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<document id="F59B3622FE81434D3BA731937DDDC209" ID-DOI="10.11646/zootaxa.5370.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10147814" ID-ZooBank="48903495-7C6C-46E4-9B1B-D6A2F2781873" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1700202702953" checkinUser="plazi" docAuthor="Harzhauser, Mathias &amp; Landau, Bernard" docDate="2023" docId="03BEE17BFFF5C051FF0EF94D78E83559" docLanguage="en" docName="zootaxa.5370.1.1.pdf" docOrigin="Zootaxa 5370 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5370.1.1/52270" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Fimbriatella Sacco 1895" docType="treatment" docVersion="1" lastPageNumber="54" masterDocId="FF879903FFC1C064FF99FFF27C023738" masterDocTitle="The Architectonicidae and Mathildidae (Gastropoda, Heterobranchia) of the Miocene Paratethys Sea-victims of the Miocene Climatic Transition" masterLastPageNumber="74" masterPageNumber="1" pageNumber="53" updateTime="1706539831396" updateUser="carolina">
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<mods:titleInfo id="0B99F510CBE2CA3FFBBAADB59BAAEC34">
<mods:title id="1E150FD590DF667F499C363D9F239E81">The Architectonicidae and Mathildidae (Gastropoda, Heterobranchia) of the Miocene Paratethys Sea-victims of the Miocene Climatic Transition</mods:title>
</mods:titleInfo>
<mods:name id="C2C85EAB67DD73B224FAA556BE1C1970" type="personal">
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<mods:namePart id="AA2ADD589DD3F250F772FE5BB2EA9342">Harzhauser, Mathias</mods:namePart>
<mods:nameIdentifier id="EB0031D5D25D8FBE1DE5158B5877510D" type="ORCID">0000-0002-4471-6655</mods:nameIdentifier>
<mods:affiliation id="1130D42921B8084B7ED75DEA384174D9">Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655;</mods:affiliation>
<mods:nameIdentifier id="502946461EF0B0FE5820F3CAC88E3217" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
</mods:name>
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<mods:roleTerm id="A52C7CB41E6D8E1352AF6A5D2CC58326">Author</mods:roleTerm>
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<mods:namePart id="7A0FC654769C349DC8D5414C3A068992">Landau, Bernard</mods:namePart>
<mods:nameIdentifier id="217862F9428E6CAE906E4C9647E7A86D" type="ORCID">0000-0002-4471-6655</mods:nameIdentifier>
<mods:affiliation id="2EE3C75D56ADECC8F6D1B668C21A5F42">Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655; &amp; Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal. bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494 &amp; Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655;</mods:affiliation>
<mods:nameIdentifier id="94B05B8B688CF2258FF493F8FB6FD22D" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="0810369422855E21077CE8C5740BB2D0">text</mods:typeOfResource>
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<mods:title id="CE39CF7407470CCF9F91376B2B5ABCAF">Zootaxa</mods:title>
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<mods:date id="7B88847CB58064EEBC4C297FBD249BCB">2023</mods:date>
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<mods:number id="307620FF26131487EB54295118437474">2023-11-14</mods:number>
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<mods:number id="B6B50396DFD203B440479F6AF0B5A8B3">5370</mods:number>
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<mods:number id="0577AF6E3CE1DA94A1BD8F3224661826">1</mods:number>
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<mods:start id="975E251202FF1CC10CE45CF23E219524">1</mods:start>
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<mods:classification id="C203780A4511049AF035FC946DCA582C">journal article</mods:classification>
<mods:identifier id="CC8026D35A6F87D637010432A697C16F" type="DOI">10.11646/zootaxa.5370.1.1</mods:identifier>
<mods:identifier id="B6EAADA6A51EA024B1A016A8A1E57825" type="ISSN">1175-5334</mods:identifier>
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<treatment id="03BEE17BFFF5C051FF0EF94D78E83559" LSID="urn:lsid:plazi:treatment:03BEE17BFFF5C051FF0EF94D78E83559" httpUri="http://treatment.plazi.org/id/03BEE17BFFF5C051FF0EF94D78E83559" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<subSubSection id="C30D03E6FFF5C050FF0EF94D7E1D31E1" box="[151,543,1727,1754]" pageId="52" pageNumber="53" type="nomenclature">
<paragraph id="8BA8506DFFF5C050FF0EF94D7E1D31E1" blockId="52.[151,543,1727,1754]" box="[151,543,1727,1754]" pageId="52" pageNumber="53">
<emphasis id="B9638C7FFFF5C050FF0EF94D7E1D31E1" bold="true" box="[151,543,1727,1754]" pageId="52" pageNumber="53">
Genus
<taxonomicName id="4C172BEEFFF5C050FF77F94D7E1D31E1" ID-CoL="BGDS4" authority="Sacco, 1895" authorityName="Sacco" authorityYear="1895" box="[238,543,1727,1754]" class="Gastropoda" family="Mathildidae" genus="Fimbriatella" kingdom="Animalia" order="Heterostropha" pageId="52" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF5C050FF77F94D7D8431E2" bold="true" box="[238,390,1727,1754]" italics="true" pageId="52" pageNumber="53">Fimbriatella</emphasis>
Sacco, 1895
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFF5C050FF0EF8FA7DFB3051" pageId="52" pageNumber="53" type="type_taxon">
<paragraph id="8BA8506DFFF5C050FF0EF8FA797C3019" blockId="52.[151,1437,1799,2042]" box="[151,1406,1799,1826]" pageId="52" pageNumber="53">
<materialsCitation id="3B7F5A30FFF5C050FF0EF8FA79783019" box="[151,1402,1799,1826]" country="Italy" location="Sacco" municipality="Miocene" pageId="52" pageNumber="53" specimenCount="1" typeStatus="holotype">
<emphasis id="B9638C7FFFF5C050FF0EF8FA7D303019" bold="true" box="[151,306,1799,1826]" pageId="52" pageNumber="53">
<typeStatus id="54ACEECFFFF5C050FF0EF8FA7CD0301A" box="[151,210,1800,1826]" pageId="52" pageNumber="53">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C172BEEFFF5C050FEA0F8F57EF1301A" authority="Michelotti, 1847" authorityName="Michelotti" authorityYear="1847" box="[313,755,1799,1826]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" kingdom="Animalia" order="Littorinimorpha" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="fimbriatum">
<emphasis id="B9638C7FFFF5C050FEA0F8F57E2C3019" box="[313,558,1799,1825]" italics="true" pageId="52" pageNumber="53">Cerithium fimbriatum</emphasis>
<bibRefCitation id="EF862D9CFFF5C050FDACF8F57EF1301A" author="Michelotti, G." box="[565,755,1799,1826]" pageId="52" pageNumber="53" pagination="1 - 408" refId="ref57076" refString="Michelotti, G. (1847) Description des fossiles des terrains miocenes de l'Italie septentrionale. Natuurkundige Verhandelingen van de Hollandsche Maatschappij der Wetenschappen te Haarlem, Series 2, 3 (2), 1 - 408. [https: // www. biodiversitylibrary. org / page / 2608445]" type="journal article" year="1847">Michelotti, 1847</bibRefCitation>
</taxonomicName>
; original designation by
<location id="8EC806B6FFF5C050FB91F8FA7849301A" LSID="urn:lsid:plazi:treatment:03BEE17BFFF5C051FF0EF94D78E83559:8EC806B6FFF5C050FB91F8FA7849301A" box="[1032,1099,1800,1826]" country="Italy" municipality="Miocene" name="Sacco" pageId="52" pageNumber="53">Sacco</location>
(1985: 36).
<collectingMunicipality id="6BCCCA17FFF5C050FB4CF8F579383019" box="[1237,1338,1799,1825]" pageId="52" pageNumber="53">Miocene</collectingMunicipality>
,
<collectingCountry id="F30010FDFFF5C050FADCF8F579783019" box="[1349,1402,1799,1825]" name="Italy" pageId="52" pageNumber="53">Italy</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA8506DFFF5C050FF0EF8DE7DFB3051" blockId="52.[151,1437,1799,2042]" pageId="52" pageNumber="53">
Note that Sacco (1895: 36 erroneously wrote “
<taxonomicName id="4C172BEEFFF5C050FD3EF8DE7FAC307D" ID-CoL="8V7ZG" authority="(Micht.)" baseAuthorityName="Micht." box="[679,942,1835,1861]" class="Gastropoda" family="Mathildidae" genus="Fimbriatella" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="fimbriatella">
<emphasis id="B9638C7FFFF5C050FD3EF8DE7F45307D" box="[679,839,1835,1861]" italics="true" pageId="52" pageNumber="53">F. fimbriatella</emphasis>
(Micht.)
</taxonomicName>
” but corrected this mistake in Sacco (1896: 81) (see also
<bibRefCitation id="EF862D9CFFF5C050FEB5F8BD7DEF3051" author="Bieler, R." box="[300,493,1871,1898]" pageId="52" pageNumber="53" pagination="595 - 641" refId="ref49179" refString="Bieler, R. (1995) Mathildidae from New Caledonia and the Loyalty Islands (Gastropoda: Heterobranchia). In: Bouchet, P. (Ed.), Resultats des campagnes MUSORSTOM 14. Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie, 167, pp. 595 - 641." type="journal article" year="1995">Bieler 1995: 600</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFF5C051FF5EF8817FE23694" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="diagnosis">
<paragraph id="8BA8506DFFF5C050FF5EF8817DEA30C1" blockId="52.[151,1437,1799,2042]" pageId="52" pageNumber="53">
<emphasis id="B9638C7FFFF5C050FF5EF8817DAB30B5" bold="true" box="[199,425,1907,1933]" pageId="52" pageNumber="53">Original diagnosis.</emphasis>
“
<emphasis id="B9638C7FFFF5C050FE24F8867FB83089" italics="true" pageId="52" pageNumber="53">
<taxonomicName id="4C172BEEFFF5C050FE24F8867E2A30B5" box="[445,552,1908,1933]" class="Gastropoda" family="Liotiidae" genus="Liotia" kingdom="Animalia" order="Trochida" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="specie">La specie</taxonomicName>
di questo Gruppo distinguonsi dale vere
<taxonomicName id="4C172BEEFFF5C050FC60F881786230B5" ID-CoL="8V5Q6" authorityName="O. Semper" authorityYear="1865" box="[1017,1120,1907,1933]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="52" pageNumber="53" phylum="Mollusca" rank="genus">Mathilda</taxonomicName>
per la forma più conica, gli anfratti fortemente angulosi, la bocca meno rotunda, subcaudata, ecc
</emphasis>
.” [species of this group are distinguished from real
<taxonomicName id="4C172BEEFFF5C050FE9CF8497D6E30ED" ID-CoL="8V5Q6" authorityName="O. Semper" authorityYear="1865" box="[261,364,1979,2005]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="52" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF5C050FE9CF8497D6E30ED" box="[261,364,1979,2005]" italics="true" pageId="52" pageNumber="53">Mathilda</emphasis>
</taxonomicName>
by its more conical shell, the strongly angular whorls, the less circular aperture, [its] subcaudate [base], etc.] (Sacco 1895: 36).
</paragraph>
<paragraph id="8BA8506DFFF4C051FF5EFF647FE23694" blockId="53.[151,1437,150,609]" pageId="53" pageNumber="54">
<emphasis id="B9638C7FFFF4C051FF5EFF647DA23788" bold="true" box="[199,416,150,176]" pageId="53" pageNumber="54">Revised diagnosis.</emphasis>
Medium-sized to large, solid, broad turritelliform with strongly angled whorls with concave profile. Protoconch heterostrophic Sculpture on first teleoconch whorl of three primary spiral cords: S1 moderately developed, placed abutting upper suture; S2 placed mid-whorl weakest; S3 strongest, placed close above abapical suture. Later whorls with carinate S3, coinciding with periphery. Secondary spiral cords may appear between suture and S1, and between S1 and S2. Axial sculpture of close-set, narrow, arcuate axial lamellae that overrun cords forming tubercles at intersections. Aperture subcircular, anteriorly slightly angulated. Columella slightly twisted. Columellar callus indistinct. No parietal callus. Siphonal canal short, wide. Differing from
<taxonomicName id="4C172BEEFFF4C051FB38FE9C790A36B0" ID-CoL="8V5Q6" authorityName="O. Semper" authorityYear="1865" box="[1185,1288,366,392]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="53" pageNumber="54" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF4C051FB38FE9C790A36B0" box="[1185,1288,366,392]" italics="true" pageId="53" pageNumber="54">Mathilda</emphasis>
</taxonomicName>
in it is weak S2, concave whorl profile and carinate S3, which coincides with periphery.
</paragraph>
</subSubSection>
<subSubSection id="C30D03E6FFF4C051FF5EFE4478E83559" pageId="53" pageNumber="54" type="discussion">
<paragraph id="8BA8506DFFF4C051FF5EFE4478E83559" blockId="53.[151,1437,150,609]" pageId="53" pageNumber="54">
<emphasis id="B9638C7FFFF4C051FF5EFE447D4E36E8" bold="true" box="[199,332,438,464]" pageId="53" pageNumber="54">Discussion.</emphasis>
<taxonomicName id="4C172BEEFFF4C051FECDFE447DE036E8" authorityName="Sacco" authorityYear="1895" box="[340,482,438,464]" class="Gastropoda" family="Mathildidae" genus="Fimbriatella" kingdom="Animalia" pageId="53" pageNumber="54" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF4C051FECDFE447DE036E8" box="[340,482,438,464]" italics="true" pageId="53" pageNumber="54">Fimbriatella</emphasis>
</taxonomicName>
was synonymized with
<taxonomicName id="4C172BEEFFF4C051FD68FE447F5A36E8" authorityName="O. Semper" authorityYear="1865" box="[753,856,438,464]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="53" pageNumber="54" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF4C051FD68FE447F5A36E8" box="[753,856,438,464]" italics="true" pageId="53" pageNumber="54">Mathilda</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF862D9CFFF4C051FC1AFE44785136E8" author="Bieler, R." box="[899,1107,438,465]" pageId="53" pageNumber="54" pagination="595 - 641" refId="ref49179" refString="Bieler, R. (1995) Mathildidae from New Caledonia and the Loyalty Islands (Gastropoda: Heterobranchia). In: Bouchet, P. (Ed.), Resultats des campagnes MUSORSTOM 14. Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie, 167, pp. 595 - 641." type="journal article" year="1995">Bieler (1995: 600)</bibRefCitation>
but in our opinion the different mode of sculpture formation and the concave whorl profile justify a separation of
<taxonomicName id="4C172BEEFFF4C051FBD5FE2878D836CC" authorityName="Sacco" authorityYear="1895" box="[1100,1242,474,500]" class="Gastropoda" family="Mathildidae" genus="Fimbriatella" kingdom="Animalia" pageId="53" pageNumber="54" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF4C051FBD5FE2878D836CC" box="[1100,1242,474,500]" italics="true" pageId="53" pageNumber="54">Fimbriatella</emphasis>
</taxonomicName>
. Extant
<taxonomicName id="4C172BEEFFF4C051FAAFFE287CEA3520" pageId="53" pageNumber="54">
<emphasis id="B9638C7FFFF4C051FAAFFE28799F36CC" box="[1334,1437,474,500]" italics="true" pageId="53" pageNumber="54">Mathilda</emphasis>
species
</taxonomicName>
such as
<taxonomicName id="4C172BEEFFF4C051FED5FE0C7E7A3520" authorityName="Kuroda" authorityYear="1958" box="[332,632,510,536]" class="Gastropoda" family="Mathildidae" genus="Mathilda" isUncertain="true" kingdom="Animalia" order="Littorinimorpha" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="quinquelirata">
<emphasis id="B9638C7FFFF4C051FED5FE0C7DB13520" box="[332,435,510,536]" italics="true" pageId="53" pageNumber="54">Mathilda</emphasis>
cf.
<emphasis id="B9638C7FFFF4C051FE79FE0C7E7A3520" box="[480,632,510,536]" italics="true" pageId="53" pageNumber="54">quinquelirata</emphasis>
</taxonomicName>
in
<bibRefCitation id="EF862D9CFFF4C051FD06FE0C7F343521" author="Bieler, R." box="[671,822,510,537]" pageId="53" pageNumber="54" pagination="595 - 641" refId="ref49179" refString="Bieler, R. (1995) Mathildidae from New Caledonia and the Loyalty Islands (Gastropoda: Heterobranchia). In: Bouchet, P. (Ed.), Resultats des campagnes MUSORSTOM 14. Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie, 167, pp. 595 - 641." type="journal article" year="1995">Bieler (1995)</bibRefCitation>
and
<taxonomicName id="4C172BEEFFF4C051FCE9FE0C793A3520" authority="(Okutani &amp; Habe, 1981)" baseAuthorityName="Okutani &amp; Habe" baseAuthorityYear="1981" box="[880,1336,510,536]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="fusca">
<emphasis id="B9638C7FFFF4C051FCE9FE0C78183520" box="[880,1050,510,536]" italics="true" pageId="53" pageNumber="54">Mathilda fusca</emphasis>
(Okutani &amp; Habe, 1981)
</taxonomicName>
differ in that the periphery is formed by S2, which migrates towards the abapical suture during ontogeny giving them a ‘
<emphasis id="B9638C7FFFF4C051FAE3FDD17CF53558" italics="true" pageId="53" pageNumber="54">Fibriatella</emphasis>
-like’ profile, but their periphery is at S2, and therefore they should be placed in
<taxonomicName id="4C172BEEFFF4C051FBE5FDB478E13558" authorityName="O. Semper" authorityYear="1865" box="[1148,1251,582,608]" class="Gastropoda" family="Mathildidae" genus="Mathilda" kingdom="Animalia" order="Littorinimorpha" pageId="53" pageNumber="54" phylum="Mollusca" rank="genus">
<emphasis id="B9638C7FFFF4C051FBE5FDB478E13558" box="[1148,1251,582,608]" italics="true" pageId="53" pageNumber="54">Mathilda</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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