distributed in the Oriental Region, records from countries other than
<collectingCountry id="F3018D93F06F4A63C4B23690B8489396" box="[978,1035,1504,1527]" name="India" pageId="3" pageNumber="299">India</collectingCountry>
,
<collectingCountry id="F3018D93F06F4A63C3783690B8C79396" box="[1048,1156,1504,1527]" name="Sri Lanka" pageId="3" pageNumber="299">
<collectingRegion id="49D203E1F06F4A63C3783690B8749396" box="[1048,1079,1504,1527]" country="United Kingdom" name="Surrey" pageId="3" pageNumber="299">Sri</collectingRegion>
Lanka
</collectingCountry>
,
<collectingCountry id="F3018D93F06F4A63C3F03690B8879396" box="[1168,1220,1504,1527]" name="Laos" pageId="3" pageNumber="299">Laos</collectingCountry>
and
<collectingCountry id="F3018D93F06F4A63C39A3690B9749396" box="[1274,1335,1504,1527]" name="Japan" pageId="3" pageNumber="299">Japan</collectingCountry>
(Ryukyu Islands) need to be re-confirmed based on a detailed study of the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C30C9E88F06F4A64C7A73555B92D973E" lastPageId="4" lastPageNumber="300" pageId="3" pageNumber="299" type="diagnosis">
<paragraph id="8BA9CD03F06F4A63C7A73555BE7A9115" blockId="3.[151,1436,152,2012]" pageId="3" pageNumber="299">
<emphasis id="B9621111F06F4A63C7A73555BD81905D" bold="true" box="[199,450,1573,1596]" pageId="3" pageNumber="299">Differential diagnosis.</emphasis>
<taxonomicName id="4C16B680F06F4A63C6AA3554BEA2905D" box="[458,737,1572,1596]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="abnormalis">
<emphasis id="B9621111F06F4A63C6AA3554BEA2905D" box="[458,737,1572,1596]" italics="true" pageId="3" pageNumber="299">Chasmogenus abnormalis</emphasis>
</taxonomicName>
may be reliably distinguished from other Oriental species of the genus mainly by the morphology of the male genitalia, especially in the shape of apex of parameres and the median lobe of the aedeagus.
<taxonomicName id="4C16B680F06F4A63C6353519BE1690E0" box="[341,597,1641,1665]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="cattienus">
<emphasis id="B9621111F06F4A63C6353519BE1690E0" box="[341,597,1641,1665]" italics="true" pageId="3" pageNumber="299">Chasmogenus cattienus</emphasis>
</taxonomicName>
from
<collectingCountry id="F3018D93F06F4A63C5FD351ABEB490E0" box="[669,759,1642,1665]" name="Vietnam" pageId="3" pageNumber="299">Vietnam</collectingCountry>
may be easily distinguished by wide median lobe with very narrow apex and simple apex of the parameres (see
<bibRefCitation id="EF87B0F2F06F4A63C5DD35FDBF1490C5" author="Hebauer" box="[701,855,1677,1700]" pageId="3" pageNumber="299" refString="Hebauer, F. (2002) New Hydrophilidae of the Old World (Coleoptera, Hydrophilidae). Acta Coleopterologica, 18 (3), 3 - 24." type="journal article" year="2002">Hebauer 2002</bibRefCitation>
:
<figureCitation id="132DD186F06F4A63C40135FDBFE290C5" box="[865,929,1677,1700]" captionStart="FIGURES 1 6" captionStartId="1.[151,258,1622,1644]" captionTargetBox="[150,1429,213,1598]" captionTargetId="figure@1.[150,1436,194,1599]" captionTargetPageId="1" captionText="FIGURES 1 6. Lectotype of Philydrus abnormalis Sharp, 1890 (= Chasmogenus abnormalis). 1: general habitus, dorsal view; 2: detail of head and the shape of clypeus; 3 5: aedeagus (3: ventral, 4: lateral, 5: dorsal); 6: labels of the lectotype." httpUri="https://zenodo.org/record/261466/files/figure.png" pageId="3" pageNumber="299">Fig. 3</figureCitation>
). Aedeagi of
<taxonomicName id="4C16B680F06F4A63C35135FDB88390C5" box="[1073,1216,1676,1700]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="rubricollis">
<emphasis id="B9621111F06F4A63C35135FDB88390C5" box="[1073,1216,1676,1700]" italics="true" pageId="3" pageNumber="299">C. rubricollis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C16B680F06F4A63C39435FDB90790C5" box="[1268,1348,1676,1700]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="larsi">
<emphasis id="B9621111F06F4A63C39435FDB90790C5" box="[1268,1348,1676,1700]" italics="true" pageId="3" pageNumber="299">C. larsi</emphasis>
</taxonomicName>
seem to be rather similar to that of
<taxonomicName id="4C16B680F06F4A63C6D435C0BE0E90A6" box="[436,589,1711,1735]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="abnormalis">
<emphasis id="B9621111F06F4A63C6D435C0BE0E90A6" box="[436,589,1711,1735]" italics="true" pageId="3" pageNumber="299">C. abnormalis</emphasis>
</taxonomicName>
by the general shape and proportions, and additional studies of these species is needed to understand the differences. Based on the drawings provided by
<bibRefCitation id="EF87B0F2F06F4A63C48135A2B8CB9088" author="Hebauer" box="[993,1160,1746,1769]" pageId="3" pageNumber="299" refString="Hebauer, F. (1992) The species of the genus Chasmogenus Sharp, 1882 (Coleoptera, Hydrophilidae). Acta Coleopterologica, 8 (2), 61 - 92." type="journal article" year="1992">Hebauer (1992</bibRefCitation>
,
<bibRefCitation id="EF87B0F2F06F4A63C3F835A2B8929088" author="Hebauer" box="[1176,1233,1746,1769]" pageId="3" pageNumber="299" refString="Hebauer, F. (1995) Neues zu den Acidocerina Hansen (Helocharae dʼOrchymont) der indomalaiischen Region (Coleoptera, Hydrophilidae). Acta Coleopterologica, 11 (3), 3 - 14." type="journal article" year="1995">1995</bibRefCitation>
), it seems that
<taxonomicName id="4C16B680F06F4A63C2E435A2BD4B916D" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="rubricollis">
<emphasis id="B9621111F06F4A63C2E435A2BD4B916D" italics="true" pageId="3" pageNumber="299">C. rubricollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C16B680F06F4A63C6F43585BE6E916D" box="[404,557,1780,1804]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="abnormalis">
<emphasis id="B9621111F06F4A63C6F43585BE6E916D" box="[404,557,1780,1804]" italics="true" pageId="3" pageNumber="299">C. abnormalis</emphasis>
</taxonomicName>
in apically narrowing parameres (apically widened in
<taxonomicName id="4C16B680F06F4A63C3113585B94A916D" box="[1137,1289,1780,1804]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="abnormalis">
<emphasis id="B9621111F06F4A63C3113585B94A916D" box="[1137,1289,1780,1804]" italics="true" pageId="3" pageNumber="299">C. abnormalis</emphasis>
</taxonomicName>
), and
<taxonomicName id="4C16B680F06F4A63C22C3585B9DF916D" box="[1356,1436,1780,1804]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="larsi">
<emphasis id="B9621111F06F4A63C22C3585B9DF916D" box="[1356,1436,1780,1804]" italics="true" pageId="3" pageNumber="299">C. larsi</emphasis>
</taxonomicName>
differs by parameres subapically sinuate on outer margin (evenly arcuate in
<taxonomicName id="4C16B680F06F4A63C4AF3468B82B914E" box="[975,1128,1815,1839]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="abnormalis">
<emphasis id="B9621111F06F4A63C4AF3468B82B914E" box="[975,1128,1815,1839]" italics="true" pageId="3" pageNumber="299">C. abnormalis</emphasis>
</taxonomicName>
). The characteristic apically widened parameres with two more or less distinct inner teeth easily diagnose
<taxonomicName id="4C16B680F06F4A63C496344AB8D29130" box="[1014,1169,1849,1873]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="abnormalis">
<emphasis id="B9621111F06F4A63C496344AB8D29130" box="[1014,1169,1849,1873]" italics="true" pageId="3" pageNumber="299">C. abnormalis</emphasis>
</taxonomicName>
from all other Oriental, African and Australian-Pacific species.
</paragraph>
<paragraph id="8BA9CD03F06F4A64C7A734F0B92D973E" blockId="3.[151,1436,152,2012]" lastBlockId="4.[151,1436,152,663]" lastPageId="4" lastPageNumber="300" pageId="3" pageNumber="299">
<emphasis id="B9621111F06F4A63C7A734F0BD8B91F6" bold="true" box="[199,456,1920,1943]" pageId="3" pageNumber="299">Lectotype designation.</emphasis>
On our request to loan the type series of
<taxonomicName id="4C16B680F06F4A63C4E934F0B86191F6" box="[905,1058,1919,1943]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="abnormalis">
<emphasis id="B9621111F06F4A63C4E934F0B86191F6" box="[905,1058,1919,1943]" italics="true" pageId="3" pageNumber="299">C. abnormalis</emphasis>
</taxonomicName>
, we received three specimens (one female and two males), which evidently are the same as those that
<bibRefCitation id="EF87B0F2F06F4A63C4E034D2B80891D8" author="Orchymont" box="[896,1099,1954,1977]" pageId="3" pageNumber="299" refString="Orchymont, A. dʼ (1939) Revision des especes du sous-genre Crephelochares dʼ Helochares. Bulletin et Annales de la Societe Entomologique de Belgique, 79, 154 - 166." type="journal article" year="1939">Orchymont (1939)</bibRefCitation>
examined in the course of his revision of
<taxonomicName id="4C16B680F06F4A63C67534B4BE1191BD" authority="Crephelochares" authorityName="Crephelochares" box="[277,594,1988,2012]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="B9621111F06F4A63C67534B4BDCC91BD" box="[277,399,1988,2012]" italics="true" pageId="3" pageNumber="299">Helochares</emphasis>
(
<emphasis id="B9621111F06F4A63C6C134B4BE0A91BD" box="[417,585,1988,2012]" italics="true" pageId="3" pageNumber="299">Crephelochares</emphasis>
)
</taxonomicName>
(=
<taxonomicName id="4C16B680F06F4A63C51334B4BF4B91BD" box="[627,776,1988,2012]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="B9621111F06F4A63C51334B4BF4B91BD" box="[627,776,1988,2012]" italics="true" pageId="3" pageNumber="299">Chasmogenus</emphasis>
</taxonomicName>
). Only two of these specimens bear the label data precisely corresponding to the original description, i.e. locality data but also the collecting date of
<date id="FFA8EBC3F0684A64C30233EAB94896D0" box="[1122,1291,152,177]" pageId="4" pageNumber="300" value="1882-04-17">17th April 1882</date>
. We consider only these two specimens as a part of the original type series, in spite of the fact that three syntypes were mentioned in the original description. Although the female specimen was considered as holotype by previous authors (
<bibRefCitation id="EF87B0F2F0684A64C2403390BC8C9778" author="Orchymont" pageId="4" pageNumber="300" refString="Orchymont, A. dʼ (1939) Revision des especes du sous-genre Crephelochares dʼ Helochares. Bulletin et Annales de la Societe Entomologique de Belgique, 79, 154 - 166." type="journal article" year="1939">Orchymont 1939</bibRefCitation>
,
<bibRefCitation id="EF87B0F2F0684A64C7B93272BD319778" author="Hebauer" box="[217,370,258,281]" pageId="4" pageNumber="300" refString="Hebauer, F. (1992) The species of the genus Chasmogenus Sharp, 1882 (Coleoptera, Hydrophilidae). Acta Coleopterologica, 8 (2), 61 - 92." type="journal article" year="1992">Hebauer 1992</bibRefCitation>
), the holotype was not specified in the original description and hence both specimens available need to be considered as syntypes. Since the morphology of the aedeagus is crucial for the species identification in
<taxonomicName id="4C16B680F0684A64C7F73237BD6F973E" box="[151,300,327,351]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="300" phylum="Arthropoda" rank="genus">
<emphasis id="B9621111F0684A64C7F73237BD6F973E" box="[151,300,327,351]" italics="true" pageId="4" pageNumber="300">Chasmogenus</emphasis>
</taxonomicName>
, we are designating here the male specimen as the lectotype, in order to fix the identity of the species.
</paragraph>
</subSubSection>
<subSubSection id="C30C9E88F0684A64C7A7321AB9CD94F6" pageId="4" pageNumber="300" type="discussion">
<paragraph id="8BA9CD03F0684A64C7A7321AB9CD94F6" blockId="4.[151,1436,152,663]" pageId="4" pageNumber="300">
<emphasis id="B9621111F0684A64C7A7321ABEB097E0" bold="true" box="[199,755,362,385]" pageId="4" pageNumber="300">Comments on variation in aedeagus morphology.</emphasis>
Based on the specimens examined for this study,
<taxonomicName id="4C16B680F0684A64C264321AB9DF97E0" box="[1284,1436,361,385]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="300" phylum="Arthropoda" rank="species" species="abnormalis">
<emphasis id="B9621111F0684A64C264321AB9DF97E0" box="[1284,1436,361,385]" italics="true" pageId="4" pageNumber="300">C. abnormalis</emphasis>
</taxonomicName>
are slightly variable in the extent of the subapical constriction of the median lobe of the aedeagus, and more variable in the detailed morphology of the apex of the paramere. The paramere is always widened apically, and bears two teeth (dorsal and ventral one). The size and shape of the teeth varies slightly even between the left and right paramere, and the teeth seem to be generally large in specimens from western part of the range (
<collectingCountry id="F3018D93F0684A64C3653285B836946D" box="[1029,1141,501,524]" name="Sri Lanka" pageId="4" pageNumber="300">
<collectingRegion id="49D203E1F0684A64C3653285B866946D" box="[1029,1061,501,524]" country="United Kingdom" name="Surrey" pageId="4" pageNumber="300">Sri</collectingRegion>
Lanka
</collectingCountry>
,
<collectingCountry id="F3018D93F0684A64C3E43285B8FF946D" box="[1156,1212,501,524]" name="India" pageId="4" pageNumber="300">India</collectingCountry>
) than in those from Southeast Asia. In the examined specimens from Ryukyu
<collectingCountry id="F3018D93F0684A64C45F3168BFC0944E" box="[831,899,536,559]" name="Iceland" pageId="4" pageNumber="300">Island</collectingCountry>
, these teeth are very small, even though still recognizable. The reexamination of the specimens from Ryukyu confirm that these specimens agree with the
<typeStatus id="54AD73A1F0684A64C27F314AB9C29430" box="[1311,1409,570,593]" pageId="4" pageNumber="300" type="lectotype">lectotype</typeStatus>
in all external characters and the morphology of the aedeagus, including the presence of two teeth at apex of the paramere; the simple apically widened parameres illustrated by
<bibRefCitation id="EF87B0F2F0684A64C5AE31F0BF9294F6" author="Watanabe" box="[718,977,640,663]" pageId="4" pageNumber="300" refString="Watanabe, N. (1987) The Japanese species of Helochares (Crephelochares) (Coleoptera: Hydrophilidae), with description of a new species from Honshu. Aquatic Insects, 9, 11 - 15. http: // dx. doi. org / 10.1080 / 01650428709361262" type="journal article" year="1987">
Watanabe (1987:
<figureCitation id="132DD186F0684A64C4EA31F0BF8A94F6" box="[906,969,640,663]" captionStart="FIGURES 1 6" captionStartId="1.[151,258,1622,1644]" captionTargetBox="[150,1429,213,1598]" captionTargetId="figure@1.[150,1436,194,1599]" captionTargetPageId="1" captionText="FIGURES 1 6. Lectotype of Philydrus abnormalis Sharp, 1890 (= Chasmogenus abnormalis). 1: general habitus, dorsal view; 2: detail of head and the shape of clypeus; 3 5: aedeagus (3: ventral, 4: lateral, 5: dorsal); 6: labels of the lectotype." httpUri="https://zenodo.org/record/261466/files/figure.png" pageId="4" pageNumber="300">Fig. 5</figureCitation>
)
</bibRefCitation>
are evidently due to inaccurate drawings.
</paragraph>
</subSubSection>
</treatment>
</document>