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<document id="2035BED71C5EFE0D09D8E6864D7FD24E" ID-DOI="10.11646/zootaxa.5370.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10147814" ID-ZooBank="48903495-7C6C-46E4-9B1B-D6A2F2781873" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1700202702953" checkinUser="plazi" docAuthor="Harzhauser, Mathias &amp; Landau, Bernard" docDate="2023" docId="03BEE17BFFF9C05DFF0EFD7C7DDD32A5" docLanguage="en" docName="zootaxa.5370.1.1.pdf" docOrigin="Zootaxa 5370 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5370.1.1/52270" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Pseudotuba Harzhauser &amp; Landau 2023, nov. gen." docType="treatment" docVersion="1" lastPageNumber="58" masterDocId="FF879903FFC1C064FF99FFF27C023738" masterDocTitle="The Architectonicidae and Mathildidae (Gastropoda, Heterobranchia) of the Miocene Paratethys Sea-victims of the Miocene Climatic Transition" masterLastPageNumber="74" masterPageNumber="1" pageNumber="57" updateTime="1706539831396" updateUser="carolina">
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<mods:title id="11EDB98DB50234A2180C251B2C3ED334">The Architectonicidae and Mathildidae (Gastropoda, Heterobranchia) of the Miocene Paratethys Sea-victims of the Miocene Climatic Transition</mods:title>
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<mods:namePart id="820D8B6213B93F35187A03BB3401B431">Harzhauser, Mathias</mods:namePart>
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<mods:affiliation id="C63E8BB6AE83C12A114C79CBD467784E">Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655;</mods:affiliation>
<mods:nameIdentifier id="DDEC6099CD2C15B91D0445DD09263D04" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:namePart id="17C3D455B85328CB2DCD1AB4F841C4D3">Landau, Bernard</mods:namePart>
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<mods:affiliation id="9150C0326F63CA065B404FE1A274711F">Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655; &amp; Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal. bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494 &amp; Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655;</mods:affiliation>
<mods:nameIdentifier id="26273B84EFF2A9588FF0B209CD68DC87" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:date id="B95EF8DFE2F71BFA88C8B73AE3197C5F">2023</mods:date>
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<subSubSection id="C30D03E6FFF9C05CFF0EFD7C7DF33590" box="[151,497,654,681]" pageId="56" pageNumber="57" type="nomenclature">
<paragraph id="8BA8506DFFF9C05CFF0EFD7C7DF33590" blockId="56.[151,497,654,681]" box="[151,497,654,681]" pageId="56" pageNumber="57">
<emphasis id="B9638C7FFFF9C05CFF0EFD7C7DF33590" bold="true" box="[151,497,654,681]" pageId="56" pageNumber="57">
Genus
<taxonomicName id="4C172BEEFFF9C05CFF77FD7C7D7E3591" ID-CoL="BXSCY" authority="Harzhauser &amp; Landau, 2023" authorityName="Harzhauser &amp; Landau" authorityYear="2023" box="[238,380,654,681]" class="Gastropoda" family="Mathildidae" genus="Pseudotuba" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Heterostropha" pageId="56" pageNumber="57" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B9638C7FFFF9C05CFF77FD7C7D7E3591" bold="true" box="[238,380,654,681]" italics="true" pageId="56" pageNumber="57">Pseudotuba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2503104FFF9C05CFE1DFD7C7DF33590" box="[388,497,654,680]" pageId="56" pageNumber="57" rank="genus">nov. gen.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFF9C05CFF0EFD2478A635C8" box="[151,1188,726,753]" pageId="56" pageNumber="57" type="type_taxon">
<paragraph id="8BA8506DFFF9C05CFF0EFD2478A635C8" blockId="56.[151,1437,726,2013]" box="[151,1188,726,753]" pageId="56" pageNumber="57">
<materialsCitation id="3B7F5A30FFF9C05CFF0EFD2478A235C8" box="[151,1184,726,753]" location="Middle Miocene" municipality="Central Paratethys Sea" pageId="56" pageNumber="57" specimenCount="1" typeStatus="holotype">
<emphasis id="B9638C7FFFF9C05CFF0EFD247D3035C8" bold="true" box="[151,306,726,752]" pageId="56" pageNumber="57">
<typeStatus id="54ACEECFFFF9C05CFF0EFD247CD035C8" box="[151,210,726,752]" pageId="56" pageNumber="57">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C172BEEFFF9C05CFEA0FD247EC935C9" authority="(Sacco, 1895)" baseAuthorityName="Sacco" baseAuthorityYear="1895" box="[313,715,726,753]" class="Gastropoda" family="Mathildidae" genus="Pseudotuba" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="badensis">
<emphasis id="B9638C7FFFF9C05CFEA0FD247E2435C8" box="[313,550,726,752]" italics="true" pageId="56" pageNumber="57">Pseudotuba badensis</emphasis>
(Sacco, 1895)
</taxonomicName>
,
<location id="8EC806B6FFF9C05CFD4EFD247F9635C8" LSID="urn:lsid:plazi:treatment:03BEE17BFFF9C05DFF0EFD7C7DDD32A5:8EC806B6FFF9C05CFD4EFD247F9635C8" box="[727,916,726,752]" municipality="Central Paratethys Sea" name="Middle Miocene" pageId="56" pageNumber="57">Middle Miocene</location>
,
<collectingMunicipality id="6BCCCA17FFF9C05CFC06FD2478A235C8" box="[927,1184,726,752]" pageId="56" pageNumber="57">Central Paratethys Sea</collectingMunicipality>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFF9C05CFF5EFD087EC33464" pageId="56" pageNumber="57" type="diagnosis">
<paragraph id="8BA8506DFFF9C05CFF5EFD087EC33464" blockId="56.[151,1437,726,2013]" pageId="56" pageNumber="57">
<emphasis id="B9638C7FFFF9C05CFF5EFD087D40342C" bold="true" box="[199,322,762,788]" pageId="56" pageNumber="57">Diagnosis.</emphasis>
Medium-sized, solid, turbiniform shell, with strongly convex whorls and slightly gradate spire; Protoconch hyperstrophic. First teleoconch whorl with six narrow, close-set, convex spiral cords crossed by numerous much weaker axial lamellae. Last whorl globular.
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFF9C05CFF5EFC94792B3344" pageId="56" pageNumber="57" type="description">
<paragraph id="8BA8506DFFF9C05CFF5EFC94792B3344" blockId="56.[151,1437,726,2013]" pageId="56" pageNumber="57">
<emphasis id="B9638C7FFFF9C05CFF5EFC947D5B34B8" bold="true" box="[199,345,870,896]" pageId="56" pageNumber="57">Description.</emphasis>
Medium-sized, solid, turbiniform shell, with strongly convex whorls and slightly gradate spire. Protoconch hyperstrophic. Teleoconch of convex whorls with deeply incised suture. Rounded periphery at about one-third whorl height. First teleoconch whorl with six narrow, close-set, convex spiral cords crossed by numerous much weaker, prosocline axial lamellae forming fine, dense cancellate pattern, with small, rounded beads at intersections. Secondary spiral cords intercalated between primary cords on later whorls. Last whorl globular, with numerous spiral cords of subequal strength and spacing; base not or weakly delimited, convex, umbilicus relatively broad to narrow, delimited by periumbilical cord in some species. Aperture subcircular. Columella concave with broad rim. Umbilicus narrow or reduced to chink. No parietal callus. Outer lip thin. Spiral lirae within aperture in some species.
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFF9C05CFF5EFB747F3F32A4" pageId="56" pageNumber="57" type="etymology">
<paragraph id="8BA8506DFFF9C05CFF5EFB747F3A3398" blockId="56.[151,1437,726,2013]" box="[199,824,1158,1187]" pageId="56" pageNumber="57">
<emphasis id="B9638C7FFFF9C05CFF5EFB747D4C3398" bold="true" box="[199,334,1158,1184]" pageId="56" pageNumber="57">Etymology.</emphasis>
Prefix
<emphasis id="B9638C7FFFF9C05CFE39FB747DED3398" box="[416,495,1158,1184]" italics="true" pageId="56" pageNumber="57">pseudo</emphasis>
- (from Greek ψευδής, false).
</paragraph>
<paragraph id="8BA8506DFFF9C05CFF5EFB587F3F32A4" blockId="56.[151,1437,726,2013]" pageId="56" pageNumber="57">
<emphasis id="B9638C7FFFF9C05CFF5EFB587D9333FC" bold="true" box="[199,401,1194,1220]" pageId="56" pageNumber="57">Included species.</emphasis>
<taxonomicName id="4C172BEEFFF9C05CFE0EFB587F5033FC" authority="Grateloup, 1828" authorityName="Grateloup" authorityYear="1828" box="[407,850,1194,1220]" class="Gastropoda" family="Mathildidae" genus="Cyclostoma" kingdom="Animalia" order="Littorinimorpha" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="cancellata">
<emphasis id="B9638C7FFFF9C05CFE0EFB587E9533FC" box="[407,663,1194,1220]" italics="true" pageId="56" pageNumber="57">Cyclostoma cancellata</emphasis>
<bibRefCitation id="EF862D9CFFF9C05CFD04FB587F5033FC" author="Grateloup, J. - P. S. de" box="[669,850,1194,1220]" pageId="56" pageNumber="57" pagination="3 - 27" refId="ref53029" refString="Grateloup, J. - P. S. de (1828) Description de plusieurs especes de coquilles fossiles des environs de Dax (Landes). Bulletin d'histoire naturelle de la societe Linneenne de Bordeaux, Series 7, 2, 3 - 27. [https: // www. biodiversitylibrary. org / item / 110491 # page / 7 / mode / 1 up]" type="journal article" year="1828">Grateloup, 1828</bibRefCitation>
</taxonomicName>
(Langhian,
<collectingCountry id="F30010FDFFF9C05CFC40FB58782533FC" box="[985,1063,1194,1220]" name="France" pageId="56" pageNumber="57">France</collectingCountry>
) (
<bibRefCitation id="EF862D9CFFF9C05CFBA5FB58799533FC" author="Cossmann, M. &amp; Peyrot, A." box="[1084,1431,1194,1221]" pageId="56" pageNumber="57" pagination="5 - 321" refId="ref50593" refString="Cossmann, M. &amp; Peyrot, A. (1922) Conchologie neogenique de l'Aquitaine. Actes de la societe Linneenne de Bordeaux, 73, 5 - 321. [https: // www. biodiversitylibrary. org / item / 98736 # page / 11 / mode / 1 up]" type="journal article" year="1922">Cossmann &amp; Peyrot 1922: 179</bibRefCitation>
, pl. 3, figs 3536,
<figureCitation id="132C4CE8FFF9C05CFEC7FB3C7DC033D0" box="[350,450,1230,1256]" captionStart="FIGURE 30" captionStartId="62.[151,250,962,986]" captionTargetBox="[151,1436,181,938]" captionTargetId="figure-30@62.[151,1436,181,938]" captionTargetPageId="62" captionText="FIGURE 30. Schematic drawings of Paratethyan Architectonicidae showing size relations amongst species; colors group genera. 1. Ammotectonica gregorovae nov. sp., 2. Ammotectonica soproniensis (Strausz, 1960), 3. Simplexollata anticollata nov. sp., 4. Simplexollata simplex (Bronn, 1831), 5. Solatisonax tavianii nov. sp., 6. Solatisonax? transversa nov. sp., 7. Solatisonax kostejana (Boettger, 1907), 8. Granosolarium semilaevis (Sacco, 1892), 9. Heliacus berthae (Boettger, 1902), 10. Heliacus globosus nov. sp., 11. Heliacus miser (Dujardin, 1837), 12. Nipteraxis deformatus nov. sp., 13. Nipteraxis exmoniliferus (Sacco, 1892), 14. Nipteraxis monilifer (Bronn, 1831), 15. Pseudotorinia grasemanni nov. sp., 16. Pseudotorinia marthae (Boettger, 1902), 17. Spirolaxis cornicula (Boettger, 1902)." figureDoi="http://doi.org/10.5281/zenodo.10147878" httpUri="https://zenodo.org/record/10147878/files/figure.png" pageId="56" pageNumber="57">Figs 30F</figureCitation>
<figureCitation id="132C4CE8FFF9C05CFE58FB137DE533D1" box="[449,487,1231,1263]" captionStart="FIGURE 1" captionStartId="5.[151,250,1522,1546]" captionTargetBox="[232,1366,234,1455]" captionTargetId="figure-58@5.[232,1366,234,1455]" captionTargetPageId="5" captionText="FIGURE 1. Descriptive terms used in descriptions of Architectonicidae (modified from Bieler 1993)." figureDoi="http://doi.org/10.5281/zenodo.10147816" httpUri="https://zenodo.org/record/10147816/files/figure.png" pageId="56" pageNumber="57">
<subScript id="17935228FFF9C05CFE58FB137DCB33D7" attach="left" box="[449,457,1249,1263]" fontSize="6" pageId="56" pageNumber="57">1</subScript>
F
</figureCitation>
<figureCitation id="132C4CE8FFF9C05CFE7EFB137DED33D7" box="[487,495,1249,1263]" captionStart="FIGURE 4" captionStartId="9.[151,250,1590,1614]" captionTargetBox="[151,1436,151,1566]" captionTargetId="figure-233@9.[151,1436,151,1569]" captionTargetPageId="9" captionText="FIGURE 4 Protoconchs of Paratethyan Architectonicidae. A. Ammotectonica gregorovae nov. sp., paratype, NHMW 1862/0029/0078c. B. Simplexollata anticollata nov. sp., paratype, NHMW 1854/0035/0331, Lăpugiu de Sus (Romania). C. Simplexollata carocollata (Lamarck, 1822), NHMW 1850/0018/0115, Léognan (France), Burdigalian. D. Simplexollata simplex (Bronn, 1831), NHMW 2013/0078/0109, Baden (Austria).E. Nipteraxis deformatusnov. sp., paratype, NHMW 2010/0004/0456a, Bad Vöslau (Austria). F. Nipteraxis exmoniliferus (Sacco, 1892), NHMW 1907/0323/1959, M̂llersdorf (Austria). G. Solatectonica kostejana (Boettger, 1907), NHMW 2012/0229/0049b, Traiskirchen (Austria). H. Nipteraxis monilifer (Bronn, 1831), NHMW 1846/0037/0427, M̂llersdorf (Austria). I. Solatisonax tavianii nov. sp., NHMW2016/0266/0002, SL: 8.0 mm, MD: 15.0 mm, Lăpugiu des Sus (Romania). J. Solatisonax transversa nov. sp., holotype, NHMW 2016/0177/0341, Lăpugiu de Sus (Roma- nia). K. Granosolarium semilaevis (Sacco, 1892), NHMW 1846/0037/0428d, M̂llersdorf (Austria). L. Heliacus globosus nov. sp., holotype NHMW 1907/0328/1959a, M̂llersdorf (Austria). M. Heliacus miser (Dujardin, 1837), NHMW 2023/0267/0001, Steinebrunn (Austria). N. Pseudotorinia grasemanni nov. sp., paratype, NHMW 2023/0267/0010, Steinebrunn (Austria). O. Pseudotorinia marthae (Boettger, 1902), NHMW 2012/0229/0049, Traiskirchen (Austria). All figures same scale." figureDoi="http://doi.org/10.5281/zenodo.10147820" httpUri="https://zenodo.org/record/10147820/files/figure.png" pageId="56" pageNumber="57">
<subScript id="17935228FFF9C05CFE7EFB137DED33D7" attach="left" box="[487,495,1249,1263]" fontSize="6" pageId="56" pageNumber="57">4</subScript>
</figureCitation>
).
<taxonomicName id="4C172BEEFFF9C05CFD9EFB3C7F6C33D1" authority="sensu Sacco, 1895" authorityName="sensu Sacco" authorityYear="1895" box="[519,878,1230,1257]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="sulcata">
<emphasis id="B9638C7FFFF9C05CFD9EFB3C7E9A33D0" box="[519,664,1230,1256]" italics="true" pageId="56" pageNumber="57">Tuba sulcata</emphasis>
sensu Sacco, 1895
</taxonomicName>
(Langhian, Colli Torinesi,
<collectingCountry id="F30010FDFFF9C05CFB39FB3C78D733D0" box="[1184,1237,1230,1256]" name="Italy" pageId="56" pageNumber="57">Italy</collectingCountry>
) (Sacco 1895: pl. 3, fig. 43);
<taxonomicName id="4C172BEEFFF9C05CFE88FB007E763235" authority="Sacco, 1895" authorityName="Sacco" authorityYear="1895" box="[273,628,1266,1293]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="pedemontana">
<emphasis id="B9638C7FFFF9C05CFE88FB007DE43234" box="[273,486,1266,1292]" italics="true" pageId="56" pageNumber="57">Tuba pedemontana</emphasis>
Sacco, 1895
</taxonomicName>
(Burdigalian/Langhian, Colli Torinesi,
<collectingCountry id="F30010FDFFF9C05CFBB2FB0078623234" box="[1067,1120,1266,1292]" name="Italy" pageId="56" pageNumber="57">Italy</collectingCountry>
) (Sacco 1895: pl. 3, fig. 45);
<taxonomicName id="4C172BEEFFF9C05CFF0EFAE47E013209" authority="d'Orbigny, 1852" authorityName="d'Orbigny" authorityYear="1852" box="[151,515,1302,1329]" class="Gastropoda" family="Turbinidae" genus="Turbo" kingdom="Animalia" order="Trochida" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="bellardii">
<emphasis id="B9638C7FFFF9C05CFF0EFAE47D433208" box="[151,321,1302,1328]" italics="true" pageId="56" pageNumber="57">Turbo bellardii</emphasis>
dOrbigny, 1852
</taxonomicName>
(Tortonian, Po Basin,
<collectingCountry id="F30010FDFFF9C05CFC9AFAE47F3A3208" box="[771,824,1302,1328]" name="Italy" pageId="56" pageNumber="57">Italy</collectingCountry>
) (Sacco 1895: pl. 3, fig. 46).
<taxonomicName id="4C172BEEFFF9C05CFB1FFAE57D4A326C" authority="Deshayes, 1861" authorityName="Deshayes" authorityYear="1861" class="Gastropoda" family="Littorinidae" genus="Littorina" kingdom="Animalia" order="Littorinimorpha" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="cyclostomoides">
<emphasis id="B9638C7FFFF9C05CFB1FFAE5799E3208" box="[1158,1436,1302,1328]" italics="true" pageId="56" pageNumber="57">Littorina cyclostomoides</emphasis>
<bibRefCitation id="EF862D9CFFF9C05CFF0EFAC87D4A326C" author="Deshayes, G. P." box="[151,328,1338,1364]" pageId="56" pageNumber="57" pagination="921 - 968" refId="ref51820" refString="Deshayes, G. P. (1861 - 1864) Description des animaux sans vertebres decouverts dans le bassin de Paris pour servir de supplement a la Description des coquilles fossiles des environs de Paris comprenant une revue generale de toutes les especes actuellement connues. Tome deuxieme. Mollusques acephales monomyaires et Brachiopodes. Mollusques cephales. Premiere Partie. Bailliere, Paris, 968 pp., 62 pls. [pp. 1 - 432, pls. 1 - 26 (1861), pp. 433 - 640, pls. 27 - 39 (1862), pp. 641 - 920, pls. 40 - 62 (1863), pp. 921 - 968 (1864).] [https: // www. biodiversitylibrary. org / page / 35623270]" type="book chapter" year="1861">Deshayes, 1861</bibRefCitation>
</taxonomicName>
(Eocene, Paris Basin,
<collectingCountry id="F30010FDFFF9C05CFDDDFAC87E90326C" box="[580,658,1338,1364]" name="France" pageId="56" pageNumber="57">France</collectingCountry>
) (
<bibRefCitation id="EF862D9CFFF9C05CFD31FAC87F55326C" author="Deshayes, G. P." box="[680,855,1338,1364]" pageId="56" pageNumber="57" pagination="921 - 968" refId="ref51820" refString="Deshayes, G. P. (1861 - 1864) Description des animaux sans vertebres decouverts dans le bassin de Paris pour servir de supplement a la Description des coquilles fossiles des environs de Paris comprenant une revue generale de toutes les especes actuellement connues. Tome deuxieme. Mollusques acephales monomyaires et Brachiopodes. Mollusques cephales. Premiere Partie. Bailliere, Paris, 968 pp., 62 pls. [pp. 1 - 432, pls. 1 - 26 (1861), pp. 433 - 640, pls. 27 - 39 (1862), pp. 641 - 920, pls. 40 - 62 (1863), pp. 921 - 968 (1864).] [https: // www. biodiversitylibrary. org / page / 35623270]" type="book chapter" year="1861">Deshayes 1861</bibRefCitation>
: pl. 16 figs 14) (own material studied).
<taxonomicName id="4C172BEEFFF9C05CFA85FAC87D943240" authority="Cossmann, 1907" authorityName="Cossmann" authorityYear="1907" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="elatospira">
<emphasis id="B9638C7FFFF9C05CFA85FAC87CD33240" italics="true" pageId="56" pageNumber="57">Tuba elatospira</emphasis>
<bibRefCitation id="EF862D9CFFF9C05CFF43FAAC7D943240" author="Cossmann, M." box="[218,406,1374,1400]" pageId="56" pageNumber="57" pagination="182 - 286" refId="ref50379" refString="Cossmann, M. (1907) Appendice No. 4 au catalogue illustre des coquilles fossils de l'Eocene des environs de Paris. Annales de la societe royale zoologique et malacologique de Belgique, 41, 182 - 286, pls. 5 - 10. [https: // www. biodiversitylibrary. org / page / 5599125]" type="journal article" year="1907">Cossmann, 1907</bibRefCitation>
</taxonomicName>
(Eocene, Paris Basin,
<collectingCountry id="F30010FDFFF9C05CFD05FAAD7EE93241" box="[668,747,1375,1401]" name="France" pageId="56" pageNumber="57">France</collectingCountry>
, (
<bibRefCitation id="EF862D9CFFF9C05CFC99FAAC7FB93240" author="Cossmann, M." box="[768,955,1374,1400]" pageId="56" pageNumber="57" pagination="182 - 286" refId="ref50379" refString="Cossmann, M. (1907) Appendice No. 4 au catalogue illustre des coquilles fossils de l'Eocene des environs de Paris. Annales de la societe royale zoologique et malacologique de Belgique, 41, 182 - 286, pls. 5 - 10. [https: // www. biodiversitylibrary. org / page / 5599125]" type="journal article" year="1907">Cossmann 1907</bibRefCitation>
: pl. 8 fig. 1293).
<taxonomicName id="4C172BEEFFF9C05CFB0AFAAC79393240" authorityName="J. de C. Sowerby" authorityYear="1823" box="[1171,1339,1374,1400]" class="Gastropoda" family="Turbinidae" genus="Turbo" kingdom="Animalia" order="Trochida" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="sculptus">
<emphasis id="B9638C7FFFF9C05CFB0AFAAC79393240" box="[1171,1339,1374,1400]" italics="true" pageId="56" pageNumber="57">Turbo sculptus</emphasis>
</taxonomicName>
J. de C. Sowerby, 1823 (Eocene,
<collectingCountry id="F30010FDFFF9C05CFE29FA707DD832A4" box="[432,474,1410,1436]" name="United Kingdom" pageId="56" pageNumber="57">UK</collectingCountry>
) (Sowerby 1823: pl. 395 fig. 2)
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFF9C05DFF5EFA547DDD32A5" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57" type="discussion">
<paragraph id="8BA8506DFFF9C05CFF5EFA547815314D" blockId="56.[151,1437,726,2013]" pageId="56" pageNumber="57">
<emphasis id="B9638C7FFFF9C05CFF5EFA547E7F32F8" bold="true" box="[199,637,1446,1472]" pageId="56" pageNumber="57">Stratigraphic and geographic range.</emphasis>
The genus was present already during the Eocene, when it was represented by several species in the Paris Basin. During the Miocene,
<taxonomicName id="4C172BEEFFF9C05CFCECFA387FFB32DC" authorityName="Harzhauser &amp; Landau" authorityYear="2023" box="[885,1017,1482,1508]" class="Gastropoda" family="Mathildidae" genus="Pseudotuba" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="56" pageNumber="57" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF9C05CFCECFA387FFB32DC" box="[885,1017,1482,1508]" italics="true" pageId="56" pageNumber="57">Pseudotuba</emphasis>
</taxonomicName>
was represented in the entire circum-Mediterranean region, including the North Sea Basin. We are not aware of Pliocene occurrences.
<emphasis id="B9638C7FFFF9C05CFB41FA1C799F3130" box="[1240,1437,1518,1544]" italics="true" pageId="56" pageNumber="57">Tuba seriaformis</emphasis>
Schr̂der, 1995, from the Early Cretaceous of
<collectingCountry id="F30010FDFFF9C05CFD3DF9E07EF73114" box="[676,757,1554,1580]" name="Poland" pageId="56" pageNumber="57">Poland</collectingCountry>
, is already quite close to
<taxonomicName id="4C172BEEFFF9C05CFB8FF9E078983114" authorityName="Harzhauser &amp; Landau" authorityYear="2023" box="[1046,1178,1554,1580]" class="Gastropoda" family="Mathildidae" genus="Pseudotuba" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="56" pageNumber="57" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF9C05CFB8FF9E078983114" box="[1046,1178,1554,1580]" italics="true" pageId="56" pageNumber="57">Pseudotuba</emphasis>
</taxonomicName>
and develops a hyperstrophic protoconch and six spiral cords on the first teleoconch whorl (see
<bibRefCitation id="EF862D9CFFF9C05CFC7CF9C478653168" author="Kaim, A." box="[997,1127,1590,1616]" pageId="56" pageNumber="57" pagination="3 - 183" refId="ref55153" refString="Kaim, A. (2004) The evolution of conch ontogeny in Mesozoic open sea gastropods. Palaeontologia polonica, 62, 3 - 183. [http: // www. palaeontologia. pan. pl / PP 62 / PP- 62 _ calosc. pdf]" type="journal article" year="2004">Kaim 2004</bibRefCitation>
: fig. 117). However, due to the huge stratigraphic gap we refrain from including this species in
<taxonomicName id="4C172BEEFFF9C05CFC15F9A87812314C" authorityName="Harzhauser &amp; Landau" authorityYear="2023" box="[908,1040,1626,1652]" class="Gastropoda" family="Mathildidae" genus="Pseudotuba" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="56" pageNumber="57" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF9C05CFC15F9A87812314C" box="[908,1040,1626,1652]" italics="true" pageId="56" pageNumber="57">Pseudotuba</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA8506DFFF9C05CFF5EF98C793A31A0" blockId="56.[151,1437,726,2013]" box="[199,1336,1662,1689]" pageId="56" pageNumber="57">
<emphasis id="B9638C7FFFF9C05CFF5EF98C7DAA31A0" bold="true" box="[199,424,1662,1688]" pageId="56" pageNumber="57">Paleoenvironment.</emphasis>
The
<typeStatus id="54ACEECFFFF9C05CFE79F98D7E1231A1" box="[480,528,1663,1689]" pageId="56" pageNumber="57">type</typeStatus>
species was found in middle to outer neritic depositional environments.
</paragraph>
<paragraph id="8BA8506DFFF9C05CFF5EF9507D373080" blockId="56.[151,1437,726,2013]" pageId="56" pageNumber="57">
<emphasis id="B9638C7FFFF9C05CFF5EF9507D4E3184" bold="true" box="[199,332,1698,1724]" pageId="56" pageNumber="57">Discussion.</emphasis>
Species placed herein in the genus
<taxonomicName id="4C172BEEFFF9C05CFD7AF9507F653184" authority="Harzhauser &amp; Landau, 2023" authorityName="Harzhauser &amp; Landau" authorityYear="2023" box="[739,871,1698,1724]" class="Gastropoda" family="Mathildidae" genus="Pseudotuba" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="56" pageNumber="57" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B9638C7FFFF9C05CFD7AF9507F653184" box="[739,871,1698,1724]" italics="true" pageId="56" pageNumber="57">Pseudotuba</emphasis>
</taxonomicName>
<emphasis id="B9638C7FFFF9C05CFCF6F9507FDB3184" bold="true" box="[879,985,1698,1724]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="A2503104FFF9C05CFCF6F9507FDB3184" box="[879,985,1698,1724]" pageId="56" pageNumber="57" rank="genus">nov. gen.</taxonomicNameLabel>
</emphasis>
have been placed so far in
<taxonomicName id="4C172BEEFFF9C05CFA8AF9507CCD31D8" authority="I. Lea, 1833" authorityName="I. Lea" authorityYear="1833" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="56" pageNumber="57" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF9C05CFA8AF95079493184" box="[1299,1355,1698,1724]" italics="true" pageId="56" pageNumber="57">Tuba</emphasis>
I.
<bibRefCitation id="EF862D9CFFF9C05CFAF2F9517CCD31D8" author="Lea, I." pageId="56" pageNumber="57" refId="ref56283" refString="Lea, I. (1833) Contributions to Geology. Carey, Lea and Blanchard, Philadelphia, Pennsylvania, 227 pp., 6 pls. [https: // www. biodiversitylibrary. org / page / 17592495]" type="book" year="1833">Lea, 1833</bibRefCitation>
</taxonomicName>
[
<typeStatus id="54ACEECFFFF9C05CFF46F9357D1031D9" box="[223,274,1735,1761]" pageId="56" pageNumber="57">type</typeStatus>
species
<taxonomicName id="4C172BEEFFF9C05CFEECF9347EB731D8" authority="I. Lea, 1833" authorityName="I. Lea" authorityYear="1833" box="[373,693,1734,1761]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="alternata">
<emphasis id="B9638C7FFFF9C05CFEECF9347E1C31D8" box="[373,542,1734,1760]" italics="true" pageId="56" pageNumber="57">Tuba alternata</emphasis>
I.
<bibRefCitation id="EF862D9CFFF9C05CFDD9F9357EB731D8" author="Lea, I." box="[576,693,1734,1761]" pageId="56" pageNumber="57" refId="ref56283" refString="Lea, I. (1833) Contributions to Geology. Carey, Lea and Blanchard, Philadelphia, Pennsylvania, 227 pp., 6 pls. [https: // www. biodiversitylibrary. org / page / 17592495]" type="book" year="1833">Lea, 1833</bibRefCitation>
</taxonomicName>
, by subsequent designation,
<bibRefCitation id="EF862D9CFFF9C05CFC66F93478E931D8" author="Cossmann M." box="[1023,1259,1734,1760]" pageId="56" pageNumber="57" refId="ref50440" refString="Cossmann M. (1912) Essais de paleoconchologie comparee. Neuvieme livraison. published by the author, Paris, 215 pp., 10 pls. [https: // www. biodiversitylibrary. org / item / 112976 # page / 13 / mode / 1 up]" type="book" year="1912">Cossmann, 1912: 13</bibRefCitation>
, Eocene, Western Atlantic (see https://science.mnhn.fr/institution/mnhn/collection/f/item/j05056?listIndex=9&amp;listCount=76 for specimens of the
<typeStatus id="54ACEECFFFF9C05CFEFDF8FD7D963011" box="[356,404,1807,1833]" pageId="56" pageNumber="57">type</typeStatus>
species)]. Synonyms of
<taxonomicName id="4C172BEEFFF9C05CFD2FF8FC7EEC3010" authorityName="I. Lea" authorityYear="1833" box="[694,750,1806,1832]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="56" pageNumber="57" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF9C05CFD2FF8FC7EEC3010" box="[694,750,1806,1832]" italics="true" pageId="56" pageNumber="57">Tuba</emphasis>
</taxonomicName>
are:
<taxonomicName id="4C172BEEFFF9C05CFCB5F8FD78393010" authority="Jeffreys, 1884" authorityName="Jeffreys" authorityYear="1884" box="[812,1083,1806,1833]" class="Gastropoda" family="Mathildidae" genus="Gegania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="56" pageNumber="57" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF9C05CFCB5F8FD7F8E3011" box="[812,908,1807,1833]" italics="true" pageId="56" pageNumber="57">Gegania</emphasis>
<bibRefCitation id="EF862D9CFFF9C05CFC0EF8FC78393010" author="Jeffreys, J. G." box="[919,1083,1806,1832]" pageId="56" pageNumber="57" pagination="341 - 372" refId="ref55069" refString="Jeffreys, J. G. (1884) On the Mollusca procured during the ' Lightning' and ' Porcupine' expeditions, 1868 - 70. (Part VIII). Proceedings of the zoological society of London, 1882, 341 - 372, pls. 26 - 28. [http: // www. biodiversitylibrary. org / page / 28690228] https: // doi. org / 10.1111 / j. 1096 - 3642.1884. tb 02837. x" type="journal article" year="1884">Jeffreys, 1884</bibRefCitation>
</taxonomicName>
,
<typeStatus id="54ACEECFFFF9C05CFBD0F8FD787B3011" box="[1097,1145,1807,1833]" pageId="56" pageNumber="57">type</typeStatus>
species
<taxonomicName id="4C172BEEFFF9C05CFB46F8FD7D3A3074" authority="Jeffreys, 1884" authorityName="Jeffreys" authorityYear="1884" class="Gastropoda" family="Mathildidae" genus="Gegania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="pinguis">
<emphasis id="B9638C7FFFF9C05CFB46F8FD799E3010" box="[1247,1436,1807,1833]" italics="true" pageId="56" pageNumber="57">Gegania pinguis</emphasis>
<bibRefCitation id="EF862D9CFFF9C05CFF0EF8C07D3A3074" author="Jeffreys, J. G." box="[151,312,1842,1868]" pageId="56" pageNumber="57" pagination="341 - 372" refId="ref55069" refString="Jeffreys, J. G. (1884) On the Mollusca procured during the ' Lightning' and ' Porcupine' expeditions, 1868 - 70. (Part VIII). Proceedings of the zoological society of London, 1882, 341 - 372, pls. 26 - 28. [http: // www. biodiversitylibrary. org / page / 28690228] https: // doi. org / 10.1111 / j. 1096 - 3642.1884. tb 02837. x" type="journal article" year="1884">Jeffreys, 1884</bibRefCitation>
</taxonomicName>
, by monotypy. Present day, Eastern Atlantic;
<taxonomicName id="4C172BEEFFF9C05CFCAAF8C178D33074" authority="Finlay &amp; Marwick, 1937" authorityName="Finlay &amp; Marwick" authorityYear="1937" box="[819,1233,1842,1868]" class="Gastropoda" family="Mathildidae" genus="Kaitangata" kingdom="Animalia" pageId="56" pageNumber="57" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF9C05CFCAAF8C17FB33074" box="[819,945,1843,1868]" italics="true" pageId="56" pageNumber="57">Kaitangata</emphasis>
<bibRefCitation id="EF862D9CFFF9C05CFC21F8C078D33074" author="Finlay, H. J. &amp; Marwick, J." box="[952,1233,1842,1868]" pageId="56" pageNumber="57" pagination="1 - 140" refId="ref52561" refString="Finlay, H. J. &amp; Marwick, J. (1937) The Wangaloan and associated molluscan faunas of Kaitangata-Green Island subdivision. New Zealand geological survey palaeontological bulletin, 15, 1 - 140." type="journal article" year="1937">Finlay &amp; Marwick, 1937</bibRefCitation>
</taxonomicName>
,
<typeStatus id="54ACEECFFFF9C05CFB45F8C1790E3075" box="[1244,1292,1843,1869]" pageId="56" pageNumber="57">type</typeStatus>
species
<taxonomicName id="4C172BEEFFF9C05CFAF3F8C17E913048" authority="Finlay &amp; Marwick, 1937" authorityName="Finlay &amp; Marwick" authorityYear="1937" class="Gastropoda" family="Mathildidae" genus="Kaitangata" kingdom="Animalia" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="hendersoni">
<emphasis id="B9638C7FFFF9C05CFAF3F8C17D723048" italics="true" pageId="56" pageNumber="57">Kaitangata hendersoni</emphasis>
<bibRefCitation id="EF862D9CFFF9C05CFEE1F8A47E913048" author="Finlay, H. J. &amp; Marwick, J." box="[376,659,1878,1904]" pageId="56" pageNumber="57" pagination="1 - 140" refId="ref52561" refString="Finlay, H. J. &amp; Marwick, J. (1937) The Wangaloan and associated molluscan faunas of Kaitangata-Green Island subdivision. New Zealand geological survey palaeontological bulletin, 15, 1 - 140." type="journal article" year="1937">Finlay &amp; Marwick, 1937</bibRefCitation>
</taxonomicName>
, by original designation. Paleocene.
<collectingCountry id="F30010FDFFF9C05CFBA9F8A578C93048" box="[1072,1227,1878,1905]" name="New Zealand" pageId="56" pageNumber="57">New Zealand</collectingCountry>
;
<emphasis id="B9638C7FFFF9C05CFB4FF8A4792A3048" box="[1238,1320,1878,1904]" italics="true" pageId="56" pageNumber="57">Tubena</emphasis>
<bibRefCitation id="EF862D9CFFF9C05CFAA9F8A47CD030AC" author="Marwick, J." pageId="56" pageNumber="57" pagination="181 - 192" refId="ref56837" refString="Marwick, J. (1943) Some Tertiary Mollusca from North Otago. Transactions of the royal society of New Zealand. 73, 181 - 192. [https: // paperspast. natlib. govt. nz / periodicals / TPRSNZ 1943 - 73.2.12.17]" type="journal article" year="1943">Marwick, 1943</bibRefCitation>
,
<typeStatus id="54ACEECFFFF9C05CFF45F8897D0E30AD" box="[220,268,1915,1941]" pageId="56" pageNumber="57">type</typeStatus>
species
<taxonomicName id="4C172BEEFFF9C05CFEF3F8897F2530AC" authority="Marwick, 1943" authorityName="Marwick" authorityYear="1943" box="[362,807,1914,1941]" class="Gastropoda" family="Mathildidae" genus="Gegania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="viola" subGenus="Tubena">
<emphasis id="B9638C7FFFF9C05CFEF3F8897DC830AD" box="[362,458,1915,1941]" italics="true" pageId="56" pageNumber="57">Gegania</emphasis>
(
<emphasis id="B9638C7FFFF9C05CFE43F8887E2E30AC" box="[474,556,1914,1940]" italics="true" pageId="56" pageNumber="57">Tubena</emphasis>
)
<emphasis id="B9638C7FFFF9C05CFDA2F8887E7130AC" box="[571,627,1914,1940]" italics="true" pageId="56" pageNumber="57">viola</emphasis>
<bibRefCitation id="EF862D9CFFF9C05CFDE3F8887F2530AC" author="Marwick, J." box="[634,807,1914,1940]" pageId="56" pageNumber="57" pagination="181 - 192" refId="ref56837" refString="Marwick, J. (1943) Some Tertiary Mollusca from North Otago. Transactions of the royal society of New Zealand. 73, 181 - 192. [https: // paperspast. natlib. govt. nz / periodicals / TPRSNZ 1943 - 73.2.12.17]" type="journal article" year="1943">Marwick, 1943</bibRefCitation>
</taxonomicName>
; original designation by
<bibRefCitation id="EF862D9CFFF9C05CFBA1F888792830AC" author="Marwick, J." box="[1080,1322,1914,1940]" pageId="56" pageNumber="57" pagination="181 - 192" refId="ref56837" refString="Marwick, J. (1943) Some Tertiary Mollusca from North Otago. Transactions of the royal society of New Zealand. 73, 181 - 192. [https: // paperspast. natlib. govt. nz / periodicals / TPRSNZ 1943 - 73.2.12.17]" type="journal article" year="1943">Marwick (1943: 188)</bibRefCitation>
. Miocene,
<collectingCountry id="F30010FDFFF9C05CFF0EF86D7D333080" box="[151,305,1950,1977]" name="New Zealand" pageId="56" pageNumber="57">New Zealand</collectingCountry>
.
</paragraph>
<paragraph id="8BA8506DFFF9C05DFF5EF831799536E9" blockId="56.[151,1437,726,2013]" lastBlockId="57.[151,1437,151,1437]" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
In his revised diagnosis of
<taxonomicName id="4C172BEEFFF9C05CFE76F8307F0030E4" authority=", Bieler (1995: 627)" authorityName=", Bieler" authorityPageNumber="627" authorityYear="1995" box="[495,770,1986,2013]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="56" pageNumber="57" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF9C05CFE76F8307E2530E4" box="[495,551,1986,2012]" italics="true" pageId="56" pageNumber="57">Tuba</emphasis>
,
<bibRefCitation id="EF862D9CFFF9C05CFDADF8307F0030E4" author="Bieler, R." box="[564,770,1986,2013]" pageId="56" pageNumber="57" pagination="595 - 641" refId="ref49179" refString="Bieler, R. (1995) Mathildidae from New Caledonia and the Loyalty Islands (Gastropoda: Heterobranchia). In: Bouchet, P. (Ed.), Resultats des campagnes MUSORSTOM 14. Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie, 167, pp. 595 - 641." type="journal article" year="1995">Bieler (1995: 627)</bibRefCitation>
</taxonomicName>
stated: “
<emphasis id="B9638C7FFFF9C05DFCFAF83078FA37C1" italics="true" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">Teleoconch […] periphery and base rounded (juveniles with double keel) […] 1 or 2 of the main spiral ribs markedly more prominent than the others; […] upper point of attachment of the following whorl at a spiral rib less prominent than at least one of the exposed ribs above; this attachment rib, with an additional rib next to it, forming a distinct double edge at outer shell base.”</emphasis>
The two more prominent primary spiral cords correspond to S2 and S
<quantity id="4CEFFD88FFF8C05DFC9AFEF17F2D3625" box="[771,815,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="57" pageNumber="58" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="4C172BEEFFF8C05DFCAFFEF17FBF3625" authorityName="Dall" authorityYear="1889" box="[822,957,259,285]" class="Gastropoda" family="Mathildidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="57" pageNumber="58" phylum="Mollusca" rank="family">Mathildidae</taxonomicName>
and give the early whorls a somewhat angulated profile. In contrast,
<taxonomicName id="4C172BEEFFF8C05DFE52FED57E4D3679" authorityName="Harzhauser &amp; Landau" authorityYear="2023" box="[459,591,295,321]" class="Gastropoda" family="Mathildidae" genus="Pseudotuba" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="57" pageNumber="58" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF8C05DFE52FED57E4D3679" box="[459,591,295,321]" italics="true" pageId="57" pageNumber="58">Pseudotuba</emphasis>
</taxonomicName>
develops six primary cords that are present from the teleoconch/protoconch boundary and are not, or hardly, differentiated. Thus, the early whorls are not angulated. In addition, the base of
<taxonomicName id="4C172BEEFFF8C05DFF0EFE9D7D1936B1" authorityName="Harzhauser &amp; Landau" authorityYear="2023" box="[151,283,367,393]" class="Gastropoda" family="Mathildidae" genus="Pseudotuba" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="57" pageNumber="58" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF8C05DFF0EFE9D7D1936B1" box="[151,283,367,393]" italics="true" pageId="57" pageNumber="58">Pseudotuba</emphasis>
</taxonomicName>
is rounded and lacks a “
<emphasis id="B9638C7FFFF8C05DFDBEFE9D7EAC36B1" box="[551,686,367,393]" italics="true" pageId="57" pageNumber="58">double edge</emphasis>
”. Lastly,
<taxonomicName id="4C172BEEFFF8C05DFC81FE9D7F9E36B1" authorityName="Harzhauser &amp; Landau" authorityYear="2023" box="[792,924,367,393]" class="Gastropoda" family="Mathildidae" genus="Pseudotuba" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="57" pageNumber="58" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF8C05DFC81FE9D7F9E36B1" box="[792,924,367,393]" italics="true" pageId="57" pageNumber="58">Pseudotuba</emphasis>
</taxonomicName>
has a hyperstrophic rather than heterostrophic protoconch. In our opinion, the consistent difference in the protoconch and different number and development of the primary cords on the early whorls calls into question a close relation between
<taxonomicName id="4C172BEEFFF8C05DFB99FE45788636E9" authorityName="Harzhauser &amp; Landau" authorityYear="2023" box="[1024,1156,439,465]" class="Gastropoda" family="Mathildidae" genus="Pseudotuba" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="57" pageNumber="58" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF8C05DFB99FE45788636E9" box="[1024,1156,439,465]" italics="true" pageId="57" pageNumber="58">Pseudotuba</emphasis>
</taxonomicName>
and
<taxonomicName id="4C172BEEFFF8C05DFB22FE4578F136E9" authorityName=", Bieler" authorityYear="1995" box="[1211,1267,439,465]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF8C05DFB22FE4578F136E9" box="[1211,1267,439,465]" italics="true" pageId="57" pageNumber="58">Tuba</emphasis>
</taxonomicName>
and
<taxonomicName id="4C172BEEFFF8C05DFAB0FE45799236E9" authorityName="O. Semper" authorityYear="1865" box="[1321,1424,439,465]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="57" pageNumber="58" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF8C05DFAB0FE45799236E9" box="[1321,1424,439,465]" italics="true" pageId="57" pageNumber="58">Mathilda</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA8506DFFF8C05DFF5EFE287E463329" blockId="57.[151,1437,151,1437]" pageId="57" pageNumber="58">
<emphasis id="B9638C7FFFF8C05DFF5EFE287DD236CD" bold="true" box="[199,464,474,501]" pageId="57" pageNumber="58">
Fossil species of
<taxonomicName id="4C172BEEFFF8C05DFE17FE287DC836CD" authorityName=", Bieler" authorityYear="1995" box="[398,458,474,501]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF8C05DFE17FE287DC836CD" bold="true" box="[398,458,474,501]" italics="true" pageId="57" pageNumber="58">Tuba</emphasis>
</taxonomicName>
.
</emphasis>
Currently, MolluscaBase eds. (2023) lists 16 fossil and extant species in
<taxonomicName id="4C172BEEFFF8C05DFA88FE29794B36CD" authorityName=", Bieler" authorityYear="1995" box="[1297,1353,475,501]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF8C05DFA88FE29794B36CD" box="[1297,1353,475,501]" italics="true" pageId="57" pageNumber="58">Tuba</emphasis>
</taxonomicName>
. Along with the
<typeStatus id="54ACEECFFFF8C05DFF60FE0D7D2B3521" box="[249,297,511,537]" pageId="57" pageNumber="58">type</typeStatus>
species of
<taxonomicName id="4C172BEEFFF8C05DFE3FFE0D7E043521" authorityName="Jeffreys" authorityYear="1884" box="[422,518,511,537]" class="Gastropoda" family="Mathildidae" genus="Gegania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF8C05DFE3FFE0D7E043521" box="[422,518,511,537]" italics="true" pageId="57" pageNumber="58">Gegania</emphasis>
</taxonomicName>
,
<taxonomicName id="4C172BEEFFF8C05DFD8DFE0D7F5D3521" authority="(Jeffreys, 1884)" baseAuthorityName="Jeffreys" baseAuthorityYear="1884" box="[532,863,511,537]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="pinguis">
<emphasis id="B9638C7FFFF8C05DFD8DFE0D7EA73520" box="[532,677,511,537]" italics="true" pageId="57" pageNumber="58">Tuba pinguis</emphasis>
(
<bibRefCitation id="EF862D9CFFF8C05DFD2DFE0D7F5A3521" author="Jeffreys, J. G." box="[692,856,511,537]" pageId="57" pageNumber="58" pagination="341 - 372" refId="ref55069" refString="Jeffreys, J. G. (1884) On the Mollusca procured during the ' Lightning' and ' Porcupine' expeditions, 1868 - 70. (Part VIII). Proceedings of the zoological society of London, 1882, 341 - 372, pls. 26 - 28. [http: // www. biodiversitylibrary. org / page / 28690228] https: // doi. org / 10.1111 / j. 1096 - 3642.1884. tb 02837. x" type="journal article" year="1884">Jeffreys, 1884</bibRefCitation>
)
</taxonomicName>
, the Recent
<taxonomicName id="4C172BEEFFF8C05DFC71FE0D79323521" authority="(Powell, 1971)" baseAuthorityName="Powell" baseAuthorityYear="1971" box="[1000,1328,511,537]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="valkyrie">
<emphasis id="B9638C7FFFF8C05DFC71FE0D78833521" box="[1000,1153,511,537]" italics="true" pageId="57" pageNumber="58">Tuba valkyrie</emphasis>
(Powell, 1971)
</taxonomicName>
,
<taxonomicName id="4C172BEEFFF8C05DFAA5FE0D7D723505" authority="Bieler, 1995" authorityName="Bieler" authorityYear="1995" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="fuscocincta">
<emphasis id="B9638C7FFFF8C05DFAA5FE0D7CD93504" italics="true" pageId="57" pageNumber="58">T. fuscocincta</emphasis>
<bibRefCitation id="EF862D9CFFF8C05DFF7AFDD17D723505" author="Bieler, R." box="[227,368,546,573]" pageId="57" pageNumber="58" pagination="595 - 641" refId="ref49179" refString="Bieler, R. (1995) Mathildidae from New Caledonia and the Loyalty Islands (Gastropoda: Heterobranchia). In: Bouchet, P. (Ed.), Resultats des campagnes MUSORSTOM 14. Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie, 167, pp. 595 - 641." type="journal article" year="1995">Bieler, 1995</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C172BEEFFF8C05DFEE2FDD17E833505" authority="(Dall, 1889)" baseAuthorityName="Dall" baseAuthorityYear="1889" box="[379,641,547,573]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="jeffreysi">
<emphasis id="B9638C7FFFF8C05DFEE2FDD17DF23505" box="[379,496,547,573]" italics="true" pageId="57" pageNumber="58">T. jeffreysi</emphasis>
(
<bibRefCitation id="EF862D9CFFF8C05DFD99FDD17E7A3505" author="Dall, W. H." box="[512,632,547,573]" pageId="57" pageNumber="58" pagination="1 - 492" refId="ref51269" refString="Dall, W. H. (1889) Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. Coast Survey Steamer &quot; Blake &quot;, Lieut. - Commander C. D. Sigsbee, U. S. N., and Commander J. R. Bartlett, U. S. N., commanding. XXIX. Report on the Mollusca. Part 2, Gastropoda and Scaphopoda. Bulletin of the museum of comparative zoology at Harvard College, 18, 1 - 492. [http: // www. biodiversitylibrary. org / item / 25505]" type="journal article" year="1889">Dall, 1889</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C172BEEFFF8C05DFD21FDD178223505" authority="(T. Nakayama, 2000)" baseAuthorityName="T. Nakayama" baseAuthorityYear="2000" box="[696,1056,547,573]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="kiiensis">
<emphasis id="B9638C7FFFF8C05DFD21FDD17F2A3505" box="[696,808,547,573]" italics="true" pageId="57" pageNumber="58">T. kiiensis</emphasis>
(T. Nakayama, 2000)
</taxonomicName>
, represent species with differentiated spiral cords on early teleoconch whorls. Of the fossil species, along with the
<typeStatus id="54ACEECFFFF8C05DFBAEFDB578653559" box="[1079,1127,583,609]" pageId="57" pageNumber="58">type</typeStatus>
species
<taxonomicName id="4C172BEEFFF8C05DFB51FDB57CCD35BD" authority="I. Lea, 1833" authorityName="I. Lea" authorityYear="1833" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="alternata">
<emphasis id="B9638C7FFFF8C05DFB51FDB579493559" box="[1224,1355,583,609]" italics="true" pageId="57" pageNumber="58">T. alternata</emphasis>
I.
<bibRefCitation id="EF862D9CFFF8C05DFAF2FDB57CCD35BD" author="Lea, I." pageId="57" pageNumber="58" refId="ref56283" refString="Lea, I. (1833) Contributions to Geology. Carey, Lea and Blanchard, Philadelphia, Pennsylvania, 227 pp., 6 pls. [https: // www. biodiversitylibrary. org / page / 17592495]" type="book" year="1833">Lea, 1833</bibRefCitation>
</taxonomicName>
(Eocene,
<collectingCountry id="F30010FDFFF8C05DFEA6FD997D7935BD" box="[319,379,619,645]" name="United States of America" pageId="57" pageNumber="58">USA</collectingCountry>
), we retain
<taxonomicName id="4C172BEEFFF8C05DFE63FD997FBD35BD" authority="(Finlay &amp; Marwick, 1937)" baseAuthorityName="Finlay &amp; Marwick" baseAuthorityYear="1937" box="[506,959,619,645]" class="Gastropoda" family="Mathildidae" genus="Kaitangata" kingdom="Animalia" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="hendersoni">
<emphasis id="B9638C7FFFF8C05DFE63FD997E9335BD" box="[506,657,619,645]" italics="true" pageId="57" pageNumber="58">T. hendersoni</emphasis>
(
<bibRefCitation id="EF862D9CFFF8C05DFD39FD997FB535BD" author="Finlay, H. J. &amp; Marwick, J." box="[672,951,619,645]" pageId="57" pageNumber="58" pagination="1 - 140" refId="ref52561" refString="Finlay, H. J. &amp; Marwick, J. (1937) The Wangaloan and associated molluscan faunas of Kaitangata-Green Island subdivision. New Zealand geological survey palaeontological bulletin, 15, 1 - 140." type="journal article" year="1937">Finlay &amp; Marwick, 1937</bibRefCitation>
)
</taxonomicName>
(Paleocene,
<collectingCountry id="F30010FDFFF8C05DFBD7FD9978EA35BD" box="[1102,1256,619,645]" name="New Zealand" pageId="57" pageNumber="58">New Zealand</collectingCountry>
),
<taxonomicName id="4C172BEEFFF8C05DFB63FD997D493591" authority="(P. A. Maxwell, 1969)" baseAuthorityName="P. A. Maxwell" baseAuthorityYear="1969" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="olsoni">
<emphasis id="B9638C7FFFF8C05DFB63FD99795B35BD" box="[1274,1369,619,645]" italics="true" pageId="57" pageNumber="58">T. olsoni</emphasis>
(P. A. Maxwell, 1969)
</taxonomicName>
(Eocene,
<collectingCountry id="F30010FDFFF8C05DFE27FD7D7E553591" box="[446,599,655,681]" name="New Zealand" pageId="57" pageNumber="58">New Zealand</collectingCountry>
) and
<taxonomicName id="4C172BEEFFF8C05DFD0EFD7D7FB13591" authority="(Marwick, 1943)" baseAuthorityName="Marwick" baseAuthorityYear="1943" box="[663,947,655,681]" class="Gastropoda" family="Mathildidae" genus="Gegania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="viola">
<emphasis id="B9638C7FFFF8C05DFD0EFD7D7EEE3591" box="[663,748,655,681]" italics="true" pageId="57" pageNumber="58">T. viola</emphasis>
(
<bibRefCitation id="EF862D9CFFF8C05DFD65FD7D7FA93591" author="Marwick, J." box="[764,939,655,681]" pageId="57" pageNumber="58" pagination="181 - 192" refId="ref56837" refString="Marwick, J. (1943) Some Tertiary Mollusca from North Otago. Transactions of the royal society of New Zealand. 73, 181 - 192. [https: // paperspast. natlib. govt. nz / periodicals / TPRSNZ 1943 - 73.2.12.17]" type="journal article" year="1943">Marwick, 1943</bibRefCitation>
)
</taxonomicName>
(Miocene,
<collectingCountry id="F30010FDFFF8C05DFBAFFD7D78CD3591" box="[1078,1231,655,681]" name="New Zealand" pageId="57" pageNumber="58">New Zealand</collectingCountry>
) in
<taxonomicName id="4C172BEEFFF8C05DFB65FD7D79363591" authorityName=", Bieler" authorityYear="1995" box="[1276,1332,655,681]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF8C05DFB65FD7D79363591" box="[1276,1332,655,681]" italics="true" pageId="57" pageNumber="58">Tuba</emphasis>
</taxonomicName>
. In addition,
<taxonomicName id="4C172BEEFFF8C05DFF49FD417DC335F5" authority="I. Lea, 1833" authorityName="I. Lea" authorityYear="1833" box="[208,449,691,717]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="striata">
<emphasis id="B9638C7FFFF8C05DFF49FD417D3635F4" box="[208,308,691,716]" italics="true" pageId="57" pageNumber="58">T. striata</emphasis>
I.
<bibRefCitation id="EF862D9CFFF8C05DFEC8FD417DC335F5" author="Lea, I." box="[337,449,691,717]" pageId="57" pageNumber="58" refId="ref56283" refString="Lea, I. (1833) Contributions to Geology. Carey, Lea and Blanchard, Philadelphia, Pennsylvania, 227 pp., 6 pls. [https: // www. biodiversitylibrary. org / page / 17592495]" type="book" year="1833">Lea, 1833</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C172BEEFFF8C05DFE6FFD417F3235F5" authority="(Conrad, 1833)" baseAuthorityName="Conrad" baseAuthorityYear="1833" box="[502,816,691,717]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="antiquata">
<emphasis id="B9638C7FFFF8C05DFE6FFD417E7E35F4" box="[502,636,691,716]" italics="true" pageId="57" pageNumber="58">T. antiquata</emphasis>
(
<bibRefCitation id="EF862D9CFFF8C05DFD12FD417F2535F5" author="Conrad, T. A." box="[651,807,691,717]" pageId="57" pageNumber="58" pagination="339 - 346" refId="ref50194" refString="Conrad, T. A. (1833) On some new fossil and Recent shells of the United States. The American journal of sciences and arts, 23 (2), 339 - 346. [https: // www. biodiversitylibrary. org / item / 97166 # page / 357 / mode / 1 up]" type="journal article" year="1833">Conrad, 1833</bibRefCitation>
)
</taxonomicName>
, both from the Eocene of the
<collectingCountry id="F30010FDFFF8C05DFBE1FD4178B135F5" box="[1144,1203,691,717]" name="United States of America" pageId="57" pageNumber="58">USA</collectingCountry>
, and
<taxonomicName id="4C172BEEFFF8C05DFB75FD417D2E35C9" authority="Olsson, 1930" authorityName="Olsson" authorityYear="1930" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="peruviana">
<emphasis id="B9638C7FFFF8C05DFB75FD41799F35F4" box="[1260,1437,691,717]" italics="true" pageId="57" pageNumber="58">Tuba peruviana</emphasis>
Olsson, 1930
</taxonomicName>
(Eocene,
<collectingCountry id="F30010FDFFF8C05DFE06FD257DD635C9" box="[415,468,727,753]" name="Peru" pageId="57" pageNumber="58">Peru</collectingCountry>
) are placed in
<taxonomicName id="4C172BEEFFF8C05DFDE4FD257EB735C9" authorityName=", Bieler" authorityYear="1995" box="[637,693,727,753]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF8C05DFDE4FD257EB735C9" box="[637,693,727,753]" italics="true" pageId="57" pageNumber="58">Tuba</emphasis>
</taxonomicName>
herein.
<taxonomicName id="4C172BEEFFF8C05DFC8BFD25794D35C9" authority="(Stilwell &amp; Zinsmeister, 1992)" baseAuthorityName="Stilwell &amp; Zinsmeister" baseAuthorityYear="1992" box="[786,1359,727,753]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="antarctodema">
<emphasis id="B9638C7FFFF8C05DFC8BFD257FEC35C9" box="[786,1006,727,753]" italics="true" pageId="57" pageNumber="58">Tuba antarctodema</emphasis>
(Stilwell &amp; Zinsmeister, 1992)
</taxonomicName>
and
<taxonomicName id="4C172BEEFFF8C05DFA11FD257E9B342D" authority="(Stilwell &amp; Zinsmeister, 1992)" baseAuthorityName="Stilwell &amp; Zinsmeister" baseAuthorityYear="1992" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="austroconvexa">
<emphasis id="B9638C7FFFF8C05DFA11FD257D3B342C" italics="true" pageId="57" pageNumber="58">T. austroconvexa</emphasis>
(Stilwell &amp; Zinsmeister, 1992)
</taxonomicName>
, might be two additional
<taxonomicName id="4C172BEEFFF8C05DFC2FFD09784B342D" box="[950,1097,763,789]" pageId="57" pageNumber="58">
<emphasis id="B9638C7FFFF8C05DFC2FFD097FEC342D" box="[950,1006,763,789]" italics="true" pageId="57" pageNumber="58">Tuba</emphasis>
species
</taxonomicName>
, documented from the Eocene of
<collectingCountry id="F30010FDFFF8C05DFF2DFCED7D2E3401" box="[180,300,799,825]" name="Antarctica" pageId="57" pageNumber="58">Antarctica</collectingCountry>
, but we have not seen specimens. Oligocene and Miocene
<taxonomicName id="4C172BEEFFF8C05DFC5BFCED78503401" box="[962,1106,799,825]" pageId="57" pageNumber="58">
<emphasis id="B9638C7FFFF8C05DFC5BFCED7FF83401" box="[962,1018,799,825]" italics="true" pageId="57" pageNumber="58">Tuba</emphasis>
species
</taxonomicName>
from the Proto-Mediterraean Sea are:
<taxonomicName id="4C172BEEFFF8C05DFF61FCB17E553465" authority="Sacco, 1895" authorityName="Sacco" authorityYear="1895" box="[248,599,834,861]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="alternicincta">
<emphasis id="B9638C7FFFF8C05DFF61FCB17DC43465" box="[248,454,835,861]" italics="true" pageId="57" pageNumber="58">Tuba alternicincta</emphasis>
Sacco, 1895
</taxonomicName>
(Rupelian, Colli Torinesi,
<collectingCountry id="F30010FDFFF8C05DFC1AFCB17FBA3465" box="[899,952,835,861]" name="Italy" pageId="57" pageNumber="58">Italy</collectingCountry>
) (Sacco 1895: pl. 3, fig. 44);
<taxonomicName id="4C172BEEFFF8C05DFA9AFCB17D6834B9" authority="(Sacco, 1895)" baseAuthorityName="Sacco" baseAuthorityYear="1895" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="miocaenica">
<emphasis id="B9638C7FFFF8C05DFA9AFCB17CC534B8" italics="true" pageId="57" pageNumber="58">Tuba miocaenica</emphasis>
(Sacco, 1895)
</taxonomicName>
(Langhian, Colli Torinesi,
<collectingCountry id="F30010FDFFF8C05DFD02FC957ED234B9" box="[667,720,871,897]" name="Italy" pageId="57" pageNumber="58">Italy</collectingCountry>
) (Sacco 1895: pl. 3, figs 4850;
<bibRefCitation id="EF862D9CFFF8C05DFBA6FC95797634B9" author="Ferrero Mortara, E. &amp; Montefameglio &amp; Novelli, M. &amp; Opesso, G. &amp; Pavia, G. &amp; Tampieri, R." box="[1087,1396,871,897]" pageId="57" pageNumber="58" pagination="1 - 489" refId="ref52430" refString="Ferrero Mortara, E., Montefameglio, Novelli, M., Opesso, G., Pavia, G. &amp; Tampieri, R. (1984) Catalogo dei tipi e degli esemplari figurati della collezione Bellardi e Sacco. Parte II. Museo regionale di scienze naturali, catalogo, 7, 1 - 489. [https: // www. biodiversitylibrary. org / item / 269689 # page / 5 / mode / 1 up]" type="journal article" year="1984">
Ferrero Mortara
<emphasis id="B9638C7FFFF8C05DFB60FC95793034B9" box="[1273,1330,871,897]" italics="true" pageId="57" pageNumber="58">et al.</emphasis>
1984
</bibRefCitation>
; pl. 41, figs 7, 9);
<taxonomicName id="4C172BEEFFF8C05DFEA2FC797EB2349D" authority="(Sacco, 1895)" baseAuthorityName="Sacco" baseAuthorityYear="1895" box="[315,688,906,933]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="percingulata">
<emphasis id="B9638C7FFFF8C05DFEA2FC797E08349D" box="[315,522,907,933]" italics="true" pageId="57" pageNumber="58">Tuba percingulata</emphasis>
(Sacco, 1895)
</taxonomicName>
(Burdigalian, Colli Torinesi,
<collectingCountry id="F30010FDFFF8C05DFB98FC797834349D" box="[1025,1078,907,933]" name="Italy" pageId="57" pageNumber="58">Italy</collectingCountry>
) (Sacco 1895: pl. 3, fig. 51). A further undescribed species of
<taxonomicName id="4C172BEEFFF8C05DFE68FC5D7E7734F1" authorityName="Harzhauser &amp; Landau" authorityYear="2023" box="[497,629,943,969]" class="Gastropoda" family="Mathildidae" genus="Pseudotuba" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="57" pageNumber="58" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF8C05DFE68FC5D7E7734F1" box="[497,629,943,969]" italics="true" pageId="57" pageNumber="58">Pseudotuba</emphasis>
</taxonomicName>
from the Upper Miocene Tortonian of Cacela Velha, southern
<collectingCountry id="F30010FDFFF8C05DFAAFFC5D799A34F1" box="[1334,1432,943,969]" name="Portugal" pageId="57" pageNumber="58">Portugal</collectingCountry>
, is present in the NHMW coll. (ex. BL coll.) that closes the geographic gap between the Miocene French Atlantic and Proto-Mediterranean occurrences.
</paragraph>
<paragraph id="8BA8506DFFF8C05DFF5EFBE97E763399" blockId="57.[151,1437,151,1437]" pageId="57" pageNumber="58">
<emphasis id="B9638C7FFFF8C05DFF5EFBE97DA0330D" box="[199,418,1051,1077]" italics="true" pageId="57" pageNumber="58">Gegania kuteiana</emphasis>
(
<bibRefCitation id="EF862D9CFFF8C05DFE2DFBE97E43330D" author="Beets, C." box="[436,577,1051,1077]" pageId="57" pageNumber="58" pagination="218 - 254" refId="ref48864" refString="Beets, C. (1942) Mollusken aus dem Tertiar des Ostindischen Archipels. Leidsche Geologische Mededeelingen, 13, 218 - 254. [https: // repository. naturalis. nl / pub / 505999]" type="journal article" year="1942">Beets, 1942</bibRefCitation>
), from the Miocene of Borneo, should be excluded from
<taxonomicName id="4C172BEEFFF8C05DFB7BFBE9796B330D" authorityName="Dall" authorityYear="1889" box="[1250,1385,1051,1077]" class="Gastropoda" family="Mathildidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="57" pageNumber="58" phylum="Mollusca" rank="family">Mathildidae</taxonomicName>
and might rather represent a
<taxonomicName id="4C172BEEFFF8C05DFE3FFBCD7E493361" authorityName="Conrad" authorityYear="1860" box="[422,587,1087,1113]" class="Gastropoda" family="Capulidae" genus="Cerithioderma" kingdom="Animalia" order="Littorinimorpha" pageId="57" pageNumber="58" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF8C05DFE3FFBCD7E493361" box="[422,587,1087,1113]" italics="true" pageId="57" pageNumber="58">Cerithioderma</emphasis>
</taxonomicName>
(
<taxonomicName id="4C172BEEFFF8C05DFDC0FBCD7ECC3361" authorityName="Fleming" authorityYear="1822" box="[601,718,1087,1113]" class="Gastropoda" family="Capulidae" kingdom="Animalia" order="Littorinimorpha" pageId="57" pageNumber="58" phylum="Mollusca" rank="family">Capulidae</taxonomicName>
) (see
<bibRefCitation id="EF862D9CFFF8C05DFC96FBCD7F8C3361" author="Beets, C." box="[783,910,1087,1113]" pageId="57" pageNumber="58" pagination="218 - 254" refId="ref48864" refString="Beets, C. (1942) Mollusken aus dem Tertiar des Ostindischen Archipels. Leidsche Geologische Mededeelingen, 13, 218 - 254. [https: // repository. naturalis. nl / pub / 505999]" type="journal article" year="1942">Beets 1942</bibRefCitation>
; pl. 16, figs 2123). Similarly,
<taxonomicName id="4C172BEEFFF8C05DFB7FFBCD7D533345" authority="W. H. Dall, 1892" authorityName="W. H. Dall" authorityYear="1892" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="acutissima">
<emphasis id="B9638C7FFFF8C05DFB7FFBCD799E3360" box="[1254,1436,1087,1113]" italics="true" pageId="57" pageNumber="58">Tuba acutissima</emphasis>
W.H.
<bibRefCitation id="EF862D9CFFF8C05DFF41FB917D533345" author="Dall, W. H." box="[216,337,1123,1149]" pageId="57" pageNumber="58" pagination="201 - 473" refId="ref51405" refString="Dall, W. H. (1892) Contributions to the Tertiary fauna of Florida, with especial reference to the Miocene silex-beds of Tampa and the Pliocene beds of the Caloosahatchie River. Part II. Streptodont and other gastropods, concluded. Transactions of the Wagner free institute of science of Philadelphia, 3 (2), 201 - 473, pls. 13 - 22. [https: // www. biodiversitylibrary. org / item / 98240 # page / 211 / mode / 1 up]" type="journal article" year="1892">Dall, 1892</bibRefCitation>
</taxonomicName>
, from the Miocene of
<collectingRegion id="49D39E8FFFF8C05DFDD3FB917E9C3345" box="[586,670,1123,1149]" country="United States of America" name="Florida" pageId="57" pageNumber="58">Florida</collectingRegion>
, should be excluded from
<taxonomicName id="4C172BEEFFF8C05DFC5FFB9178523345" authorityName="Dall" authorityYear="1889" box="[966,1104,1123,1149]" class="Gastropoda" family="Mathildidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="57" pageNumber="58" phylum="Mollusca" rank="family">Mathildidae</taxonomicName>
, due to its non-heterostrophic protoconch (see
<bibRefCitation id="EF862D9CFFF8C05DFEC9FB757DC13399" author="Dall, W. H." box="[336,451,1159,1185]" pageId="57" pageNumber="58" pagination="201 - 473" refId="ref51405" refString="Dall, W. H. (1892) Contributions to the Tertiary fauna of Florida, with especial reference to the Miocene silex-beds of Tampa and the Pliocene beds of the Caloosahatchie River. Part II. Streptodont and other gastropods, concluded. Transactions of the Wagner free institute of science of Philadelphia, 3 (2), 201 - 473, pls. 13 - 22. [https: // www. biodiversitylibrary. org / item / 98240 # page / 211 / mode / 1 up]" type="journal article" year="1892">Dall 1892</bibRefCitation>
: pl. 18, fig. 4a).
</paragraph>
<paragraph id="8BA8506DFFF8C05DFF5EFB597DDD32A5" blockId="57.[151,1437,151,1437]" pageId="57" pageNumber="58">
<taxonomicName id="4C172BEEFFF8C05DFF5EFB597CFD33FD" authorityName=", Bieler" authorityYear="1995" box="[199,255,1195,1221]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF8C05DFF5EFB597CFD33FD" box="[199,255,1195,1221]" italics="true" pageId="57" pageNumber="58">Tuba</emphasis>
</taxonomicName>
might have appeared already during the Late Cretaceous and is documented by numerous species;
<emphasis id="B9638C7FFFF8C05DFAF4FB59799733FC" box="[1389,1429,1195,1220]" italics="true" pageId="57" pageNumber="58">e.g.</emphasis>
,
<taxonomicName id="4C172BEEFFF8C05DFF0EFB3D7E3133D1" authority="d'Orbigny 1843" authorityName="d'Orbigny" authorityYear="1843" box="[151,563,1231,1257]" class="Gastropoda" family="Mathildidae" genus="Gegania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="vibrayeana">
<emphasis id="B9638C7FFFF8C05DFF0EFB3D7D7933D1" box="[151,379,1231,1257]" italics="true" pageId="57" pageNumber="58">Gegania vibrayeana</emphasis>
dOrbigny 1843
</taxonomicName>
(
<collectingCountry id="F30010FDFFF8C05DFDD9FB3D7E9033D1" box="[576,658,1231,1257]" name="France" pageId="57" pageNumber="58">France</collectingCountry>
),
<taxonomicName id="4C172BEEFFF8C05DFD3DFB3D7FB733D1" authority="Conrad, 1860" authorityName="Conrad" authorityYear="1860" box="[676,949,1231,1257]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="bella">
<emphasis id="B9638C7FFFF8C05DFD3DFB3D7F1B33D1" box="[676,793,1231,1257]" italics="true" pageId="57" pageNumber="58">Tuba bella</emphasis>
<bibRefCitation id="EF862D9CFFF8C05DFC86FB3D7FB733D1" author="Conrad, T. A." box="[799,949,1231,1257]" pageId="57" pageNumber="58" pagination="275 - 298" refId="ref50255" refString="Conrad, T. A. (1860) Description of new species of Cretaceous and Eocene fossils of Mississippi and Alabama. Journal of the academy of natural sciences of Philadelphia, Series 2, 4, 275 - 298. [https: // www. biodiversitylibrary. org / page / 36862163]" type="journal article" year="1860">Conrad, 1860</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F30010FDFFF8C05DFC5DFB3D780233D1" box="[964,1024,1231,1257]" name="United States of America" pageId="57" pageNumber="58">USA</collectingCountry>
),
<taxonomicName id="4C172BEEFFF8C05DFB8BFB3D794733D1" authority="Wade, 1926" authorityName="Wade" authorityYear="1926" box="[1042,1349,1231,1257]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="parabella">
<emphasis id="B9638C7FFFF8C05DFB8BFB3D78BF33D1" box="[1042,1213,1231,1257]" italics="true" pageId="57" pageNumber="58">Tuba parabella</emphasis>
Wade, 1926
</taxonomicName>
(
<collectingCountry id="F30010FDFFF8C05DFACDFB3D799233D1" box="[1364,1424,1231,1257]" name="United States of America" pageId="57" pageNumber="58">USA</collectingCountry>
),
<taxonomicName id="4C172BEEFFF8C05DFF0EFB017E173235" authority="Stephenson, 1941" authorityName="Stephenson" authorityYear="1941" box="[151,533,1267,1293]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="manzaneti">
<emphasis id="B9638C7FFFF8C05DFF0EFB017D4A3234" box="[151,328,1267,1293]" italics="true" pageId="57" pageNumber="58">Tuba manzaneti</emphasis>
Stephenson, 1941
</taxonomicName>
(
<collectingCountry id="F30010FDFFF8C05DFDBCFB017E633235" box="[549,609,1267,1293]" name="United States of America" pageId="57" pageNumber="58">USA</collectingCountry>
),
<taxonomicName id="4C172BEEFFF8C05DFDEAFB0178383235" authority="Dockery, 1993" authorityName="Dockery" authorityYear="1993" box="[627,1082,1267,1293]" class="Gastropoda" family="Mathildidae" genus="Gegania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="mississippiensis">
<emphasis id="B9638C7FFFF8C05DFDEAFB017F8F3234" box="[627,909,1267,1293]" italics="true" pageId="57" pageNumber="58">Gegania mississippiensis</emphasis>
<bibRefCitation id="EF862D9CFFF8C05DFC0AFB0178383235" author="Dockery, D. T. III" box="[915,1082,1267,1293]" pageId="57" pageNumber="58" pagination="1 - 191" refId="ref52043" refString="Dockery, D. T. III (1993) The streptoneuran gastropods, exclusive of the Stenoglossa, of the Coffee Sand (Campanian) of northeastern Mississippi. Mississippi department of environmental quality, office of Geology, Bulletin, 129, 1 - 191." type="journal article" year="1993">Dockery, 1993</bibRefCitation>
</taxonomicName>
. The placement of these species in
<taxonomicName id="4C172BEEFFF8C05DFF2AFAE57CE93209" authorityName=", Bieler" authorityYear="1995" box="[179,235,1303,1329]" class="Gastropoda" family="Mathildidae" genus="Tuba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="57" pageNumber="58" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF8C05DFF2AFAE57CE93209" box="[179,235,1303,1329]" italics="true" pageId="57" pageNumber="58">Tuba</emphasis>
</taxonomicName>
, however, will need verification because they represent large and solid shells with conical spires, partly with nearly turreted early teleoconch whorls (
<emphasis id="B9638C7FFFF8C05DFDC7FAC97E84326C" box="[606,646,1339,1364]" italics="true" pageId="57" pageNumber="58">e.g.</emphasis>
,
<bibRefCitation id="EF862D9CFFF8C05DFD0DFAC97F45326D" author="Kollmann, H. A." box="[660,839,1338,1365]" pageId="57" pageNumber="58" pagination="156 - 7" refId="ref55374" refString="Kollmann, H. A. (2005) Revision critique de la Paleontologie francaise d'Alcide d'Orbigny. Volume 3: Gastropodes cretaces (translated and adapted by Jean-Claude Fischer). Backhuys, Leiden, 239 pp. + reprint of the corresponding volume (Terrains cretaces, Gastropodes) of Paleontologie francaise by Alcide d'Orbigny (1842 - 1843) + ii + 454 pp., pls. 149 - 236. [ISBN 90 - 5782 - 156 - 7]" type="book chapter" year="2005">Kollmann 2005</bibRefCitation>
: pl. 17 figs 11ab, https://science.mnhn.fr/institution/ mnhn/collection/f/item/s07143?listIndex=8&amp;listCount=19). The depositional setting of most of these species seems to have been shallow marine.
</paragraph>
</subSubSection>
</treatment>
</document>